treatments-xml/data/03/D7/A5/03D7A552F403082DADC45EC6C9CCB3C1.xml
2024-06-21 12:22:17 +02:00

298 lines
61 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="FC498342CD5C560F89399257A961FBC5" ID-CLB-Dataset="31348" ID-DOI="10.11646/zootaxa.4353.2.3" ID-GBIF-Dataset="5dcff406-a135-4ba0-bac3-d56b4f629887" ID-ISSN="1175-5326" ID-Zenodo-Dep="1065166" ID-ZooBank="05D61D56-6043-4A11-B2B5-CBF864299B4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511423831672" checkinUser="plazi" docAuthor="Sankaran, Pradeep M., Malamel, Jobi J., Joseph, Mathew M. &amp; Sebastian, Pothalil A." docDate="2017" docId="03D7A552F403082DADC45EC6C9CCB3C1" docLanguage="en" docName="zootaxa.4353.2.3.pdf" docOrigin="Zootaxa 4353 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tylorida Simon 1894" docType="treatment" docVersion="11" lastPageNumber="296" masterDocId="FFEEDD2AF402082FAD535B47CA5EB761" masterDocTitle="On the genus Tylorida Simon, 1894 with the first record of the genus Atelidea Simon, 1895 from India (Araneae: Tetragnathidae, Leucauginae)" masterLastPageNumber="326" masterPageNumber="294" pageNumber="295" updateTime="1698504197812" updateUser="ExternalLinkService">
<mods:mods id="57A9103A99AA3CAF8FFF936D1840FF89" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7814EA3C7AAB7EB4C0A25E156729CA21">
<mods:title id="794088ED84A9DBFEB4957B1217B00D10">On the genus Tylorida Simon, 1894 with the first record of the genus Atelidea Simon, 1895 from India (Araneae: Tetragnathidae, Leucauginae)</mods:title>
</mods:titleInfo>
<mods:name id="CBE0C784D4DF1E021C0D6F198D161EA2" type="personal">
<mods:role id="6A2F5191AC1E667230E16D6D1AFE447D">
<mods:roleTerm id="76F396338F5F7A3979119315DB95774D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="17149FB673945E7641AEC4BF2C095E6C">Sankaran, Pradeep M.</mods:namePart>
</mods:name>
<mods:name id="713B47A7A728E16BCA37C78CEF5C8A7F" type="personal">
<mods:role id="021B61D640E735894FAC333FE1020677">
<mods:roleTerm id="DF54A2CB594C991C05E96AA251CF2E6D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="161738B4386E2135950EDD12A31D5C3F">Malamel, Jobi J.</mods:namePart>
</mods:name>
<mods:name id="0DF947C6F00222D7E0047284B53E61F0" type="personal">
<mods:role id="BB96BD9A8E33618BB06129704798FA30">
<mods:roleTerm id="79F8390933461ED263359C7EF9F94F86">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="94E6167CD9406229D9985BE238167B09">Joseph, Mathew M.</mods:namePart>
</mods:name>
<mods:name id="E98A0AC78A14558EBEF0C4B0D93063A0" type="personal">
<mods:role id="860D18FD16015EBBAE4946AFA750B730">
<mods:roleTerm id="DC2819CD1932B182DF35D76652AECF37">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EDB87D5E44BB7409CA1FA7668F1432BF">Sebastian, Pothalil A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D5D86ABE6C9131C5A2F30D193A660F9B">text</mods:typeOfResource>
<mods:relatedItem id="341BE24AD115467FA083FE8F749D755F" type="host">
<mods:titleInfo id="4ADD12B678421EBBC62B621182D4548F">
<mods:title id="14C6A28B2FB4A7444C72F37D304044E0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="30F34A4DFDFA09CCD4C77AA82130DDA3">
<mods:date id="4871BD9E5EF2E0CD4B8F95493569FB99">2017</mods:date>
<mods:detail id="7AE98C03A431611DBB1F6536A87BA7D8" type="pubDate">
<mods:number id="73B65432379E93F6596209EB17ED7388">2017-11-23</mods:number>
</mods:detail>
<mods:detail id="03BC5F094BD1A046C937EAA2F8550483" type="volume">
<mods:number id="7FC37986BADA4EA0DFD37C7D4BB33F3D">4353</mods:number>
</mods:detail>
<mods:detail id="AE23AF75717A67173DAF5E8D9CB5F272" type="issue">
<mods:number id="C2035AA8CA0444D5B1CF5B9E5BAE297F">2</mods:number>
</mods:detail>
<mods:extent id="BD8CD4D8A759FF56AD66202FBB695260" unit="page">
<mods:start id="97B2FCA2F5BFD56A052A38F7152934C0">294</mods:start>
<mods:end id="AD90A83274941A8DDCD53CCEAC714F9D">326</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5A6C12825102C32BE0AFDAED6470934D">journal article</mods:classification>
<mods:identifier id="0CDD4A56274BD0D2E019E4DB9A2C960A" type="CLB-Dataset">31348</mods:identifier>
<mods:identifier id="D2B1AC4A75464B85736A516DDFC15142" type="DOI">10.11646/zootaxa.4353.2.3</mods:identifier>
<mods:identifier id="208D7BA5662A299596B3A4B7C6562E4D" type="GBIF-Dataset">5dcff406-a135-4ba0-bac3-d56b4f629887</mods:identifier>
<mods:identifier id="8D58BFC024C183BE0A1D1EDF3D98A94B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="54F9A6368E94B777D800921750E01950" type="Zenodo-Dep">1065166</mods:identifier>
<mods:identifier id="B8F7CC2B37BBA5A597F2C64333513D1C" type="ZooBank">05D61D56-6043-4A11-B2B5-CBF864299B4F</mods:identifier>
</mods:mods>
<treatment id="03D7A552F403082DADC45EC6C9CCB3C1" ID-DOI="http://doi.org/10.5281/zenodo.6051862" ID-GBIF-Taxon="137541616" ID-Zenodo-Dep="6051862" LSID="urn:lsid:plazi:treatment:03D7A552F403082DADC45EC6C9CCB3C1" httpUri="http://treatment.plazi.org/id/03D7A552F403082DADC45EC6C9CCB3C1" lastPageId="2" lastPageNumber="296" pageId="1" pageNumber="295">
<subSubSection id="C36447CFF403082EADC45EC6CBC3B2FA" box="[151,413,1409,1435]" pageId="1" pageNumber="295" type="nomenclature">
<paragraph id="8BC11444F403082EADC45EC6CBC3B2FA" blockId="1.[151,413,1409,1435]" box="[151,413,1409,1435]" pageId="1" pageNumber="295">
<heading id="D089A328F403082EADC45EC6CBC3B2FA" bold="true" box="[151,413,1409,1435]" fontSize="11" level="1" pageId="1" pageNumber="295" reason="1">
<taxonomicName id="4C7E6FC7F403082EADC45EC6CBC3B2FA" ID-CoL="844C" ID-ENA="473179" authority="Simon, 1894" authorityName="Simon" authorityYear="1894" box="[151,413,1409,1435]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F403082EADC45EC6CBC3B2FA" bold="true" box="[151,413,1409,1435]" pageId="1" pageNumber="295">
<emphasis id="B90AC856F403082EADC45EC6CAA2B2FA" bold="true" box="[151,252,1409,1435]" italics="true" pageId="1" pageNumber="295">Tylorida</emphasis>
<bibRefCitation id="EFEF69B5F403082EAC575EC6CBC3B2FA" author="Simon" box="[260,413,1409,1435]" pageId="1" pageNumber="295" refString="Simon, E. (1894) Histoire naturelle des araignees. Tome 1. Roret, Paris, 272 pp. [pp. 489 - 760]" type="book chapter" year="1894">Simon, 1894</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36447CFF403082EADC45E81CBB2B19E" pageId="1" pageNumber="295" type="materials_examined">
<paragraph id="8BC11444F403082EADC45E81CE81B2BE" blockId="1.[151,1436,1478,2007]" box="[151,1247,1478,1503]" pageId="1" pageNumber="295">
<emphasis id="B90AC856F403082EADC45E81CB6AB2BE" bold="true" box="[151,308,1478,1503]" pageId="1" pageNumber="295">Type species:</emphasis>
<taxonomicName id="4C7E6FC7F403082EAC685E81C8C0B2BE" authority="(Thorell, 1877)" authorityName="" authorityYear="1877" baseAuthorityName="Thorell" baseAuthorityYear="1877" box="[315,670,1478,1503]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="295" phylum="Arthropoda" rank="species" species="striata">
<emphasis id="B90AC856F403082EAC685E81CBB4B2BE" box="[315,490,1478,1503]" italics="true" pageId="1" pageNumber="295">Tylorida striata</emphasis>
(
<bibRefCitation id="EFEF69B5F403082EACAB5E81C8CBB2BE" author="Thorell" box="[504,661,1478,1503]" pageId="1" pageNumber="295" refString="Thorell, T. (1877) Studi sui Ragni Malesi e Papuani. I. Ragni di Selebes raccolti nel 1874 dal Dott. O. Beccari. Annali del Museo Civico di Storia Naturale di Genova, 10, 341 - 637." type="journal article" year="1877">Thorell, 1877</bibRefCitation>
)
</taxonomicName>
, originally described as
<taxonomicName id="4C7E6FC7F403082EAEFD5E8FCE84B2BE" authority="Thorell, 1877" authorityName="Thorell" authorityYear="1877" box="[942,1242,1478,1503]" class="Arachnida" family="Tetragnathidae" genus="Meta" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="295" phylum="Arthropoda" rank="species" species="striata">
<emphasis id="B90AC856F403082EAEFD5E8FCE66B2BE" box="[942,1080,1480,1503]" italics="true" pageId="1" pageNumber="295">Meta striata</emphasis>
<bibRefCitation id="EFEF69B5F403082EA96C5E81CE84B2BE" author="Thorell" box="[1087,1242,1478,1503]" pageId="1" pageNumber="295" refString="Thorell, T. (1877) Studi sui Ragni Malesi e Papuani. I. Ragni di Selebes raccolti nel 1874 dal Dott. O. Beccari. Annali del Museo Civico di Storia Naturale di Genova, 10, 341 - 637." type="journal article" year="1877">Thorell, 1877</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC11444F403082EAD945EADCBB2B19E" blockId="1.[151,1436,1478,2007]" pageId="1" pageNumber="295">
<emphasis id="B90AC856F403082EAD945EADCB03B162" bold="true" box="[199,349,1514,1539]" pageId="1" pageNumber="295">Rediagnosis.</emphasis>
Genus
<taxonomicName id="4C7E6FC7F403082EACEF5EADC846B162" authorityName="Simon" authorityYear="1894" box="[444,536,1514,1539]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F403082EACEF5EADC846B162" box="[444,536,1514,1539]" italics="true" pageId="1" pageNumber="295">Tylorida</emphasis>
</taxonomicName>
can be separated from other described tetragnathid genera by the following combination of characters: femora IV with smooth trichobothrial shaft, cymbial dorso-basal process shorter than half the cymbial width and perpendicular to the cymbium longitudinal axis, short, acute embolus with a broad, twisted embolic base, short, lamelliform conductor lying dorsal to embolus, ventrally swollen tegulum with a distomedian triangular process, long, tubular fertilization ducts connected to both copulatory ducts and spermathecal bulb, copulatory and fertilization ducts running parallel before entering into spermathecal bulb (
<bibRefCitation id="EFEF69B5F403082EA99E5DD9CB81B19E" author="Alvarez-Padilla" pageId="1" pageNumber="295" refString="Alvarez-Padilla, F. &amp; Hormiga, G. (2011) Morphological and phylogenetic atlas of the orb-weaving spider family Tetragnathidae (Araneae: Araneoidea). Zoological Journal of the Linnean Society, 162, 713 - 879. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00692. x" type="journal article" year="2011">
Álvarez-Padilla &amp; Hormiga 2011: figs 115F, 116C, 117A, B, F; present data:
<figureCitation id="134508C1F403082EAE785D85C9D9B1BA" box="[811,903,1730,1755]" captionStart="FIGURE 4" captionStartId="8.[151,250,1880,1902]" captionTargetBox="[199,1406,209,1844]" captionTargetId="figure@8.[177,1421,193,1869]" captionTargetPageId="8" captionText="FIGURE 4AH. Tylorida flava sp. nov.. A Subadult male prosoma, retrolateral; B Female prosoma, retrolateral; C Subadult male left chelicera, prolateral; D Female left chelicera, prolateral; EF Epigynum, ventral; GH Internal female genitalia, dorsal.Abbreviations:CD copulatory duct, EP epigynal plate,FD fertilization duct,ll lateral lobeof thepostero-median lobeof epigynal plate, ml median lobe of the postero-median lobe of epigynal plate, pEP postero-median lobe of epigynal plate, SB spermathecal bulb.Scale bars AB, 1 mm; CD, 0.5 mm; EH 0.2 mm." httpUri="https://zenodo.org/record/1065174/files/figure.png" pageId="1" pageNumber="295">Figs 4H</figureCitation>
,
<figureCitation id="134508C1F403082EAEC05D84C984B1BA" box="[915,986,1731,1755]" captionStart="FIGURE 8" captionStartId="12.[151,250,1600,1622]" captionTargetBox="[166,1418,221,1564]" captionTargetId="figure@12.[151,1436,193,1579]" captionTargetPageId="12" captionText="FIGURE 8AH. Tylorida marmorea (Pocock, 1901) comb. nov., male left pedipalp. AD Specimen from Ponmudi (High altitude form): A prolateral; B ventral; C retrolateral; D dorsal; EH Specimen from Akambadam (Low altitude form): E prolateral; F ventral; G retrolateral; H dorsal.Scale bar AH, 1 mm." httpUri="https://zenodo.org/record/1065182/files/figure.png" pageId="1" pageNumber="295">8AH</figureCitation>
,
<figureCitation id="134508C1F403082EAEB55D84CE65B1BA" box="[998,1083,1731,1755]" captionStart="FIGURE 10" captionStartId="14.[151,250,1562,1584]" captionTargetBox="[218,1381,203,1547]" captionTargetId="figure@14.[206,1392,185,1553]" captionTargetPageId="14" captionText="FIGURE 10AI. Tylorida marmorea (Pocock, 1901) comb. nov., male left pedipalp. AD Specimen from Ponmudi (High altitude form): A prolateral; B ventral; C retrolateral; D dorsal; EI Specimen from Akambadam (Low altitude form): E prolateral; F ventral; G &amp; I retrolateral; H dorsal.Abbreviations: C conductor,CDBP cymbial dorso-basal process, dmtT distomediantriangular process of tegulum, E embolus,EB embolicbase, P paracymbium, ST subtegulum, T tegulum, Tr trochanter. Scale bar AI, 1 mm." httpUri="https://zenodo.org/record/1065186/files/figure.png" pageId="1" pageNumber="295">10AH</figureCitation>
,
<figureCitation id="134508C1F403082EA9145D84CE88B1BB" box="[1095,1238,1731,1755]" captionStart="FIGURE 11" captionStartId="15.[151,250,1919,1941]" captionTargetBox="[226,1370,666,1886]" captionTargetId="figure@15.[204,1392,647,1902]" captionTargetPageId="15" captionText="FIGURE 11AF. Tylorida marmorea (Pocock, 1901) comb. nov., female genitalia. AB, E Specimen from Ponmudi (High altitude form): A Epigynum, ventral; B &amp; E Internal genitalia, dorsal; CD, F Specimen from Akambadam (Low altitude form): C Epigynum, ventral; D &amp; F Internal genitalia, dorsal. Abbreviations: CD copulatory duct, EP epigynal plate, FD fertilization duct, pEP postero-median lobe of epigynal plate, SB spermathecal bulb.Scale bar AF, 1 mm." httpUri="https://zenodo.org/record/1065188/files/figure.png" pageId="1" pageNumber="295">11B, D, EF</figureCitation>
,
<figureCitation id="134508C1F403082EA9B25D84CF0CB1BB" box="[1249,1362,1731,1755]" captionStart="FIGURE 14" captionStartId="19.[151,250,1904,1926]" captionTargetBox="[182,1408,202,1866]" captionTargetId="figure@19.[173,1417,193,1886]" captionTargetPageId="19" captionText="FIGURE 14AF. Tylorida striata (Thorell, 1877) from Thalacode, Ernakulam. AD Male left pedipalp: A prolateral; B ventral; C retrolateral; D dorsal; EF Female genitalia: E Epigynum, ventral; F Internal genitalia, dorsal; G Original drawings of Tylorida nicobarensis (Tikader,1977) (=Tylorida striata (Thorell, 1877)) adopted from Tikader(1977) without modification. Scale bars AD, 0.5 mm; EF, 0.2 mm." httpUri="https://zenodo.org/record/1065194/files/figure.png" pageId="1" pageNumber="295">14AD, F</figureCitation>
,
<figureCitation id="134508C1F403082EA80D5D84CB79B19E" captionStart="FIGURE 17" captionStartId="22.[151,250,1841,1863]" captionTargetBox="[191,1395,529,1822]" captionTargetId="figure@22.[186,1410,519,1831]" captionTargetPageId="22" captionText="FIGURE 17AO. Tylorida ventralis (Thorell, 1877): male left pedipalp illustrating intraspecific variations and left chelicerae. AE variety I from Edappady, Kottayam:AD Left pedipalp:A prolateral; B ventral; C retrolateral; D dorsal; E Left chelicera, prolateral; FJ variety II from Pathiramanal Island, Alappuzha: FI Left pedipalp: F prolateral; G ventral; H retrolateral; I dorsal; J Left chelicera, prolateral; variety III from Perumbalam, Alappuzha: KN Left pedipalp: K prolateral; L ventral; M retrolateral; N dorsal; O Left chelicera, prolateral. Scale bars AD, 0.5 mm; E, 1 mm;FO, 0.5 mm." httpUri="https://zenodo.org/record/1065200/files/figure.png" pageId="1" pageNumber="295">17A D, FI, KN</figureCitation>
,
<figureCitation id="134508C1F403082EAC615DA0CB81B19E" box="[306,479,1767,1791]" captionStart="FIGURE 18" captionStartId="23.[151,250,1845,1867]" captionTargetBox="[312,1268,209,1815]" captionTargetId="figure@23.[290,1301,193,1828]" captionTargetPageId="23" captionText="FIGURE 18AO. Tylorida ventralis (Thorell, 1877): female genitalia illustrating intraspecific variations. AF variety II, female with distinct caudal tubercle (AC from Kumarakom, Kottayam and DF from Kulathupuzha, Kollam): AB, DE Epigyna, ventral; C, F Internal female genitalia, dorsal; GI, MO variety III, female without caudal tubercle (GI another female from Kulathupuzha, Kollam and MO from Perumbalam, Alappuzha): GH, MN Epigyna, ventral; I, O Internal female genitalia, dorsal; JL variety I, female with indistinct caudal tubercle (from Edappady, Kottayam): JK Epigyna, ventral; L Internal female genitalia, dorsal. Scale bars AC, JL, 0.2 mm; DI, MO, 0.5 mm." httpUri="https://zenodo.org/record/1065202/files/figure.png" pageId="1" pageNumber="295">18 C, F, I, L, O</figureCitation>
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36447CFF403082DAD945C4DC9CCB3C1" lastPageId="2" lastPageNumber="296" pageId="1" pageNumber="295" type="description">
<paragraph id="8BC11444F403082DAD945C4DC89EB799" blockId="1.[151,1436,1478,2007]" lastBlockId="2.[151,1436,151,1184]" lastPageId="2" lastPageNumber="296" pageId="1" pageNumber="295">
<emphasis id="B90AC856F403082EAD945C4DCB3BB042" bold="true" box="[199,357,1802,1827]" pageId="1" pageNumber="295">Relationship.</emphasis>
Genus
<taxonomicName id="4C7E6FC7F403082EACED5C4DC844B042" authorityName="Simon" authorityYear="1894" box="[446,538,1802,1827]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F403082EACED5C4DC844B042" box="[446,538,1802,1827]" italics="true" pageId="1" pageNumber="295">Tylorida</emphasis>
</taxonomicName>
is closely allied to the genus
<taxonomicName id="4C7E6FC7F403082EAE265C4CC9BDB043" authorityName="Thorell" authorityYear="1890" box="[885,995,1803,1826]" class="Arachnida" family="Tetragnathidae" genus="Orsinome" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F403082EAE265C4CC9BDB043" box="[885,995,1803,1826]" italics="true" pageId="1" pageNumber="295">Orsinome</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFEF69B5F403082EAEA75C4DCF72B042" author="Alvarez-Padilla" box="[1012,1324,1802,1827]" pageId="1" pageNumber="295" refString="Alvarez-Padilla, F., Dimitrov, D., Giribet, G., Hormiga, G. (2009) Phylogenetic relationships of the spider family Tetragnathidae (Araneae, Araneoidea) based on morphological and DNA sequence data. Cladistics, 25, 109 - 146. https: // doi. org / 10.1111 / j. 1096 - 0031.2008.00242. x" type="journal article" year="2009">
Álvarez-Padilla
<emphasis id="B90AC856F403082EA9FD5C4BCEB6B042" box="[1198,1256,1802,1827]" italics="true" pageId="1" pageNumber="295">et al.</emphasis>
2009
</bibRefCitation>
; Álvarez-
<bibRefCitation id="EFEF69B5F403082EADC45C69CB9AB026" author="Padilla" box="[151,452,1838,1863]" pageId="1" pageNumber="295" refString="Alvarez-Padilla, F. &amp; Hormiga, G. (2011) Morphological and phylogenetic atlas of the orb-weaving spider family Tetragnathidae (Araneae: Araneoidea). Zoological Journal of the Linnean Society, 162, 713 - 879. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00692. x" type="journal article" year="2011">Padilla &amp; Hormiga 2011</bibRefCitation>
;
<bibRefCitation id="EFEF69B5F403082EAC845C69C940B026" author="Dimitrov" box="[471,798,1838,1863]" pageId="1" pageNumber="295" refString="Dimitrov, D. &amp; Hormiga, G. (2011) An extraordinary new genus of spiders from Western Australia with an expanded hypothesis on the phylogeny of Tetragnathidae (Araneae). Zoological Journal of the Linnean Society, 161, 735 - 768. https: // doi. org / 10.1111 / j. 1096 - 3642.2010.00662. x" type="journal article" year="2011">Dimitrov &amp; Hormiga 2011</bibRefCitation>
). Members of both these genera are similar in size, colouration (compare
<figureCitation id="134508C1F403082EACC55C15C84CB00A" box="[406,530,1874,1899]" captionStart="FIGURE 5" captionStartId="9.[151,250,1967,1989]" captionTargetBox="[233,1353,204,1937]" captionTargetId="figure@9.[219,1372,193,1949]" captionTargetPageId="9" captionText="FIGURE 5AF. Tylorida marmorea (Pocock, 1901) comb. nov. from Ponmudi (High altitude form). AC Male habitus: A dorsal; B ventral; C prolateral; DF Female habitus:D dorsal; E ventral; F prolateral.Scale bar AF,2 mm." httpUri="https://zenodo.org/record/1065176/files/figure.png" pageId="1" pageNumber="295">Figs 5AF</figureCitation>
,
<figureCitation id="134508C1F403082EAF715C15C837B00A" box="[546,617,1874,1899]" captionStart="FIGURE6" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[163,1423,966,1735]" captionTargetId="figure@10.[151,1436,949,1752]" captionTargetPageId="10" captionText="FIGURE6AD. Tylorida marmorea (Pocock,1901) comb.nov. from Akambadam (Low altitude form).AB Male habitus:A dorsal; B prolateral; CD Femalehabitus:C dorsal; D retrolateral. Scale bar AD, 2 mm." httpUri="https://zenodo.org/record/1065178/files/figure.png" pageId="1" pageNumber="295">6AD</figureCitation>
,
<figureCitation id="134508C1F403082EAF2B5C15C898B00A" box="[632,710,1874,1899]" captionStart="FIGURE 15" captionStartId="20.[151,250,1941,1963]" captionTargetBox="[316,1273,460,1916]" captionTargetId="figure@20.[297,1290,450,1925]" captionTargetPageId="20" captionText="FIGURE 15AF. Tylorida ventralis (Thorell, 1877): variety I, female with indistinct caudal tubercle from Edappady, Kottayam.AC Male habitus: A dorsal; B ventral; C retrolateral; DF Female habitus:D dorsal; E ventral, F retrolateral.Scale bar AF, 2 mm." httpUri="https://zenodo.org/record/1065196/files/figure.png" pageId="1" pageNumber="295">15AF</figureCitation>
with 20AF) and web building behaviour (
<bibRefCitation id="EFEF69B5F403082EA9995C15CBE3B0EE" author="Alvarez-Padilla" pageId="1" pageNumber="295" refString="Alvarez-Padilla, F. &amp; Hormiga, G. (2011) Morphological and phylogenetic atlas of the orb-weaving spider family Tetragnathidae (Araneae: Araneoidea). Zoological Journal of the Linnean Society, 162, 713 - 879. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00692. x" type="journal article" year="2011">Álvarez-Padilla &amp; Hormiga 2011: fig. 5C, E</bibRefCitation>
), making them misplaced within both the genera. Females of both genera share similar cheliceral dentition: three promarginal and four retromarginal teeth (compare
<figureCitation id="134508C1F403082EA9725CDDCEB5B0D3" box="[1057,1259,1946,1971]" captionStart="FIGURE 12" captionStartId="16.[151,250,1073,1095]" captionTargetBox="[168,1421,214,1028]" captionTargetId="figure@16.[151,1436,193,1052]" captionTargetPageId="16" captionText="FIGURE 12AK. Left chelicerae of Tylorida spp. and Orsinome cf. vethi (Hasselt, 1882). AE Left chelicerae of Tylorida marmorea (Pocock, 1901) comb. nov.: A Male (High altitude form), prolateral; B Female (High altitude form), prolateral; C Male (Low altitude form), prolateral; D Female (Low altitude form), prolateral; E Orginal drawings of male and female cheliceraeadopted from Gravely(1921) without modification; FG Left chelicerae of Tylorida striata (Thorell, 1877):F Male, retrolateral; G Female, prolateral; HI Left chelicerae of Tylorida ventralis (Thorell, 1877): H Male, prolateral, arrow indicates tiny tubercle; I Female,ventro-retrolateral;JK Left cheliceraeof Orsinome cf.vethi: J Male, retrolateral; K Female,prolateral. Scale bars AB, HI, 1 mm; CD, 0.5 mm; FG, 0.2 mm; JK, 0.5 mm; E, not to scale." httpUri="https://zenodo.org/record/1065190/files/figure.png" pageId="1" pageNumber="295">Fig. 12B, D, G, I</figureCitation>
with K). Male chelicerae of
<taxonomicName id="4C7E6FC7F403082EAC6B5CF8CBF8B0B7" authorityName="Thorell" authorityYear="1890" box="[312,422,1983,2006]" class="Arachnida" family="Tetragnathidae" genus="Orsinome" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F403082EAC6B5CF8CBF8B0B7" box="[312,422,1983,2006]" italics="true" pageId="1" pageNumber="295">Orsinome</emphasis>
</taxonomicName>
spp., however, significantly differ from that of
<taxonomicName id="4C7E6FC7F403082EAEB05CF9CE61B0B6" authorityName="Simon" authorityYear="1894" box="[995,1087,1982,2007]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F403082EAEB05CF9CE61B0B6" box="[995,1087,1982,2007]" italics="true" pageId="1" pageNumber="295">Tylorida</emphasis>
</taxonomicName>
spp. In the former, the male chelicerae are massive, nearly as wide as long with cheliceral apophysis and less number of teeth, while in the latter, it is longer than wide with numerous teeth and without any apophysis (compare
<figureCitation id="134508C1F400082DA9335BFBCF7CB7B5" box="[1120,1314,188,213]" captionStart="FIGURE 12" captionStartId="16.[151,250,1073,1095]" captionTargetBox="[168,1421,214,1028]" captionTargetId="figure@16.[151,1436,193,1052]" captionTargetPageId="16" captionText="FIGURE 12AK. Left chelicerae of Tylorida spp. and Orsinome cf. vethi (Hasselt, 1882). AE Left chelicerae of Tylorida marmorea (Pocock, 1901) comb. nov.: A Male (High altitude form), prolateral; B Female (High altitude form), prolateral; C Male (Low altitude form), prolateral; D Female (Low altitude form), prolateral; E Orginal drawings of male and female cheliceraeadopted from Gravely(1921) without modification; FG Left chelicerae of Tylorida striata (Thorell, 1877):F Male, retrolateral; G Female, prolateral; HI Left chelicerae of Tylorida ventralis (Thorell, 1877): H Male, prolateral, arrow indicates tiny tubercle; I Female,ventro-retrolateral;JK Left cheliceraeof Orsinome cf.vethi: J Male, retrolateral; K Female,prolateral. Scale bars AB, HI, 1 mm; CD, 0.5 mm; FG, 0.2 mm; JK, 0.5 mm; E, not to scale." httpUri="https://zenodo.org/record/1065190/files/figure.png" pageId="2" pageNumber="296">Fig. 12A, C, F, H</figureCitation>
with J and
<bibRefCitation id="EFEF69B5F400082DADC45B98C84EB799" author="Alvarez-Padilla" box="[151,528,223,248]" pageId="2" pageNumber="296" refString="Alvarez-Padilla, F. &amp; Hormiga, G. (2011) Morphological and phylogenetic atlas of the orb-weaving spider family Tetragnathidae (Araneae: Araneoidea). Zoological Journal of the Linnean Society, 162, 713 - 879. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00692. x" type="journal article" year="2011">Álvarez-Padilla &amp; Hormiga 2011</bibRefCitation>
: fig. 101D, H).
</paragraph>
<paragraph id="8BC11444F400082DAD945A43CE0BB5E5" blockId="2.[151,1436,151,1184]" pageId="2" pageNumber="296">
Members of both the genera differ significantly in male and female genitalia. Whereas in
<taxonomicName id="4C7E6FC7F400082DA9875A43CF6EB67C" authorityName="Simon" authorityYear="1894" box="[1236,1328,260,285]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F400082DA9875A43CF6EB67C" box="[1236,1328,260,285]" italics="true" pageId="2" pageNumber="296">Tylorida</emphasis>
</taxonomicName>
spp., the pedipalp possesses a short embolus, short, lamelliform conductor, that clearly separated from embolus, distomedian triangular process of tegulum and basally placed cymbial dorso-basal process, the pedipalp of
<taxonomicName id="4C7E6FC7F400082DA87E5A0ACFC5B605" authorityName="Thorell" authorityYear="1890" box="[1325,1435,333,356]" class="Arachnida" family="Tetragnathidae" genus="Orsinome" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F400082DA87E5A0ACFC5B605" box="[1325,1435,333,356]" italics="true" pageId="2" pageNumber="296">Orsinome</emphasis>
</taxonomicName>
spp. share features such as long embolus, long conductor, which appears contiguous to embolus, tegulum without disto-median triangular process and a more or less distally placed cymbial dorso-basal process (compare
<figureCitation id="134508C1F400082DA8745AD3CAFCB6AE" captionStart="FIGURE 10" captionStartId="14.[151,250,1562,1584]" captionTargetBox="[218,1381,203,1547]" captionTargetId="figure@14.[206,1392,185,1553]" captionTargetPageId="14" captionText="FIGURE 10AI. Tylorida marmorea (Pocock, 1901) comb. nov., male left pedipalp. AD Specimen from Ponmudi (High altitude form): A prolateral; B ventral; C retrolateral; D dorsal; EI Specimen from Akambadam (Low altitude form): E prolateral; F ventral; G &amp; I retrolateral; H dorsal.Abbreviations: C conductor,CDBP cymbial dorso-basal process, dmtT distomediantriangular process of tegulum, E embolus,EB embolicbase, P paracymbium, ST subtegulum, T tegulum, Tr trochanter. Scale bar AI, 1 mm." httpUri="https://zenodo.org/record/1065186/files/figure.png" pageId="2" pageNumber="296">Figs 10A I</figureCitation>
,
<figureCitation id="134508C1F400082DADE15AFFCB59B6B1" box="[178,263,440,464]" captionStart="FIGURE 14" captionStartId="19.[151,250,1904,1926]" captionTargetBox="[182,1408,202,1866]" captionTargetId="figure@19.[173,1417,193,1886]" captionTargetPageId="19" captionText="FIGURE 14AF. Tylorida striata (Thorell, 1877) from Thalacode, Ernakulam. AD Male left pedipalp: A prolateral; B ventral; C retrolateral; D dorsal; EF Female genitalia: E Epigynum, ventral; F Internal genitalia, dorsal; G Original drawings of Tylorida nicobarensis (Tikader,1977) (=Tylorida striata (Thorell, 1877)) adopted from Tikader(1977) without modification. Scale bars AD, 0.5 mm; EF, 0.2 mm." httpUri="https://zenodo.org/record/1065194/files/figure.png" pageId="2" pageNumber="296">14AD</figureCitation>
,
<figureCitation id="134508C1F400082DAC455AFFCBB5B6AE" box="[278,491,440,464]" captionStart="FIGURE 17" captionStartId="22.[151,250,1841,1863]" captionTargetBox="[191,1395,529,1822]" captionTargetId="figure@22.[186,1410,519,1831]" captionTargetPageId="22" captionText="FIGURE 17AO. Tylorida ventralis (Thorell, 1877): male left pedipalp illustrating intraspecific variations and left chelicerae. AE variety I from Edappady, Kottayam:AD Left pedipalp:A prolateral; B ventral; C retrolateral; D dorsal; E Left chelicera, prolateral; FJ variety II from Pathiramanal Island, Alappuzha: FI Left pedipalp: F prolateral; G ventral; H retrolateral; I dorsal; J Left chelicera, prolateral; variety III from Perumbalam, Alappuzha: KN Left pedipalp: K prolateral; L ventral; M retrolateral; N dorsal; O Left chelicera, prolateral. Scale bars AD, 0.5 mm; E, 1 mm;FO, 0.5 mm." httpUri="https://zenodo.org/record/1065200/files/figure.png" pageId="2" pageNumber="296">17AD, FI, KN</figureCitation>
, with 21AE,
<bibRefCitation id="EFEF69B5F400082DAFCB5AF0CE45B6B1" author="Alvarez-Padilla" box="[664,1051,439,464]" pageId="2" pageNumber="296" refString="Alvarez-Padilla, F. &amp; Hormiga, G. (2011) Morphological and phylogenetic atlas of the orb-weaving spider family Tetragnathidae (Araneae: Araneoidea). Zoological Journal of the Linnean Society, 162, 713 - 879. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00692. x" type="journal article" year="2011">Álvarez-Padilla &amp; Hormiga 2011</bibRefCitation>
: fig. 103A, F). In contrast to the epigyna of
<taxonomicName id="4C7E6FC7F400082DAC485A9BCB29B694" authorityName="Simon" authorityYear="1894" box="[283,375,476,501]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F400082DAC485A9BCB29B694" box="[283,375,476,501]" italics="true" pageId="2" pageNumber="296">Tylorida</emphasis>
</taxonomicName>
spp., which have prominent epigynal plate, widely separated copulatory openings and very long, tubular fertilization ducts, the epigyna of
<taxonomicName id="4C7E6FC7F400082DAFE35947C941B576" authorityName="Thorell" authorityYear="1890" box="[688,799,512,535]" class="Arachnida" family="Tetragnathidae" genus="Orsinome" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F400082DAFE35947C941B576" box="[688,799,512,535]" italics="true" pageId="2" pageNumber="296">Orsinome</emphasis>
</taxonomicName>
spp. are characterised by less prominent epigynal plate, short, thick fertilization ducts and having closely placed/contiguous copulatory openings placed in a common chamber (genital opening) at the centre of the epigynal plate (compare
<figureCitation id="134508C1F400082DAEE65900CE64B501" box="[949,1082,583,608]" captionStart="FIGURE 11" captionStartId="15.[151,250,1919,1941]" captionTargetBox="[226,1370,666,1886]" captionTargetId="figure@15.[204,1392,647,1902]" captionTargetPageId="15" captionText="FIGURE 11AF. Tylorida marmorea (Pocock, 1901) comb. nov., female genitalia. AB, E Specimen from Ponmudi (High altitude form): A Epigynum, ventral; B &amp; E Internal genitalia, dorsal; CD, F Specimen from Akambadam (Low altitude form): C Epigynum, ventral; D &amp; F Internal genitalia, dorsal. Abbreviations: CD copulatory duct, EP epigynal plate, FD fertilization duct, pEP postero-median lobe of epigynal plate, SB spermathecal bulb.Scale bar AF, 1 mm." httpUri="https://zenodo.org/record/1065188/files/figure.png" pageId="2" pageNumber="296">Figs 11AF</figureCitation>
,
<figureCitation id="134508C1F400082DA915590FCECFB501" box="[1094,1169,584,608]" captionStart="FIGURE 14" captionStartId="19.[151,250,1904,1926]" captionTargetBox="[182,1408,202,1866]" captionTargetId="figure@19.[173,1417,193,1886]" captionTargetPageId="19" captionText="FIGURE 14AF. Tylorida striata (Thorell, 1877) from Thalacode, Ernakulam. AD Male left pedipalp: A prolateral; B ventral; C retrolateral; D dorsal; EF Female genitalia: E Epigynum, ventral; F Internal genitalia, dorsal; G Original drawings of Tylorida nicobarensis (Tikader,1977) (=Tylorida striata (Thorell, 1877)) adopted from Tikader(1977) without modification. Scale bars AD, 0.5 mm; EF, 0.2 mm." httpUri="https://zenodo.org/record/1065194/files/figure.png" pageId="2" pageNumber="296">14EF</figureCitation>
,
<figureCitation id="134508C1F400082DA9CF590FCEB1B501" box="[1180,1263,584,608]" captionStart="FIGURE 18" captionStartId="23.[151,250,1845,1867]" captionTargetBox="[312,1268,209,1815]" captionTargetId="figure@23.[290,1301,193,1828]" captionTargetPageId="23" captionText="FIGURE 18AO. Tylorida ventralis (Thorell, 1877): female genitalia illustrating intraspecific variations. AF variety II, female with distinct caudal tubercle (AC from Kumarakom, Kottayam and DF from Kulathupuzha, Kollam): AB, DE Epigyna, ventral; C, F Internal female genitalia, dorsal; GI, MO variety III, female without caudal tubercle (GI another female from Kulathupuzha, Kollam and MO from Perumbalam, Alappuzha): GH, MN Epigyna, ventral; I, O Internal female genitalia, dorsal; JL variety I, female with indistinct caudal tubercle (from Edappady, Kottayam): JK Epigyna, ventral; L Internal female genitalia, dorsal. Scale bars AC, JL, 0.2 mm; DI, MO, 0.5 mm." httpUri="https://zenodo.org/record/1065202/files/figure.png" pageId="2" pageNumber="296">18AO</figureCitation>
with
<figureCitation id="134508C1F400082DA87C5900CAF5B5E5" captionStart="FIGURE 21" captionStartId="26.[151,250,1564,1586]" captionTargetBox="[192,1388,214,1533]" captionTargetId="figure@26.[166,1419,187,1548]" captionTargetPageId="26" captionText="FIGURE 21AG. Orsinome cf. vethi (Hasselt, 1882). AE Male left pedipalp: A prolateral; B ventral; C, E retrolateral; D dorsal; FG Female genitalia: F Epigynum, ventral; G Internal genitalia, dorsal. Abbreviations: C conductor, CD copulatory duct, CDBP cymbial dorso-basal process, CO copulatory opening, E embolus, EB embolic base, FD fertilization duct, GO genital opening, P paracymbium, SB spermathecal bulb, ST subtegulum, T tegulum, Tr trochanter.Scale bar AG 0.5 mm." httpUri="https://zenodo.org/record/1065208/files/figure.png" pageId="2" pageNumber="296">Fig. 21F G</figureCitation>
,
<bibRefCitation id="EFEF69B5F400082DADE5592BCB3BB5E5" author="Zhu" box="[182,357,620,645]" pageId="2" pageNumber="296" refString="Zhu, M. S., Song, D. X. &amp; Zhang, J. X. (2003) Fauna Sinica: Invertebrata. Vol. 35. Arachnida: Araneae: Tetragnathidae. Science Press, Beijing, vii + 418 pp." type="book" year="2003">
Zhu
<emphasis id="B90AC856F400082DADB9592ACB7CB5E4" box="[234,290,620,645]" italics="true" pageId="2" pageNumber="296">et al.</emphasis>
2003
</bibRefCitation>
: fig. 164D, E,
<bibRefCitation id="EFEF69B5F400082DAF5B592BC9DCB5E5" author="Alvarez-Padilla" box="[520,898,620,645]" pageId="2" pageNumber="296" refString="Alvarez-Padilla, F. &amp; Hormiga, G. (2011) Morphological and phylogenetic atlas of the orb-weaving spider family Tetragnathidae (Araneae: Araneoidea). Zoological Journal of the Linnean Society, 162, 713 - 879. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00692. x" type="journal article" year="2011">Álvarez-Padilla &amp; Hormiga 2011</bibRefCitation>
: figs 102A, 104E).
</paragraph>
<paragraph id="8BC11444F400082DAD9459C8C9CCB3C1" blockId="2.[151,1436,151,1184]" pageId="2" pageNumber="296">
<emphasis id="B90AC856F400082DAD9459C8C92DB5C9" bold="true" box="[199,883,655,680]" pageId="2" pageNumber="296">
Taxonomic significance of male chelicerae of
<taxonomicName id="4C7E6FC7F400082DAE5F59C8C933B5C9" authorityName="Simon" authorityYear="1894" box="[780,877,655,680]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F400082DAE5F59C8C933B5C9" bold="true" box="[780,877,655,680]" italics="true" pageId="2" pageNumber="296">Tylorida</emphasis>
</taxonomicName>
.
</emphasis>
Most tetragnathid males are characterised by conspicuously enlarged chelicerae, often armoured with numerous teeth and specialized cheliceral apophyses, which are used for
<emphasis id="B90AC856F400082DAC275990C876B591" box="[372,552,727,752]" italics="true" pageId="2" pageNumber="296">cheliceral clasp</emphasis>
during courtship and mating (
<bibRefCitation id="EFEF69B5F400082DAED05990CE70B591" author="Bristowe" box="[899,1070,727,752]" pageId="2" pageNumber="296" refString="Bristowe, W. S. (1958) The World of Spiders. Collins, London, xii + 304 pp." type="book" year="1958">Bristowe 1958</bibRefCitation>
;
<bibRefCitation id="EFEF69B5F400082DA9685990CF0DB591" author="Eberhard" box="[1083,1363,727,752]" pageId="2" pageNumber="296" refString="Eberhard, W. G., Huber, B. A. (1998) Courtship, copulation and sperm transfer in Leucauge mariana (Araneae, Tetragnathidae). Journal of Arachnology, 26, 342 - 368." type="journal article" year="1998">Eberhard &amp; Huber 1998</bibRefCitation>
). Like members of the genus
<taxonomicName id="4C7E6FC7F400082DACCB59BBC88FB475" authority="Latreille, 1804" authorityName="Latreille" authorityYear="1804" box="[408,721,764,789]" class="Arachnida" family="Tetragnathidae" genus="Tetragnatha" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F400082DACCB59BBC87EB474" box="[408,544,764,789]" italics="true" pageId="2" pageNumber="296">Tetragnatha</emphasis>
Latreille, 1804
</taxonomicName>
, whose chelicerae are taxonomically significant in systematics, the chelicerae of male
<taxonomicName id="4C7E6FC7F400082DACC75858CBAEB459" authorityName="Simon" authorityYear="1894" box="[404,496,799,824]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F400082DACC75858CBAEB459" box="[404,496,799,824]" italics="true" pageId="2" pageNumber="296">Tylorida</emphasis>
</taxonomicName>
spp. are useful in species identification and in the case of
<taxonomicName id="4C7E6FC7F400082DA92D5858CE84B459" authorityName="Simon" authorityYear="1894" box="[1150,1242,799,824]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F400082DA92D5858CE84B459" box="[1150,1242,799,824]" italics="true" pageId="2" pageNumber="296">Tylorida</emphasis>
</taxonomicName>
, it is particularly significant as the members are extremely similar to each other in both male (
<figureCitation id="134508C1F400082DAEA45803CE29B43D" box="[1015,1143,836,861]" captionStart="FIGURE 8" captionStartId="12.[151,250,1600,1622]" captionTargetBox="[166,1418,221,1564]" captionTargetId="figure@12.[151,1436,193,1579]" captionTargetPageId="12" captionText="FIGURE 8AH. Tylorida marmorea (Pocock, 1901) comb. nov., male left pedipalp. AD Specimen from Ponmudi (High altitude form): A prolateral; B ventral; C retrolateral; D dorsal; EH Specimen from Akambadam (Low altitude form): E prolateral; F ventral; G retrolateral; H dorsal.Scale bar AH, 1 mm." httpUri="https://zenodo.org/record/1065182/files/figure.png" pageId="2" pageNumber="296">Figs 8AH</figureCitation>
,
<figureCitation id="134508C1F400082DA9D05803CE86B43D" box="[1155,1240,836,860]" captionStart="FIGURE 14" captionStartId="19.[151,250,1904,1926]" captionTargetBox="[182,1408,202,1866]" captionTargetId="figure@19.[173,1417,193,1886]" captionTargetPageId="19" captionText="FIGURE 14AF. Tylorida striata (Thorell, 1877) from Thalacode, Ernakulam. AD Male left pedipalp: A prolateral; B ventral; C retrolateral; D dorsal; EF Female genitalia: E Epigynum, ventral; F Internal genitalia, dorsal; G Original drawings of Tylorida nicobarensis (Tikader,1977) (=Tylorida striata (Thorell, 1877)) adopted from Tikader(1977) without modification. Scale bars AD, 0.5 mm; EF, 0.2 mm." httpUri="https://zenodo.org/record/1065194/files/figure.png" pageId="2" pageNumber="296">14AD</figureCitation>
,
<figureCitation id="134508C1F400082DA9B75803CAF3B4E1" captionStart="FIGURE 17" captionStartId="22.[151,250,1841,1863]" captionTargetBox="[191,1395,529,1822]" captionTargetId="figure@22.[186,1410,519,1831]" captionTargetPageId="22" captionText="FIGURE 17AO. Tylorida ventralis (Thorell, 1877): male left pedipalp illustrating intraspecific variations and left chelicerae. AE variety I from Edappady, Kottayam:AD Left pedipalp:A prolateral; B ventral; C retrolateral; D dorsal; E Left chelicera, prolateral; FJ variety II from Pathiramanal Island, Alappuzha: FI Left pedipalp: F prolateral; G ventral; H retrolateral; I dorsal; J Left chelicera, prolateral; variety III from Perumbalam, Alappuzha: KN Left pedipalp: K prolateral; L ventral; M retrolateral; N dorsal; O Left chelicera, prolateral. Scale bars AD, 0.5 mm; E, 1 mm;FO, 0.5 mm." httpUri="https://zenodo.org/record/1065200/files/figure.png" pageId="2" pageNumber="296">17AD, FI, K N</figureCitation>
) and female (
<figureCitation id="134508C1F400082DAC155820CB98B4E1" box="[326,454,871,896]" captionStart="FIGURE 9" captionStartId="13.[151,250,1723,1745]" captionTargetBox="[187,1419,448,1670]" captionTargetId="figure@13.[151,1436,419,1687]" captionTargetPageId="13" captionText="FIGURE 9AD. Tylorida marmorea (Pocock, 1901) comb. nov., female genitalia. AB Specimen from Ponmudi (High altitude form): A Epigynum, ventral; B Internal genitalia, dorsal; CD Specimen from Akambadam (Low altitude form): C Epigynum, ventral; D Internal genitalia, dorsal.Scale bars AB, 1 mm; CD, 0.5 mm." httpUri="https://zenodo.org/record/1065184/files/figure.png" pageId="2" pageNumber="296">Figs 9AD</figureCitation>
,
<figureCitation id="134508C1F400082DAC82582FC842B4E1" box="[465,540,872,896]" captionStart="FIGURE 14" captionStartId="19.[151,250,1904,1926]" captionTargetBox="[182,1408,202,1866]" captionTargetId="figure@19.[173,1417,193,1886]" captionTargetPageId="19" captionText="FIGURE 14AF. Tylorida striata (Thorell, 1877) from Thalacode, Ernakulam. AD Male left pedipalp: A prolateral; B ventral; C retrolateral; D dorsal; EF Female genitalia: E Epigynum, ventral; F Internal genitalia, dorsal; G Original drawings of Tylorida nicobarensis (Tikader,1977) (=Tylorida striata (Thorell, 1877)) adopted from Tikader(1977) without modification. Scale bars AD, 0.5 mm; EF, 0.2 mm." httpUri="https://zenodo.org/record/1065194/files/figure.png" pageId="2" pageNumber="296">14EF</figureCitation>
,
<figureCitation id="134508C1F400082DAF7B582FC823B4E1" box="[552,637,872,896]" captionStart="FIGURE 18" captionStartId="23.[151,250,1845,1867]" captionTargetBox="[312,1268,209,1815]" captionTargetId="figure@23.[290,1301,193,1828]" captionTargetPageId="23" captionText="FIGURE 18AO. Tylorida ventralis (Thorell, 1877): female genitalia illustrating intraspecific variations. AF variety II, female with distinct caudal tubercle (AC from Kumarakom, Kottayam and DF from Kulathupuzha, Kollam): AB, DE Epigyna, ventral; C, F Internal female genitalia, dorsal; GI, MO variety III, female without caudal tubercle (GI another female from Kulathupuzha, Kollam and MO from Perumbalam, Alappuzha): GH, MN Epigyna, ventral; I, O Internal female genitalia, dorsal; JL variety I, female with indistinct caudal tubercle (from Edappady, Kottayam): JK Epigyna, ventral; L Internal female genitalia, dorsal. Scale bars AC, JL, 0.2 mm; DI, MO, 0.5 mm." httpUri="https://zenodo.org/record/1065202/files/figure.png" pageId="2" pageNumber="296">18AO</figureCitation>
,
<bibRefCitation id="EFEF69B5F400082DAFDB582FC986B4E1" author="Jager" box="[648,984,871,896]" pageId="2" pageNumber="296" refString="Jager, P. &amp; Praxaysombath, B. (2009) Spiders from Laos: new species and new records (Arachnida: Araneae). Acta Arachnologica, Tokyo, 58, 27 - 51. https: // doi. org / 10.2476 / asjaa. 58.27" type="journal article" year="2009">Jäger &amp; Praxaysombath 2009</bibRefCitation>
: fig. 25) genitalic aspects and exhibiting intense intraspecific variations, both somatic as well as genitalic (
<figureCitation id="134508C1F400082DAE2858CBC9AFB4C5" box="[891,1009,908,933]" captionStart="FIGURE 1" captionStartId="5.[151,250,1861,1883]" captionTargetBox="[188,1398,206,1835]" captionTargetId="figure@5.[181,1405,195,1844]" captionTargetPageId="5" captionText="FIGURE 1AF. Field photographs of Tylorida spp. A Female Tylorida flava sp. nov. from Illithodu, Ernakulam (ADSH100454); BC Tylorida marmorea (Pocock, 1901) comb. nov. from Ponmudi, Trivandrum (ADSH100458): B Male; C Female;D Femaleof Tylorida striata (Thorell, 1877)from MuriyadKol Wetland, Thrissur,Kerala (CATE99462); EFFemales of Tylorida ventralis (Thorell, 1877): E Silver morph from Pathiramanal Island, Alappuzha (ADSH1004515); F Yellow morph from Kanakappalam, Kottayam (ADSH1004518). Photo credit A &amp; E, Pradeep M. Sankaran; B &amp; C, Jimmy Paul; D, Nafin K.S.; F, Dhruv A. Prajapati." httpUri="https://zenodo.org/record/1065168/files/figure.png" pageId="2" pageNumber="296">Figs 1EF</figureCitation>
,
<figureCitation id="134508C1F400082DAEAE58CBCE60B4C4" box="[1021,1086,908,933]" captionStart="FIGURE 5" captionStartId="9.[151,250,1967,1989]" captionTargetBox="[233,1353,204,1937]" captionTargetId="figure@9.[219,1372,193,1949]" captionTargetPageId="9" captionText="FIGURE 5AF. Tylorida marmorea (Pocock, 1901) comb. nov. from Ponmudi (High altitude form). AC Male habitus: A dorsal; B ventral; C prolateral; DF Female habitus:D dorsal; E ventral; F prolateral.Scale bar AF,2 mm." httpUri="https://zenodo.org/record/1065176/files/figure.png" pageId="2" pageNumber="296">5AF</figureCitation>
,
<figureCitation id="134508C1F400082DA91958CBCECFB4C4" box="[1098,1169,908,933]" captionStart="FIGURE6" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[163,1423,966,1735]" captionTargetId="figure@10.[151,1436,949,1752]" captionTargetPageId="10" captionText="FIGURE6AD. Tylorida marmorea (Pocock,1901) comb.nov. from Akambadam (Low altitude form).AB Male habitus:A dorsal; B prolateral; CD Femalehabitus:C dorsal; D retrolateral. Scale bar AD, 2 mm." httpUri="https://zenodo.org/record/1065178/files/figure.png" pageId="2" pageNumber="296">6AD</figureCitation>
,
<figureCitation id="134508C1F400082DA9CE58CBCEBDB4C5" box="[1181,1251,908,932]" captionStart="FIGURE 8" captionStartId="12.[151,250,1600,1622]" captionTargetBox="[166,1418,221,1564]" captionTargetId="figure@12.[151,1436,193,1579]" captionTargetPageId="12" captionText="FIGURE 8AH. Tylorida marmorea (Pocock, 1901) comb. nov., male left pedipalp. AD Specimen from Ponmudi (High altitude form): A prolateral; B ventral; C retrolateral; D dorsal; EH Specimen from Akambadam (Low altitude form): E prolateral; F ventral; G retrolateral; H dorsal.Scale bar AH, 1 mm." httpUri="https://zenodo.org/record/1065182/files/figure.png" pageId="2" pageNumber="296">8AH</figureCitation>
,
<figureCitation id="134508C1F400082DA9A358CBCF69B4C5" box="[1264,1335,908,932]" captionStart="FIGURE 9" captionStartId="13.[151,250,1723,1745]" captionTargetBox="[187,1419,448,1670]" captionTargetId="figure@13.[151,1436,419,1687]" captionTargetPageId="13" captionText="FIGURE 9AD. Tylorida marmorea (Pocock, 1901) comb. nov., female genitalia. AB Specimen from Ponmudi (High altitude form): A Epigynum, ventral; B Internal genitalia, dorsal; CD Specimen from Akambadam (Low altitude form): C Epigynum, ventral; D Internal genitalia, dorsal.Scale bars AB, 1 mm; CD, 0.5 mm." httpUri="https://zenodo.org/record/1065184/files/figure.png" pageId="2" pageNumber="296">9AD</figureCitation>
,
<figureCitation id="134508C1F400082DA81058CBCFC6B4C5" box="[1347,1432,908,932]" captionStart="FIGURE 10" captionStartId="14.[151,250,1562,1584]" captionTargetBox="[218,1381,203,1547]" captionTargetId="figure@14.[206,1392,185,1553]" captionTargetPageId="14" captionText="FIGURE 10AI. Tylorida marmorea (Pocock, 1901) comb. nov., male left pedipalp. AD Specimen from Ponmudi (High altitude form): A prolateral; B ventral; C retrolateral; D dorsal; EI Specimen from Akambadam (Low altitude form): E prolateral; F ventral; G &amp; I retrolateral; H dorsal.Abbreviations: C conductor,CDBP cymbial dorso-basal process, dmtT distomediantriangular process of tegulum, E embolus,EB embolicbase, P paracymbium, ST subtegulum, T tegulum, Tr trochanter. Scale bar AI, 1 mm." httpUri="https://zenodo.org/record/1065186/files/figure.png" pageId="2" pageNumber="296">10AH</figureCitation>
,
<figureCitation id="134508C1F400082DADC458F7CABAB4A9" box="[151,228,944,968]" captionStart="FIGURE 11" captionStartId="15.[151,250,1919,1941]" captionTargetBox="[226,1370,666,1886]" captionTargetId="figure@15.[204,1392,647,1902]" captionTargetPageId="15" captionText="FIGURE 11AF. Tylorida marmorea (Pocock, 1901) comb. nov., female genitalia. AB, E Specimen from Ponmudi (High altitude form): A Epigynum, ventral; B &amp; E Internal genitalia, dorsal; CD, F Specimen from Akambadam (Low altitude form): C Epigynum, ventral; D &amp; F Internal genitalia, dorsal. Abbreviations: CD copulatory duct, EP epigynal plate, FD fertilization duct, pEP postero-median lobe of epigynal plate, SB spermathecal bulb.Scale bar AF, 1 mm." httpUri="https://zenodo.org/record/1065188/files/figure.png" pageId="2" pageNumber="296">11AF</figureCitation>
,
<figureCitation id="134508C1F400082DADA058F7CB16B4A9" box="[243,328,944,968]" captionStart="FIGURE 12" captionStartId="16.[151,250,1073,1095]" captionTargetBox="[168,1421,214,1028]" captionTargetId="figure@16.[151,1436,193,1052]" captionTargetPageId="16" captionText="FIGURE 12AK. Left chelicerae of Tylorida spp. and Orsinome cf. vethi (Hasselt, 1882). AE Left chelicerae of Tylorida marmorea (Pocock, 1901) comb. nov.: A Male (High altitude form), prolateral; B Female (High altitude form), prolateral; C Male (Low altitude form), prolateral; D Female (Low altitude form), prolateral; E Orginal drawings of male and female cheliceraeadopted from Gravely(1921) without modification; FG Left chelicerae of Tylorida striata (Thorell, 1877):F Male, retrolateral; G Female, prolateral; HI Left chelicerae of Tylorida ventralis (Thorell, 1877): H Male, prolateral, arrow indicates tiny tubercle; I Female,ventro-retrolateral;JK Left cheliceraeof Orsinome cf.vethi: J Male, retrolateral; K Female,prolateral. Scale bars AB, HI, 1 mm; CD, 0.5 mm; FG, 0.2 mm; JK, 0.5 mm; E, not to scale." httpUri="https://zenodo.org/record/1065190/files/figure.png" pageId="2" pageNumber="296">12AD</figureCitation>
,
<figureCitation id="134508C1F400082DAC0558E8CBFCB4A9" box="[342,418,943,968]" captionStart="FIGURE 16" captionStartId="21.[151,250,1423,1445]" captionTargetBox="[195,1390,195,1369]" captionTargetId="figure@21.[184,1405,182,1378]" captionTargetPageId="21" captionText="FIGURE 16AJ. Tylorida ventralis(Thorell, 1877):variety II, female with distinct caudal tubercle (Figs AE) and variety III, female without caudal tubercle (Figs FJ).AB, FG Male habitus:A, F dorsal; B,G retrolateral; CE, HJ Femalehabitus:C, H dorsal; D prolateral;E, IJ retrolateral.ADfrom Pathiramanal Island, Alappuzha, Efrom Kumarakom,Kottayam, FIfrom Perumbalam, Alappuzha, J from Kulathupuzha, Kollam.Scale bar AJ, 2 mm." httpUri="https://zenodo.org/record/1065198/files/figure.png" pageId="2" pageNumber="296">16AJ</figureCitation>
,
<figureCitation id="134508C1F400082DACE258F7C8D8B4A6" box="[433,646,944,968]" captionStart="FIGURE 17" captionStartId="22.[151,250,1841,1863]" captionTargetBox="[191,1395,529,1822]" captionTargetId="figure@22.[186,1410,519,1831]" captionTargetPageId="22" captionText="FIGURE 17AO. Tylorida ventralis (Thorell, 1877): male left pedipalp illustrating intraspecific variations and left chelicerae. AE variety I from Edappady, Kottayam:AD Left pedipalp:A prolateral; B ventral; C retrolateral; D dorsal; E Left chelicera, prolateral; FJ variety II from Pathiramanal Island, Alappuzha: FI Left pedipalp: F prolateral; G ventral; H retrolateral; I dorsal; J Left chelicera, prolateral; variety III from Perumbalam, Alappuzha: KN Left pedipalp: K prolateral; L ventral; M retrolateral; N dorsal; O Left chelicera, prolateral. Scale bars AD, 0.5 mm; E, 1 mm;FO, 0.5 mm." httpUri="https://zenodo.org/record/1065200/files/figure.png" pageId="2" pageNumber="296">17AD, FI, KN</figureCitation>
,
<figureCitation id="134508C1F400082DAFC658F7C8B4B4A9" box="[661,746,944,968]" captionStart="FIGURE 18" captionStartId="23.[151,250,1845,1867]" captionTargetBox="[312,1268,209,1815]" captionTargetId="figure@23.[290,1301,193,1828]" captionTargetPageId="23" captionText="FIGURE 18AO. Tylorida ventralis (Thorell, 1877): female genitalia illustrating intraspecific variations. AF variety II, female with distinct caudal tubercle (AC from Kumarakom, Kottayam and DF from Kulathupuzha, Kollam): AB, DE Epigyna, ventral; C, F Internal female genitalia, dorsal; GI, MO variety III, female without caudal tubercle (GI another female from Kulathupuzha, Kollam and MO from Perumbalam, Alappuzha): GH, MN Epigyna, ventral; I, O Internal female genitalia, dorsal; JL variety I, female with indistinct caudal tubercle (from Edappady, Kottayam): JK Epigyna, ventral; L Internal female genitalia, dorsal. Scale bars AC, JL, 0.2 mm; DI, MO, 0.5 mm." httpUri="https://zenodo.org/record/1065202/files/figure.png" pageId="2" pageNumber="296">18AO</figureCitation>
,
<bibRefCitation id="EFEF69B5F400082DAFAA58F7CE0FB4A9" author="Jager" box="[761,1105,943,968]" pageId="2" pageNumber="296" refString="Jager, P. &amp; Praxaysombath, B. (2009) Spiders from Laos: new species and new records (Arachnida: Araneae). Acta Arachnologica, Tokyo, 58, 27 - 51. https: // doi. org / 10.2476 / asjaa. 58.27" type="journal article" year="2009">Jäger &amp; Praxaysombath 2009</bibRefCitation>
: figs 24, 2831, 33, 3637), which will make species identification hard. We have examined
<specimenCount id="9D78DFCDF400082DAECE5893C9B1B48C" box="[925,1007,980,1005]" pageId="2" pageNumber="296" type="male">4 male</specimenCount>
specimens of
<taxonomicName id="4C7E6FC7F400082DA9F25892CF6BB48D" authorityName="Sankaran &amp; Malamel &amp; Joseph &amp; Sebastian" authorityYear="2017" baseAuthorityName="Pocock" baseAuthorityYear="1901" box="[1185,1333,981,1004]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="marmorea">
<emphasis id="B90AC856F400082DA9F25892CF6BB48D" box="[1185,1333,981,1004]" italics="true" pageId="2" pageNumber="296">T. marmorea</emphasis>
</taxonomicName>
,
<specimenCount id="9D78DFCDF400082DA8185893CFC5B48C" box="[1355,1435,980,1005]" pageId="2" pageNumber="296" type="male">1 male</specimenCount>
specimen of
<taxonomicName id="4C7E6FC7F400082DAC7B58BECBD3B371" authorityName="Thorell" authorityYear="1877" box="[296,397,1017,1040]" class="Arachnida" family="Tetragnathidae" genus="Meta" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="striata">
<emphasis id="B90AC856F400082DAC7B58BECBD3B371" box="[296,397,1017,1040]" italics="true" pageId="2" pageNumber="296">T. striata</emphasis>
</taxonomicName>
and twenty one male specimens of
<taxonomicName id="4C7E6FC7F400082DAE7758BEC9FDB371" authorityName="Thorell" authorityYear="1877" box="[804,931,1015,1040]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="B90AC856F400082DAE7758BEC9FDB371" box="[804,931,1015,1040]" italics="true" pageId="2" pageNumber="296">T. ventralis</emphasis>
</taxonomicName>
. Despite the small sample size, it was found that the male chelicerae of all the Indian
<taxonomicName id="4C7E6FC7F400082DAF325F5BC8E3B354" authorityName="Simon" authorityYear="1894" box="[609,701,1052,1077]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90AC856F400082DAF325F5BC8E3B354" box="[609,701,1052,1077]" italics="true" pageId="2" pageNumber="296">Tylorida</emphasis>
</taxonomicName>
spp. lack specialized apophysis/apophyses, but possess teeth and tubercles that are varying in number, shape, size and mutual spacing from species to species (
<figureCitation id="134508C1F400082DA98F5F78CFCEB339" box="[1244,1424,1087,1112]" captionStart="FIGURE 12" captionStartId="16.[151,250,1073,1095]" captionTargetBox="[168,1421,214,1028]" captionTargetId="figure@16.[151,1436,193,1052]" captionTargetPageId="16" captionText="FIGURE 12AK. Left chelicerae of Tylorida spp. and Orsinome cf. vethi (Hasselt, 1882). AE Left chelicerae of Tylorida marmorea (Pocock, 1901) comb. nov.: A Male (High altitude form), prolateral; B Female (High altitude form), prolateral; C Male (Low altitude form), prolateral; D Female (Low altitude form), prolateral; E Orginal drawings of male and female cheliceraeadopted from Gravely(1921) without modification; FG Left chelicerae of Tylorida striata (Thorell, 1877):F Male, retrolateral; G Female, prolateral; HI Left chelicerae of Tylorida ventralis (Thorell, 1877): H Male, prolateral, arrow indicates tiny tubercle; I Female,ventro-retrolateral;JK Left cheliceraeof Orsinome cf.vethi: J Male, retrolateral; K Female,prolateral. Scale bars AB, HI, 1 mm; CD, 0.5 mm; FG, 0.2 mm; JK, 0.5 mm; E, not to scale." httpUri="https://zenodo.org/record/1065190/files/figure.png" pageId="2" pageNumber="296">Figs 12A, F, H</figureCitation>
). Unlike males, females have chelicerae armed with similar number of teeth: three promarginal teeth, which are slightly larger than the four retromarginal teeth (
<figureCitation id="134508C1F400082DAFE75FC0C94CB3C1" box="[692,786,1159,1184]" captionStart="FIGURE 4" captionStartId="8.[151,250,1880,1902]" captionTargetBox="[199,1406,209,1844]" captionTargetId="figure@8.[177,1421,193,1869]" captionTargetPageId="8" captionText="FIGURE 4AH. Tylorida flava sp. nov.. A Subadult male prosoma, retrolateral; B Female prosoma, retrolateral; C Subadult male left chelicera, prolateral; D Female left chelicera, prolateral; EF Epigynum, ventral; GH Internal female genitalia, dorsal.Abbreviations:CD copulatory duct, EP epigynal plate,FD fertilization duct,ll lateral lobeof thepostero-median lobeof epigynal plate, ml median lobe of the postero-median lobe of epigynal plate, pEP postero-median lobe of epigynal plate, SB spermathecal bulb.Scale bars AB, 1 mm; CD, 0.5 mm; EH 0.2 mm." httpUri="https://zenodo.org/record/1065174/files/figure.png" pageId="2" pageNumber="296">Figs 4D</figureCitation>
,
<figureCitation id="134508C1F400082DAE4E5FCFC9DDB3C1" box="[797,899,1160,1184]" captionStart="FIGURE 12" captionStartId="16.[151,250,1073,1095]" captionTargetBox="[168,1421,214,1028]" captionTargetId="figure@16.[151,1436,193,1052]" captionTargetPageId="16" captionText="FIGURE 12AK. Left chelicerae of Tylorida spp. and Orsinome cf. vethi (Hasselt, 1882). AE Left chelicerae of Tylorida marmorea (Pocock, 1901) comb. nov.: A Male (High altitude form), prolateral; B Female (High altitude form), prolateral; C Male (Low altitude form), prolateral; D Female (Low altitude form), prolateral; E Orginal drawings of male and female cheliceraeadopted from Gravely(1921) without modification; FG Left chelicerae of Tylorida striata (Thorell, 1877):F Male, retrolateral; G Female, prolateral; HI Left chelicerae of Tylorida ventralis (Thorell, 1877): H Male, prolateral, arrow indicates tiny tubercle; I Female,ventro-retrolateral;JK Left cheliceraeof Orsinome cf.vethi: J Male, retrolateral; K Female,prolateral. Scale bars AB, HI, 1 mm; CD, 0.5 mm; FG, 0.2 mm; JK, 0.5 mm; E, not to scale." httpUri="https://zenodo.org/record/1065190/files/figure.png" pageId="2" pageNumber="296">12B, G, I</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>