421 lines
54 KiB
XML
421 lines
54 KiB
XML
<document id="CE266CE9B97721E2BDFE78255F39AC4E" ID-CLB-Dataset="3917" ID-DOI="10.5281/zenodo.1196029" ID-GBIF-Dataset="f34e9863-9f62-4f2a-8819-7e6e8fc09403" ID-ISSN="0035-418" ID-Zenodo-Dep="1196029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634783645596" checkinUser="plazi" docAuthor="Schuchert, Peter" docDate="2018" docId="03FE406DFFE0E249FC32FC9EDD1A1C87" docLanguage="en" docName="8_Schuchert.pdf" docOrigin="Revue suisse de Zoologie 125 (1)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Leuckartiara longicalcar Schuchert 2018, n. spec." docType="treatment" docVersion="6" lastPageNumber="123" masterDocId="FFC73815FFF3E25FFF85FFAED9671F0B" masterDocTitle="DNA barcoding of some Pandeidae species (Cnidaria, Hydrozoa, Anthoathecata)" masterLastPageNumber="127" masterPageNumber="101" pageNumber="120" updateTime="1699463991308" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="C11D886C28FB8E334C65AF1F3118A08E" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="F002DDAAFED4FFB3B5851AB89FF2CA8E">
|
||
<mods:title id="47C01E90DCB4B46C21E25F3C478C10BA">DNA barcoding of some Pandeidae species (Cnidaria, Hydrozoa, Anthoathecata)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="BF5B71EBF4FCF7508D90D9000CDFBF26" type="personal">
|
||
<mods:role id="3F3C00BC760C2825D158D6E6B4BA0C30">
|
||
<mods:roleTerm id="E3A0508CCDC5CC4FE38CA0D73B930D6E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7E96FE26D94075B7381CA5C49730723D">Schuchert, Peter</mods:namePart>
|
||
<mods:affiliation id="1D83A004044CFFD4180C9B4BA14C6F92">Muséum d'histoire naturelle, CP 6434, CH- 1211 Genève 6, Switzerland, E-mail: peter. schuchert @ ville-ge. ch</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="BDDA786EC7D5208C7F5F7B99045E2276">text</mods:typeOfResource>
|
||
<mods:relatedItem id="23E6B0ECE73D99AB5A5EF2C53AB33517" type="host">
|
||
<mods:titleInfo id="E8B9E40922CD713851FAC25053FEA967">
|
||
<mods:title id="B28202D5DA7BCD665A5E229FE1B9D583">Revue suisse de Zoologie</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="405F374B9943B9EC2CC8DC2BF9283FF1">
|
||
<mods:date id="6439435A030C4DB5F3ED027556919D3C">2018</mods:date>
|
||
<mods:detail id="828B3948DC58CC1D62C436C44C51303A" type="pubDate">
|
||
<mods:number id="C643635C9FFA144B220E7567C9A0A501">2018-03-31</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="BD93D52A165B32C26E00108CFCDC2B41" type="volume">
|
||
<mods:number id="5C7E5FA06020D33327949EF28B33A214">125</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="E8892292A2B55C94FCC98B30CBF6A401" type="issue">
|
||
<mods:number id="1A53F8EBC6704C179383E6F1D480AB13">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="04DE59D65B44825ECF7C9D91D21B4181" unit="page">
|
||
<mods:start id="E357A91E90C3087A5E15EA7A1D3B8AB7">101</mods:start>
|
||
<mods:end id="2EC1D2E6DEFC72D29BDB31C6D59E9388">127</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="B5B9907979085BB53FD9B2D7F733B83D">journal article</mods:classification>
|
||
<mods:identifier id="45A8C6EB6459E12F671132CC27F810ED" type="CLB-Dataset">3917</mods:identifier>
|
||
<mods:identifier id="5DA11DD8283A29A23C68602DD484CF77" type="DOI">10.5281/zenodo.1196029</mods:identifier>
|
||
<mods:identifier id="40CC149B073F302E9ACA9DCFB84F4E39" type="GBIF-Dataset">f34e9863-9f62-4f2a-8819-7e6e8fc09403</mods:identifier>
|
||
<mods:identifier id="82986CFC1624B9C13734F04AA9231A0B" type="ISSN">0035-418</mods:identifier>
|
||
<mods:identifier id="E0F8D187B281F5F65B62DA81B4603F81" type="Zenodo-Dep">1196029</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03FE406DFFE0E249FC32FC9EDD1A1C87" ID-GBIF-Taxon="189160419" LSID="urn:lsid:plazi:treatment:03FE406DFFE0E249FC32FC9EDD1A1C87" httpUri="http://treatment.plazi.org/id/03FE406DFFE0E249FC32FC9EDD1A1C87" lastPageId="22" lastPageNumber="123" pageId="19" pageNumber="120">
|
||
<subSubSection id="C34DA2F0FFE0E24CFC32FC9EDC421C43" box="[951,1317,816,840]" pageId="19" pageNumber="120" type="nomenclature">
|
||
<paragraph id="8BE8F17BFFE0E24CFC32FC9EDC421C43" blockId="19.[951,1317,816,872]" box="[951,1317,816,840]" pageId="19" pageNumber="120">
|
||
<heading id="D0A04617FFE0E24CFC32FC9EDC421C43" bold="true" box="[951,1317,816,840]" centered="true" level="2" pageId="19" pageNumber="120" reason="2">
|
||
<emphasis id="B9232D69FFE0E24CFC32FC9EDC421C43" bold="true" box="[951,1317,816,840]" pageId="19" pageNumber="120">
|
||
<taxonomicName id="4C578AF8FFE0E24CFC32FC9EDDAD1C43" ID-CoL="6PRRK" authority="Schuchert, 2018" authorityName="Schuchert" authorityYear="2018" box="[951,1226,816,840]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="19" pageNumber="120" phylum="Cnidaria" rank="species" species="longicalcar" status="sp. nov.">
|
||
<emphasis id="B9232D69FFE0E24CFC32FC9EDDAD1C43" bold="true" box="[951,1226,816,840]" italics="true" pageId="19" pageNumber="120">Leuckartiara longicalcar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2109012FFE0E24CFB54FC9FDC421C43" box="[1233,1317,817,840]" pageId="19" pageNumber="120" rank="species">n. spec.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DA2F0FFE0E24CFB82FCFFDDB21C63" box="[1031,1237,849,872]" pageId="19" pageNumber="120" type="description">
|
||
<paragraph id="8BE8F17BFFE0E24CFB82FCFFDDB21C63" blockId="19.[951,1317,816,872]" box="[1031,1237,849,872]" pageId="19" pageNumber="120">
|
||
<figureCitation id="136CEDFEFFE0E24CFB82FCFFDD3E1C63" box="[1031,1113,849,872]" captionStart="Fig" captionStartId="19.[834,869,2027,2048]" captionTargetBox="[833,1438,1150,1986]" captionTargetId="figure-415@19.[833,1438,1150,1986]" captionTargetPageId="19" captionText="Fig. 14. Leuckartiara longicalcar, n. spec., living animal (paratype), total height about 15 mm." figureDoi="http://doi.org/10.5281/zenodo.5590770" httpUri="https://zenodo.org/record/5590770/files/figure.png" pageId="19" pageNumber="120">Figs 14</figureCitation>
|
||
,
|
||
<figureCitation id="136CEDFEFFE0E24CFBE1FCFFDD1A1C63" box="[1124,1149,849,872]" captionStart="Fig" captionStartId="20.[151,186,2025,2046]" captionTargetBox="[151,1436,771,1975]" captionTargetId="figure-187@20.[151,1436,771,1975]" captionTargetPageId="20" captionText="Fig. 15. Leuckartiara longicalcar, n. spec., living animal. (A) Details of manubrium with gonad folds. (B) Lateral view of a perradial tentacle with the characteristic, long, abaxial spur. (C) Adradial tentacles and rudimentary bulbs, note red ocelli." figureDoi="http://doi.org/10.5281/zenodo.5590772" httpUri="https://zenodo.org/record/5590772/files/figure.png" pageId="19" pageNumber="120">15</figureCitation>
|
||
A-C, 16
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DA2F0FFE0E24CFCBAFCD1DAA81CE1" pageId="19" pageNumber="120" type="reference_group">
|
||
<paragraph id="8BE8F17BFFE0E24CFCBAFCD1DDF71CBB" blockId="19.[831,1436,895,1002]" pageId="19" pageNumber="120">
|
||
in part
|
||
<emphasis id="B9232D69FFE0E24CFC04FCD1DD3C1C98" box="[897,1115,895,916]" italics="true" pageId="19" pageNumber="120">
|
||
<taxonomicName id="4C578AF8FFE0E24CFC04FCD1DD201C9F" ID-CoL="723RK" box="[897,1095,895,916]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="19" pageNumber="120" phylum="Cnidaria" rank="species" species="octona">Leuckartiara octona</taxonomicName>
|
||
. ‒
|
||
</emphasis>
|
||
<bibRefCitation id="EFC68C8AFFE0E24CFBE5FCD1DC091C9F" author="Mackie G. O. & Mackie G. V." box="[1120,1390,895,916]" pageId="19" pageNumber="120" pagination="63 - 84" refId="ref17349" refString="Mackie G. O., Mackie G. V. 1963. Systematic and biological notes on living hydromedusae from Puget Sound. Bulletin (National Museum of Canada). Biological series 199: 63 - 84." type="book chapter" year="1963">Mackie & Mackie, 1963: 68</bibRefCitation>
|
||
. [not
|
||
<taxonomicName id="4C578AF8FFE0E24CFC0EFC35DDED1CBB" ID-CoL="723RK" authority="(Fleming, 1823)" baseAuthorityName="Fleming" baseAuthorityYear="1823" box="[907,1162,923,944]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="19" pageNumber="120" phylum="Cnidaria" rank="species" species="octona">
|
||
<emphasis id="B9232D69FFE0E24CFC0EFC35DA821CBB" box="[907,997,923,944]" italics="true" pageId="19" pageNumber="120">L. octona</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC68C8AFFE0E24CFC77FC35DDE41CBB" author="Fleming J." box="[1010,1155,923,944]" pageId="19" pageNumber="120" pagination="294 - 303" refId="ref16195" refString="Fleming J. 1823. Gleamings of natural history, gathered on the coast of Scotland during a voyage in 1821. Edinburgh Philosophical Journal 8: 294 - 303." type="journal article" year="1823">Fleming, 1823</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
]
|
||
</paragraph>
|
||
<paragraph id="8BE8F17BFFE0E24CFCBAFC16DAA81CE1" blockId="19.[831,1436,895,1002]" pageId="19" pageNumber="120">
|
||
<treatmentCitationGroup id="AB47D655FFE0E24CFCBAFC16DAA81CE1" pageId="19" pageNumber="120">
|
||
<taxonomicName id="4C578AF8FFE0E24CFCBAFC16DD741CC6" box="[831,1043,952,973]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="19" pageNumber="120" phylum="Cnidaria" rank="species" species="undetermined">
|
||
<emphasis id="B9232D69FFE0E24CFCBAFC16DADB1CC6" box="[831,956,952,973]" italics="true" pageId="19" pageNumber="120">Leuckartiara</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
‒
|
||
<treatmentCitation id="0AF6D76AFFE0E24CFBB4FC16DCFF1CC6" author="Arai M. N. & Brinckmann-Voss A." box="[1073,1432,952,973]" page="56" pageId="19" pageNumber="120" year="1980">
|
||
<bibRefCitation id="EFC68C8AFFE0E24CFBB4FC16DCFF1CC6" author="Arai M. N. & Brinckmann-Voss A." box="[1073,1432,952,973]" pageId="19" pageNumber="120" pagination="1 - 192" refId="ref15698" refString="Arai M. N., Brinckmann-Voss A. 1980. Hydromedusae of British Columbia and Puget Sound. Canadian Bulletin of Fisheries and Aquatic Sciences 204: 1 - 192." type="journal article" year="1980">Arai & Brinckmann-Voss, 1980: 56</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 30.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DA2F0FFE0E24BFCBAFBACDD891E32" lastPageId="20" lastPageNumber="121" pageId="19" pageNumber="120" type="materials_examined">
|
||
<paragraph id="8BE8F17BFFE0E24BFCBAFBACDB191E31" blockId="19.[831,1436,1026,1081]" lastBlockId="20.[151,756,258,314]" lastPageId="20" lastPageNumber="121" pageId="19" pageNumber="120">
|
||
<materialsCitation id="3B3FFB26FFE0E24CFCBAFBACDCFB1B32" ID-GBIF-Occurrence="3391165311" country="United States of America" location="San Juan Island" municipality="Friday Harbor" pageId="19" pageNumber="120" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B9232D69FFE0E24CFCBAFBACDAD51B12" bold="true" box="[831,946,1026,1049]" pageId="19" pageNumber="120">
|
||
<typeStatus id="54EC4FD9FFE0E24CFCBAFBACDACB1B12" box="[831,940,1026,1049]" pageId="19" pageNumber="120" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
MHNG-INVE-98638; female;
|
||
<collectingCountry id="F340B1EBFFE0E24CFAA3FBACDC381B12" box="[1318,1375,1026,1049]" name="United States of America" pageId="19" pageNumber="120">USA</collectingCountry>
|
||
,
|
||
<location id="8E88A7A0FFE0E24CFAF1FBACDAA21B32" LSID="urn:lsid:plazi:treatment:03FE406DFFE0E249FC32FC9EDD1A1C87:8E88A7A0FFE0E24CFAF1FBACDAA21B32" country="United States of America" municipality="Friday Harbor" name="San Juan Island" pageId="19" pageNumber="120">San Juan Island</location>
|
||
,
|
||
<collectingMunicipality id="6B8C6B01FFE0E24CFC5DFB8CDD1B1B32" box="[984,1148,1058,1081]" pageId="19" pageNumber="120">Friday Harbor</collectingMunicipality>
|
||
, 48.54514° -123.01206°,
|
||
</materialsCitation>
|
||
depth
|
||
<quantity id="4CAF5C9EFFE7E24BFF5BFEACD8441E12" box="[222,291,258,281]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="121" unit="m" value="0.5">0.5 m</quantity>
|
||
; collection date
|
||
<date id="FFE9D7BBFFE7E24BFE65FEACDB3C1E12" box="[480,603,258,281]" pageId="20" pageNumber="121" value="2011-05-20">20.05.2011</date>
|
||
; preserved in formalin, subsequently transferred to ethanol.
|
||
</paragraph>
|
||
<caption id="DF28A1F3FFE0E24CFCC7F845DC601717" ID-DOI="http://doi.org/10.5281/zenodo.5590770" ID-Zenodo-Dep="5590770" httpUri="https://zenodo.org/record/5590770/files/figure.png" pageId="19" pageNumber="120" startId="19.[834,869,2027,2048]" targetBox="[833,1438,1150,1986]" targetPageId="19">
|
||
<paragraph id="8BE8F17BFFE0E24CFCC7F845DC601717" blockId="19.[834,1438,2027,2076]" pageId="19" pageNumber="120">
|
||
Fig. 14.
|
||
<taxonomicName id="4C578AF8FFE0E24CFC25F845DDFC170B" authority="Schuchert, 2018" authorityName="Schuchert" authorityYear="2018" box="[928,1179,2027,2048]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="19" pageNumber="120" phylum="Cnidaria" rank="species" species="longicalcar" status="sp. nov.">
|
||
<emphasis id="B9232D69FFE0E24CFC25F845DDFC170B" box="[928,1179,2027,2048]" italics="true" pageId="19" pageNumber="120">Leuckartiara longicalcar</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2109012FFE0E24CFB35F845DC63170B" box="[1200,1284,2027,2048]" pageId="19" pageNumber="120" rank="species">n. spec.</taxonomicNameLabel>
|
||
, living animal (paratype), total height about 15 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE8F17BFFE0E24CFD65F7A8DB831716" blockId="19.[151,756,2021,2077]" box="[736,740,2054,2077]" pageId="19" pageNumber="120">.</paragraph>
|
||
<paragraph id="8BE8F17BFFE7E24BFF12FEFDD8E41D7F" blockId="20.[151,756,339,628]" pageId="20" pageNumber="121">
|
||
<materialsCitation id="3B3FFB26FFE7E24BFF12FEFDD8781EA6" ID-GBIF-Occurrence="3391165301" country="United States of America" elevation="5" location="Friday Harbor" municipality="San Juan Island" pageId="20" pageNumber="121" specimenCount="9" typeStatus="paratype">
|
||
<emphasis id="B9232D69FFE7E24BFF12FEFDD8741E61" bold="true" box="[151,275,339,362]" pageId="20" pageNumber="121">
|
||
<typeStatus id="54EC4FD9FFE7E24BFF12FEFDD8691E61" box="[151,270,339,362]" pageId="20" pageNumber="121" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
MHNG-INVE-78922,
|
||
<specimenCount id="9D513AF2FFE7E24BFD9FFEFDDBC01E61" box="[538,679,339,362]" pageId="20" pageNumber="121" type="generic">9 specimens</specimenCount>
|
||
;
|
||
<collectingCountry id="F340B1EBFFE7E24BFD32FEFDDB881E61" box="[695,751,339,362]" name="United States of America" pageId="20" pageNumber="121">USA</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6B8C6B01FFE7E24BFF12FEDBD8251E87" box="[151,322,373,396]" pageId="20" pageNumber="121">San Juan Island</collectingMunicipality>
|
||
,
|
||
<location id="8E88A7A0FFE7E24BFEC8FEDBD8801E87" LSID="urn:lsid:plazi:treatment:03FE406DFFE0E249FC32FC9EDD1A1C87:8E88A7A0FFE7E24BFEC8FEDBD8801E87" box="[333,487,373,396]" country="United States of America" municipality="San Juan Island" name="Friday Harbor" pageId="20" pageNumber="121">Friday Harbor</location>
|
||
, 48.54514° -123.01206°, depth
|
||
<quantity id="4CAF5C9EFFE7E24BFF59FE38D8781EA6" box="[220,287,406,429]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="121" unit="m" value="0.5">
|
||
<elevation id="007A1648FFE7E24BFF59FE38D8781EA6" box="[220,287,406,429]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="121" unit="m" value="0.5">0.5 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B3FFB26FFE7E24BFEA8FE38D8281D58" ID-GBIF-Occurrence="3391165307" collectingDate="2011-05-20" collectorName="Moria Galbraith" country="Canada" elevation="25" location="Salish Sea" municipality="GenBank" pageId="20" pageNumber="121" specimenCount="3" stateProvince="British Columbia" typeStatus="paratype">
|
||
collection date
|
||
<date id="FFE9D7BBFFE7E24BFE52FE38DB351EA6" box="[471,594,406,429]" pageId="20" pageNumber="121" value="2011-05-20">
|
||
<collectingDate id="EFAD2E53FFE7E24BFE52FE38DB351EA6" box="[471,594,406,429]" pageId="20" pageNumber="121" value="2011-05-20">20.05.2011</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCount id="9D513AF2FFE7E24BFDDAFE38DB931EA6" box="[607,756,406,429]" pageId="20" pageNumber="121" type="generic">one specimen</specimenCount>
|
||
used to isolate DNA 869; for
|
||
<collectingMunicipality id="6B8C6B01FFE7E24BFE7AFE19DB021EC5" box="[511,613,439,462]" pageId="20" pageNumber="121">GenBank</collectingMunicipality>
|
||
numbers of sequences see
|
||
<tableCitation id="C6D5C4C0FFE7E24BFEC8FE76D8CC1EE4" box="[333,427,472,495]" captionStart="Table 1" captionStartId="3.[144,197,540,561]" captionTargetBox="[154,1936,636,1788]" captionTargetPageId="3" captionText="Table 1. Sample data, voucher numbers, and GenBank data of specimens examined by the author. The sample data of other sequences retrieved from GenBank and shown in Figs 8-9 can be obtained by searching the accession numbers in GenBank. nd = no data." httpUri="http://table.plazi.org/id/DF28A1F3FFF0E25CFF15FDB2DAC41D45" pageId="20" pageNumber="121" tableUuid="DF28A1F3FFF0E25CFF15FDB2DAC41D45">Table 1</tableCitation>
|
||
. ‒ MHNG-INVE-82312,
|
||
<specimenCount id="9D513AF2FFE7E24BFD62FE76D86B1D1A" pageId="20" pageNumber="121" type="generic">2 specimens</specimenCount>
|
||
;
|
||
<collectingCountry id="F340B1EBFFE7E24BFEA6FE54D8101D1A" box="[291,375,506,529]" name="Canada" pageId="20" pageNumber="121">Canada</collectingCountry>
|
||
,
|
||
<collectingRegion id="49933F99FFE7E24BFE0BFE54DB3E1D1A" box="[398,601,506,529]" country="Canada" name="British Columbia" pageId="20" pageNumber="121">British Columbia</collectingRegion>
|
||
,
|
||
<location id="8E88A7A0FFE7E24BFDF5FE54DB881D1A" LSID="urn:lsid:plazi:treatment:03FE406DFFE0E249FC32FC9EDD1A1C87:8E88A7A0FFE7E24BFDF5FE54DB881D1A" box="[624,751,506,529]" country="Canada" municipality="GenBank" name="Salish Sea" pageId="20" pageNumber="121" stateProvince="British Columbia">Salish Sea</location>
|
||
, 49.2505° -123.74867°, depth
|
||
<quantity id="4CAF5C9EFFE7E24BFE72FDB5DB361D39" box="[503,593,539,562]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="20" pageNumber="121" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">
|
||
<elevation id="007A1648FFE7E24BFE72FDB5DB361D39" box="[503,593,539,562]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="20" pageNumber="121" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">0-50 m</elevation>
|
||
</quantity>
|
||
; collected by
|
||
<collectorName id="26A294ADFFE7E24BFF12FD92D8281D58" box="[151,335,572,595]" pageId="20" pageNumber="121">Moria Galbraith</collectorName>
|
||
</materialsCitation>
|
||
; preserved in formalin, subsequently transferred to ethanol.
|
||
</paragraph>
|
||
<paragraph id="8BE8F17BFFE7E24BFF12FD20DD891E32" blockId="20.[151,756,654,710]" lastBlockId="20.[831,1436,258,313]" pageId="20" pageNumber="121">
|
||
<emphasis id="B9232D69FFE7E24BFF12FD20D8001DAE" bold="true" box="[151,359,654,677]" pageId="20" pageNumber="121">Additional data:</emphasis>
|
||
Several photographs of living medusae taken by Kevin Lee off the coast of Palos Verdes,
|
||
<collectingRegion id="49933F99FFE7E24BFC1FFEACDD6D1E12" box="[922,1034,258,281]" country="United States of America" name="California" pageId="20" pageNumber="121">California</collectingRegion>
|
||
,
|
||
<collectingCountry id="F340B1EBFFE7E24BFB9CFEACDD361E12" box="[1049,1105,258,281]" name="United States of America" pageId="20" pageNumber="121">USA</collectingCountry>
|
||
, 33.8211° -118.4569°, one of the photos is reproduced here in
|
||
<figureCitation id="136CEDFEFFE7E24BFB19FE8CDD8D1E32" box="[1180,1258,290,313]" captionStart="Fig" captionStartId="21.[151,186,1154,1175]" captionTargetBox="[151,1436,257,1115]" captionTargetId="figure-378@21.[151,1439,257,1116]" captionTargetPageId="21" captionText="Fig. 16. Leuckartiara longicalcar, n. spec., living animal, height 13-18 mm, photographed by Kevin Lee off Los Angeles, California, USA. Note, this photograph is copyright protected and permission to use it here has been obtained by paying a royalty fee to the author and copyright holder Kevin Lee (www.diverkevin.com)." figureDoi="http://doi.org/10.5281/zenodo.5590776" httpUri="https://zenodo.org/record/5590776/files/figure.png" pageId="20" pageNumber="121">Fig. 16</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DA2F0FFE7E24BFCBAFEFFDAE61EA3" pageId="20" pageNumber="121" type="etymology">
|
||
<paragraph id="8BE8F17BFFE7E24BFCBAFEFFDAE61EA3" blockId="20.[831,1437,337,424]" pageId="20" pageNumber="121">
|
||
<emphasis id="B9232D69FFE7E24BFCBAFEFFDAA31E63" bold="true" box="[831,964,337,360]" pageId="20" pageNumber="121">Etymology:</emphasis>
|
||
From the Latin
|
||
<emphasis id="B9232D69FFE7E24BFB01FEFFDDAB1E63" box="[1156,1228,337,360]" italics="true" pageId="20" pageNumber="121">longus</emphasis>
|
||
, long, and
|
||
<emphasis id="B9232D69FFE7E24BFAD4FEFFDCF11E63" box="[1361,1430,337,360]" italics="true" pageId="20" pageNumber="121">calcar</emphasis>
|
||
, spur, referring to the long abaxial spurs of the tentacle bulbs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DA2F0FFE7E24BFCBAFE6EDD051EFC" pageId="20" pageNumber="121" type="materials_examined">
|
||
<paragraph id="8BE8F17BFFE7E24BFCBAFE6EDD051EFC" blockId="20.[831,1436,448,503]" pageId="20" pageNumber="121">
|
||
<emphasis id="B9232D69FFE7E24BFCBAFE6EDA871EDC" bold="true" box="[831,992,448,471]" pageId="20" pageNumber="121">Type locality:</emphasis>
|
||
<materialsCitation id="3B3FFB26FFE7E24BFC69FE6EDD391EFC" ID-GBIF-Occurrence="3391165304" country="United States of America" latitude="48.54514" location="San Juan Island" longLatPrecision="1" longitude="-123.01206" municipality="Friday Harbor" pageId="20" pageNumber="121" specimenCount="1">
|
||
<collectingCountry id="F340B1EBFFE7E24BFC69FE6EDD431EDC" box="[1004,1060,448,471]" name="United States of America" pageId="20" pageNumber="121">USA</collectingCountry>
|
||
,
|
||
<location id="8E88A7A0FFE7E24BFBB1FE6EDD8F1EDC" LSID="urn:lsid:plazi:treatment:03FE406DFFE0E249FC32FC9EDD1A1C87:8E88A7A0FFE7E24BFBB1FE6EDD8F1EDC" box="[1076,1256,448,471]" country="United States of America" latitude="48.54514" longLatPrecision="1" longitude="-123.01206" municipality="Friday Harbor" name="San Juan Island" pageId="20" pageNumber="121">San Juan Island</location>
|
||
,
|
||
<collectingMunicipality id="6B8C6B01FFE7E24BFB7CFE6EDCFF1EDC" box="[1273,1432,448,471]" pageId="20" pageNumber="121">Friday Harbor</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="EE6397BCFFE7E24BFCBAFE4EDAA61EFC" box="[831,961,480,503]" degrees="48.54514" direction="north" orientation="latitude" pageId="20" pageNumber="121" precision="1" value="48.54514">48.54514°N</geoCoordinate>
|
||
<geoCoordinate id="EE6397BCFFE7E24BFC4DFE4EDD391EFC" box="[968,1118,480,503]" degrees="123.01206" direction="west" orientation="longitude" pageId="20" pageNumber="121" precision="1" value="-123.01206">123.01206°W</geoCoordinate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DA2F0FFE7E24AFCBAFDA1D9841AE7" lastPageId="21" lastPageNumber="122" pageId="20" pageNumber="121" type="diagnosis">
|
||
<paragraph id="8BE8F17BFFE7E24AFCBAFDA1D9841AE7" blockId="20.[831,1443,527,710]" lastBlockId="21.[151,757,1301,1516]" lastPageId="21" lastPageNumber="122" pageId="20" pageNumber="121">
|
||
<emphasis id="B9232D69FFE7E24BFCBAFDA1DADE1D2D" bold="true" box="[831,953,527,550]" pageId="20" pageNumber="121">Diagnosis:</emphasis>
|
||
<taxonomicName id="4C578AF8FFE7E24BFC4BFDA1DD3B1D2D" authorityName="Hartlaub" authorityYear="1914" box="[974,1116,527,550]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="20" pageNumber="121" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B9232D69FFE7E24BFC4BFDA1DD3B1D2D" box="[974,1116,527,550]" italics="true" pageId="20" pageNumber="121">Leuckartiara</emphasis>
|
||
</taxonomicName>
|
||
medusa
|
||
<quantity id="4CAF5C9EFFE7E24BFB5CFDA1DC301D2D" box="[1241,1367,527,550]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="20" pageNumber="121" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15-20 mm</quantity>
|
||
total height, with large pointed apical process of about 1/3 to 2/5 of total bell height, umbrella higher than wide; up to 16-24 tentacles, between each tentacle pair 1-3 small, rudimentary bulbs, perradial and interradial tentacles with conspicuous, long, pointed abaxial spurs reaching up to 1/6 of the bell height; tentacles and bulbs lacking tentacles usually with small red abaxial ocelli. Manubrium about 1/2 of subumbrellar height, paleorange, with long mesenteries, mouth cruciform, mouth margin moderately ruffled. Gonads in adradial series of horizontal folds, distinct interradial connecting fold absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF28A1F3FFE7E24BFF12F847DC4E1710" ID-DOI="http://doi.org/10.5281/zenodo.5590772" ID-Zenodo-Dep="5590772" httpUri="https://zenodo.org/record/5590772/files/figure.png" pageId="20" pageNumber="121" startId="20.[151,186,2025,2046]" targetBox="[151,1436,771,1975]" targetPageId="20">
|
||
<paragraph id="8BE8F17BFFE7E24BFF12F847DC4E1710" blockId="20.[151,1434,2025,2075]" pageId="20" pageNumber="121">
|
||
Fig. 15.
|
||
<taxonomicName id="4C578AF8FFE7E24BFF73F847D88018F5" authority="Schuchert, 2018" authorityName="Schuchert" authorityYear="2018" box="[246,487,2025,2046]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="20" pageNumber="121" phylum="Cnidaria" rank="species" species="longicalcar" status="sp. nov.">
|
||
<emphasis id="B9232D69FFE7E24BFF73F847D88018F5" box="[246,487,2025,2046]" italics="true" pageId="20" pageNumber="121">Leuckartiara longicalcar</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2109012FFE7E24BFE77F847DB5A18F5" box="[498,573,2025,2046]" pageId="20" pageNumber="121" rank="species">n. spec.</taxonomicNameLabel>
|
||
, living animal. (A) Details of manubrium with gonad folds. (B) Lateral view of a perradial tentacle with the characteristic, long, abaxial spur. (C) Adradial tentacles and rudimentary bulbs, note red ocelli.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF28A1F3FFE6E24AFF12FB2CDA1D1BDB" ID-DOI="http://doi.org/10.5281/zenodo.5590776" ID-Zenodo-Dep="5590776" httpUri="https://zenodo.org/record/5590776/files/figure.png" pageId="21" pageNumber="122" startId="21.[151,186,1154,1175]" targetBox="[151,1436,257,1115]" targetPageId="21">
|
||
<paragraph id="8BE8F17BFFE6E24AFF12FB2CDA1D1BDB" blockId="21.[151,1437,1154,1232]" pageId="21" pageNumber="122">
|
||
Fig. 16.
|
||
<taxonomicName id="4C578AF8FFE6E24AFF73FB2CD88F1B9C" authority="Schuchert, 2018" authorityName="Schuchert" authorityYear="2018" box="[246,488,1154,1175]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="21" pageNumber="122" phylum="Cnidaria" rank="species" species="longicalcar" status="sp. nov.">
|
||
<emphasis id="B9232D69FFE6E24AFF73FB2CD88F1B9C" box="[246,488,1154,1175]" italics="true" pageId="21" pageNumber="122">Leuckartiara longicalcar</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2109012FFE6E24AFE76FB2CDB591B9C" box="[499,574,1154,1175]" pageId="21" pageNumber="122" rank="species">n. spec.</taxonomicNameLabel>
|
||
, living animal, height 13-18 mm, photographed by Kevin Lee off Los Angeles, California, USA. Note, this photograph is copyright protected and permission to use it here has been obtained by paying a royalty fee to the author and copyright holder Kevin Lee (www.diverkevin.com).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34DA2F0FFE6E24AFF12F9ABDAF418C6" pageId="21" pageNumber="122" type="description">
|
||
<paragraph id="8BE8F17BFFE6E24AFF12F9ABDBE61997" blockId="21.[151,756,1541,2077]" pageId="21" pageNumber="122">
|
||
<emphasis id="B9232D69FFE6E24AFF12F9ABD8421917" bold="true" box="[151,293,1541,1564]" pageId="21" pageNumber="122">Description:</emphasis>
|
||
<taxonomicName id="4C578AF8FFE6E24AFEABF9ABD8DD1917" authorityName="Hartlaub" authorityYear="1914" box="[302,442,1541,1564]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="21" pageNumber="122" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B9232D69FFE6E24AFEABF9ABD8DD1917" box="[302,442,1541,1564]" italics="true" pageId="21" pageNumber="122">Leuckartiara</emphasis>
|
||
</taxonomicName>
|
||
medusa up to
|
||
<quantity id="4CAF5C9EFFE6E24AFDE6F9ABDBB11917" box="[611,726,1541,1564]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="21" pageNumber="122" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15-20 mm</quantity>
|
||
in height and about
|
||
<quantity id="4CAF5C9EFFE6E24AFED1F98BD8F81937" box="[340,415,1573,1596]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="122" unit="mm" value="10.0">10 mm</quantity>
|
||
in diameter when mature, with a large, pointed apical process of about 1/3 to 2/5 of total bell height, umbrella higher than wide. Interradial, subumbrellar pockets of variable size present.
|
||
</paragraph>
|
||
<paragraph id="8BE8F17BFFE6E24AFF12F90BD8BC1837" blockId="21.[151,756,1541,2077]" pageId="21" pageNumber="122">Manubrium about half the height of the subumbrellar height, shaped like inverted vase, connected to radial canals via long mesenteries (about 1/3 of manubrium height). Manubrium base and mouth opening cruciform, mouth rim moderately ruffled.</paragraph>
|
||
<paragraph id="8BE8F17BFFE6E24AFF12F8EBDA871AC7" blockId="21.[151,756,1541,2077]" lastBlockId="21.[831,1437,1301,1997]" pageId="21" pageNumber="122">
|
||
Gonad tissue in 8 series of broad, adradial, horizontal folds, 8-12 folds in an adradial series, many folds with a central depression and resembling a loop or simply bifurcated (
|
||
<figureCitation id="136CEDFEFFE6E24AFE91F80BD81D18B7" box="[276,378,1957,1980]" captionStart="Fig" captionStartId="20.[151,186,2025,2046]" captionTargetBox="[151,1436,771,1975]" captionTargetId="figure-187@20.[151,1436,771,1975]" captionTargetPageId="20" captionText="Fig. 15. Leuckartiara longicalcar, n. spec., living animal. (A) Details of manubrium with gonad folds. (B) Lateral view of a perradial tentacle with the characteristic, long, abaxial spur. (C) Adradial tentacles and rudimentary bulbs, note red ocelli." figureDoi="http://doi.org/10.5281/zenodo.5590772" httpUri="https://zenodo.org/record/5590772/files/figure.png" pageId="21" pageNumber="122">Fig. 15A</figureCitation>
|
||
). The two series of gonad folds of one quadrant usually not connected by a fold across the interradial region as in other congeners (thus without the H-form of the gonad folds, often described as “horse-shoe shape” in older publications, comp.
|
||
<figureCitation id="136CEDFEFFE6E24AFAA2FABBDCF71A27" box="[1319,1424,1301,1324]" captionStart="Fig" captionStartId="22.[151,186,2025,2046]" captionTargetBox="[151,1436,1232,1989]" captionTargetId="figure-456@22.[151,1436,1232,1989]" captionTargetPageId="22" captionText="Fig. 17. Leuckartiara spec., living animal. (A) Whole medusa. (B) Bell margin with tentacle bases. (C) Details of manubrium with gonad folds (immature?)." figureDoi="http://doi.org/10.5281/zenodo.5590778" httpUri="https://zenodo.org/record/5590778/files/figure.png" pageId="21" pageNumber="122">Fig. 17C</figureCitation>
|
||
). Sometimes an inconspicuous interradial connection of the two rows of folds may be present at the aboral end of the manubrium. No gonadal pits. Egg size about
|
||
<quantity id="4CAF5C9EFFE6E24AFAC0FADBDCFE1A87" box="[1349,1433,1397,1420]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="122" unit="mm" value="0.1">0.1 mm</quantity>
|
||
. Radial canals slightly jagged and broad. Ring canal smooth, broad.
|
||
</paragraph>
|
||
<paragraph id="8BE8F17BFFE6E24AFCBAFA7BDADA1846" blockId="21.[831,1437,1301,1997]" pageId="21" pageNumber="122">
|
||
Tentacles usually 16, sometimes up to 24, between each tentacle pair 1-3 small, rudimentary bulbs without tentacles. Bases of tentacles laterally compressed, clasping bell margin. Perradial and interradial tentacle bases (oldest tentacles) with long, pointed abaxial spurs, reaching up to 1/6 of the bell height (
|
||
<figureCitation id="136CEDFEFFE6E24AFAAEF9D8DCE81986" box="[1323,1423,1654,1677]" captionStart="Fig" captionStartId="20.[151,186,2025,2046]" captionTargetBox="[151,1436,771,1975]" captionTargetId="figure-187@20.[151,1436,771,1975]" captionTargetPageId="20" captionText="Fig. 15. Leuckartiara longicalcar, n. spec., living animal. (A) Details of manubrium with gonad folds. (B) Lateral view of a perradial tentacle with the characteristic, long, abaxial spur. (C) Adradial tentacles and rudimentary bulbs, note red ocelli." figureDoi="http://doi.org/10.5281/zenodo.5590772" httpUri="https://zenodo.org/record/5590772/files/figure.png" pageId="21" pageNumber="122">Fig. 15B</figureCitation>
|
||
), shorter (younger) tentacles with short spur or no spur. Spurs appear solid, without internal canal. A small red ocellus present on most tentacles and also rudimentary bulbs, situated on abaxial side at interface of tentacle to exumbrella, in tentacles with long abaxial spurs ocelli at end of spur.
|
||
</paragraph>
|
||
<paragraph id="8BE8F17BFFE6E24AFCBAF8F8DC141886" blockId="21.[831,1437,1301,1997]" pageId="21" pageNumber="122">Colour: Manubrium pale orange, proximal parts of tentacles pale orange to yellowish, ocelli orange-red.</paragraph>
|
||
<paragraph id="8BE8F17BFFE6E24AFCBAF838DAF418C6" blockId="21.[831,1437,1301,1997]" pageId="21" pageNumber="122">Nematocysts of tentacles microbasic heteronemes, ca. 4 x 7 μm.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DA2F0FFE6E24AFCBAF848DD1A1716" pageId="21" pageNumber="122" type="distribution">
|
||
<paragraph id="8BE8F17BFFE6E24AFCBAF848DD1A1716" blockId="21.[831,1436,2022,2077]" pageId="21" pageNumber="122">
|
||
<emphasis id="B9232D69FFE6E24AFCBAF848DAB118F6" bold="true" box="[831,982,2022,2045]" pageId="21" pageNumber="122">Distribution:</emphasis>
|
||
North-eastern Pacific, from Vancouver Island to Southern
|
||
<collectingRegion id="49933F99FFE6E24AFB8FF7A8DD1D1716" box="[1034,1146,2054,2077]" country="United States of America" name="California" pageId="21" pageNumber="122">California</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DA2F0FFE5E249FF12FEACDD1A1C87" pageId="22" pageNumber="123" type="discussion">
|
||
<paragraph id="8BE8F17BFFE5E249FF12FEACDB051EB0" blockId="22.[151,757,258,1188]" pageId="22" pageNumber="123">
|
||
<emphasis id="B9232D69FFE5E249FF12FEACD86E1E12" bold="true" box="[151,265,258,281]" pageId="22" pageNumber="123">Remarks:</emphasis>
|
||
This species was described by
|
||
<bibRefCitation id="EFC68C8AFFE5E249FD18FEACD8BB1E32" author="Arai M. N. & Brinckmann-Voss A." pageId="22" pageNumber="123" pagination="1 - 192" refId="ref15698" refString="Arai M. N., Brinckmann-Voss A. 1980. Hydromedusae of British Columbia and Puget Sound. Canadian Bulletin of Fisheries and Aquatic Sciences 204: 1 - 192." type="journal article" year="1980">Arai & Brinckmann-Voss (1980: 56)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="4C578AF8FFE5E249FD89FE8CDB931E32" box="[524,756,290,313]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="undetermined">
|
||
<emphasis id="B9232D69FFE5E249FD89FE8CDBFE1E32" box="[524,665,290,313]" italics="true" pageId="22" pageNumber="123">Leuckartiara</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
distinct from
|
||
<taxonomicName id="4C578AF8FFE5E249FEBFFEEDD8C01E51" baseAuthorityName="Fleming" baseAuthorityYear="1823" box="[314,423,323,346]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="octona">
|
||
<emphasis id="B9232D69FFE5E249FEBFFEEDD8C01E51" box="[314,423,323,346]" italics="true" pageId="22" pageNumber="123">L. octona</emphasis>
|
||
</taxonomicName>
|
||
. Dr Anita Brinckmann-Voss (pers. com., 2013) told me that she initially intended to name it in a subsequent publication, but was now unable to do it and encouraged me to do it myself.
|
||
</paragraph>
|
||
<paragraph id="8BE8F17BFFE5E249FF12FE6AD8CE1D76" blockId="22.[151,757,258,1188]" pageId="22" pageNumber="123">
|
||
<taxonomicName id="4C578AF8FFE5E249FF12FE6AD8CB1ED0" authorityName="Schuchert" authorityYear="2018" box="[151,428,452,475]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="longicalcar">
|
||
<emphasis id="B9232D69FFE5E249FF12FE6AD8CB1ED0" box="[151,428,452,475]" italics="true" pageId="22" pageNumber="123">Leuckartiara longicalcar</emphasis>
|
||
</taxonomicName>
|
||
does not match any of the known species (
|
||
<bibRefCitation id="EFC68C8AFFE5E249FEC0FE4BD8B01EF7" author="Kramp P. L." box="[325,471,485,508]" pageId="22" pageNumber="123" pagination="1 - 200" refId="ref16990" refString="Kramp P. L. 1968. The hydromedusae of the Pacific and Indian Oceans. Sections II and III. Dana Report 72: 1 - 200." type="book chapter" year="1968">Kramp, 1968</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC68C8AFFE5E249FE60FE4BDBCD1EF7" author="Pages F. & Gili J. M. & Bouillon J." box="[485,682,485,508]" pageId="22" pageNumber="123" pagination="1 - 64" refId="ref17594" refString="Pages F., Gili J. M., Bouillon J. 1992. Medusae (Hydrozoa, Scyphozoa, Cubozoa) of the Benguela Current (southeastern Atlantic). Scientia Marina 56 (Suppl. 1): 1 - 64." type="journal article" year="1992">
|
||
Pages
|
||
<emphasis id="B9232D69FFE5E249FDAEFE4BDB3B1EF7" box="[555,604,485,508]" italics="true" pageId="22" pageNumber="123">et al</emphasis>
|
||
., 1992
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC68C8AFFE5E249FD33FE4BD84F1D17" author="Xu Z. - Z. & Huang J. - Q." pageId="22" pageNumber="123" pagination="549 - 562" refId="ref18757" refString="Xu Z. - Z., Huang J. - Q. 2004. A survey on Anthomedusae (Hydrozoa: Hydroidomedusae) from the Taiwan Strait with description of new species and new combinations. Acta Oceanologica Sinica 23 (3): 549 - 562." type="journal article" year="2004">Xu & Huang, 2004</bibRefCitation>
|
||
; Schuchert, 2017). It has previously been misidentified as
|
||
<taxonomicName id="4C578AF8FFE5E249FED6FD88DBE61D36" authority="(Fleming, 1823)" baseAuthorityName="Fleming" baseAuthorityYear="1823" box="[339,641,550,573]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="octona">
|
||
<emphasis id="B9232D69FFE5E249FED6FD88D8D81D36" box="[339,447,550,573]" italics="true" pageId="22" pageNumber="123">L. octona</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC68C8AFFE5E249FE51FD88DB1E1D36" author="Fleming J." box="[468,633,550,573]" pageId="22" pageNumber="123" pagination="294 - 303" refId="ref16195" refString="Fleming J. 1823. Gleamings of natural history, gathered on the coast of Scotland during a voyage in 1821. Edinburgh Philosophical Journal 8: 294 - 303." type="journal article" year="1823">Fleming, 1823</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and been considered related to
|
||
<taxonomicName id="4C578AF8FFE5E249FEFFFDE8DB131D56" authority="Bigelow, 1940" authorityName="Bigelow" authorityYear="1940" box="[378,628,582,605]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="zacae">
|
||
<emphasis id="B9232D69FFE5E249FEFFFDE8D8B41D56" box="[378,467,582,605]" italics="true" pageId="22" pageNumber="123">L. zacae</emphasis>
|
||
<bibRefCitation id="EFC68C8AFFE5E249FE5CFDE8DB131D56" author="Bigelow H. B." box="[473,628,582,605]" pageId="22" pageNumber="123" pagination="281 - 321" refId="ref15928" refString="Bigelow H. B. 1940. Eastern Pacific Expeditions of the New York Zoological Society. XX. Medusae of the Templeton Crocker and Eastern Pacific Zaca Expeditions, 1936 - 1938. Zoologica, N. Y. 25 (3): 281 - 321." type="journal article" year="1940">Bigelow, 1940</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFC68C8AFFE5E249FD2FFDE8D8FA1D76" author="Arai M. N. & Brinckmann-Voss A." pageId="22" pageNumber="123" pagination="1 - 192" refId="ref15698" refString="Arai M. N., Brinckmann-Voss A. 1980. Hydromedusae of British Columbia and Puget Sound. Canadian Bulletin of Fisheries and Aquatic Sciences 204: 1 - 192." type="journal article" year="1980">Arai & Brinckmann-Voss, 1980</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE8F17BFFE5E249FF12FD29DB0C1C4B" blockId="22.[151,757,258,1188]" pageId="22" pageNumber="123">
|
||
<taxonomicName id="4C578AF8FFE5E249FF12FD29D81E1D95" baseAuthorityName="Fleming" baseAuthorityYear="1823" box="[151,377,647,670]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="octona">
|
||
<emphasis id="B9232D69FFE5E249FF12FD29D81E1D95" box="[151,377,647,670]" italics="true" pageId="22" pageNumber="123">Leuckartiara octona</emphasis>
|
||
</taxonomicName>
|
||
is indeed similar in appearance, but lacks the long abaxial spurs and regularly has a fold across the interradial region connecting the adradial series of folds. The 16S and COI sequence data (
|
||
<figureCitation id="136CEDFEFFE5E249FD40FD46D9DC1C14" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[151,186,1968,1989]" captionStartId-1="13.[151,186,1913,1934]" captionTargetId-0="graphics-9@12.[87,1710,443,1593]" captionTargetId-1="graphics-6@13.[102,1709,536,1563]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Fig. 8. 16S maximum likelihood phylogenetic tree of Pandeidae species obtained with PhyML (GTR+G+I model) and based on 595 bp positions of the mitochondrial 16S gene. Node-support values are bootstrap values of 100 pseudoreplicates (shown only if > 70%). For more details see text and Table 1. Samples based on the polyp stage are indicated, all others are medusa samples." captionText-1="Fig. 9. COI maximum likelihood phylogenetic tree of Pandeidae species obtained with PhyML (GTR+G+I model) and based on 664 bp positions of the mitochondrial COI gene. Node-support values are bootstrap values of 100 pseudoreplicates (shown only if> 70%). For more details see text and Table 1. Samples based on the polyp stage are indicated, all others are medusa samples." figureDoi-0="http://doi.org/10.5281/zenodo.5590756" figureDoi-1="http://doi.org/10.5281/zenodo.5590759" httpUri-0="https://zenodo.org/record/5590756/files/figure.png" httpUri-1="https://zenodo.org/record/5590759/files/figure.png" pageId="22" pageNumber="123">Figs 8-9</figureCitation>
|
||
) clearly separated
|
||
<taxonomicName id="4C578AF8FFE5E249FE15FCA6DB4C1C14" authorityName="Schuchert" authorityYear="2018" box="[400,555,776,799]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="longicalcar">
|
||
<emphasis id="B9232D69FFE5E249FE15FCA6DB4C1C14" box="[400,555,776,799]" italics="true" pageId="22" pageNumber="123">L. longicalcar</emphasis>
|
||
</taxonomicName>
|
||
from the Atlantic
|
||
<taxonomicName id="4C578AF8FFE5E249FF12FC87D99B1C4B" baseAuthorityName="Fleming" baseAuthorityYear="1823" box="[151,252,809,832]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="octona">
|
||
<emphasis id="B9232D69FFE5E249FF12FC87D99B1C4B" box="[151,252,809,832]" italics="true" pageId="22" pageNumber="123">L. octona</emphasis>
|
||
</taxonomicName>
|
||
, although they are closely related.
|
||
</paragraph>
|
||
<paragraph id="8BE8F17BFFE5E249FF12FCE7DD171E50" blockId="22.[151,757,258,1188]" lastBlockId="22.[832,1437,258,908]" pageId="22" pageNumber="123">
|
||
<taxonomicName id="4C578AF8FFE5E249FF12FCE7DB451C6B" authority="Bigelow, 1940" authorityName="Bigelow" authorityYear="1940" box="[151,546,841,864]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="zacae">
|
||
<emphasis id="B9232D69FFE5E249FF12FCE7D8081C6B" box="[151,367,841,864]" italics="true" pageId="22" pageNumber="123">Leuckartiara zacae</emphasis>
|
||
<bibRefCitation id="EFC68C8AFFE5E249FEFBFCE7DB451C6B" author="Bigelow H. B." box="[382,546,841,864]" pageId="22" pageNumber="123" pagination="281 - 321" refId="ref15928" refString="Bigelow H. B. 1940. Eastern Pacific Expeditions of the New York Zoological Society. XX. Medusae of the Templeton Crocker and Eastern Pacific Zaca Expeditions, 1936 - 1938. Zoologica, N. Y. 25 (3): 281 - 321." type="journal article" year="1940">Bigelow, 1940</bibRefCitation>
|
||
</taxonomicName>
|
||
is a rare species first found in the Gulf of
|
||
<collectingCountry id="F340B1EBFFE5E249FE29FCC4DB631C8A" box="[428,516,874,897]" name="Panama" pageId="22" pageNumber="123">Panama</collectingCountry>
|
||
. It is somewhat larger than
|
||
<taxonomicName id="4C578AF8FFE5E249FF4AFC24D80D1CAA" authorityName="Schuchert" authorityYear="2018" box="[207,362,906,929]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="longicalcar">
|
||
<emphasis id="B9232D69FFE5E249FF4AFC24D80D1CAA" box="[207,362,906,929]" italics="true" pageId="22" pageNumber="123">L. longicalcar</emphasis>
|
||
</taxonomicName>
|
||
and has about the same number of tentacles. The most prominent difference is the length of the tentacle spurs: they are much longer and extend up to 2/3 of the bell height.
|
||
<bibRefCitation id="EFC68C8AFFE5E249FE24FC45DB2D1B09" author="Bigelow H. B." box="[417,586,1003,1026]" pageId="22" pageNumber="123" pagination="281 - 321" refId="ref15928" refString="Bigelow H. B. 1940. Eastern Pacific Expeditions of the New York Zoological Society. XX. Medusae of the Templeton Crocker and Eastern Pacific Zaca Expeditions, 1936 - 1938. Zoologica, N. Y. 25 (3): 281 - 321." type="journal article" year="1940">Bigelow (1940)</bibRefCitation>
|
||
described them as exumbrellar ribs containing a thin gastrodermal canal. Additionally,
|
||
<taxonomicName id="4C578AF8FFE5E249FEAFFB82D8E31B48" authorityName="Bigelow" authorityYear="1940" box="[298,388,1068,1091]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="zacae">
|
||
<emphasis id="B9232D69FFE5E249FEAFFB82D8E31B48" box="[298,388,1068,1091]" italics="true" pageId="22" pageNumber="123">L. zacae</emphasis>
|
||
</taxonomicName>
|
||
has no apical process (but
|
||
<bibRefCitation id="EFC68C8AFFE5E249FD2FFB82D9B91B6F" author="Kramp P. L." pageId="22" pageNumber="123" pagination="1 - 162" refId="ref16966" refString="Kramp P. L. 1965. The hydromedusae of the Pacific and Indian Oceans. Dana Report 63: 1 - 162." type="book chapter" year="1965">Kramp (1965)</bibRefCitation>
|
||
observed a small process in a juvenile specimen from
|
||
<collectingCountry id="F340B1EBFFE5E249FF50FBC3D8581B8F" box="[213,319,1133,1156]" name="Indonesia" pageId="22" pageNumber="123">Indonesia</collectingCountry>
|
||
, the identity of this material is perhaps questionable), the umbrella without the process is larger (21 versus
|
||
<quantity id="4CAF5C9EFFE5E249FC32FEACDD601E12" box="[951,1031,258,281]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="123" unit="mm" value="12.0">12 mm</quantity>
|
||
), the manubrium is more voluminous and has more gonadal folds. It is only known to occur in tropical seas (
|
||
<bibRefCitation id="EFC68C8AFFE5E249FC57FEEADD031E50" author="Kramp P. L." box="[978,1124,324,347]" pageId="22" pageNumber="123" pagination="1 - 162" refId="ref16966" refString="Kramp P. L. 1965. The hydromedusae of the Pacific and Indian Oceans. Dana Report 63: 1 - 162." type="book chapter" year="1965">Kramp, 1965</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE8F17BFFE5E249FCC5FECBDAC31D0B" blockId="22.[832,1437,258,908]" pageId="22" pageNumber="123">
|
||
Other
|
||
<taxonomicName id="4C578AF8FFE5E249FC0BFECBDD1E1E77" box="[910,1145,357,380]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="undetermined">
|
||
<emphasis id="B9232D69FFE5E249FC0BFECBDD7E1E77" box="[910,1049,357,380]" italics="true" pageId="22" pageNumber="123">Leuckartiara</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with tentacle spurs are
|
||
<taxonomicName id="4C578AF8FFE5E249FCC5FE28DD001E96" authority="Browne, 1916" authorityName="Browne" authorityYear="1916" box="[832,1127,390,413]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="gardineri">
|
||
<emphasis id="B9232D69FFE5E249FCC5FE28DA281E96" box="[832,847,390,413]" italics="true" pageId="22" pageNumber="123">L</emphasis>
|
||
.
|
||
<emphasis id="B9232D69FFE5E249FCDBFE28DAA51E96" box="[862,962,390,413]" italics="true" pageId="22" pageNumber="123">gardineri</emphasis>
|
||
<bibRefCitation id="EFC68C8AFFE5E249FC4EFE28DD001E96" author="Browne E." box="[971,1127,390,413]" pageId="22" pageNumber="123" pagination="169 - 209" refId="ref16067" refString="Browne E. T 1916. Medusae from the Indian Ocean. Transactions of the Linnean Society of London, Zoology 17 (2): 169 - 209." type="journal article" year="1916">Browne, 1916</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C578AF8FFE5E249FBF1FE28DD2E1EB5" authority="Brinckmann-Voss, Arai & Nagasawa, 2005" authorityName="Brinckmann-Voss, Arai & Nagasawa" authorityYear="2005" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="acuta">
|
||
<emphasis id="B9232D69FFE5E249FBF1FE28DDA91E96" box="[1140,1230,390,413]" italics="true" pageId="22" pageNumber="123">L. acuta</emphasis>
|
||
<bibRefCitation id="EFC68C8AFFE5E249FB52FE28DD2E1EB5" author="Brinckmann-Voss A. & Arai M. N. & Nagasawa K." pageId="22" pageNumber="123" pagination="131 - 139" refId="ref16017" refString="Brinckmann-Voss A., Arai M. N., Nagasawa K. 2005. Leuckartiara acuta (Hydrozoa, Anthoathecatae, Pandeidae), a new species from the Pacific. Publications of the Seto Marine Biological Laboratory 40 (3 - 4): 131 - 139." type="journal article" year="2005">Brinckmann-Voss, Arai & Nagasawa, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C578AF8FFE5E249FB00FE09DA9D1ED4" authority="Huang, Xu, Lin & Qiu, 2008" authorityName="Huang, Xu, Lin & Qiu" authorityYear="2008" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="fujianensis">
|
||
<emphasis id="B9232D69FFE5E249FB00FE09DC701EB5" box="[1157,1303,423,446]" italics="true" pageId="22" pageNumber="123">L. fujianensis</emphasis>
|
||
<bibRefCitation id="EFC68C8AFFE5E249FA9AFE09DA9D1ED4" author="Huang J. Q. & Xu Z. Z. & Lin J. Z. & Qiu M. F." pageId="22" pageNumber="123" pagination="408 - 412" refId="ref16752" refString="Huang J. Q., Xu Z. Z., Lin J. Z., Qiu M. F. 2008. Three new species of Anthomedusae (Hydrozoa, Hydroidomedusae) from the Fujian sea water. Journal of Xiamen University Natural Science 47 (3): 408 - 412." type="journal article" year="2008">Huang, Xu, Lin & Qiu, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
. All three have only four fully formed tentacles.
|
||
</paragraph>
|
||
<paragraph id="8BE8F17BFFE5E249FCC5FDA4DD1A1C87" blockId="22.[832,1437,258,908]" pageId="22" pageNumber="123">
|
||
Kevin
|
||
<bibRefCitation id="EFC68C8AFFE5E249FC15FDA4DD6A1D2A" author="Lee K." box="[912,1037,522,545]" pageId="22" pageNumber="123" refId="ref17197" refString="Lee K. 2017. Leuckartiara spp. (?) Blue water dive, miles offshore Palos Verdes, California USA. Available at:" type="book" year="2017">Lee (2017)</bibRefCitation>
|
||
published a series of magnificent photos of
|
||
<taxonomicName id="4C578AF8FFE5E249FC47FD85DD2A1D49" authorityName="Hartlaub" authorityYear="1914" box="[962,1101,555,578]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B9232D69FFE5E249FC47FD85DD2A1D49" box="[962,1101,555,578]" italics="true" pageId="22" pageNumber="123">Leuckartiara</emphasis>
|
||
</taxonomicName>
|
||
medusae observed off Los Angeles,
|
||
<collectingRegion id="49933F99FFE5E249FC2EFDE2DD7C1D68" box="[939,1051,588,611]" country="United States of America" name="California" pageId="22" pageNumber="123">California</collectingRegion>
|
||
. One of them is reproduced here (
|
||
<figureCitation id="136CEDFEFFE5E249FCC2FDC3DAFB1D8F" box="[839,924,621,644]" captionStart="Fig" captionStartId="22.[151,186,2025,2046]" captionTargetBox="[151,1436,1232,1989]" captionTargetId="figure-456@22.[151,1436,1232,1989]" captionTargetPageId="22" captionText="Fig. 17. Leuckartiara spec., living animal. (A) Whole medusa. (B) Bell margin with tentacle bases. (C) Details of manubrium with gonad folds (immature?)." figureDoi="http://doi.org/10.5281/zenodo.5590778" httpUri="https://zenodo.org/record/5590778/files/figure.png" pageId="22" pageNumber="123">Fig. 17</figureCitation>
|
||
). These medusae must clearly be referred to
|
||
<taxonomicName id="4C578AF8FFE5E249FCC5FD20DAB21DAE" authority="Schuchert, 2018" authorityName="Schuchert" authorityYear="2018" box="[832,981,654,677]" class="Hydrozoa" family="Pandeidae" genus="Leuckartiara" kingdom="Animalia" order="Anthoathecata" pageId="22" pageNumber="123" phylum="Cnidaria" rank="species" species="longicalcar" status="sp. nov.">
|
||
<emphasis id="B9232D69FFE5E249FCC5FD20DAB21DAE" box="[832,981,654,677]" italics="true" pageId="22" pageNumber="123">L. longicalcar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2109012FFE5E249FC5EFD20DD4C1DAE" box="[987,1067,654,677]" pageId="22" pageNumber="123" rank="species">n. spec.</taxonomicNameLabel>
|
||
Some of the individuals are almost identical to the ones shown here, while others (
|
||
<figureCitation id="136CEDFEFFE5E249FAC0FD01DCF31DCD" box="[1349,1428,687,710]" captionStart="Fig" captionStartId="22.[151,186,2025,2046]" captionTargetBox="[151,1436,1232,1989]" captionTargetId="figure-456@22.[151,1436,1232,1989]" captionTargetPageId="22" captionText="Fig. 17. Leuckartiara spec., living animal. (A) Whole medusa. (B) Bell margin with tentacle bases. (C) Details of manubrium with gonad folds (immature?)." figureDoi="http://doi.org/10.5281/zenodo.5590778" httpUri="https://zenodo.org/record/5590778/files/figure.png" pageId="22" pageNumber="123">Fig. 17</figureCitation>
|
||
) appear somewhat larger, with up to 24 tentacles, and a more voluminous stomach. Some of the individuals have a more intense colour, appearing more reddish, and also the tentacle bases show some reddish pigments. The distribution of the species extends thus from Vancouver Island to Southern
|
||
<collectingRegion id="49933F99FFE5E249FB8FFCDBDD1D1C87" box="[1034,1146,885,908]" country="United States of America" name="California" pageId="22" pageNumber="123">California</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |