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<mods:title id="DA8FB21536AB01DC6F8B39DEBE311B0F">Taxonomy of the ' Afroeudesis group' of glandulariine ant-like stone beetles (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<treatment id="03EC517274432451BBEFFF1DB4F2FE91" ID-DOI="http://doi.org/10.5281/zenodo.5942767" ID-GBIF-Taxon="157293832" ID-Zenodo-Dep="5942767" LSID="urn:lsid:plazi:treatment:03EC517274432451BBEFFF1DB4F2FE91" httpUri="http://treatment.plazi.org/id/03EC517274432451BBEFFF1DB4F2FE91" lastPageId="14" lastPageNumber="219" pageId="13" pageNumber="218">
<subSubSection id="C35FB3EF74432452BBEFFF1DB768FF66" box="[151,377,153,180]" pageId="13" pageNumber="218" type="nomenclature">
<paragraph id="8BFAE06474432452BBEFFF1DB768FF66" blockId="13.[151,377,153,180]" box="[151,377,153,180]" pageId="13" pageNumber="218">
<heading id="D0B2570874432452BBEFFF1DB768FF66" bold="true" box="[151,377,153,180]" fontSize="11" level="1" pageId="13" pageNumber="218" reason="1">
<taxonomicName id="4C459BE774432452BBEFFF1DB768FF66" ID-CoL="8XJCK" authority="Franz" authorityName="Franz" authorityYear="1963" box="[151,377,153,180]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452BBEFFF1DB768FF66" bold="true" box="[151,377,153,180]" pageId="13" pageNumber="218">
<emphasis id="B9313C7674432452BBEFFF1DB737FF66" bold="true" box="[151,294,153,180]" italics="true" pageId="13" pageNumber="218">Afroeudesis</emphasis>
Franz
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35FB3EF74432452BBEFFF5AB5D8FEE1" pageId="13" pageNumber="218" type="reference_group">
<paragraph id="8BFAE06474432452BBEFFF5AB5D8FEE1" blockId="13.[151,1437,221,307]" pageId="13" pageNumber="218">
<treatmentCitationGroup id="AB55C74A74432452BBEFFF5AB5D8FEE1" infered="true" pageId="13" pageNumber="218">
<taxonomicName id="4C459BE774432452BBEFFF5AB7A3FF24" ID-CoL="8XJCK" authority="Franz, 1963: 16" authorityName="Franz" authorityPageNumber="16" authorityYear="1963" box="[151,434,221,246]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452BBEFFF5AB71FFF24" box="[151,270,222,246]" italics="true" pageId="13" pageNumber="218">Afroeudesis</emphasis>
<treatmentCitation id="0AE4C67574432452BA6CFF5AB7A3FF24" author="Franz, H." box="[276,434,221,246]" page="16" pageId="13" pageNumber="218" year="1963">
<bibRefCitation id="EFD49D9574432452BA6CFF5AB7A3FF24" author="Franz, H." box="[276,434,221,246]" pageId="13" pageNumber="218" pagination="1 - 93" refId="ref7193" refString="Franz, H. (1963) Scydmaeniden (Coleoptera) aus Ostafrika, dem Kongo und der Tschadrepublik (vorwiegend aus dem im Sommer 1962 vom Verfasser selbst gesammelten Ausbeuten). Stuttgarter Beitrage zur Naturkunde, 120, 1 - 93." type="journal article" year="1963">Franz, 1963: 16</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(as subgenus of
<taxonomicName id="4C459BE774432452B924FF5AB4E0FF24" ID-CoL="8XPCS" authorityName="Binaghi" authorityYear="1948" box="[604,753,222,246]" class="Insecta" family="Staphylinidae" genus="Pseudoeudesis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452B924FF5AB4E0FF24" box="[604,753,222,246]" italics="true" pageId="13" pageNumber="218">Pseudoeudesis</emphasis>
</taxonomicName>
).
<typeStatus id="54FE5EC674432452B87DFF5AB526FF24" box="[773,823,222,246]" pageId="13" pageNumber="218">Type</typeStatus>
species:
<taxonomicName id="4C459BE774432452B8EBFF5AB333FF24" authority="Franz, 1963" authorityName="Franz" authorityYear="1963" box="[915,1314,221,246]" class="Insecta" family="Staphylinidae" genus="Pseudoeudesis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis id="B9313C7674432452B8EBFF5AB2B2FF24" box="[915,1187,222,246]" italics="true" pageId="13" pageNumber="218">Pseudoeudesis basilewskyi</emphasis>
<bibRefCitation id="EFD49D9574432452BFD1FF5AB333FF24" author="Franz, H." box="[1193,1314,221,246]" pageId="13" pageNumber="218" pagination="1 - 93" refId="ref7193" refString="Franz, H. (1963) Scydmaeniden (Coleoptera) aus Ostafrika, dem Kongo und der Tschadrepublik (vorwiegend aus dem im Sommer 1962 vom Verfasser selbst gesammelten Ausbeuten). Stuttgarter Beitrage zur Naturkunde, 120, 1 - 93." type="journal article" year="1963">Franz, 1963</bibRefCitation>
</taxonomicName>
(des. orig.). Recognized as a member of Glandulariini and elevated to genus rank by
<bibRefCitation id="EFD49D9574432452B8C9FF78B26CFEC7" author="Jaloszynski, P." box="[945,1149,252,277]" pageId="13" pageNumber="218" pagination="493 - 507" refId="ref7691" refString="Jaloszynski, P. (2015 a) Confused identity: subgenera of Pseudoeudesis Binaghi belong in two tribes (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4048 (4), 493 - 507. https: // doi. org / 10.11646 / zootaxa. 4048.4.2" type="journal article" year="2015">Jałoszyński (2015a)</bibRefCitation>
, who also gave an emended diagnosis and divided the genus into subgenera (
<bibRefCitation id="EFD49D9574432452B9CBFE9EB561FEE1" author="Jaloszynski, P." box="[691,880,282,307]" pageId="13" pageNumber="218" pagination="493 - 507" refId="ref7691" refString="Jaloszynski, P. (2015 a) Confused identity: subgenera of Pseudoeudesis Binaghi belong in two tribes (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4048 (4), 493 - 507. https: // doi. org / 10.11646 / zootaxa. 4048.4.2" type="journal article" year="2015">Jałoszyński 2015a</bibRefCitation>
,
<bibRefCitation id="EFD49D9574432452B802FE9EB5AFFEE1" author="Jaloszynski, P." box="[890,958,282,307]" pageId="13" pageNumber="218" pagination="163 - 173" refId="ref7905" refString="Jaloszynski, P. (2016 b) Microscydmus trianguliceps Franz and Pseudoeudesis sulcata Franz transferred to Afroeudesis Franz (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4173 (2), 163 - 173. https: // doi. org / 10.11646 / zootaxa. 4173.2.7" type="journal article" year="2016">2016b</bibRefCitation>
).
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</paragraph>
</subSubSection>
<subSubSection id="C35FB3EF74432452BBEFFEE5B703FCFE" pageId="13" pageNumber="218" type="diagnosis">
<paragraph id="8BFAE06474432452BBEFFEE5B703FCFE" blockId="13.[151,1437,353,1243]" pageId="13" pageNumber="218">
<emphasis id="B9313C7674432452BBEFFEE5B767FEAE" bold="true" box="[151,374,353,380]" pageId="13" pageNumber="218">Revised diagnosis.</emphasis>
<taxonomicName id="4C459BE774432452BAF9FEE5B414FEA9" authorityName="Franz" authorityYear="1963" box="[385,517,353,379]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452BAF9FEE5B414FEA9" box="[385,517,353,379]" italics="true" pageId="13" pageNumber="218">Afroeudesis</emphasis>
</taxonomicName>
differs from all genera of Glandulariini in a unique autapomorphy: frons and vertex with a subtriangular elevated 'platform' covered with scale-like microsculpture and with longitudinal median groove, the platform is anteriorly demarcated by a deep frontal impression (
<figureCitation id="137EFCE174432452B884FE2DB240FE11" box="[1020,1105,425,451]" captionStart="FIGURES 912" captionStartId="4.[151,264,1612,1637]" captionTargetBox="[151,1436,330,1600]" captionTargetId="figure@4.[151,1436,330,1600]" captionTargetPageId="4" captionText="FIGURES 912. Examples of the symmetrical setal pattern on frons and vertex in representatives of the 'Afroeudesis lineage' (insertions of setae, including those broken off, marked by circles).Meridaphes venezolanus (9); Stenichnoconnus huijbregtsensis (10); Pseudoraphes minor (11); Afroeudesis (Nanoscydmus) baoshana (12)." httpUri="https://zenodo.org/record/3234302/files/figure.png" pageId="13" pageNumber="218">Fig. 12</figureCitation>
). Additionally, the following combination of synapomorphies characterizes
<taxonomicName id="4C459BE774432452B9E7FE49B532FE35" authorityName="Franz" authorityYear="1963" box="[671,803,461,487]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452B9E7FE49B532FE35" box="[671,803,461,487]" italics="true" pageId="13" pageNumber="218">Afroeudesis</emphasis>
</taxonomicName>
: body (Figs 58) moderately stout, distinctly constricted between head and prothorax and between prothorax and elytra; head (
<figureCitation id="137EFCE174432452B8C0FE75B21AFDD9" box="[952,1035,497,523]" captionStart="FIGURES 912" captionStartId="4.[151,264,1612,1637]" captionTargetBox="[151,1436,330,1600]" captionTargetId="figure@4.[151,1436,330,1600]" captionTargetPageId="4" captionText="FIGURES 912. Examples of the symmetrical setal pattern on frons and vertex in representatives of the 'Afroeudesis lineage' (insertions of setae, including those broken off, marked by circles).Meridaphes venezolanus (9); Stenichnoconnus huijbregtsensis (10); Pseudoraphes minor (11); Afroeudesis (Nanoscydmus) baoshana (12)." httpUri="https://zenodo.org/record/3234302/files/figure.png" pageId="13" pageNumber="218">Fig. 12</figureCitation>
) about as long as broad or slightly elongate, with the most convex site of eye closer to occipital constriction than to mandibular bases; posteromedian impression on vertex present, setae on frons and vertex forming a largely symmetrical pattern; lateral margins of anterior portion of clypeus divergent anterad; antennal insertions distant from mandibular bases; pronotum broadest in front of middle or subequal in width between posterior and anterior third, lacking lateral and sublateral carinae, with transverse antebasal groove or impression connecting a pair of inner lateral pits and with a pair of outer lateral pits; bristles on head and pronotum absent; mesoventral intercoxal process carinate, narrow; anterior metaventral process present; metaventral intermetacoxal process with a pair of long spines; and aedeagus with free, slender parameres.
</paragraph>
</subSubSection>
<subSubSection id="C35FB3EF74432451BBBFFCB2B4F2FE91" lastPageId="14" lastPageNumber="219" pageId="13" pageNumber="218" type="discussion">
<paragraph id="8BFAE06474432452BBBFFCB2B30AFBD6" blockId="13.[151,1437,353,1243]" pageId="13" pageNumber="218">
<emphasis id="B9313C7674432452BBBFFCB2B72AFC82" bold="true" box="[199,315,822,848]" pageId="13" pageNumber="218">Remarks.</emphasis>
Species of
<taxonomicName id="4C459BE774432452BABBFCB1B456FC9D" authorityName="Franz" authorityYear="1963" box="[451,583,821,847]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452BABBFCB1B456FC9D" box="[451,583,821,847]" italics="true" pageId="13" pageNumber="218">Afroeudesis</emphasis>
</taxonomicName>
are similar in the general appearance, but show a significant variability in some structures. They can have the mandibles planar or with a dorsal preapical tooth, the hypostomal ridges complete or incomplete, the hypomeral ridges present or absent; the anterior metaventral process broad or narrow, the lateral metaventral carinae present or absent, the mesoscutellum exposed or hidden under the posterior pronotal margin; one, two or even none basal elytral foveae, and an even or odd number of the pronotal antebasal pits. Differences in these characters are used in subgeneric diagnoses and were compiled in a table by
<bibRefCitation id="EFD49D9574432452BF4BFC6DB306FBD6" author="Jaloszynski, P." box="[1075,1303,1001,1028]" pageId="13" pageNumber="218" pagination="163 - 173" refId="ref7905" refString="Jaloszynski, P. (2016 b) Microscydmus trianguliceps Franz and Pseudoeudesis sulcata Franz transferred to Afroeudesis Franz (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4173 (2), 163 - 173. https: // doi. org / 10.11646 / zootaxa. 4173.2.7" type="journal article" year="2016">Jałoszyński (2016b)</bibRefCitation>
.
</paragraph>
<paragraph id="8BFAE06474432452BBBFFB89B299FB09" blockId="13.[151,1437,353,1243]" pageId="13" pageNumber="218">
Interestingly, differences between subgenera of
<taxonomicName id="4C459BE774432452B9A6FB89B573FBF5" authorityName="Franz" authorityYear="1963" box="[734,866,1037,1063]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452B9A6FB89B573FBF5" box="[734,866,1037,1063]" italics="true" pageId="13" pageNumber="218">Afroeudesis</emphasis>
</taxonomicName>
seem to be bigger than those between
<taxonomicName id="4C459BE774432452BE77FB89B384FBF5" box="[1295,1429,1037,1063]" class="Insecta" family="Staphylinidae" genus="Meridaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452BE77FB89B384FBF5" box="[1295,1429,1037,1063]" italics="true" pageId="13" pageNumber="218">Meridaphes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C459BE774432452BBEFFBB5B727FB99" authorityName="Franz" authorityYear="1980" box="[151,310,1073,1099]" class="Insecta" family="Staphylinidae" genus="Pseudoraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452BBEFFBB5B727FB99" box="[151,310,1073,1099]" italics="true" pageId="13" pageNumber="218">Pseudoraphes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C459BE774432452BA13FBB5B438FB99" authorityName="Franz" authorityYear="1985" box="[363,553,1073,1099]" class="Insecta" family="Staphylinidae" genus="Stenichnoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452BA13FBB5B438FB99" box="[363,553,1073,1099]" italics="true" pageId="13" pageNumber="218">Stenichnoconnus</emphasis>
</taxonomicName>
, maintained here as separate genera. It would be possible to divide
<taxonomicName id="4C459BE774432452BE60FBB5B38DFB99" authorityName="Franz" authorityYear="1963" box="[1304,1436,1073,1099]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452BE60FBB5B38DFB99" box="[1304,1436,1073,1099]" italics="true" pageId="13" pageNumber="218">Afroeudesis</emphasis>
</taxonomicName>
into four genera, especially that the split between the sub-Himalayan, African and South American components of this clade may be very ancient. However, the subgenera of
<taxonomicName id="4C459BE774432452B85FFBFDB5BAFB41" authorityName="Franz" authorityYear="1963" box="[807,939,1145,1171]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452B85FFBFDB5BAFB41" box="[807,939,1145,1171]" italics="true" pageId="13" pageNumber="218">Afroeudesis</emphasis>
</taxonomicName>
share the unique elongate, subtriangular and unusually microsculptured 'platform' on the frons and vertex, a structure that has clearly evolved only once within the Glandulariini. The subgeneric classification of
<taxonomicName id="4C459BE774432452B9A8FB45B545FB09" authorityName="Franz" authorityYear="1963" box="[720,852,1217,1243]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452B9A8FB45B545FB09" box="[720,852,1217,1243]" italics="true" pageId="13" pageNumber="218">Afroeudesis</emphasis>
</taxonomicName>
species is here maintained.
</paragraph>
<caption id="DF3AB0EC74432452BBEFF84FB49DF836" box="[151,652,1995,2020]" httpUri="https://zenodo.org/record/3234314/files/figure.png" pageId="13" pageNumber="218" startId="13.[151,250,1995,2020]" targetBox="[161,1417,1304,1961]" targetPageId="13">
<paragraph id="8BFAE06474432452BBEFF84FB49DF836" blockId="13.[151,652,1995,2020]" box="[151,652,1995,2020]" pageId="13" pageNumber="218">
<emphasis id="B9313C7674432452BBEFF84FB731F836" bold="true" box="[151,288,1995,2020]" pageId="13" pageNumber="218">FIGURE 41.</emphasis>
Distribution of '
<taxonomicName id="4C459BE774432452BABFF848B42EF836" authorityName="Franz" authorityYear="1963" box="[455,575,1996,2020]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674432452BABFF848B42EF836" box="[455,575,1996,2020]" italics="true" pageId="13" pageNumber="218">Afroeudesis</emphasis>
</taxonomicName>
group'.
</paragraph>
</caption>
<paragraph id="8BFAE06474402451BBBFFF1CB4F2FE91" blockId="14.[151,1437,152,323]" pageId="14" pageNumber="219">
<emphasis id="B9313C7674402451BBBFFF1CB421FF61" bold="true" box="[199,560,152,179]" pageId="14" pageNumber="219">Composition and distribution.</emphasis>
<taxonomicName id="4C459BE774402451B940FF1DB4ADFF61" authorityName="Franz" authorityYear="1963" box="[568,700,153,179]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B9313C7674402451B940FF1DB4ADFF61" box="[568,700,153,179]" italics="true" pageId="14" pageNumber="219">Afroeudesis</emphasis>
</taxonomicName>
comprises five species in four subgenera:
<taxonomicName id="4C459BE774402451BFE3FF1DB30EFF61" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[1179,1311,153,179]" class="Insecta" family="Staphylinidae" genus="Afroeudesis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B9313C7674402451BFE3FF1DB30EFF61" box="[1179,1311,153,179]" italics="true" pageId="14" pageNumber="219">Afroeudesis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202810D74402451BE5FFF1DB372FF61" box="[1319,1379,153,179]" pageId="14" pageNumber="219" sensu="stricto">s. str.</taxonomicNameLabel>
with one species (
<collectingCountry id="F352A0F474402451BA50FF39B784FF0A" box="[296,405,189,216]" name="Tanzania" pageId="14" pageNumber="219">Tanzania</collectingCountry>
); subgen.
<emphasis id="B9313C7674402451B97EFF39B460FF05" box="[518,625,189,215]" italics="true" pageId="14" pageNumber="219">Eburneus</emphasis>
<bibRefCitation id="EFD49D9574402451B901FF39B55EFF05" author="Jaloszynski, P." box="[633,847,189,215]" pageId="14" pageNumber="219" pagination="163 - 173" refId="ref7905" refString="Jaloszynski, P. (2016 b) Microscydmus trianguliceps Franz and Pseudoeudesis sulcata Franz transferred to Afroeudesis Franz (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4173 (2), 163 - 173. https: // doi. org / 10.11646 / zootaxa. 4173.2.7" type="journal article" year="2016">Jałoszyński, 2016b</bibRefCitation>
with one species (
<collectingCountry id="F352A0F474402451BF58FF39B2BAFF05" box="[1056,1195,189,215]" name="Ivory Coast" pageId="14" pageNumber="219">Ivory Coast</collectingCountry>
); subgen.
<emphasis id="B9313C7674402451BE65FF39B38CFF05" box="[1309,1437,189,215]" italics="true" pageId="14" pageNumber="219">Eudesoides</emphasis>
<bibRefCitation id="EFD49D9574402451BBEFFF65B762FF29" author="Jaloszynski, P." box="[151,371,225,251]" pageId="14" pageNumber="219" pagination="163 - 173" refId="ref7905" refString="Jaloszynski, P. (2016 b) Microscydmus trianguliceps Franz and Pseudoeudesis sulcata Franz transferred to Afroeudesis Franz (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4173 (2), 163 - 173. https: // doi. org / 10.11646 / zootaxa. 4173.2.7" type="journal article" year="2016">Jałoszyński, 2016b</bibRefCitation>
with one species (
<collectingCountry id="F352A0F474402451B920FF65B4B3FF2E" box="[600,674,225,252]" name="Brazil" pageId="14" pageNumber="219">Brazil</collectingCountry>
); and subgen.
<emphasis id="B9313C7674402451B82CFF65B5E1FF29" box="[852,1008,225,251]" italics="true" pageId="14" pageNumber="219">Nanoscydmus</emphasis>
<bibRefCitation id="EFD49D9574402451B884FF65B2D8FF29" author="Jaloszynski, P." box="[1020,1225,225,251]" pageId="14" pageNumber="219" pagination="173 - 178" refId="ref7549" refString="Jaloszynski, P. (2009) Nanoscydmus gen. n. from China and Nepal (Coleoptera: Scydmaenidae). Koleopterologische Rundschau, 79, 173 - 178." type="journal article" year="2009">Jałoszyński, 2009</bibRefCitation>
with two species (
<collectingCountry id="F352A0F474402451BBE7FE81B6F9FECD" box="[159,232,261,287]" name="China" pageId="14" pageNumber="219">China</collectingCountry>
:
<collectingRegion id="49812E8674402451BB8BFE81B75FFECD" box="[243,334,261,287]" country="China" name="Yunnan" pageId="14" pageNumber="219">Yunnan</collectingRegion>
, and
<collectingCountry id="F352A0F474402451BAF2FE81B7DEFEF2" box="[394,463,261,288]" name="Nepal" pageId="14" pageNumber="219">Nepal</collectingCountry>
) (
<figureCitation id="137EFCE174402451BA9EFE81B42BFECD" box="[486,570,261,287]" captionStart="FIGURE 41" captionStartId="13.[151,250,1995,2020]" captionTargetBox="[161,1417,1304,1961]" captionTargetId="figure@13.[151,1436,1269,1969]" captionTargetPageId="13" captionText="FIGURE 41. Distribution of 'Afroeudesis group'." httpUri="https://zenodo.org/record/3234314/files/figure.png" pageId="14" pageNumber="219">Fig. 41</figureCitation>
). A specimen from the northern, sub-Himalayan region of
<collectingCountry id="F352A0F474402451BFAAFE81B31DFECD" box="[1234,1292,261,287]" name="India" pageId="14" pageNumber="219">India</collectingCountry>
(Darjeeling) representing an undetermined species was also seen.
</paragraph>
</subSubSection>
</treatment>
</document>