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<document id="C4DF51F6C0C63E34CCB1A63525D178C7" ID-CLB-Dataset="29419" ID-DOI="10.11646/zootaxa.4482.3.1" ID-GBIF-Dataset="5916d93e-11d4-49c5-8085-8bcd37d3739b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440674" ID-ZooBank="10258391-162F-4C7D-AA5E-1A03A4F3FE19" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538374587817" checkinUser="plazi" docAuthor="Pal, Saunak, Vijayakumar, S. P., Shanker, Kartik, Jayarajan, Aditi &amp; Deepak, V." docDate="2018" docId="038E021DFFACFFB84FA9FB8AFDEEFB95" docLanguage="en" docName="zootaxa.4482.3.1.pdf" docOrigin="Zootaxa 4482 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monilesaurus ellioti Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak 2018, comb. nov." docType="treatment" docVersion="8" lastPageNumber="431" masterDocId="FFB77A65FFB0FFA64F3EFF9CFFEFFFE8" masterDocTitle="A systematic revision of Calotes Cuvier, 1817 (Squamata: Agamidae) from the Western Ghats adds two genera and reveals two new species" masterLastPageNumber="450" masterPageNumber="401" pageNumber="429" updateTime="1698559486032" updateUser="ExternalLinkService">
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<mods:title id="CFA0BB2BCD714AED4462DD53A4365648">A systematic revision of Calotes Cuvier, 1817 (Squamata: Agamidae) from the Western Ghats adds two genera and reveals two new species</mods:title>
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<mods:namePart id="7E487BE2F9A5F6EE916F6E38C1123EAB">Pal, Saunak</mods:namePart>
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<mods:namePart id="7BB4F15CD2C807FC8987EC0BFCB62679">Vijayakumar, S. P.</mods:namePart>
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<mods:namePart id="2D3D39FD9E3BCA5876946AEE4C4F0A93">Jayarajan, Aditi</mods:namePart>
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<treatment id="038E021DFFACFFB84FA9FB8AFDEEFB95" ID-DOI="http://doi.org/10.5281/zenodo.5996701" ID-GBIF-Taxon="148421701" ID-Zenodo-Dep="5996701" LSID="urn:lsid:plazi:treatment:038E021DFFACFFB84FA9FB8AFDEEFB95" httpUri="http://treatment.plazi.org/id/038E021DFFACFFB84FA9FB8AFDEEFB95" lastPageId="30" lastPageNumber="431" pageId="28" pageNumber="429">
<subSubSection id="C33DE080FFACFFBA4FA9FB8AFB6EFB9B" pageId="28" pageNumber="429" type="nomenclature">
<paragraph id="8B98B30BFFACFFBA4FA9FB8AFD0AFBD8" blockId="28.[151,741,1046,1072]" box="[151,741,1046,1072]" pageId="28" pageNumber="429">
<heading id="D0D00467FFACFFBA4FA9FB8AFD0AFBD8" bold="true" box="[151,741,1046,1072]" fontSize="11" level="1" pageId="28" pageNumber="429" reason="1">
<emphasis id="B9536F19FFACFFBA4FA9FB8AFD0AFBD8" bold="true" box="[151,741,1046,1072]" pageId="28" pageNumber="429">
<taxonomicName id="4C27C888FFACFFBA4FA9FB8AFDB8FBD8" authority="(Gunther, 1864)" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[151,599,1046,1072]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="species" species="ellioti" status="comb. nov.">
<emphasis id="B9536F19FFACFFBA4FA9FB8AFE65FBD8" bold="true" box="[151,394,1046,1072]" italics="true" pageId="28" pageNumber="429">Monilesaurus ellioti</emphasis>
(
<bibRefCitation id="EFB6CEFAFFACFFBA4EA7FB8AFDA0FBD8" box="[409,591,1046,1072]" pageId="28" pageNumber="429" refId="ref35460" refString="Gunther, A. (1864) The Reptiles of British India. Taylor &amp; Francis, London, 452 pp." type="book">Günther, 1864</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A260D262FFACFFBA4D60FB8AFD0AFBD8" box="[606,741,1046,1072]" pageId="28" pageNumber="429">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B98B30BFFACFFBA4FA9FBC1FB6EFB9B" blockId="28.[151,1153,1117,1231]" box="[151,1153,1117,1139]" pageId="28" pageNumber="429">
<emphasis id="B9536F19FFACFFBA4FA9FBC1FED2FB9B" box="[151,317,1117,1139]" italics="true" pageId="28" pageNumber="429">
<taxonomicName id="4C27C888FFACFFBA4FA9FBC1FF0BFB9B" box="[151,228,1117,1139]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="genus">Calotes</taxonomicName>
rouxii—
</emphasis>
(not of
<bibRefCitation id="EFB6CEFAFFACFFBA4EB9FBC2FDBAFB9B" box="[391,597,1117,1139]" pageId="28" pageNumber="429" refId="ref34976" refString="Dumril, A. M. C. &amp; Bibron, G. (1837) Erpetologie Generale ou Histoire Naturelle Complete des Reptiles. Fol. 4. Librairie Encyclopedique Roret, Paris, 570 pp." type="book">Dum. &amp; Bibr., 1837</bibRefCitation>
), Jerdon, 1853. J. Asiat. Soc. Beng. (2) xxii, 1853: 471
</paragraph>
</subSubSection>
<subSubSection id="C33DE080FFACFFBA4FA9FBE0FEFEF893" pageId="28" pageNumber="429" type="description">
<paragraph id="8B98B30BFFACFFBA4FA9FBE0FD06FB7A" blockId="28.[151,1153,1117,1231]" box="[151,745,1148,1170]" pageId="28" pageNumber="429">
<taxonomicName id="4C27C888FFACFFBA4FA9FBE0FEC9FB7A" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" baseAuthorityYear="2018" box="[151,294,1148,1170]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="species" species="ellioti">
<emphasis id="B9536F19FFACFFBA4FA9FBE0FEC9FB7A" box="[151,294,1148,1170]" italics="true" pageId="28" pageNumber="429">Calotes ellioti</emphasis>
</taxonomicName>
<bibRefCitation id="EFB6CEFAFFACFFBA4E01FBE0FE3CFB7A" box="[319,467,1148,1170]" pageId="28" pageNumber="429" refId="ref35460" refString="Gunther, A. (1864) The Reptiles of British India. Taylor &amp; Francis, London, 452 pp." type="book">Günther, 1864</bibRefCitation>
. Rept. Brit. Ind. 1864: 142.
</paragraph>
<paragraph id="8B98B30BFFACFFBA4FA9FB06FCB2FB58" blockId="28.[151,1153,1117,1231]" box="[151,861,1178,1200]" pageId="28" pageNumber="429">
<taxonomicName id="4C27C888FFACFFBA4FA9FB06FEB3FB58" authorityName="Theobald" authorityYear="1876" box="[151,348,1178,1200]" class="Reptilia" family="Agamidae" genus="Bronchocela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="species" species="indica">
<emphasis id="B9536F19FFACFFBA4FA9FB06FEB3FB58" box="[151,348,1178,1200]" italics="true" pageId="28" pageNumber="429">Bronchocela indica</emphasis>
</taxonomicName>
—Theobald, 1876. Cat. Rept. Brit. Ind. 1876: 105.
</paragraph>
<paragraph id="8B98B30BFFACFFBA4FA9FB25FCADFB27" blockId="28.[151,1153,1117,1231]" box="[151,834,1209,1231]" pageId="28" pageNumber="429">
<taxonomicName id="4C27C888FFACFFBA4FA9FB25FEC2FB27" authorityName="Smith" authorityYear="1935" box="[151,301,1209,1231]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="species" species="elliotti">
<emphasis id="B9536F19FFACFFBA4FA9FB25FEC2FB27" box="[151,301,1209,1231]" italics="true" pageId="28" pageNumber="429">Calotes elliotti</emphasis>
</taxonomicName>
<bibRefCitation id="EFB6CEFAFFACFFBA4E79FB25FE2AFB27" box="[327,453,1209,1231]" pageId="28" pageNumber="429" refId="ref37909" refString="Smith, M. A. (1935) The Fauna of British India, Including Ceylon and Burma; Reptilia and Amphibia. Fol. II. Sauria. Taylor and Francis, London, 440 pp." type="book">Smith, 1935</bibRefCitation>
. Fauna of
<collectingCountry id="F330F39BFFACFFBA4D10FB25FD40FB27" box="[558,687,1209,1231]" name="India" pageId="28" pageNumber="429">British India</collectingCountry>
, ii, 1935: 207.
</paragraph>
<paragraph id="8B98B30BFFACFFBA4FA9FB60FA7FFAFC" blockId="28.[151,1436,1276,2023]" box="[151,1424,1276,1301]" pageId="28" pageNumber="429">
<emphasis id="B9536F19FFACFFBA4FA9FB60FE7FFAFD" bold="true" box="[151,400,1276,1301]" pageId="28" pageNumber="429">Original description.</emphasis>
<bibRefCitation id="EFB6CEFAFFACFFBA4EA9FB60FDB1FAFD" box="[407,606,1276,1301]" pageId="28" pageNumber="429" refId="ref35460" refString="Gunther, A. (1864) The Reptiles of British India. Taylor &amp; Francis, London, 452 pp." type="book">Günther, A. 1864</bibRefCitation>
. The
<taxonomicName id="4C27C888FFACFFBA4DA4FB60FD19FAFD" authorityName="Linnæus" authorityYear="1758" box="[666,758,1276,1301]" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptiles" pageId="28" pageNumber="429" phylum="Chordata" rank="order">Reptiles</taxonomicName>
of
<collectingCountry id="F330F39BFFACFFBA4C25FB60FC44FAFD" box="[795,939,1276,1301]" name="India" pageId="28" pageNumber="429">British India</collectingCountry>
. London (Taylor &amp; Francis), xxvii, 452 pp.
</paragraph>
<paragraph id="8B98B30BFFACFFBA4FF9FA83FE0FF8D8" blockId="28.[151,1436,1276,2023]" pageId="28" pageNumber="429">
<emphasis id="B9536F19FFACFFBA4FF9FA83FE3BFAD0" bold="true" box="[199,468,1311,1336]" pageId="28" pageNumber="429">Taxonomic comments.</emphasis>
<taxonomicName id="4C27C888FFACFFBA4EE0FABCFD2CFAD0" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[478,707,1312,1336]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="species" species="ellioti">
<emphasis id="B9536F19FFACFFBA4EE0FABCFD2CFAD0" box="[478,707,1312,1336]" italics="true" pageId="28" pageNumber="429">Monilesaurus ellioti</emphasis>
</taxonomicName>
was described, based on Jerdons observation of
<taxonomicName id="4C27C888FFACFFBA4A3DFABCFA87FADF" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1837" box="[1283,1384,1312,1335]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="species" species="rouxii">
<emphasis id="B9536F19FFACFFBA4A3DFABCFA87FADF" box="[1283,1384,1312,1335]" italics="true" pageId="28" pageNumber="429">C. rouxii</emphasis>
</taxonomicName>
like animals from “Malabar” in his list of
<taxonomicName id="4C27C888FFACFFBA4D01FAD8FD74FAB5" authorityName="Linnæus" authorityYear="1758" box="[575,667,1348,1373]" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptiles" pageId="28" pageNumber="429" phylum="Chordata" rank="order">Reptiles</taxonomicName>
of
<collectingRegion id="49E37DE9FFACFFBA4DFFFAD8FCC9FAB5" box="[705,806,1348,1373]" country="Sri Lanka" name="Southern" pageId="28" pageNumber="429">Southern</collectingRegion>
<collectingCountry id="F330F39BFFACFFBA4C10FAD8FC85FAB5" box="[814,874,1348,1373]" name="India" pageId="28" pageNumber="429">India</collectingCountry>
, 1853 and a collection of drawings of this species in the possession of W. Elliot. Historically, southern
<collectingRegion id="49E37DE9FFACFFBA4C34FAFBFC92FA68" box="[778,893,1383,1408]" country="India" name="Karnataka" pageId="28" pageNumber="429">Karnataka</collectingRegion>
and northern
<collectingRegion id="49E37DE9FFACFFBA4B19FAFBFB9DFA68" box="[1063,1138,1383,1408]" country="India" name="Kerala" pageId="28" pageNumber="429">Kerala</collectingRegion>
have been referred to as Malabar.
<bibRefCitation id="EFB6CEFAFFACFFBA4E3AFA10FE23FA4D" box="[260,460,1420,1445]" pageId="28" pageNumber="429" refId="ref34283" refString="Boulenger, G. A. (1885) Catalogue of lizards in the British museum. Fol. 1. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae, Second edition. British Museum (Natural History), London, 436 pp." type="book">Boulenger (1885)</bibRefCitation>
gave a short description of this species based on the collections made by Col. R.H. Beddome from Anamalai hills, Sivagiri Ghat, Tirunelveli and Malabar. As there are no
<typeStatus id="549C0DA9FFACFFBA4B4AFA2CFB4BFA20" box="[1140,1188,1456,1480]" pageId="28" pageNumber="429">type</typeStatus>
specimens associated with this species, we hereby designate a
<typeStatus id="549C0DA9FFACFFBA4D62FA49FD5AFA05" box="[604,693,1493,1517]" pageId="28" pageNumber="429" type="neotype">neotype</typeStatus>
for this species. The selection of the
<typeStatus id="549C0DA9FFACFFBA4B6BFA49FB41FA05" box="[1109,1198,1493,1517]" pageId="28" pageNumber="429" type="neotype">neotype</typeStatus>
is roughly congruent with the original
<typeStatus id="549C0DA9FFACFFBA4E68FA64FE69F9F8" box="[342,390,1528,1552]" pageId="28" pageNumber="429">type</typeStatus>
locality of this species “Malabar” as mentioned by Jerdon (1853). Murthy (1978) described a subspecies
<taxonomicName id="4C27C888FFACFFBA4E2BF980FD79F9DC" authorityName="Murthy" authorityYear="1978" box="[277,662,1564,1588]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="subSpecies" species="ellioti" subSpecies="amarambalamensis">
<emphasis id="B9536F19FFACFFBA4E2BF980FD79F9DC" box="[277,662,1564,1588]" italics="true" pageId="28" pageNumber="429">Calotes ellioti amarambalamensis</emphasis>
</taxonomicName>
the
<typeStatus id="549C0DA9FFACFFBA4DF8F981FD19F9DD" box="[710,758,1565,1589]" pageId="28" pageNumber="429">type</typeStatus>
locality of which is Amarambalam, Nilambur. The
<typeStatus id="549C0DA9FFACFFBA4A04F980FA73F9DD" box="[1338,1436,1564,1589]" pageId="28" pageNumber="429" type="holotype">holotype</typeStatus>
of this species is now at ZSI Madras (ZSI Madras 159) (Das
<emphasis id="B9536F19FFACFFBA4C7BF9DDFC99F9B0" box="[837,886,1600,1624]" italics="true" pageId="28" pageNumber="429">et al</emphasis>
. 1998). We were unable to trace the specimen in ZSI collections despite repeated attempts. For now, the specimen may be considered as lost. Hence, we compare another specimen collected from the same locality (BNHS 1033), a female individual whose characters match well with other
<taxonomicName id="4C27C888FFACFFBA4E2DF931FE94F92C" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" baseAuthorityYear="2018" box="[275,379,1708,1732]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="species" species="ellioti">
<emphasis id="B9536F19FFACFFBA4E2DF931FE94F92C" box="[275,379,1708,1732]" italics="true" pageId="28" pageNumber="429">M. ellioti</emphasis>
</taxonomicName>
and the description provided by Murthy. Also, the locality is within the distribution range of
<taxonomicName id="4C27C888FFACFFBA4FA9F94DFEEFF900" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" baseAuthorityYear="2018" box="[151,256,1744,1768]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="species" species="ellioti">
<emphasis id="B9536F19FFACFFBA4FA9F94DFEEFF900" box="[151,256,1744,1768]" italics="true" pageId="28" pageNumber="429">M. ellioti</emphasis>
</taxonomicName>
and contiguous with the
<typeStatus id="549C0DA9FFACFFBA4D15F94CFDB4F900" box="[555,603,1744,1768]" pageId="28" pageNumber="429">type</typeStatus>
locality. Therefore, we consider
<taxonomicName id="4C27C888FFACFFBA4CE3F94CFA8CF900" authorityName="Murthy" authorityYear="1978" box="[989,1379,1744,1768]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="subSpecies" species="ellioti" subSpecies="amarambalamensis">
<emphasis id="B9536F19FFACFFBA4CE3F94CFA8CF900" box="[989,1379,1744,1768]" italics="true" pageId="28" pageNumber="429">Calotes ellioti amarambalamensis</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C27C888FFACFFBA4E52F969FD63F8E5" authority="(Gunther, 1864)" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[364,652,1780,1805]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="species" species="ellioti">
<emphasis id="B9536F19FFACFFBA4E52F969FE3CF8E4" box="[364,467,1780,1804]" italics="true" pageId="28" pageNumber="429">M. ellioti</emphasis>
(
<bibRefCitation id="EFB6CEFAFFACFFBA4EDEF968FD6CF8E5" box="[480,643,1780,1805]" pageId="28" pageNumber="429" refId="ref35460" refString="Gunther, A. (1864) The Reptiles of British India. Taylor &amp; Francis, London, 452 pp." type="book">Günther, 1864</bibRefCitation>
)
</taxonomicName>
. ZSI Madras 159 is wrongly mentioned as a
<typeStatus id="549C0DA9FFACFFBA4BBFF969FB5EF8E5" box="[1153,1201,1781,1805]" pageId="28" pageNumber="429">type</typeStatus>
specimen of
<taxonomicName id="4C27C888FFACFFBA4A78F968FE33F8D8" authority="(Uetz &amp; Hosek, 2016)" baseAuthorityName="Uetz &amp; Hosek" baseAuthorityYear="2016" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="429" phylum="Chordata" rank="species" species="ellioti">
<emphasis id="B9536F19FFACFFBA4A78F968FF37F8D8" italics="true" pageId="28" pageNumber="429">Calotes ellioti</emphasis>
(
<bibRefCitation id="EFB6CEFAFFACFFBA4FD9F884FE3BF8D8" box="[231,468,1815,1840]" pageId="28" pageNumber="429" refId="ref38353" refString="Uetz, P. &amp; Hosek, J. (2016) The Reptile Database. Available from: http: // reptile-database. reptarium. cz (accessed 6 May 2016)" type="url">Uetz &amp; Hošek, 2016</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B98B30BFFACFFBA4FF9F8A2FEFEF893" blockId="28.[151,1436,1276,2023]" pageId="28" pageNumber="429">
<emphasis id="B9536F19FFACFFBA4FF9F8A2FEDFF8BF" bold="true" box="[199,304,1854,1879]" pageId="28" pageNumber="429">
<typeStatus id="549C0DA9FFACFFBA4FF9F8A2FEC4F8BF" box="[199,299,1854,1879]" pageId="28" pageNumber="429" type="neotype">Neotype</typeStatus>
.
</emphasis>
CESL 0 45, adult male collected from Chembra reserve forest,
<collectingRegion id="49E37DE9FFACFFBA4CCBF8A2FBAFF8BF" box="[1013,1088,1854,1879]" country="India" name="Kerala" pageId="28" pageNumber="429">Kerala</collectingRegion>
by SPP, MVP and SPV on
<date id="FF9995CBFFACFFBA4A4AF8A3FEE2F893" pageId="28" pageNumber="429" value="2010-06-10">10th June 2010</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C33DE080FFACFFBB4FF9F81AFC90FE60" lastPageId="29" lastPageNumber="430" pageId="28" pageNumber="429" type="materials_examined">
<paragraph id="8B98B30BFFACFFBB4FF9F81AFC90FE60" blockId="28.[151,1436,1276,2023]" lastBlockId="29.[151,1437,151,2013]" lastPageId="29" lastPageNumber="430" pageId="28" pageNumber="429">
<materialsCitation id="3B4FB956FFACFFBA4FF9F81AFAA4F877" ID-GBIF-Occurrence="1913889719" box="[199,1355,1926,1951]" country="India" location="Southern" pageId="28" pageNumber="429" specimenCount="1" stateProvince="Southern">
<emphasis id="B9536F19FFACFFBA4FF9F81AFE10F877" bold="true" box="[199,511,1926,1951]" pageId="28" pageNumber="429">Other material examined.</emphasis>
ZSI 4325 gravid female &amp; ZSI 4328 male collected from
<collectingRegion id="49E37DE9FFACFFBA4BA1F81AFAEBF877" box="[1183,1284,1926,1951]" country="Sri Lanka" name="Southern" pageId="28" pageNumber="429">Southern</collectingRegion>
<collectingCountry id="F330F39BFFACFFBA4A30F81AFAA4F877" box="[1294,1355,1926,1951]" name="India" pageId="28" pageNumber="429">India</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B4FB956FFACFFBA4A69F81BFBC7F82B" ID-GBIF-Occurrence="1913889714" collectionCode="CESL" country="India" location="Kerala" pageId="28" pageNumber="429" specimenCount="42" specimenCount-adult="42" stateProvince="Kerala">
<collectionCode id="ED362BCEFFACFFBA4A69F81BFA74F877" box="[1367,1435,1927,1951]" pageId="28" pageNumber="429">CESL</collectionCode>
0
<specimenCount id="9D217882FFACFFBA4F9AF837FEEDF82B" box="[164,258,1962,1987]" pageId="28" pageNumber="429" type="adult">42 adult</specimenCount>
male collected from Manikunjamalai, Wayanad reserve forest,
<collectingRegion id="49E37DE9FFACFFBA4CE4F836FBC7F82B" box="[986,1064,1962,1987]" country="India" name="Kerala" pageId="28" pageNumber="429">Kerala</collectingRegion>
</materialsCitation>
;
<materialsCitation id="3B4FB956FFACFFBA4B08F837FDABF80F" ID-GBIF-Occurrence="1913889716" collectionCode="CESL" country="India" location="Chembra" pageId="28" pageNumber="429" specimenCount="45" specimenCount-adult="45" stateProvince="Kerala">
<collectionCode id="ED362BCEFFACFFBA4B08F837FB95F82B" box="[1078,1146,1963,1987]" pageId="28" pageNumber="429">CESL</collectionCode>
0
<specimenCount id="9D217882FFACFFBA4BAFF836FB01F82B" box="[1169,1262,1962,1987]" pageId="28" pageNumber="429" type="adult">45 adult</specimenCount>
male collected from
<location id="8EF8E5D0FFACFFBA4FE6F852FED1F80F" LSID="urn:lsid:plazi:treatment:038E021DFFACFFB84FA9FB8AFDEEFB95:8EF8E5D0FFACFFBA4FE6F852FED1F80F" box="[216,318,1998,2023]" country="India" name="Chembra" pageId="28" pageNumber="429" stateProvince="Kerala">Chembra</location>
reserve forest,
<collectingRegion id="49E37DE9FFACFFBA4EC8F852FDABF80F" box="[502,580,1998,2023]" country="India" name="Kerala" pageId="28" pageNumber="429">Kerala</collectingRegion>
</materialsCitation>
;
<materialsCitation id="3B4FB956FFACFFBB4D6DF853FF0AFF58" ID-GBIF-Occurrence="1913889727" collectionCode="CESL" country="India" lastPageId="29" lastPageNumber="430" location="Kerala" pageId="28" pageNumber="429" specimenCount="47" specimenCount-adult="47" stateProvince="Kerala">
<collectionCode id="ED362BCEFFACFFBA4D6DF853FD78F80F" box="[595,663,1999,2023]" pageId="28" pageNumber="429">CESL</collectionCode>
0
<specimenCount id="9D217882FFACFFBA4D8EF853FCE0F80F" box="[688,783,1998,2023]" pageId="28" pageNumber="429" type="adult">47 adult</specimenCount>
female collected from Vaithiri, Wayanad reserve forest,
<collectingRegion id="49E37DE9FFADFFBB4FA9FF0BFF0AFF58" box="[151,229,151,176]" country="India" name="Kerala" pageId="29" pageNumber="430">Kerala</collectingRegion>
</materialsCitation>
; CESL 0 57,
<materialsCitation id="3B4FB956FFADFFBB4E47FF04FD04FF3D" ID-GBIF-Occurrence="1913889717" collectionCode="CESL" country="India" location="Kerala" pageId="29" pageNumber="430" specimenCount="121" specimenCount-adult="121" stateProvince="Kerala">
<collectionCode id="ED362BCEFFADFFBB4E47FF04FE52FF58" box="[377,445,152,176]" pageId="29" pageNumber="430">CESL</collectionCode>
0 59 and
<collectionCode id="ED362BCEFFADFFBB4D14FF04FD81FF58" box="[554,622,152,176]" pageId="29" pageNumber="430">CESL</collectionCode>
0
<specimenCount id="9D217882FFADFFBB4DB8FF0BFD0DFF58" box="[646,738,151,176]" pageId="29" pageNumber="430" type="adult">61 adult</specimenCount>
females and
<collectionCode id="ED362BCEFFADFFBB4C45FF04FC50FF58" box="[891,959,152,176]" pageId="29" pageNumber="430">CESL</collectionCode>
0 58 and
<collectionCode id="ED362BCEFFADFFBB4B12FF04FB9FFF58" box="[1068,1136,152,176]" pageId="29" pageNumber="430">CESL</collectionCode>
0
<specimenCount id="9D217882FFADFFBB4BB9FF0BFB0BFF58" box="[1159,1252,151,176]" pageId="29" pageNumber="430" type="adult">60 adult</specimenCount>
males collected from Vallerimala, Wayanad reserve forest,
<collectingRegion id="49E37DE9FFADFFBB4DA3FF20FD04FF3D" box="[669,747,188,213]" country="India" name="Kerala" pageId="29" pageNumber="430">Kerala</collectingRegion>
</materialsCitation>
;
<materialsCitation id="3B4FB956FFADFFBB4DC0FF20FE8EFF10" ID-GBIF-Occurrence="1913889729" collectionCode="CESL" country="India" location="Arlam Wildlife Sanctuary" pageId="29" pageNumber="430" specimenCount="1" stateProvince="Kerala">
<collectionCode id="ED362BCEFFADFFBB4DC0FF20FCADFF3C" box="[766,834,188,212]" pageId="29" pageNumber="430">CESL</collectionCode>
0 77, adult female collected from
<location id="8EF8E5D0FFADFFBB4BD4FF20FEE7FF10" LSID="urn:lsid:plazi:treatment:038E021DFFACFFB84FA9FB8AFDEEFB95:8EF8E5D0FFADFFBB4BD4FF20FEE7FF10" country="India" name="Arlam Wildlife Sanctuary" pageId="29" pageNumber="430" stateProvince="Kerala">Arlam Wildlife Sanctuary</location>
,
<collectingRegion id="49E37DE9FFADFFBB4E2DFF43FE8EFF10" box="[275,353,223,248]" country="India" name="Kerala" pageId="29" pageNumber="430">Kerala</collectingRegion>
</materialsCitation>
;
<materialsCitation id="3B4FB956FFADFFBB4E52FF7CFAFAFF10" ID-GBIF-Occurrence="1913889718" box="[364,1301,223,248]" collectionCode="CESL" country="India" location="Peppara Wildlife Sanctuary" municipality="Bonacord" pageId="29" pageNumber="430" specimenCount="33" specimenCount-adult="33" stateProvince="Kerala">
<collectionCode id="ED362BCEFFADFFBB4E52FF7CFE5FFF10" box="[364,432,224,248]" pageId="29" pageNumber="430">CESL</collectionCode>
0
<specimenCount id="9D217882FFADFFBB4EFBFF7CFDCFFF10" box="[453,544,223,248]" pageId="29" pageNumber="430" type="adult">33 adult</specimenCount>
male collected from
<collectingMunicipality id="6BFC2971FFADFFBB4C33FF43FC92FF10" box="[781,893,223,248]" pageId="29" pageNumber="430">Bonacord</collectingMunicipality>
,
<location id="8EF8E5D0FFADFFBB4CB9FF7CFB53FF10" LSID="urn:lsid:plazi:treatment:038E021DFFACFFB84FA9FB8AFDEEFB95:8EF8E5D0FFADFFBB4CB9FF7CFB53FF10" box="[903,1212,223,248]" country="India" municipality="Bonacord" name="Peppara Wildlife Sanctuary" pageId="29" pageNumber="430" stateProvince="Kerala">Peppara Wildlife Sanctuary</location>
,
<collectingRegion id="49E37DE9FFADFFBB4BF9FF43FAFAFF10" box="[1223,1301,223,248]" country="India" name="Kerala" pageId="29" pageNumber="430">Kerala</collectingRegion>
</materialsCitation>
;
<materialsCitation id="3B4FB956FFADFFBB4A1EFF7CFC56FEF5" ID-GBIF-Occurrence="1913889723" collectionCode="CESL" country="India" location="Kerala" pageId="29" pageNumber="430" specimenCode="CESL 162" specimenCount="1" stateProvince="Kerala">
<specimenCode id="DB811B70FFADFFBB4A1EFF7CFA78FF10" box="[1312,1431,223,248]" collectionCode="CESL" pageId="29" pageNumber="430">CESL 162</specimenCode>
, adult male collected from Sholyar, Vazhachal reserve forest,
<collectingRegion id="49E37DE9FFADFFBB4C55FE98FC56FEF5" box="[875,953,260,285]" country="India" name="Kerala" pageId="29" pageNumber="430">Kerala</collectingRegion>
</materialsCitation>
;
<materialsCitation id="3B4FB956FFADFFBB4CF7FE98FC90FE60" ID-GBIF-Occurrence="1913889713" collectionCode="CESL" country="India" county="Nilambur" location="Appendix" municipality="GenBank" pageId="29" pageNumber="430" specimenCode="CESL 204, BNHS 1033" specimenCount="369" specimenCount-adult="369" stateProvince="Kerala">
<specimenCode id="DB811B70FFADFFBB4CF7FE98FBA8FEF4" box="[969,1095,260,284]" collectionCode="CESL" pageId="29" pageNumber="430">CESL 204</specimenCode>
, adult female collected from Pamba, Periyar tiger reserve,
<collectingRegion id="49E37DE9FFADFFBB4ED5FEBBFDD9FEA8" box="[491,566,295,320]" country="India" name="Kerala" pageId="29" pageNumber="430">Kerala</collectingRegion>
and
<collectionCode id="ED362BCEFFADFFBB4D4FFEB4FD5AFEA8" box="[625,693,296,320]" pageId="29" pageNumber="430">CESL</collectionCode>
<specimenCount id="9D217882FFADFFBB4D80FEBBFCC5FEA8" box="[702,810,295,320]" pageId="29" pageNumber="430" type="adult">369 adult</specimenCount>
male collected from Peppara, Cardemom hills,
<collectingRegion id="49E37DE9FFADFFBB4A6FFEBBFA73FEA8" box="[1361,1436,295,320]" country="India" name="Kerala" pageId="29" pageNumber="430">Kerala</collectingRegion>
and
<specimenCode id="DB811B70FFADFFBB4FF2FED0FEB1FE8C" box="[204,350,332,356]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="29" pageNumber="430">BNHS 1033</specimenCode>
, adult female collected from
<collectingCounty id="62F9CB87FFADFFBB4DFFFED0FCC3FE8D" box="[705,812,332,357]" pageId="29" pageNumber="430">Nilambur</collectingCounty>
reserve forest,
<collectingRegion id="49E37DE9FFADFFBB4CD4FED0FBD7FE8D" box="[1002,1080,332,357]" country="India" name="Kerala" pageId="29" pageNumber="430">Kerala</collectingRegion>
. Details of collection locality, specimen voucher and
<collectingMunicipality id="6BFC2971FFADFFBB4EA6FEF3FDEDFE60" box="[408,514,367,392]" pageId="29" pageNumber="430">GenBank</collectingMunicipality>
accession number in
<location id="8EF8E5D0FFADFFBB4DCBFEF3FC8CFE60" LSID="urn:lsid:plazi:treatment:038E021DFFACFFB84FA9FB8AFDEEFB95:8EF8E5D0FFADFFBB4DCBFEF3FC8CFE60" box="[757,867,367,392]" country="India" county="Nilambur" municipality="GenBank" name="Appendix" pageId="29" pageNumber="430" stateProvince="Kerala">Appendix</location>
1.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33DE080FFADFFBB4FF9FE08FAF1FBF8" pageId="29" pageNumber="430" type="diagnosis">
<paragraph id="8B98B30BFFADFFBB4FF9FE08FD2CFD6D" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
<emphasis id="B9536F19FFADFFBB4FF9FE08FDF7FE45" bold="true" box="[199,536,404,429]" pageId="29" pageNumber="430">Diagnosis and comparison.</emphasis>
A small sized
<taxonomicName id="4C27C888FFADFFBB4DE1FE08FC96FE44" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[735,889,404,428]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="444" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B9536F19FFADFFBB4DE1FE08FC96FE44" box="[735,889,404,428]" italics="true" pageId="29" pageNumber="430">Monilesaurus</emphasis>
</taxonomicName>
(SVL up to
<quantity id="4CDF1EEEFFADFFBB4B18FE08FB7CFE44" box="[1062,1171,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="7.38" pageId="29" pageNumber="430" unit="mm" value="73.8">73.8 mm</quantity>
) characterized by the posteroventral orientation of lateral scales; antehumeral fold well developed, extending across the throat; 5258 midbody scale rows; nuchal crest composed of 34 long, well developed spines; two separated supratympanic spines; a long, distinct isolated postorbital spine; dorsal and lateral scales keeled, ventral scales strongly keeled; paired postmentals, first pair separated by a single scale; 2428 subdigital lamellae under fourth finger, 2634 subdigital lamellae under fourth toe; 910 supralabials and 89 infralabials; olive brown above with angular darker cross bars on dorsum, a white spot below the eye.
</paragraph>
<paragraph id="8B98B30BFFADFFBB4FF9FD13FAF1FBF8" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
<taxonomicName id="4C27C888FFADFFBB4FF9FD13FE1CFD40" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[199,499,655,680]" pageId="29" pageNumber="430" rank="subSpecies" subSpecies="ellioti">
Morphologically
<emphasis id="B9536F19FFADFFBB4EB5FD0DFE1CFD40" box="[395,499,656,680]" italics="true" pageId="29" pageNumber="430">M. ellioti</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName id="4C27C888FFADFFBB4C25FD0DFC41FD40" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[795,942,657,680]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="montanus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4C25FD0DFC41FD40" box="[795,942,657,680]" italics="true" pageId="29" pageNumber="430">M. montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4C8BFD13FB8AFD4F" box="[949,1125,655,680]" pageId="29" pageNumber="430" rank="species">
<emphasis id="B9536F19FFADFFBB4C8BFD13FBB1FD40" bold="true" box="[949,1118,655,680]" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<taxonomicName id="4C27C888FFADFFBB4B4AFD0DFB33FD4F" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1837" box="[1140,1244,656,680]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="rouxii" status="comb. nov.">
<emphasis id="B9536F19FFADFFBB4B4AFD0DFB33FD4F" box="[1140,1244,656,680]" italics="true" pageId="29" pageNumber="430">M. rouxii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4BDDFD13FA8CFD4F" box="[1251,1379,655,680]" pageId="29" pageNumber="430">
<emphasis id="B9536F19FFADFFBB4BDDFD13FAB2FD40" bold="true" box="[1251,1373,655,680]" pageId="29" pageNumber="430">comb. nov</emphasis>
.
</taxonomicNameLabel>
; and
<taxonomicName id="4C27C888FFADFFBB4FA9FD29FE95FD24" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[151,378,692,716]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="acanthocephalus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4FA9FD29FE95FD24" box="[151,378,692,716]" italics="true" pageId="29" pageNumber="430">M. acanthocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4EBFFD28FDC2FD24" box="[385,557,692,717]" pageId="29" pageNumber="430" rank="species">
<emphasis id="B9536F19FFADFFBB4EBFFD28FDC8FD25" bold="true" box="[385,551,692,717]" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, but can be distinguished by a combination of the following characters: 5258 midbody scale rows (vs.
<quantity id="4CDF1EEEFFADFFBB4E92FD4BFDE0FD18" box="[428,527,727,752]" metricMagnitude="0" metricUnit="m" metricValue="1.2446" metricValueMax="1.3208" metricValueMin="1.1684" pageId="29" pageNumber="430" unit="in" value="49.0" valueMax="52.0" valueMin="46.0">4652 in</quantity>
<taxonomicName id="4C27C888FFADFFBB4D2BFD45FD48FD18" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[533,679,729,752]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="montanus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4D2BFD45FD48FD18" box="[533,679,729,752]" italics="true" pageId="29" pageNumber="430">M. montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4D93FD4BFCB5FD07" box="[685,858,727,752]" pageId="29" pageNumber="430" rank="species">
<emphasis id="B9536F19FFADFFBB4D93FD4BFCBCFD18" bold="true" box="[685,851,727,752]" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<quantity id="4CDF1EEEFFADFFBB4C59FD4BFC25FD18" box="[871,970,727,752]" metricMagnitude="0" metricUnit="m" metricValue="1.6002" metricValueMax="1.6256" metricValueMin="1.5748" pageId="29" pageNumber="430" unit="in" value="63.0" valueMax="64.0" valueMin="62.0">6264 in</quantity>
<taxonomicName id="4C27C888FFADFFBB4CEEFD45FB5CFD18" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[976,1203,728,752]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="acanthocephalus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4CEEFD45FB5CFD18" box="[976,1203,728,752]" italics="true" pageId="29" pageNumber="430">M. acanthocephalus</emphasis>
</taxonomicName>
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<emphasis id="B9536F19FFADFFBB4B87FD4BFAB0FD18" bold="true" box="[1209,1375,727,752]" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, and
<quantity id="4CDF1EEEFFADFFBB4FA9FD60FF12FCFD" box="[151,253,764,789]" metricMagnitude="0" metricUnit="m" metricValue="1.3716" metricValueMax="1.4224" metricValueMin="1.3208" pageId="29" pageNumber="430" unit="in" value="54.0" valueMax="56.0" valueMin="52.0">5256 in</quantity>
<taxonomicName id="4C27C888FFADFFBB4E39FD61FE9EFCFC" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1837" box="[263,369,765,788]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="rouxii">
<emphasis id="B9536F19FFADFFBB4E39FD61FE9EFCFC" box="[263,369,765,788]" italics="true" pageId="29" pageNumber="430">M. rouxii</emphasis>
</taxonomicName>
) presence of a long, distinct isolated spine in the posterior corner of orbit (vs. absent in
<taxonomicName id="4C27C888FFADFFBB4A43FD61FF37FCDF" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1837" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="rouxii">
<emphasis id="B9536F19FFADFFBB4A43FD61FF37FCDF" italics="true" pageId="29" pageNumber="430">M. rouxii</emphasis>
</taxonomicName>
; very small, indistinct tubercle like in
<taxonomicName id="4C27C888FFADFFBB4DB9FCBDFCF6FCD0" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[647,793,801,824]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="montanus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4DB9FCBDFCF6FCD0" box="[647,793,801,824]" italics="true" pageId="29" pageNumber="430">M. montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4C1EFC83FC23FCDF" box="[800,972,799,824]" pageId="29" pageNumber="430" rank="species">
<emphasis id="B9536F19FFADFFBB4C1EFC83FC29FCD0" bold="true" box="[800,966,799,824]" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, and much longer in
<taxonomicName id="4C27C888FFADFFBB4B87FCBDFA73FCD0" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[1209,1436,800,824]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="acanthocephalus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4B87FCBDFA73FCD0" box="[1209,1436,800,824]" italics="true" pageId="29" pageNumber="430">M. acanthocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4FA9FCD8FEA6FCB4" box="[151,329,836,861]" pageId="29" pageNumber="430" rank="species">
<emphasis id="B9536F19FFADFFBB4FA9FCD8FEAAFCB5" bold="true" box="[151,325,836,861]" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
); 34 long nuchal spines (vs. 36 small nuchal spines in
<taxonomicName id="4C27C888FFADFFBB4CDAFCD9FB96FCB4" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[996,1145,837,860]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="montanus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4CDAFCD9FB96FCB4" box="[996,1145,837,860]" italics="true" pageId="29" pageNumber="430">M. montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4BBCFCD8FAD9FCB4" box="[1154,1334,836,861]" pageId="29" pageNumber="430" rank="species">
<emphasis id="B9536F19FFADFFBB4BBCFCD8FAC0FCB5" bold="true" box="[1154,1327,836,861]" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, 6 much longer nuchal spines in
<taxonomicName id="4C27C888FFADFFBB4E8CFCF5FD76FC68" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[434,665,872,896]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="acanthocephalus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4E8CFCF5FD76FC68" box="[434,665,872,896]" italics="true" pageId="29" pageNumber="430">M. acanthocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4D9AFCFBFCB2FC97" box="[676,861,871,896]" pageId="29" pageNumber="430" rank="species">
<emphasis id="B9536F19FFADFFBB4D9AFCFBFCB7FC68" bold="true" box="[676,856,871,896]" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, 78 smaller nuchal spines in
<taxonomicName id="4C27C888FFADFFBB4BF2FCF4FADDFC97" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1837" box="[1228,1330,872,895]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="rouxii">
<emphasis id="B9536F19FFADFFBB4BF2FCF4FADDFC97" box="[1228,1330,872,895]" italics="true" pageId="29" pageNumber="430">C. rouxii</emphasis>
</taxonomicName>
); longer, prominent isolated spine on the back of head and above tympanum (vs. much smaller in
<taxonomicName id="4C27C888FFADFFBB4BB6FC11FAF4FC4C" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[1160,1307,909,932]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="montanus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4BB6FC11FAF4FC4C" box="[1160,1307,909,932]" italics="true" pageId="29" pageNumber="430">M. montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4A1DFC10FF29FC2F" pageId="29" pageNumber="430" rank="species">
<emphasis id="B9536F19FFADFFBB4A1DFC10FF2EFC20" bold="true" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, and
<taxonomicName id="4C27C888FFADFFBB4E3AFC2DFE84FC2F" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1837" box="[260,363,944,968]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="rouxii">
<emphasis id="B9536F19FFADFFBB4E3AFC2DFE84FC2F" box="[260,363,944,968]" italics="true" pageId="29" pageNumber="430">M. rouxii</emphasis>
</taxonomicName>
) and presence of a white spot below the eye (vs. absent in
<taxonomicName id="4C27C888FFADFFBB4CC5FC2DFB8DFC2F" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1837" box="[1019,1122,944,968]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="rouxii">
<emphasis id="B9536F19FFADFFBB4CC5FC2DFB8DFC2F" box="[1019,1122,944,968]" italics="true" pageId="29" pageNumber="430">M. rouxii</emphasis>
</taxonomicName>
; in the form of a band in
<taxonomicName id="4C27C888FFADFFBB4A43FC2DFEEBFC04" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="montanus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4A43FC2DFEEBFC04" italics="true" pageId="29" pageNumber="430">M. montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4E34FC48FE5AFC04" box="[266,437,980,1005]" pageId="29" pageNumber="430" rank="species">
<emphasis id="B9536F19FFADFFBB4E34FC48FE5FFC05" bold="true" box="[266,432,980,1005]" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
) and smaller body size: adult SVL
<quantity id="4CDF1EEEFFADFFBB4C7CFC48FC09FC04" box="[834,998,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="6.659999999999999" metricValueMax="7.38" metricValueMin="5.9399999999999995" pageId="29" pageNumber="430" unit="mm" value="66.6" valueMax="73.8" valueMin="59.4">59.473.8 mm</quantity>
, n=9 (vs.
<taxonomicName id="4C27C888FFADFFBB4B6AFC48FB0EFC04" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[1108,1249,980,1004]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="montanus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4B6AFC48FB0EFC04" box="[1108,1249,980,1004]" italics="true" pageId="29" pageNumber="430">C. montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4BD6FC48FA7AFC04" box="[1256,1429,980,1005]" pageId="29" pageNumber="430" rank="species">
<emphasis id="B9536F19FFADFFBB4BD6FC48FA61FC05" bold="true" box="[1256,1422,980,1005]" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
, adult SVL
<quantity id="4CDF1EEEFFADFFBB4E2CFC6BFE4DFBE7" box="[274,418,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="7.22" metricValueMax="8.34" metricValueMin="6.1" pageId="29" pageNumber="430" unit="mm" value="72.2" valueMax="83.4" valueMin="61.0">6183.4 mm</quantity>
, n=8; and
<taxonomicName id="4C27C888FFADFFBB4D29FC65FD15FBF8" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[535,762,1016,1040]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="430" phylum="Chordata" rank="species" species="acanthocephalus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFADFFBB4D29FC65FD15FBF8" box="[535,762,1016,1040]" italics="true" pageId="29" pageNumber="430">M. acanthocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFADFFBB4C3FFC6BFC40FBE7" box="[769,943,1015,1040]" pageId="29" pageNumber="430" rank="species">
<emphasis id="B9536F19FFADFFBB4C3FFC6BFC47FBF8" bold="true" box="[769,936,1015,1040]" pageId="29" pageNumber="430">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
adult SVL
<quantity id="4CDF1EEEFFADFFBB4B0FFC6BFB39FBE7" box="[1073,1238,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="7.075" metricValueMax="7.26" metricValueMin="6.89" pageId="29" pageNumber="430" unit="mm" value="70.75" valueMax="72.6" valueMin="68.9">68.972.6 mm</quantity>
, n=3).
</paragraph>
</subSubSection>
<subSubSection id="C33DE080FFADFFB84FF9FB80FEDBFD40" lastPageId="30" lastPageNumber="431" pageId="29" pageNumber="430" type="description">
<paragraph id="8B98B30BFFADFFBB4FF9FB80FDCDFA28" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
<emphasis id="B9536F19FFADFFBB4FF9FB80FEB6FBDD" bold="true" box="[199,345,1052,1077]" pageId="29" pageNumber="430">Description.</emphasis>
Based on CESL 0 45. A medium sized adult male (SVL-
<quantity id="4CDF1EEEFFADFFBB4CD8FB80FBBEFBDC" box="[998,1105,1052,1077]" metricMagnitude="-2" metricUnit="m" metricValue="6.37" pageId="29" pageNumber="430" unit="mm" value="63.7">63.7 mm</quantity>
), Morphometric and meristic data are summarised in Appendix 2 &amp; 3. General habitus moderately compressed. Head moderately large (HL/SVL ratio 0.29), elongate (HW/HL ratio 0.63), maximum height less than maximum width; snout pointed; rostral broader than high; nostrils in single nasal shield, which is separated from rostral by a single scale; mental shield narrower than rostral; two postmentals, first pair in contact with each other; genials keeled; gular scales strongly keeled, slightly smaller than genials; scales on top of snout smooth except median row, which is keeled; scales on top of head heterogenous in size and shape, keeled; supraorbital scales keeled; canthus-rostralis and supraciliary edge sharp; a spine at the posterior corner of the orbit; two separated long spines on posterior end of head, the anterior slightly longer, midway between nuchal crest and tympanum, posterior above tympanum; orbit diameter 70% of distance between anterior border of orbit and snout tip; tympanum exposed, its greatest diameter 54% horizontal diameter of orbit; enlarged keeled scale between tympanum and orbit; posterior region of jaws swollen; supralabials 10/10; infralabials 9/9.
</paragraph>
<paragraph id="8B98B30BFFADFFBB4FF9FA50FDF7F955" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">Nuchal crest well developed, composed of four primary, long conical spines, the first being the smallest and the third longest; the remaining vertebral scales slightly enlarged relative to adjacent rows and possess a more pronounced median keel forming a serrated ridge like the dorsal crest which continues till the tail base; 54 longitudinal scale rows around midbody; scales on dorsum keeled, oriented postero-dorsally, while lateral ones oriented postero-ventrally; lateral scales smaller than dorsal, keeled; ventrals strongly keeled, irregular, slightly smaller than dorsals but of similar size as laterals, genials and gular scales; a strong, oblique antehumeral fold, nearly extending across the throat.</paragraph>
<paragraph id="8B98B30BFFADFFBB4FF9F95BFADCF898" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
Limbs slender and covered with strongly keeled scales, larger than laterals, forming parallel longitudinal rows; scales under thighs weakly keeled; length of hindlimb ca. 87% SVL; relative length of fingers 4&gt;3&gt;2&gt;5&gt;1; relative lengths of toes 4&gt;3&gt;5&gt;2&gt;1; fourth toe longer than fifth finger; 23 subdigital lamellae under fourth finger; 27 subdigital lamellae under fourth toe; subdigital lamellae with sharp keels, bicarinate; tail slender, swollen at the base; scales on dorsal and ventral surface of tail with sharp keels, larger than laterals; tail length
<quantity id="4CDF1EEEFFADFFBB4BF1F8C4FAC0F898" box="[1231,1327,1880,1904]" metricMagnitude="-1" metricUnit="m" metricValue="1.72" pageId="29" pageNumber="430" unit="mm" value="172.0">172 mm</quantity>
.
</paragraph>
<paragraph id="8B98B30BFFADFFB84FF9F8E0FAA0FEA8" blockId="29.[151,1437,151,2013]" lastBlockId="30.[151,1437,151,1149]" lastPageId="30" lastPageNumber="431" pageId="29" pageNumber="430">
<emphasis id="B9536F19FFADFFBB4FF9F8E0FEB0F87D" bold="true" box="[199,351,1916,1941]" pageId="29" pageNumber="430">Colouration.</emphasis>
In life: dorsum and head blackish-brown with irregular lighter patches on the back; a light brown band from above the shoulder till the dorsal crest forming a v shape; head laterally dark blackish with a white spot below the posterior end of eye; stripe from above nostril to anterior margin of orbit extending till the tympanum from the posterior margin of orbit in the form of a black band; tympanum pale grey, lip scales white; a blackish triangular patch behind the tympanum continuing to the antehumeral fold; ventral uniformly lighter, pale grey; gular pouch with a reddish stripe extending beyond the throat; tail with alternating dark and light blotches forming irregular bands towards the end. Representative image showing live colouration (
<figureCitation id="131CAF8EFFAEFFB84B51FE98FB25FEF5" box="[1135,1226,260,285]" captionStart="FIGURE 6" captionStartId="15.[151,250,1511,1533]" captionTargetBox="[340,1247,193,1476]" captionTargetId="figure@15.[340,1247,193,1476]" captionTargetPageId="15" captionText="FIGURE 6. Lateral photograph showing live coloration of A. adult male Monilesaurus rouxii comb. nov. and B. adult male Monilesaurus ellioti comb. nov." httpUri="https://zenodo.org/record/1440686/files/figure.png" pageId="30" pageNumber="431">Fig. 6b</figureCitation>
). In preservative: colouration pattern mostly similar to that in life, except overall paler; bands on the head dull greyish brown.
</paragraph>
<paragraph id="8B98B30BFFAEFFB84FF9FED0FF24FDF0" blockId="30.[151,1437,151,1149]" pageId="30" pageNumber="431">
<emphasis id="B9536F19FFAEFFB84FF9FED0FD15FE8D" bold="true" box="[199,762,332,357]" pageId="30" pageNumber="431">Variation and secondary sexual characteristics.</emphasis>
The other specimens examined agree with the CESL 0
<quantity id="4CDF1EEEFFAEFFB84A5DFED0FA73FE8D" box="[1379,1436,332,357]" metricMagnitude="0" metricUnit="m" metricValue="1.143" pageId="30" pageNumber="431" unit="in" value="45.0">45 in</quantity>
general morphology and scalation except for some differences that are summarised in Appendix 2 &amp; 3. All the examined female specimens (CESL 0 47, CESL 0 58 and CESL 061) have much smaller nuchal spines compared to the males, reduced postero-orbital spine; lack a dorsal crest and gular sac; overall colouration olive-brown, lighter head and vertebral region, a dark band along the side of the head to the neck and presence of a black antehumeral fold.
</paragraph>
<paragraph id="8B98B30BFFAEFFB84FF9FDB8FEDBFD40" blockId="30.[151,1437,151,1149]" pageId="30" pageNumber="431">
<emphasis id="B9536F19FFAEFFB84FF9FDB8FE7BFDD5" bold="true" box="[199,404,548,573]" pageId="30" pageNumber="431">Genetic distance.</emphasis>
<taxonomicName id="4C27C888FFAEFFB84EA2FDB9FDECFDD4" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" baseAuthorityYear="2018" box="[412,515,548,572]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="ellioti" status="comb. nov.">
<emphasis id="B9536F19FFAEFFB84EA2FDB9FDECFDD4" box="[412,515,548,572]" italics="true" pageId="30" pageNumber="431">M. ellioti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFAEFFB84D32FDB8FD62FDD4" box="[524,653,548,573]" pageId="30" pageNumber="431">
<emphasis id="B9536F19FFAEFFB84D32FDB8FD69FDD5" bold="true" box="[524,646,548,573]" pageId="30" pageNumber="431">comb. nov</emphasis>
.
</taxonomicNameLabel>
shows 12% intraspecific genetic divergence in the 16S gene; 47% interspecific genetic divergence from
<taxonomicName id="4C27C888FFAEFFB84D66FDD5FD29FDB7" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1837" box="[600,710,584,608]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="rouxii" status="comb. nov.">
<emphasis id="B9536F19FFAEFFB84D66FDD5FD29FDB7" box="[600,710,584,608]" italics="true" pageId="30" pageNumber="431">M. rouxii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFAEFFB84DEDFDDBFCB6FDB7" box="[723,857,583,608]" pageId="30" pageNumber="431">
<emphasis id="B9536F19FFAEFFB84DEDFDDBFCBDFD88" bold="true" box="[723,850,583,608]" pageId="30" pageNumber="431">comb. nov</emphasis>
.
</taxonomicNameLabel>
; 57% interspecific genetic divergence from
<taxonomicName id="4C27C888FFAEFFB84A43FDD5FEBAFD6C" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="acanthocephalus" status="gen. et sp. nov">
<emphasis id="B9536F19FFAEFFB84A43FDD5FEBAFD6C" italics="true" pageId="30" pageNumber="431">M. acanthocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFAEFFB84E60FDF0FDE0FD6D" box="[350,527,620,645]" pageId="30" pageNumber="431" rank="species">
<emphasis id="B9536F19FFAEFFB84E60FDF0FDE0FD6D" bold="true" box="[350,527,620,645]" pageId="30" pageNumber="431">gen. et sp. nov</emphasis>
</taxonomicNameLabel>
and 48% interspecific genetic divergence from
<taxonomicName id="4C27C888FFAEFFB84B76FDF1FB32FD6C" authority="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak, 2018" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" box="[1096,1245,621,644]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="montanus" status="gen. et sp. nov.">
<emphasis id="B9536F19FFAEFFB84B76FDF1FB32FD6C" box="[1096,1245,621,644]" italics="true" pageId="30" pageNumber="431">M. montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFAEFFB84BD8FDF0FA73FD6C" box="[1254,1436,620,645]" pageId="30" pageNumber="431" rank="species">
<emphasis id="B9536F19FFAEFFB84BD8FDF0FA7AFD6D" bold="true" box="[1254,1429,620,645]" pageId="30" pageNumber="431">gen. et sp. nov</emphasis>
.
</taxonomicNameLabel>
(Appendix 5).
</paragraph>
</subSubSection>
<subSubSection id="C33DE080FFAEFFB84FF9FD28FD04FCB5" pageId="30" pageNumber="431" type="distribution">
<paragraph id="8B98B30BFFAEFFB84FF9FD28FD04FCB5" blockId="30.[151,1437,151,1149]" pageId="30" pageNumber="431">
<emphasis id="B9536F19FFAEFFB84FF9FD28FE8FFD25" bold="true" box="[199,352,692,717]" pageId="30" pageNumber="431">Distribution.</emphasis>
<taxonomicName id="4C27C888FFAEFFB84E57FD28FDA2FD24" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[361,589,692,716]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="ellioti" status="comb. nov.">
<emphasis id="B9536F19FFAEFFB84E57FD28FDA2FD24" box="[361,589,692,716]" italics="true" pageId="30" pageNumber="431">Monilesaurus ellioti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFAEFFB84D68FD28FD37FD24" box="[598,728,692,717]" pageId="30" pageNumber="431">
<emphasis id="B9536F19FFAEFFB84D68FD28FD3EFD25" bold="true" box="[598,721,692,717]" pageId="30" pageNumber="431">comb. nov</emphasis>
.
</taxonomicNameLabel>
is endemic to the
<collectingRegion id="49E37DE9FFAEFFB84C91FD29FBE5FD24" box="[943,1034,693,716]" country="American Samoa" name="Western" pageId="30" pageNumber="431">Western</collectingRegion>
Ghats and is distributed across the low and medium elevation forests (up to
<quantity id="4CDF1EEEFFAEFFB84D5EFD44FD5BFD07" box="[608,692,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="431" unit="m" value="1000.0">1000 m</quantity>
asl) of southern parts of central
<collectingRegion id="49E37DE9FFAEFFB84B23FD44FB97FD07" box="[1053,1144,728,751]" country="American Samoa" name="Western" pageId="30" pageNumber="431">Western</collectingRegion>
Ghats (Coorg plateau and south) and the southern
<collectingRegion id="49E37DE9FFAEFFB84E99FD61FDEDFCFC" box="[423,514,765,788]" country="American Samoa" name="Western" pageId="30" pageNumber="431">Western</collectingRegion>
Ghats. This is one of the most common evergreen forest dwelling agamid lizards in this region. During this study,
<taxonomicName id="4C27C888FFAEFFB84D24FCBDFD6BFCD0" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[538,644,800,824]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="ellioti" status="comb. nov.">
<emphasis id="B9536F19FFAEFFB84D24FCBDFD6BFCD0" box="[538,644,800,824]" italics="true" pageId="30" pageNumber="431">M. ellioti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFAEFFB84DB0FC83FCECFCD0" box="[654,771,799,824]" pageId="30" pageNumber="431">comb.nov.</taxonomicNameLabel>
was recorded in various sites in the central and southern
<collectingRegion id="49E37DE9FFAEFFB84FA9FCD9FF1DFCB4" box="[151,242,837,860]" country="American Samoa" name="Western" pageId="30" pageNumber="431">Western</collectingRegion>
Ghats (See
<figureCitation id="131CAF8EFFAEFFB84E44FCD8FE54FCB4" box="[378,443,836,861]" captionStart="FIGURE 3" captionStartId="12.[151,250,1924,1946]" captionTargetBox="[434,1141,804,1819]" captionTargetId="figure@12.[324,1231,696,1979]" captionTargetPageId="12" captionText="FIGURE 3. Map showing distribution of members of Monilesaurus gen. nov. across the Western Ghats, India. Coloured symbols indicate selected sampling localities." httpUri="https://zenodo.org/record/1440680/files/figure.png" pageId="30" pageNumber="431">Fig. 3</figureCitation>
&amp; Appendix 1 for details).
</paragraph>
</subSubSection>
<subSubSection id="C33DE080FFAEFFB84FF9FCFBFDEEFB95" pageId="30" pageNumber="431" type="biology_ecology">
<paragraph id="8B98B30BFFAEFFB84FF9FCFBFDEEFB95" blockId="30.[151,1437,151,1149]" pageId="30" pageNumber="431">
<emphasis id="B9536F19FFAEFFB84FF9FCFBFDF7FC68" bold="true" box="[199,536,871,896]" pageId="30" pageNumber="431">Ecology and natural history.</emphasis>
<taxonomicName id="4C27C888FFAEFFB84D20FCF4FD10FC68" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[542,767,872,896]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="ellioti">
<emphasis id="B9536F19FFAEFFB84D20FCF4FD10FC68" box="[542,767,872,896]" italics="true" pageId="30" pageNumber="431">Monilesaurus ellioti</emphasis>
</taxonomicName>
is a diurnal lizard, semi-arboreal to arboreal in habit, and so far, has been recorded mostly in semi-evergreen and evergreen forests. Individuals were mostly seen perching on shrubs, branches and actively moving on tree trunks. In some instances, it has also been observed in coffee plantations surrounded by evergreen forests. In some sites like Parambikulam, Vazhachal reserve forest and Brahmagiri hills, it was observed to occur syntopically in the same habitat as
<taxonomicName id="4C27C888FFAEFFB84B17FC65FB7BFBE7" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1837" box="[1065,1172,1016,1040]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="rouxii" status="comb. nov.">
<emphasis id="B9536F19FFAEFFB84B17FC65FB7BFBE7" box="[1065,1172,1016,1040]" italics="true" pageId="30" pageNumber="431">M. rouxii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFAEFFB84BA1FC6BFACEFBE7" box="[1183,1313,1015,1040]" pageId="30" pageNumber="431">
<emphasis id="B9536F19FFAEFFB84BA1FC6BFAF3FBF8" bold="true" box="[1183,1308,1015,1040]" pageId="30" pageNumber="431">comb. nov</emphasis>
.
</taxonomicNameLabel>
, but more abundant within dense forests, while
<taxonomicName id="4C27C888FFAEFFB84D0BFB81FD73FBDC" authorityName="Pal &amp; Vijayakumar &amp; Shanker &amp; Jayarajan &amp; Deepak" authorityYear="2018" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1837" box="[565,668,1053,1076]" class="Reptilia" family="Agamidae" genus="Monilesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="rouxii" status="comb. nov.">
<emphasis id="B9536F19FFAEFFB84D0BFB81FD73FBDC" box="[565,668,1053,1076]" italics="true" pageId="30" pageNumber="431">M. rouxii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260D262FFAEFFB84D9CFB80FCCDFBDC" box="[674,802,1052,1077]" pageId="30" pageNumber="431">
<emphasis id="B9536F19FFAEFFB84D9CFB80FCF4FBDD" bold="true" box="[674,795,1052,1077]" pageId="30" pageNumber="431">comb. nov</emphasis>
.
</taxonomicNameLabel>
is generally restricted to forest edges. In some instances, gravid females were recorded during pre-and mid monsoon (JuneAugust), which hints that monsoon might be a breeding season for this species.
</paragraph>
</subSubSection>
</treatment>
</document>