326 lines
57 KiB
XML
326 lines
57 KiB
XML
<document id="29FB986B3426E643ACF47E68D8BEFCFB" ID-CLB-Dataset="46333" ID-DOI="10.11646/zootaxa.3768.5.3" ID-GBIF-Dataset="3e360f06-2719-439e-9536-9f4dd74dc7a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="228011" ID-ZooBank="8A447FE6-C25D-4542-9DDB-37A41687C35E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460390133935" checkinUser="plazi" docAuthor="Yu, Daoyuan, Deharveng, Louis & Zhang, Feng" docDate="2014" docId="03BE075BFFCDFFA717B8FBC2FF75B7D0" docLanguage="en" docName="zt03768p575.pdf" docOrigin="Zootaxa 3768 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monodontocerus absens Yu, Deharveng & Zhang, 2014, sp. nov." docType="treatment" docVersion="10" lastPageNumber="563" masterDocId="FF877F23FFCFFFA1172FFFB8FFBFB00C" masterDocTitle="New species of Monodontocerus (Collembola: Tomoceridae) from southern China with diagnostic notes on the genus and introduction of new taxonomic characters" masterLastPageNumber="575" masterPageNumber="557" pageNumber="559" updateTime="1698298113494" updateUser="ExternalLinkService">
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<mods:mods id="C15425B6C1ADA6264E97100732826A31" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EEE0BA8D17501356C9F4F64D470CF39A">
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<mods:title id="60821EE55884FE9D99B855019335B330">New species of Monodontocerus (Collembola: Tomoceridae) from southern China with diagnostic notes on the genus and introduction of new taxonomic characters</mods:title>
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</mods:titleInfo>
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<mods:name id="8261F5F4117CF4C054BAB2986EED10B6" type="personal">
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<mods:role id="EC84647E76ED5A0268435662C4A283B0">
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<mods:roleTerm id="A8946BC068CF5489B698F75AABB977CD">Author</mods:roleTerm>
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<mods:namePart id="033F323A4C073C6B68ADF82C0E91929C">Yu, Daoyuan</mods:namePart>
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</mods:name>
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<mods:name id="EDB88BEAA222BA20674DC90B4BEEEF72" type="personal">
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<mods:roleTerm id="B7C4D11133E9C2331886071DCED64D01">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A60851078B88228500949B613B3CB8EE">Deharveng, Louis</mods:namePart>
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</mods:name>
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<mods:name id="14C75A96F403485C1784850A18626C75" type="personal">
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<mods:roleTerm id="23207DFBE584A3BA91C6CAE990350559">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="376A5B210A4C1BB5C5E9A7DD2FD2B006">Zhang, Feng</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3082A87E4B75776DA9D5668960ECEDED">text</mods:typeOfResource>
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<mods:titleInfo id="ECD32A032585D7F58919F6114C8E9D5F">
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<mods:title id="7B0663E2699D614426EBCFA5108233C5">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="4DDCC2397BA9C672DBCC551CCB3DFE33">
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<mods:date id="E2895965706E167447873D8E890C5664">2014</mods:date>
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<mods:detail id="00D209091DBD42CF63F69562690EE1C9" type="volume">
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<mods:number id="A92C02B972F0BAE35AD1953BC1D9CE10">3768</mods:number>
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<mods:detail id="86A70B42BC0D75CC9088ABE2981FA609" type="issue">
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<mods:number id="F676EC50A608F1FBF9A87C1BFA5E5629">5</mods:number>
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<mods:classification id="F4FA3579F8CADDD88D370C6829317F3D">journal article</mods:classification>
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<mods:identifier id="D1BE2633BC32C63CD28ACEA621ADD4F4" type="CLB-Dataset">46333</mods:identifier>
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<mods:identifier id="FE542F495A61BF4D03720B64D63BBAEB" type="DOI">10.11646/zootaxa.3768.5.3</mods:identifier>
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<mods:identifier id="FA3CC782ABE0E927B87D2F098B70F40E" type="GBIF-Dataset">3e360f06-2719-439e-9536-9f4dd74dc7a3</mods:identifier>
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<mods:identifier id="2928D39856632442264E0164B77EA122" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03BE075BFFCDFFA717B8FBC2FF75B7D0" ID-DOI="http://doi.org/10.5281/zenodo.5678993" ID-GBIF-Taxon="119416543" ID-Zenodo-Dep="5678993" LSID="urn:lsid:plazi:treatment:03BE075BFFCDFFA717B8FBC2FF75B7D0" httpUri="http://treatment.plazi.org/id/03BE075BFFCDFFA717B8FBC2FF75B7D0" lastPageId="6" lastPageNumber="563" pageId="2" pageNumber="559">
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<subSubSection id="C30DE5C6FFCDFFA317B8FBC2FE4CB4BB" pageId="2" pageNumber="559" type="nomenclature">
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<paragraph id="8BA8B64DFFCDFFA317B8FBC2FDAFB49F" blockId="2.[151,528,1146,1207]" box="[151,528,1146,1171]" pageId="2" pageNumber="559">
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<heading id="D0E00121FFCDFFA317B8FBC2FDAFB49F" bold="true" box="[151,528,1146,1171]" fontSize="10" level="2" pageId="2" pageNumber="559" reason="2">
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<emphasis id="B9636A5FFFCDFFA317B8FBC2FDAFB49F" bold="true" box="[151,528,1146,1171]" pageId="2" pageNumber="559">
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<taxonomicName id="4C17CDCEFFCDFFA317B8FBC2FE0FB49F" box="[151,432,1146,1171]" class="Entognatha" family="Tomoceridae" genus="Monodontocerus" kingdom="Animalia" order="Collembola" pageId="2" pageNumber="559" phylum="Arthropoda" rank="species" species="absens" status="sp. nov.">
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<emphasis id="B9636A5FFFCDFFA317B8FBC2FE0FB49F" bold="true" box="[151,432,1146,1171]" italics="true" pageId="2" pageNumber="559">Monodontocerus absens</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A250D724FFCDFFA31699FBC2FDAFB49F" box="[438,528,1146,1171]" pageId="2" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA8B64DFFCDFFA317B8FB26FE4CB4BB" blockId="2.[151,528,1146,1207]" box="[151,499,1182,1207]" pageId="2" pageNumber="559">
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<figureCitation id="132CAAC8FFCDFFA317B8FB26FF46B4BB" box="[151,249,1182,1207]" captionStart="FIGURES 1 – 5. 1 – 3" captionStartId="3.[151,264,1933,1956]" captionTargetBox="[151,1436,373,1911]" captionTargetId="figure@3.[151,1436,373,1912]" captionTargetPageId="3" captionText="FIGURES 1 – 5. 1 – 3, Monodontocerus absens sp. nov. 1, habitus; 2, spotted pigmentation on Abd. III; 3, head, arrows pointing to the pigmented sockets of cephalic posterior macrochaetae; 4, habitus of M. mulunensis sp. nov.; 5, habitus of M. trigrandis sp. nov. Scale bar: 1, 200 Μm; 2, 3, 25 Μm; 4, 5, 100 Μm." httpUri="https://zenodo.org/record/228012/files/figure.png" pageId="2" pageNumber="559">Figs 1–3</figureCitation>
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,
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<figureCitation id="132CAAC8FFCDFFA31629FB26FEFEB4BB" box="[262,321,1182,1207]" captionStart-0="FIGURES 6 – 14" captionStart-1="FIGURES 15 – 22" captionStart-2="FIGURES 23 – 29" captionStartId-0="4.[151,264,1919,1942]" captionStartId-1="5.[151,264,1571,1594]" captionStartId-2="7.[151,264,1916,1939]" captionTargetBox-0="[243,1341,223,1893]" captionTargetBox-1="[182,1426,185,1546]" captionTargetBox-2="[175,1415,206,1886]" captionTargetId-0="figure@4.[220,1357,193,1912]" captionTargetId-1="figure@5.[167,1431,165,1555]" captionTargetId-2="figure@7.[158,1422,195,1894]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionText-0="FIGURES 6 – 14. Monodontocerus absens sp. nov.: 6 – 9, dorsal side of antenna. 6, one distal subsegment of Ant. III; 7, most distal part of Ant. III; 8, one middle subsegment of Ant. IV; 9, apex of Ant. IV; 10, labrum; 11, mandible; 12, maxillary lamella 5; 13, maxillary outer lobe; 14, labial palp (papillae A – E and H labeled). Scale bar: 6 – 9, 12, 14, 20 Μm; 10, 11, 13, 50 Μm." captionText-1="FIGURES 15 – 22. Monodontocerus absens sp. nov.: 15, labial chaetotaxy; 16, trochanteral-femoral organ; 17, femur and tibiotarsus; 18, dorsal view of tibiotarsal distal chaetae on front leg (TH: tenent hair, the same below); 19, anterior view of tibiotarsal distal chaetae on hind leg; 20, pattern of tibiotarsal distal chaetae (hind leg, distal view); 21, claw; 22, tenaculum. Scale bar: 15, 19, 21, 22, 50 Μm; 16, 120 Μm; 17, 200 Μm; 18, 40 Μm." captionText-2="FIGURES 23 – 29. Monodontocerus absens sp. nov.: 23, manubrium (" T " shaped marks: sockets of scales; circle with a slash: pseudopore); 24, disto-external corner of manubrium and corner chaeta; 25, dental spines; 26, feathered dental chaeta; 27, mucro, apical tooth hypothesized; 28, mature female genital plate; 29, immature female genital plate. Scale bar: 23, 25, 100 Μm; 24, 26 – 29, 50 Μm." httpUri-0="https://zenodo.org/record/228013/files/figure.png" httpUri-1="https://zenodo.org/record/228014/files/figure.png" httpUri-2="https://zenodo.org/record/228015/files/figure.png" pageId="2" pageNumber="559">6–29</figureCitation>
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,
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<figureCitation id="132CAAC8FFCDFFA31663FB26FE2BB4BB" box="[332,404,1182,1207]" captionStart="FIGURES 54 – 59" captionStartId="14.[151,264,1812,1835]" captionTargetBox="[168,1426,206,1779]" captionTargetId="figure@14.[151,1436,193,1791]" captionTargetPageId="14" captionText="FIGURES 54 – 59. Dorsal cephalic and body chaetotaxy: 54 & 57, M. absens sp. nov.; 55 & 58, M. mulunensis sp. nov.; 56 & 59, M. trigrandis sp. nov. Large circle: macrochaeta; small circle: mesochaeta; wavy line: bothriotricha; s: normal s-chaeta; ms: s-microchaeta; rs: rod-like s-chaeta on body; circle with a slash: pseudopore." httpUri="https://zenodo.org/record/228019/files/figure.png" pageId="2" pageNumber="559">54, 57</figureCitation>
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,
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<tableCitation id="C69583F6FFCDFFA3168FFB26FE4CB4BB" box="[416,499,1182,1207]" captionStart="TABLE 2" captionStartId="8.[151,239,152,175]" captionTargetBox="[151,1391,234,594]" captionTargetPageId="8" captionText="TABLE 2. Variability of characters among Monodontocerus. A, anterior area; IO, interocular area; PO, postocular area; P, posterior area;?, unknown data." httpUri="http://table.plazi.org/id/DF68E6C5FFC7FFA917B8FF20FDB8B0C3" pageId="2" pageNumber="559" tableUuid="DF68E6C5FFC7FFA917B8FF20FDB8B0C3">Table 2</tableCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30DE5C6FFCDFFA317B8FB5EFE42B566" pageId="2" pageNumber="559" type="materials_examined">
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<paragraph id="8BA8B64DFFCDFFA317B8FB5EFE42B566" blockId="2.[151,1436,1254,2035]" pageId="2" pageNumber="559">
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<emphasis id="B9636A5FFFCDFFA317B8FB5EFEF9B4F3" bold="true" box="[151,326,1254,1279]" pageId="2" pageNumber="559">
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<typeStatus id="54AC08EFFFCDFFA317B8FB5EFF6DB4F3" box="[151,210,1254,1279]" pageId="2" pageNumber="559">Type</typeStatus>
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material.
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</emphasis>
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||
<typeStatus id="54AC08EFFFCDFFA31662FB5EFE0AB4F3" box="[333,437,1254,1279]" pageId="2" pageNumber="559" type="holotype">Holotype</typeStatus>
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female and two
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<typeStatus id="54AC08EFFFCDFFA3155CFB5FFD6BB4F3" box="[627,724,1255,1279]" pageId="2" pageNumber="559" type="paratype">paratype</typeStatus>
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females on slides, collected in Ba Xian Dong (cave), Ba Xian, Duan County, Hechi, Guangxi Zhuang Autonomous Region,
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<collectingCountry id="F300F6DDFFCDFFA31469FAB2FC33B52F" box="[838,908,1290,1315]" name="China" pageId="2" pageNumber="559">China</collectingCountry>
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, x=
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<geoCoordinate id="EE23D08AFFCDFFA31493FAB3FBDFB52F" box="[956,1120,1291,1315]" direction="east" orientation="longitude" pageId="2" pageNumber="559" precision="1" value="108.13281">108.132811°E</geoCoordinate>
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||
; y=
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<geoCoordinate id="EE23D08AFFCDFFA313BEFAB3FA94B52F" box="[1169,1323,1291,1315]" direction="north" orientation="latitude" pageId="2" pageNumber="559" precision="1" value="23.91404">23.914039°N</geoCoordinate>
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, elevation
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<quantity id="4CEF1BA8FFCDFFA317B8FA96FF5FB54A" box="[151,224,1326,1351]" metricMagnitude="2" metricUnit="m" metricValue="1.51" pageId="2" pageNumber="559" unit="m" value="151.0">151 m</quantity>
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,
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<date id="FFA9908DFFCDFFA317C3FA96FEDAB54B" box="[236,357,1326,1351]" pageId="2" pageNumber="559" value="2010-04-26">26.iv.2010</date>
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, by Louis Deharveng & Li Youbang (sample code: CHIgx10.67).
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<typeStatus id="54AC08EFFFCDFFA31360FA96FB08B54B" box="[1103,1207,1326,1351]" pageId="2" pageNumber="559" type="holotype">Holotype</typeStatus>
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and one
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<typeStatus id="54AC08EFFFCDFFA31231FA97FAC0B54B" box="[1310,1407,1327,1351]" pageId="2" pageNumber="559" type="paratype">paratype</typeStatus>
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in
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<collectionCode id="ED062E88FFCDFFA317B8FAEBFF5DB567" box="[151,226,1363,1387]" pageId="2" pageNumber="559">NJAU</collectionCode>
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, one
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<typeStatus id="54AC08EFFFCDFFA31633FAEBFEC3B567" box="[284,380,1363,1387]" pageId="2" pageNumber="559" type="paratype">paratype</typeStatus>
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in
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<collectionCode id="ED062E88FFCDFFA3168FFAEBFE46B566" LSID="urn:lsid:biocol.org:col:34988" box="[416,505,1363,1386]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="559">MNHN</collectionCode>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30DE5C6FFCDFFA717E8FACEFB82B6EC" lastPageId="6" lastPageNumber="563" pageId="2" pageNumber="559" type="description">
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<paragraph id="8BA8B64DFFCDFFA317E8FACEFBC6B5F7" blockId="2.[151,1436,1254,2035]" pageId="2" pageNumber="559">
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<emphasis id="B9636A5FFFCDFFA317E8FACEFEE6B583" bold="true" box="[199,345,1398,1423]" pageId="2" pageNumber="559">Description.</emphasis>
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Body length
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<quantity id="4CEF1BA8FFCDFFA316DFFACFFDC5B582" box="[496,634,1399,1423]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.8" metricValueMin="3.1" pageId="2" pageNumber="559" unit="mm" value="3.45" valueMax="3.8" valueMin="3.1">3.1–3.8 mm</quantity>
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. Ground color pale (
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<figureCitation id="132CAAC8FFCDFFA31470FACEFC1EB583" box="[863,929,1398,1423]" captionStart="FIGURES 1 – 5. 1 – 3" captionStartId="3.[151,264,1933,1956]" captionTargetBox="[151,1436,373,1911]" captionTargetId="figure@3.[151,1436,373,1912]" captionTargetPageId="3" captionText="FIGURES 1 – 5. 1 – 3, Monodontocerus absens sp. nov. 1, habitus; 2, spotted pigmentation on Abd. III; 3, head, arrows pointing to the pigmented sockets of cephalic posterior macrochaetae; 4, habitus of M. mulunensis sp. nov.; 5, habitus of M. trigrandis sp. nov. Scale bar: 1, 200 Μm; 2, 3, 25 Μm; 4, 5, 100 Μm." httpUri="https://zenodo.org/record/228012/files/figure.png" pageId="2" pageNumber="559">Fig. 1</figureCitation>
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); head and body covered with spotted brown pigment (
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<figureCitation id="132CAAC8FFCDFFA3162CFA22FEF7B5BF" box="[259,328,1434,1459]" captionStart="FIGURES 1 – 5. 1 – 3" captionStartId="3.[151,264,1933,1956]" captionTargetBox="[151,1436,373,1911]" captionTargetId="figure@3.[151,1436,373,1912]" captionTargetPageId="3" captionText="FIGURES 1 – 5. 1 – 3, Monodontocerus absens sp. nov. 1, habitus; 2, spotted pigmentation on Abd. III; 3, head, arrows pointing to the pigmented sockets of cephalic posterior macrochaetae; 4, habitus of M. mulunensis sp. nov.; 5, habitus of M. trigrandis sp. nov. Scale bar: 1, 200 Μm; 2, 3, 25 Μm; 4, 5, 100 Μm." httpUri="https://zenodo.org/record/228012/files/figure.png" pageId="2" pageNumber="559">Fig. 2</figureCitation>
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); sockets of macrochaetae and large mesochaetae, antero-dorsal part of head and anterior margins of clypeus and Th. II darker than other parts (
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<figureCitation id="132CAAC8FFCDFFA315B4FA06FD60B5DB" box="[667,735,1470,1495]" captionStart="FIGURES 1 – 5. 1 – 3" captionStartId="3.[151,264,1933,1956]" captionTargetBox="[151,1436,373,1911]" captionTargetId="figure@3.[151,1436,373,1912]" captionTargetPageId="3" captionText="FIGURES 1 – 5. 1 – 3, Monodontocerus absens sp. nov. 1, habitus; 2, spotted pigmentation on Abd. III; 3, head, arrows pointing to the pigmented sockets of cephalic posterior macrochaetae; 4, habitus of M. mulunensis sp. nov.; 5, habitus of M. trigrandis sp. nov. Scale bar: 1, 200 Μm; 2, 3, 25 Μm; 4, 5, 100 Μm." httpUri="https://zenodo.org/record/228012/files/figure.png" pageId="2" pageNumber="559">Fig. 3</figureCitation>
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); eye patches black, relatively small; legs, ventral tube, furca, Ant. I, Ant. II and base of Ant. III with diffuse pigment, remaining parts of antenna pale.
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</paragraph>
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<paragraph id="8BA8B64DFFCDFFA317E8F9BEFC9BB717" blockId="2.[151,1436,1254,2035]" pageId="2" pageNumber="559">
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Postantennal organ absent. Ommatidia 6+6, relatively smaller than in edaphic Tomocerinae. Antennae about 1.1–1.2 times as long as body, antennal length ratio as I: II: III: IV=1.0: 2.2: 17.6–17.8: 3.3. Ant. I and Ant. II dorsally scaled, ventrally with dense s-chaetae. Ant. III and Ant. IV annulated as normal for Tomocerinae, with six
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<typeStatus id="54AC08EFFFCDFFA317B8F9CBFF6DB687" box="[151,210,1651,1675]" pageId="2" pageNumber="559">types</typeStatus>
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of chaetae identified (
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<tableCitation id="C69583F6FFCDFFA316FEF9CAFD98B687" box="[465,551,1650,1675]" captionStart="TABLE 1" captionStartId="1.[151,239,726,749]" captionText="TABLE 1. Nomenclature of antennal chaetae in Monodontocerus. –, absent." pageId="2" pageNumber="559">Table 1</tableCitation>
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,
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<figureCitation id="132CAAC8FFCDFFA31519F9CAFD22B687" box="[566,669,1650,1675]" captionStart="FIGURES 6 – 14" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[243,1341,223,1893]" captionTargetId="figure@4.[220,1357,193,1912]" captionTargetPageId="4" captionText="FIGURES 6 – 14. Monodontocerus absens sp. nov.: 6 – 9, dorsal side of antenna. 6, one distal subsegment of Ant. III; 7, most distal part of Ant. III; 8, one middle subsegment of Ant. IV; 9, apex of Ant. IV; 10, labrum; 11, mandible; 12, maxillary lamella 5; 13, maxillary outer lobe; 14, labial palp (papillae A – E and H labeled). Scale bar: 6 – 9, 12, 14, 20 Μm; 10, 11, 13, 50 Μm." httpUri="https://zenodo.org/record/228013/files/figure.png" pageId="2" pageNumber="559">Figs 6–9</figureCitation>
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). Ant. III dorso-basally scaled, each subsegment near the middle of Ant. III with a whorl of about 30 chaetae; ordinary chaetae
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<typeStatus id="54AC08EFFFCDFFA31413F92FFCD3B6A3" box="[828,876,1687,1711]" pageId="2" pageNumber="559">type</typeStatus>
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a (SE) and s-chaetae of
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<typeStatus id="54AC08EFFFCDFFA313ACF92FFB0CB6A3" box="[1155,1203,1687,1711]" pageId="2" pageNumber="559">type</typeStatus>
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b,
|
||
<typeStatus id="54AC08EFFFCDFFA313F6F92FFAB6B6A3" box="[1241,1289,1687,1711]" pageId="2" pageNumber="559">type</typeStatus>
|
||
c (RLS) and
|
||
<typeStatus id="54AC08EFFFCDFFA317B8F903FF78B6DF" box="[151,199,1723,1747]" pageId="2" pageNumber="559">type</typeStatus>
|
||
d (RSS) uniformly distributed; dorso-apically, with one (SH) chaeta of
|
||
<typeStatus id="54AC08EFFFCDFFA314C9F903FBA9B6DF" box="[998,1046,1723,1747]" pageId="2" pageNumber="559">type</typeStatus>
|
||
f (TRS), one of
|
||
<typeStatus id="54AC08EFFFCDFFA313E3F903FB43B6DF" box="[1228,1276,1723,1747]" pageId="2" pageNumber="559">type</typeStatus>
|
||
e, two of
|
||
<typeStatus id="54AC08EFFFCDFFA31244F903FA24B6DF" box="[1387,1435,1723,1747]" pageId="2" pageNumber="559">type</typeStatus>
|
||
c and two of
|
||
<typeStatus id="54AC08EFFFCDFFA3161EF967FEDEB6FB" box="[305,353,1759,1783]" pageId="2" pageNumber="559">type</typeStatus>
|
||
b (
|
||
<figureCitation id="132CAAC8FFCDFFA316A6F966FE70B6FB" box="[393,463,1758,1783]" captionStart="FIGURES 6 – 14" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[243,1341,223,1893]" captionTargetId="figure@4.[220,1357,193,1912]" captionTargetPageId="4" captionText="FIGURES 6 – 14. Monodontocerus absens sp. nov.: 6 – 9, dorsal side of antenna. 6, one distal subsegment of Ant. III; 7, most distal part of Ant. III; 8, one middle subsegment of Ant. IV; 9, apex of Ant. IV; 10, labrum; 11, mandible; 12, maxillary lamella 5; 13, maxillary outer lobe; 14, labial palp (papillae A – E and H labeled). Scale bar: 6 – 9, 12, 14, 20 Μm; 10, 11, 13, 50 Μm." httpUri="https://zenodo.org/record/228013/files/figure.png" pageId="2" pageNumber="559">Fig. 7</figureCitation>
|
||
). Ant. IV unscaled, chaetotaxy similar to that of Ant. III, with one subapical papilla, apex with one
|
||
<typeStatus id="54AC08EFFFCDFFA31614F8BBFED4B717" box="[315,363,1795,1819]" pageId="2" pageNumber="559">type</typeStatus>
|
||
f chaeta and one
|
||
<typeStatus id="54AC08EFFFCDFFA3151FF8BBFDDFB717" box="[560,608,1795,1819]" pageId="2" pageNumber="559">type</typeStatus>
|
||
e chaeta (
|
||
<figureCitation id="132CAAC8FFCDFFA315FEF8BAFCAAB717" box="[721,789,1794,1819]" captionStart="FIGURES 6 – 14" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[243,1341,223,1893]" captionTargetId="figure@4.[220,1357,193,1912]" captionTargetPageId="4" captionText="FIGURES 6 – 14. Monodontocerus absens sp. nov.: 6 – 9, dorsal side of antenna. 6, one distal subsegment of Ant. III; 7, most distal part of Ant. III; 8, one middle subsegment of Ant. IV; 9, apex of Ant. IV; 10, labrum; 11, mandible; 12, maxillary lamella 5; 13, maxillary outer lobe; 14, labial palp (papillae A – E and H labeled). Scale bar: 6 – 9, 12, 14, 20 Μm; 10, 11, 13, 50 Μm." httpUri="https://zenodo.org/record/228013/files/figure.png" pageId="2" pageNumber="559">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA8B64DFFCDFFA217E8F89EFE77B14C" blockId="2.[151,1436,1254,2035]" lastBlockId="3.[151,1436,151,320]" lastPageId="3" lastPageNumber="560" pageId="2" pageNumber="559">
|
||
Labral formula 4/5, 5, 4; prelabral and labral chaetae pointed, smooth; labral chaetae with thick basal papillae fused to cuticle, proximal two rows of chaetae subequal in thickness and length, distal four chaetae distinctly stronger; distal edge of labrum with four spiny papillae with rounded apices; ventro-distal margin of labrum densely ciliated as a brush (
|
||
<figureCitation id="132CAAC8FFCDFFA316FBF82AFD97B7A7" box="[468,552,1938,1963]" captionStart="FIGURES 6 – 14" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[243,1341,223,1893]" captionTargetId="figure@4.[220,1357,193,1912]" captionTargetPageId="4" captionText="FIGURES 6 – 14. Monodontocerus absens sp. nov.: 6 – 9, dorsal side of antenna. 6, one distal subsegment of Ant. III; 7, most distal part of Ant. III; 8, one middle subsegment of Ant. IV; 9, apex of Ant. IV; 10, labrum; 11, mandible; 12, maxillary lamella 5; 13, maxillary outer lobe; 14, labial palp (papillae A – E and H labeled). Scale bar: 6 – 9, 12, 14, 20 Μm; 10, 11, 13, 50 Μm." httpUri="https://zenodo.org/record/228013/files/figure.png" pageId="2" pageNumber="559">Fig. 10</figureCitation>
|
||
). Mandible heads asymmetrical, left mandible with four teeth, right mandible with five teeth; molar plate with numerous strong teeth, including a distal tapered one on left mandible (
|
||
<figureCitation id="132CAAC8FFCDFFA31214F80EFA31B7C3" box="[1339,1422,1974,1999]" captionStart="FIGURES 6 – 14" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[243,1341,223,1893]" captionTargetId="figure@4.[220,1357,193,1912]" captionTargetPageId="4" captionText="FIGURES 6 – 14. Monodontocerus absens sp. nov.: 6 – 9, dorsal side of antenna. 6, one distal subsegment of Ant. III; 7, most distal part of Ant. III; 8, one middle subsegment of Ant. IV; 9, apex of Ant. IV; 10, labrum; 11, mandible; 12, maxillary lamella 5; 13, maxillary outer lobe; 14, labial palp (papillae A – E and H labeled). Scale bar: 6 – 9, 12, 14, 20 Μm; 10, 11, 13, 50 Μm." httpUri="https://zenodo.org/record/228013/files/figure.png" pageId="2" pageNumber="559">Fig. 11</figureCitation>
|
||
). Maxilla head with three teeth and six distinct lamellae; lamella 5 broad, curved with prolonged basal teeth, distinct beard absent (
|
||
<figureCitation id="132CAAC8FFCCFFA2161BFF2FFE38B0BC" box="[308,391,151,176]" captionStart="FIGURES 6 – 14" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[243,1341,223,1893]" captionTargetId="figure@4.[220,1357,193,1912]" captionTargetPageId="4" captionText="FIGURES 6 – 14. Monodontocerus absens sp. nov.: 6 – 9, dorsal side of antenna. 6, one distal subsegment of Ant. III; 7, most distal part of Ant. III; 8, one middle subsegment of Ant. IV; 9, apex of Ant. IV; 10, labrum; 11, mandible; 12, maxillary lamella 5; 13, maxillary outer lobe; 14, labial palp (papillae A – E and H labeled). Scale bar: 6 – 9, 12, 14, 20 Μm; 10, 11, 13, 50 Μm." httpUri="https://zenodo.org/record/228013/files/figure.png" pageId="3" pageNumber="560">Fig. 12</figureCitation>
|
||
). Maxillary outer lobe with trifurcate palp, one basal chaeta and four subequal sublobal hairs (
|
||
<figureCitation id="132CAAC8FFCCFFA217B0FF04FF4CB0D8" box="[159,243,188,213]" captionStart="FIGURES 6 – 14" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[243,1341,223,1893]" captionTargetId="figure@4.[220,1357,193,1912]" captionTargetPageId="4" captionText="FIGURES 6 – 14. Monodontocerus absens sp. nov.: 6 – 9, dorsal side of antenna. 6, one distal subsegment of Ant. III; 7, most distal part of Ant. III; 8, one middle subsegment of Ant. IV; 9, apex of Ant. IV; 10, labrum; 11, mandible; 12, maxillary lamella 5; 13, maxillary outer lobe; 14, labial palp (papillae A – E and H labeled). Scale bar: 6 – 9, 12, 14, 20 Μm; 10, 11, 13, 50 Μm." httpUri="https://zenodo.org/record/228013/files/figure.png" pageId="3" pageNumber="560">Fig. 13</figureCitation>
|
||
). Labial palp with about 20 proximal chaetae, papillae A–E respectively with 0, 5, 0, 4, 7 (a1 present but attached to papilla B
|
||
<bibRefCitation id="EF86CBBCFFCCFFA216AAFF67FDFBB0F4" author="Fjellberg" box="[389,580,223,248]" pageId="3" pageNumber="560" refString="Fjellberg, A. (1999) The labial palp in Collembola. Zoologischer Anzeiger, 237, 309 - 330." type="journal article" year="1999">(Fjellberg 1999))</bibRefCitation>
|
||
guard chaetae, hypostomal chaeta H rod-like, with h1 stronger than h2 (
|
||
<figureCitation id="132CAAC8FFCCFFA21241FF67FF09B110" captionStart="FIGURES 6 – 14" captionStartId="4.[151,264,1919,1942]" captionTargetBox="[243,1341,223,1893]" captionTargetId="figure@4.[220,1357,193,1912]" captionTargetPageId="4" captionText="FIGURES 6 – 14. Monodontocerus absens sp. nov.: 6 – 9, dorsal side of antenna. 6, one distal subsegment of Ant. III; 7, most distal part of Ant. III; 8, one middle subsegment of Ant. IV; 9, apex of Ant. IV; 10, labrum; 11, mandible; 12, maxillary lamella 5; 13, maxillary outer lobe; 14, labial palp (papillae A – E and H labeled). Scale bar: 6 – 9, 12, 14, 20 Μm; 10, 11, 13, 50 Μm." httpUri="https://zenodo.org/record/228013/files/figure.png" pageId="3" pageNumber="560">Fig. 14</figureCitation>
|
||
). Mentum with five large chaetae, submentum with approximately 25 chaetae of different sizes, 9–12 of them distinctly smaller (
|
||
<figureCitation id="132CAAC8FFCCFFA21647FE9FFE04B14C" box="[360,443,295,320]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1571,1594]" captionTargetBox="[182,1426,185,1546]" captionTargetId="figure@5.[167,1431,165,1555]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Monodontocerus absens sp. nov.: 15, labial chaetotaxy; 16, trochanteral-femoral organ; 17, femur and tibiotarsus; 18, dorsal view of tibiotarsal distal chaetae on front leg (TH: tenent hair, the same below); 19, anterior view of tibiotarsal distal chaetae on hind leg; 20, pattern of tibiotarsal distal chaetae (hind leg, distal view); 21, claw; 22, tenaculum. Scale bar: 15, 19, 21, 22, 50 Μm; 16, 120 Μm; 17, 200 Μm; 18, 40 Μm." httpUri="https://zenodo.org/record/228014/files/figure.png" pageId="3" pageNumber="560">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF68E6C5FFCCFFA217B8F835FD7DB7ED" httpUri="https://zenodo.org/record/228012/files/figure.png" pageId="3" pageNumber="560" targetBox="[151,1436,373,1911]" targetPageId="3">
|
||
<paragraph id="8BA8B64DFFCCFFA217B8F835FD7DB7ED" blockId="3.[151,1436,1933,2017]" pageId="3" pageNumber="560">
|
||
<emphasis id="B9636A5FFFCCFFA217B8F835FE85B7AF" bold="true" box="[151,314,1933,1956]" pageId="3" pageNumber="560">FIGURES 1–5.</emphasis>
|
||
1–3,
|
||
<taxonomicName id="4C17CDCEFFCCFFA2165DF835FDD6B7AF" box="[370,617,1933,1955]" class="Entognatha" family="Tomoceridae" genus="Monodontocerus" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="560" phylum="Arthropoda" rank="species" species="absens" status="sp. nov.">
|
||
<emphasis id="B9636A5FFFCCFFA2165DF835FDD6B7AF" box="[370,617,1933,1955]" italics="true" pageId="3" pageNumber="560">Monodontocerus absens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A250D724FFCCFFA21540F836FD00B7AF" box="[623,703,1934,1955]" pageId="3" pageNumber="560" rank="species">
|
||
<emphasis id="B9636A5FFFCCFFA21540F836FD00B7AF" bold="true" box="[623,703,1934,1955]" pageId="3" pageNumber="560">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
1, habitus; 2, spotted pigmentation on Abd. III; 3, head, arrows pointing to the pigmented sockets of cephalic posterior macrochaetae; 4, habitus of
|
||
<taxonomicName id="4C17CDCEFFCCFFA214BCF815FB98B7CE" box="[915,1063,1964,1986]" class="Entognatha" family="Tomoceridae" genus="Monodontocerus" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="560" phylum="Arthropoda" rank="species" species="mulunensis" status="sp. nov.">
|
||
<emphasis id="B9636A5FFFCCFFA214BCF815FB98B7CE" box="[915,1063,1964,1986]" italics="true" pageId="3" pageNumber="560">M. mulunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A250D724FFCCFFA21301F815FBC0B7CE" box="[1070,1151,1965,1986]" pageId="3" pageNumber="560" rank="species">
|
||
<emphasis id="B9636A5FFFCCFFA21301F815FBC0B7CE" bold="true" box="[1070,1151,1965,1986]" pageId="3" pageNumber="560">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; 5, habitus of
|
||
<taxonomicName id="4C17CDCEFFCCFFA2123BF815FA23B7CE" box="[1300,1436,1964,1986]" class="Entognatha" family="Tomoceridae" genus="Monodontocerus" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="560" phylum="Arthropoda" rank="species" species="trigrandis" status="sp. nov.">
|
||
<emphasis id="B9636A5FFFCCFFA2123BF815FA23B7CE" box="[1300,1436,1964,1986]" italics="true" pageId="3" pageNumber="560">M. trigrandis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A250D724FFCCFFA217B8F873FF58B7EC" box="[151,231,1995,2016]" pageId="3" pageNumber="560" rank="species">
|
||
<emphasis id="B9636A5FFFCCFFA217B8F873FF58B7EC" bold="true" box="[151,231,1995,2016]" pageId="3" pageNumber="560">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Scale bar: 1, 200Μm; 2, 3, 25Μm; 4, 5, 100Μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF68E6C5FFCBFFA517B8F8C7FADEB7DE" httpUri="https://zenodo.org/record/228013/files/figure.png" pageId="4" pageNumber="561" targetBox="[243,1341,223,1893]" targetPageId="4">
|
||
<paragraph id="8BA8B64DFFCBFFA517B8F8C7FADEB7DE" blockId="4.[151,1436,1919,2002]" pageId="4" pageNumber="561">
|
||
<emphasis id="B9636A5FFFCBFFA517B8F8C7FEF7B798" bold="true" box="[151,328,1919,1942]" pageId="4" pageNumber="561">FIGURES 6–14.</emphasis>
|
||
<taxonomicName id="4C17CDCEFFCBFFA51660F8C7FDF8B799" box="[335,583,1919,1941]" class="Entognatha" family="Tomoceridae" genus="Monodontocerus" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="561" phylum="Arthropoda" rank="species" species="absens" status="sp. nov.">
|
||
<emphasis id="B9636A5FFFCBFFA51660F8C7FDF8B799" box="[335,583,1919,1941]" italics="true" pageId="4" pageNumber="561">Monodontocerus absens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A250D724FFCBFFA51560F838FD1EB799" box="[591,673,1920,1941]" pageId="4" pageNumber="561" rank="species">
|
||
<emphasis id="B9636A5FFFCBFFA51560F838FD1EB799" bold="true" box="[591,673,1920,1941]" pageId="4" pageNumber="561">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: 6–9, dorsal side of antenna. 6, one distal subsegment of Ant. III; 7, most distal part of Ant. III; 8, one middle subsegment of Ant. IV; 9, apex of Ant. IV; 10, labrum; 11, mandible; 12, maxillary lamella 5; 13, maxillary outer lobe; 14, labial palp (papillae A–E and H labeled). Scale bar: 6–9, 12, 14, 20Μm; 10, 11, 13, 50Μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF68E6C5FFCAFFA417B8F99BFC81B69A" httpUri="https://zenodo.org/record/228014/files/figure.png" pageId="5" pageNumber="562" targetBox="[182,1426,185,1546]" targetPageId="5">
|
||
<paragraph id="8BA8B64DFFCAFFA417B8F99BFC81B69A" blockId="5.[151,1437,1571,1686]" pageId="5" pageNumber="562">
|
||
<emphasis id="B9636A5FFFCAFFA417B8F99BFEE7B635" bold="true" box="[151,344,1571,1594]" pageId="5" pageNumber="562">FIGURES 15–22.</emphasis>
|
||
<taxonomicName id="4C17CDCEFFCAFFA4164EF99CFDE3B636" box="[353,604,1572,1594]" class="Entognatha" family="Tomoceridae" genus="Monodontocerus" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="562" phylum="Arthropoda" rank="species" species="absens" status="sp. nov.">
|
||
<emphasis id="B9636A5FFFCAFFA4164EF99CFDE3B636" box="[353,604,1572,1594]" italics="true" pageId="5" pageNumber="562">Monodontocerus absens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A250D724FFCAFFA41549F99CFD04B635" box="[614,699,1572,1593]" pageId="5" pageNumber="562" rank="species">
|
||
<emphasis id="B9636A5FFFCAFFA41549F99CFD04B635" bold="true" box="[614,699,1572,1593]" pageId="5" pageNumber="562">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: 15, labial chaetotaxy; 16, trochanteral-femoral organ; 17, femur and tibiotarsus; 18, dorsal view of tibiotarsal distal chaetae on front leg (TH: tenent hair, the same below); 19, anterior view of tibiotarsal distal chaetae on hind leg; 20, pattern of tibiotarsal distal chaetae (hind leg, distal view); 21, claw; 22, tenaculum. Scale bar: 15, 19, 21, 22, 50Μm; 16, 120Μm; 17, 200Μm; 18, 40Μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA8B64DFFCAFFA417E8F905FE0AB76A" blockId="5.[151,1436,1725,2038]" pageId="5" pageNumber="562">
|
||
Head dorsally scaled, a file of smooth chaetae around the base of antenna, anterior area with 2, 2 and interocular area with five macrochaetae arranged as 2, 1, 2, the axial chaeta located more anteriorly than in other Tomocerinae species; postocular chaetae 2+2; posterior chaetae two; numerous microchaetae uniformly distributed; posterior margin of head with 10+10 chaetae (
|
||
<figureCitation id="132CAAC8FFCAFFA41403F891FC3EB74E" box="[812,897,1833,1858]" captionStart="FIGURES 54 – 59" captionStartId="14.[151,264,1812,1835]" captionTargetBox="[168,1426,206,1779]" captionTargetId="figure@14.[151,1436,193,1791]" captionTargetPageId="14" captionText="FIGURES 54 – 59. Dorsal cephalic and body chaetotaxy: 54 & 57, M. absens sp. nov.; 55 & 58, M. mulunensis sp. nov.; 56 & 59, M. trigrandis sp. nov. Large circle: macrochaeta; small circle: mesochaeta; wavy line: bothriotricha; s: normal s-chaeta; ms: s-microchaeta; rs: rod-like s-chaeta on body; circle with a slash: pseudopore." httpUri="https://zenodo.org/record/228019/files/figure.png" pageId="5" pageNumber="562">Fig. 54</figureCitation>
|
||
). Head ventrally scaled, with numerous smooth chaetae of different sizes.
|
||
</paragraph>
|
||
<paragraph id="8BA8B64DFFCAFFA717E8F8C9FD77B184" blockId="5.[151,1436,1725,2038]" lastBlockId="6.[151,1437,151,2012]" lastPageId="6" lastPageNumber="563" pageId="5" pageNumber="562">
|
||
Trochanteral-femoral organ with 1, 1 slender smooth larger chaetae and several additional smaller chaetae on trochanter (
|
||
<figureCitation id="132CAAC8FFCAFFA41635F82DFECEB7A2" box="[282,369,1941,1966]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1571,1594]" captionTargetBox="[182,1426,185,1546]" captionTargetId="figure@5.[167,1431,165,1555]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Monodontocerus absens sp. nov.: 15, labial chaetotaxy; 16, trochanteral-femoral organ; 17, femur and tibiotarsus; 18, dorsal view of tibiotarsal distal chaetae on front leg (TH: tenent hair, the same below); 19, anterior view of tibiotarsal distal chaetae on hind leg; 20, pattern of tibiotarsal distal chaetae (hind leg, distal view); 21, claw; 22, tenaculum. Scale bar: 15, 19, 21, 22, 50 Μm; 16, 120 Μm; 17, 200 Μm; 18, 40 Μm." httpUri="https://zenodo.org/record/228014/files/figure.png" pageId="5" pageNumber="562">Fig. 16</figureCitation>
|
||
). Fore, middle and hind femur respectively with at least 3, 1, 1 long outstanding chaetae on inner side. Fore, middle and hind tibiotarsus distally with 0, 0, 2 blunt spiny inner chaetae (
|
||
<figureCitation id="132CAAC8FFCAFFA413DEF801FAF5B7DE" box="[1265,1354,1977,2002]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1571,1594]" captionTargetBox="[182,1426,185,1546]" captionTargetId="figure@5.[167,1431,165,1555]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Monodontocerus absens sp. nov.: 15, labial chaetotaxy; 16, trochanteral-femoral organ; 17, femur and tibiotarsus; 18, dorsal view of tibiotarsal distal chaetae on front leg (TH: tenent hair, the same below); 19, anterior view of tibiotarsal distal chaetae on hind leg; 20, pattern of tibiotarsal distal chaetae (hind leg, distal view); 21, claw; 22, tenaculum. Scale bar: 15, 19, 21, 22, 50 Μm; 16, 120 Μm; 17, 200 Μm; 18, 40 Μm." httpUri="https://zenodo.org/record/228014/files/figure.png" pageId="5" pageNumber="562">Fig. 17</figureCitation>
|
||
). Each tibiotarsus with a distal whorl of 11 chaetae, ventral six as ordinary chaetae, dorsal five including one small, pointed tenent hair, 1+1 small accessory chaetae on either side of tenent hair and 1+1 long, slender, pointed chaetae external to tenent hair and accessory chaetae; on front leg these two small accessory chaetae subequal in size (
|
||
<figureCitation id="132CAAC8FFC9FFA71241FF04FF0BB0F4" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1571,1594]" captionTargetBox="[182,1426,185,1546]" captionTargetId="figure@5.[167,1431,165,1555]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Monodontocerus absens sp. nov.: 15, labial chaetotaxy; 16, trochanteral-femoral organ; 17, femur and tibiotarsus; 18, dorsal view of tibiotarsal distal chaetae on front leg (TH: tenent hair, the same below); 19, anterior view of tibiotarsal distal chaetae on hind leg; 20, pattern of tibiotarsal distal chaetae (hind leg, distal view); 21, claw; 22, tenaculum. Scale bar: 15, 19, 21, 22, 50 Μm; 16, 120 Μm; 17, 200 Μm; 18, 40 Μm." httpUri="https://zenodo.org/record/228014/files/figure.png" pageId="6" pageNumber="563">Fig. 18</figureCitation>
|
||
) but on mid and hind leg the posterior small chaeta is larger than the anterior one (
|
||
<figureCitation id="132CAAC8FFC9FFA71379FF67FB65B0F4" box="[1110,1242,223,248]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1571,1594]" captionTargetBox="[182,1426,185,1546]" captionTargetId="figure@5.[167,1431,165,1555]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Monodontocerus absens sp. nov.: 15, labial chaetotaxy; 16, trochanteral-femoral organ; 17, femur and tibiotarsus; 18, dorsal view of tibiotarsal distal chaetae on front leg (TH: tenent hair, the same below); 19, anterior view of tibiotarsal distal chaetae on hind leg; 20, pattern of tibiotarsal distal chaetae (hind leg, distal view); 21, claw; 22, tenaculum. Scale bar: 15, 19, 21, 22, 50 Μm; 16, 120 Μm; 17, 200 Μm; 18, 40 Μm." httpUri="https://zenodo.org/record/228014/files/figure.png" pageId="6" pageNumber="563">Figs 19, 20</figureCitation>
|
||
). Unguis slender, outer edge 0.24, 0.21, 0.16 times length of tibiotarsus, respectively, for front, mid and hind leg (
|
||
<figureCitation id="132CAAC8FFC9FFA713EFFEBCFAADB110" box="[1216,1298,260,285]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1571,1594]" captionTargetBox="[182,1426,185,1546]" captionTargetId="figure@5.[167,1431,165,1555]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Monodontocerus absens sp. nov.: 15, labial chaetotaxy; 16, trochanteral-femoral organ; 17, femur and tibiotarsus; 18, dorsal view of tibiotarsal distal chaetae on front leg (TH: tenent hair, the same below); 19, anterior view of tibiotarsal distal chaetae on hind leg; 20, pattern of tibiotarsal distal chaetae (hind leg, distal view); 21, claw; 22, tenaculum. Scale bar: 15, 19, 21, 22, 50 Μm; 16, 120 Μm; 17, 200 Μm; 18, 40 Μm." httpUri="https://zenodo.org/record/228014/files/figure.png" pageId="6" pageNumber="563">Fig. 17</figureCitation>
|
||
), with basointernal ridging visible in lateral view; lateral teeth pointed, of moderate size; inner side of unguis with five teeth, the subbasal tooth strongest. Unguiculus slender, two-thirds length of inner edge of unguis, its inner side with one tooth (
|
||
<figureCitation id="132CAAC8FFC9FFA717CFFED7FE8CB184" box="[224,307,367,392]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1571,1594]" captionTargetBox="[182,1426,185,1546]" captionTargetId="figure@5.[167,1431,165,1555]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Monodontocerus absens sp. nov.: 15, labial chaetotaxy; 16, trochanteral-femoral organ; 17, femur and tibiotarsus; 18, dorsal view of tibiotarsal distal chaetae on front leg (TH: tenent hair, the same below); 19, anterior view of tibiotarsal distal chaetae on hind leg; 20, pattern of tibiotarsal distal chaetae (hind leg, distal view); 21, claw; 22, tenaculum. Scale bar: 15, 19, 21, 22, 50 Μm; 16, 120 Μm; 17, 200 Μm; 18, 40 Μm." httpUri="https://zenodo.org/record/228014/files/figure.png" pageId="6" pageNumber="563">Fig. 21</figureCitation>
|
||
). Prestarsus with 1+1 microchaetae.
|
||
</paragraph>
|
||
<paragraph id="8BA8B64DFFC9FFA717E8FE2CFE7CB1F8" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="563">
|
||
Ventral tube scaled on both faces, anterior face with 50–70 chaetae on each side, posterior face with 110–150 chaetae; lateral flap unscaled, each side with about 75 chaetae. Rami of tenaculum with 4+4 teeth, corpus without scales or chaetae (
|
||
<figureCitation id="132CAAC8FFC9FFA7164DFE64FE09B1F8" box="[354,438,476,501]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1571,1594]" captionTargetBox="[182,1426,185,1546]" captionTargetId="figure@5.[167,1431,165,1555]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Monodontocerus absens sp. nov.: 15, labial chaetotaxy; 16, trochanteral-femoral organ; 17, femur and tibiotarsus; 18, dorsal view of tibiotarsal distal chaetae on front leg (TH: tenent hair, the same below); 19, anterior view of tibiotarsal distal chaetae on hind leg; 20, pattern of tibiotarsal distal chaetae (hind leg, distal view); 21, claw; 22, tenaculum. Scale bar: 15, 19, 21, 22, 50 Μm; 16, 120 Μm; 17, 200 Μm; 18, 40 Μm." httpUri="https://zenodo.org/record/228014/files/figure.png" pageId="6" pageNumber="563">Fig. 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA8B64DFFC9FFA717E8FDB8FE24B2C0" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="563">
|
||
Furca about half as long as body, ratio manubrium: dens: mucro as 4.7: 6.7: 1.0. Manubrial width: length about 1.0: 1.7, ventrally scaled, laterally with large round scales and a row of about 10 chaetae, distal chaetae strong and serrated, proximal chaetae smaller. Each side of manubrium with a dorsal stripe of about 190 smooth chaetae of various sizes mixed with numerous scales; external to the stripe, 26–28 pseudopores present in a longitudinal row (
|
||
<figureCitation id="132CAAC8FFC9FFA717B0FD37FF4DB2A4" box="[159,242,655,680]" captionStart="FIGURES 23 – 29" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[175,1415,206,1886]" captionTargetId="figure@7.[158,1422,195,1894]" captionTargetPageId="7" captionText="FIGURES 23 – 29. Monodontocerus absens sp. nov.: 23, manubrium (" T " shaped marks: sockets of scales; circle with a slash: pseudopore); 24, disto-external corner of manubrium and corner chaeta; 25, dental spines; 26, feathered dental chaeta; 27, mucro, apical tooth hypothesized; 28, mature female genital plate; 29, immature female genital plate. Scale bar: 23, 25, 100 Μm; 24, 26 – 29, 50 Μm." httpUri="https://zenodo.org/record/228015/files/figure.png" pageId="6" pageNumber="563">Fig. 23</figureCitation>
|
||
); one chaeta at each externo-distal corner of manubrium as long as a medium-size mesochaeta in the dorsal chaetal stripe (
|
||
<figureCitation id="132CAAC8FFC9FFA71614FD0CFE30B2C0" box="[315,399,692,717]" captionStart="FIGURES 23 – 29" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[175,1415,206,1886]" captionTargetId="figure@7.[158,1422,195,1894]" captionTargetPageId="7" captionText="FIGURES 23 – 29. Monodontocerus absens sp. nov.: 23, manubrium (" T " shaped marks: sockets of scales; circle with a slash: pseudopore); 24, disto-external corner of manubrium and corner chaeta; 25, dental spines; 26, feathered dental chaeta; 27, mucro, apical tooth hypothesized; 28, mature female genital plate; 29, immature female genital plate. Scale bar: 23, 25, 100 Μm; 24, 26 – 29, 50 Μm." httpUri="https://zenodo.org/record/228015/files/figure.png" pageId="6" pageNumber="563">Fig. 24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA8B64DFFC9FFA717E8FD60FD71B3E1" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="563">
|
||
Dens internally divided into three subsegments, length of basal part: medial part: distal part about 1.0: 1.4: 3.8. Dental spine formula 4,
|
||
<date id="FFA9908DFFC9FFA71695FD44FE63B319" box="[442,476,764,789]" bridgedPair="/" pageId="6" pageNumber="563">I/5</date>
|
||
, I, 3, I in a single row (
|
||
<figureCitation id="132CAAC8FFC9FFA71427FD44FCDEB319" box="[776,865,764,789]" captionStart="FIGURES 23 – 29" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[175,1415,206,1886]" captionTargetId="figure@7.[158,1422,195,1894]" captionTargetPageId="7" captionText="FIGURES 23 – 29. Monodontocerus absens sp. nov.: 23, manubrium (" T " shaped marks: sockets of scales; circle with a slash: pseudopore); 24, disto-external corner of manubrium and corner chaeta; 25, dental spines; 26, feathered dental chaeta; 27, mucro, apical tooth hypothesized; 28, mature female genital plate; 29, immature female genital plate. Scale bar: 23, 25, 100 Μm; 24, 26 – 29, 50 Μm." httpUri="https://zenodo.org/record/228015/files/figure.png" pageId="6" pageNumber="563">Fig. 25</figureCitation>
|
||
). Each dental spine consisting of a strong trunk surrounded by 2–6 denticles near the base; largest dental spine about 0.08 times length of dens. Three
|
||
<typeStatus id="54AC08EFFFC9FFA7126FFC98FAC4B334" box="[1344,1403,800,824]" pageId="6" pageNumber="563">types</typeStatus>
|
||
of dorsal chaetae on dens besides dental spines:
|
||
<typeStatus id="54AC08EFFFC9FFA715B5FCFDFD75B351" box="[666,714,837,861]" pageId="6" pageNumber="563">type</typeStatus>
|
||
1 long, slender and smooth;
|
||
<typeStatus id="54AC08EFFFC9FFA7133BFCFDFBFBB351" box="[1044,1092,837,861]" pageId="6" pageNumber="563">type</typeStatus>
|
||
2 similar to
|
||
<typeStatus id="54AC08EFFFC9FFA713FAFCFDFABAB351" box="[1237,1285,837,861]" pageId="6" pageNumber="563">type</typeStatus>
|
||
1 but shorter and spiny;
|
||
<typeStatus id="54AC08EFFFC9FFA7160FFCD0FEEFB38C" box="[288,336,872,896]" pageId="6" pageNumber="563">type</typeStatus>
|
||
3 feather-like, ciliated heavily on outer side and slightly on internal side (
|
||
<figureCitation id="132CAAC8FFC9FFA713C7FCDFFAFCB38C" box="[1256,1347,871,896]" captionStart="FIGURES 23 – 29" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[175,1415,206,1886]" captionTargetId="figure@7.[158,1422,195,1894]" captionTargetPageId="7" captionText="FIGURES 23 – 29. Monodontocerus absens sp. nov.: 23, manubrium (" T " shaped marks: sockets of scales; circle with a slash: pseudopore); 24, disto-external corner of manubrium and corner chaeta; 25, dental spines; 26, feathered dental chaeta; 27, mucro, apical tooth hypothesized; 28, mature female genital plate; 29, immature female genital plate. Scale bar: 23, 25, 100 Μm; 24, 26 – 29, 50 Μm." httpUri="https://zenodo.org/record/228015/files/figure.png" pageId="6" pageNumber="563">Fig. 26</figureCitation>
|
||
). Basal subsegment with only
|
||
<typeStatus id="54AC08EFFFC9FFA716B0FC35FE70B3A9" box="[415,463,909,933]" pageId="6" pageNumber="563">type</typeStatus>
|
||
1 chaetae; medial subsegment with
|
||
<typeStatus id="54AC08EFFFC9FFA71454FC35FC14B3A9" box="[891,939,909,933]" pageId="6" pageNumber="563">type</typeStatus>
|
||
1 and
|
||
<typeStatus id="54AC08EFFFC9FFA7132DFC35FB8DB3A9" box="[1026,1074,909,933]" pageId="6" pageNumber="563">type</typeStatus>
|
||
2 chaetae, the later distributed externally; distal subsegment with all three
|
||
<typeStatus id="54AC08EFFFC9FFA715B8FC08FD6DB3C4" box="[663,722,944,968]" pageId="6" pageNumber="563">types</typeStatus>
|
||
of chaetae,
|
||
<typeStatus id="54AC08EFFFC9FFA71449FC08FC29B3C4" box="[870,918,944,968]" pageId="6" pageNumber="563">type</typeStatus>
|
||
3 mainly along the outer and inner margin. Ventral side of dens covered only with oval scales.
|
||
</paragraph>
|
||
<paragraph id="8BA8B64DFFC9FFA717E8FC40FADAB454" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="563">
|
||
Mucro elongated, bearing numerous smooth chaetae with elongated sockets. Two dorsal lamellae beginning from base of mucro and ending in subapical tooth; single basal tooth with one proximal lamella; two intermediate teeth unequal with the distal tooth larger; apex of apical tooth broken in all specimens, not observed (
|
||
<figureCitation id="132CAAC8FFC9FFA7122AFB87FAE6B454" box="[1285,1369,1087,1112]" captionStart="FIGURES 23 – 29" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[175,1415,206,1886]" captionTargetId="figure@7.[158,1422,195,1894]" captionTargetPageId="7" captionText="FIGURES 23 – 29. Monodontocerus absens sp. nov.: 23, manubrium (" T " shaped marks: sockets of scales; circle with a slash: pseudopore); 24, disto-external corner of manubrium and corner chaeta; 25, dental spines; 26, feathered dental chaeta; 27, mucro, apical tooth hypothesized; 28, mature female genital plate; 29, immature female genital plate. Scale bar: 23, 25, 100 Μm; 24, 26 – 29, 50 Μm." httpUri="https://zenodo.org/record/228015/files/figure.png" pageId="6" pageNumber="563">Fig. 27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA8B64DFFC9FFA717E8FBDCFCAAB621" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="563">
|
||
Macrochaetae distributed densely along anterior margin of Th. II and sparingly in posterior rows on terga. On each side, Th. II–Abd. V with 3, 3/ 3, 3, 4, 2, 4 macrochaetae in posterior rows; Th. II with three anterior macrochaetae in a near-longitudinal line near the Th. II midline; two anterior macrochaetae on Abd. III, subequal in size; two antero-lateral macrochaetae on Abd. IV. On Abd. VI, seven macrochaetae present on upper flap. Sizes of body macrochaetae unequal, those on anterior segments generally smaller than those on posterior segments and especially for those in posterior rows; on Abd. V, chaeta p3 larger than the others and shifted slightly anterior to p1 and p2 (
|
||
<figureCitation id="132CAAC8FFC9FFA717DFFA84FEFBB559" box="[240,324,1340,1365]" captionStart="FIGURES 54 – 59" captionStartId="14.[151,264,1812,1835]" captionTargetBox="[168,1426,206,1779]" captionTargetId="figure@14.[151,1436,193,1791]" captionTargetPageId="14" captionText="FIGURES 54 – 59. Dorsal cephalic and body chaetotaxy: 54 & 57, M. absens sp. nov.; 55 & 58, M. mulunensis sp. nov.; 56 & 59, M. trigrandis sp. nov. Large circle: macrochaeta; small circle: mesochaeta; wavy line: bothriotricha; s: normal s-chaeta; ms: s-microchaeta; rs: rod-like s-chaeta on body; circle with a slash: pseudopore." httpUri="https://zenodo.org/record/228019/files/figure.png" pageId="6" pageNumber="563">Fig. 57</figureCitation>
|
||
). Bothriotricha extremely thin, ciliated, pattern 2, 1/ 0, 0, 1, 2, 0, 0 from Th. II to Abd. VI as in other Tomocerinae. Mesochaetae of various sizes mainly distributed laterally on terga and posteriorly on Abd. VI. Microchaetae uniformly distributed on terga, pattern not studied. Numerous s-chaetae on terga, but patterns difficult to see; the following have been observed: on Th. II two rod-like s-chaetae, one s-microchaeta and one normal s-chaeta (?) close to bothriotricha; on Abd. III one s-microchaeta posterior to antero-external macrochaeta and three normal s-chaetae close to bothriotricha; on Abd. IV long s-chaetae uniformly distributed, their number uncertain; Abd. V with a single anterior normal s-chaeta.
|
||
</paragraph>
|
||
<paragraph id="8BA8B64DFFC9FFA717E8F98FFD4DB695" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="563">
|
||
Pseudopores near the middle line of terga, 1, 1, 1, 1, 1, 1, 0, 0 on each side (
|
||
<figureCitation id="132CAAC8FFC9FFA71315F98FFB2FB65C" box="[1082,1168,1591,1616]" captionStart="FIGURES 54 – 59" captionStartId="14.[151,264,1812,1835]" captionTargetBox="[168,1426,206,1779]" captionTargetId="figure@14.[151,1436,193,1791]" captionTargetPageId="14" captionText="FIGURES 54 – 59. Dorsal cephalic and body chaetotaxy: 54 & 57, M. absens sp. nov.; 55 & 58, M. mulunensis sp. nov.; 56 & 59, M. trigrandis sp. nov. Large circle: macrochaeta; small circle: mesochaeta; wavy line: bothriotricha; s: normal s-chaeta; ms: s-microchaeta; rs: rod-like s-chaeta on body; circle with a slash: pseudopore." httpUri="https://zenodo.org/record/228019/files/figure.png" pageId="6" pageNumber="563">Fig. 57</figureCitation>
|
||
). On Th. II pseudopore close to the posterior most medial macrochaeta; on Th. III–Abd. IV pseudopores more near middle line and on Abd. I and Abd. II more posterior than on other terga.
|
||
</paragraph>
|
||
<paragraph id="8BA8B64DFFC9FFA717E8F91DFB82B6EC" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="563">
|
||
Two
|
||
<typeStatus id="54AC08EFFFC9FFA717D0F91DFE85B6B1" box="[255,314,1701,1725]" pageId="6" pageNumber="563">types</typeStatus>
|
||
of female genital plate observed, one highly wrinkled with 2+2 microchaetae (
|
||
<figureCitation id="132CAAC8FFC9FFA71385F91CFB44B6B0" box="[1194,1275,1700,1725]" captionStart="FIGURES 23 – 29" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[175,1415,206,1886]" captionTargetId="figure@7.[158,1422,195,1894]" captionTargetPageId="7" captionText="FIGURES 23 – 29. Monodontocerus absens sp. nov.: 23, manubrium (" T " shaped marks: sockets of scales; circle with a slash: pseudopore); 24, disto-external corner of manubrium and corner chaeta; 25, dental spines; 26, feathered dental chaeta; 27, mucro, apical tooth hypothesized; 28, mature female genital plate; 29, immature female genital plate. Scale bar: 23, 25, 100 Μm; 24, 26 – 29, 50 Μm." httpUri="https://zenodo.org/record/228015/files/figure.png" pageId="6" pageNumber="563">Fig. 28</figureCitation>
|
||
) and the other less wrinkled without chaetae (
|
||
<figureCitation id="132CAAC8FFC9FFA716DDF97FFDFAB6EC" box="[498,581,1735,1760]" captionStart="FIGURES 23 – 29" captionStartId="7.[151,264,1916,1939]" captionTargetBox="[175,1415,206,1886]" captionTargetId="figure@7.[158,1422,195,1894]" captionTargetPageId="7" captionText="FIGURES 23 – 29. Monodontocerus absens sp. nov.: 23, manubrium (" T " shaped marks: sockets of scales; circle with a slash: pseudopore); 24, disto-external corner of manubrium and corner chaeta; 25, dental spines; 26, feathered dental chaeta; 27, mucro, apical tooth hypothesized; 28, mature female genital plate; 29, immature female genital plate. Scale bar: 23, 25, 100 Μm; 24, 26 – 29, 50 Μm." httpUri="https://zenodo.org/record/228015/files/figure.png" pageId="6" pageNumber="563">Fig. 29</figureCitation>
|
||
), the latter probably from an immature form.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30DE5C6FFC9FFA717E8F954FAF9B709" box="[199,1350,1772,1797]" pageId="6" pageNumber="563" type="etymology">
|
||
<paragraph id="8BA8B64DFFC9FFA717E8F954FAF9B709" blockId="6.[151,1437,151,2012]" box="[199,1350,1772,1797]" pageId="6" pageNumber="563">
|
||
<emphasis id="B9636A5FFFC9FFA717E8F954FEF9B709" bold="true" box="[199,326,1772,1797]" pageId="6" pageNumber="563">Etymology</emphasis>
|
||
. Named after the absence of chaetae on tenaculum, from the Latin
|
||
<taxonomicName id="4C17CDCEFFC9FFA71318F954FB3CB709" box="[1079,1155,1772,1797]" class="Entognatha" family="Tomoceridae" genus="Monodontocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="563" phylum="Arthropoda" rank="species" species="absens">
|
||
<emphasis id="B9636A5FFFC9FFA71318F954FB3CB709" box="[1079,1155,1772,1797]" italics="true" pageId="6" pageNumber="563">absens</emphasis>
|
||
</taxonomicName>
|
||
: absent, missing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30DE5C6FFC9FFA717E8F8B7FF75B7D0" pageId="6" pageNumber="563" type="discussion">
|
||
<paragraph id="8BA8B64DFFC9FFA717E8F8B7FF75B7D0" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="563">
|
||
<emphasis id="B9636A5FFFC9FFA717E8F8B7FE84B724" bold="true" box="[199,315,1807,1832]" pageId="6" pageNumber="563">Remarks.</emphasis>
|
||
This species is different from other
|
||
<emphasis id="B9636A5FFFC9FFA715CBF8B7FC1DB724" box="[740,930,1807,1832]" italics="true" pageId="6" pageNumber="563">Monodontocerus</emphasis>
|
||
species mainly in the pattern of interocular chaetotaxy (2, 1, 2 while the others have 2, 3 or 2, 5) and the absence of chaetae on tenaculum (at least one present in other species). Besides, its antenna is distinctly longer than body, while other species have antennae shorter than body (
|
||
<tableCitation id="C69583F6FFC9FFA717F2F8C4FE8BB798" box="[221,308,1916,1941]" captionStart="TABLE 2" captionStartId="8.[151,239,152,175]" captionTargetBox="[151,1391,234,594]" captionTargetPageId="8" captionText="TABLE 2. Variability of characters among Monodontocerus. A, anterior area; IO, interocular area; PO, postocular area; P, posterior area;?, unknown data." httpUri="http://table.plazi.org/id/DF68E6C5FFC7FFA917B8FF20FDB8B0C3" pageId="6" pageNumber="563" tableUuid="DF68E6C5FFC7FFA917B8FF20FDB8B0C3">Table 2</tableCitation>
|
||
). The most distal spine on the basal part of dens is nearly as large as large spines on the medial part, but the difference in size between small and large spines on the basal part is not as distinct as it is on the medial part.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |