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<mods:title id="E3133570C5E7C57C983641D876924821">Taxonomic revision of the family Eosentomidae (Hexapoda: Protura) from Japan 2701</mods:title>
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<taxonomicName id="4C1397E09B43FFEA469A3B3BF2F57946" authority="Imadate" authorityName="Imadate" box="[151,483,610,636]" class="Entognatha" family="Eosentomidae" genus="Eosentomon" kingdom="Animalia" order="Protura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="tokui">
<emphasis id="B96730719B43FFEA469A3B3BF2F57946" bold="true" box="[151,483,610,636]" pageId="96" pageNumber="97">
<emphasis id="B96730719B43FFEA469A3B3BF2657946" bold="true" box="[151,371,610,636]" italics="true" pageId="96" pageNumber="97">Eosentomon tokui</emphasis>
Imadaté
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C309BFE89B43FFEA469A3BDCF3F079A7" box="[151,230,645,669]" pageId="96" pageNumber="97" type="description">
<paragraph id="8BACEC639B43FFEA469A3BDCF3F079A7" blockId="96.[151,483,610,669]" box="[151,230,645,669]" pageId="96" pageNumber="97">
<figureCitation id="1328F0E69B43FFEA469A3BDCF3F079A7" box="[151,230,645,669]" captionStart="FIGURE 54" captionStartId="96.[151,255,1891,1915]" captionTargetBox="[195,1373,775,1876]" captionTargetId="figure-263@96.[190,1379,770,1885]" captionTargetPageId="96" captionText="FIGURE 54. Eosentomon tokui Imadaté from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm." pageId="96" pageNumber="97">Fig. 54</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C309BFE89B43FFEB469A3B9EF2AC792D" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97" type="reference_group">
<paragraph id="8BACEC639B43FFEB469A3B9EF03B7BCD" blockId="96.[151,1029,710,733]" lastBlockId="97.[151,1437,151,608]" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97">
<treatmentCitationGroup id="AB03CB4D9B43FFEB469A3B9EF03B7BCD" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97">
<treatmentCitation id="0AB2CA729B43FFEA469A3B9EF13279E6" author="Imadate, G." box="[151,548,710,733]" page="597" pageId="96" pageNumber="97" year="1974">
<taxonomicName id="4C1397E09B43FFEA469A3B9EF13279E6" authority="Imadate, 1974 a: 597" authorityName="Imadate" authorityPageNumber="597" authorityYear="1974" box="[151,548,710,733]" class="Entognatha" family="Eosentomidae" genus="Eosentomon" kingdom="Animalia" order="Protura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="tokui">
<emphasis id="B96730719B43FFEA469A3B9EF25B79E7" box="[151,333,711,733]" italics="true" pageId="96" pageNumber="97">Eosentomon tokui</emphasis>
<bibRefCitation id="EF8291929B43FFEA475E3B9FF13279E6" author="Imadate, G." box="[339,548,710,732]" pageId="96" pageNumber="97" pagination="603 - 628" refId="ref63091" refString="Imadate, G. (1974 a) Contributions towards a revision of Japanese Protura. Revue d'Ecologie et du Biologie du Sol, 10 (for 1973), 603 - 628." type="journal article" year="1974">Imadaté, 1974a: 597</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 12;
<treatmentCitation id="0AB2CA729B43FFEA44983B9FF07E79E6" author="Imadate, G." box="[661,872,710,732]" page="328" pageId="96" pageNumber="97" year="1974">
<bibRefCitation id="EF8291929B43FFEA44983B9FF07E79E6" author="Imadate, G." box="[661,872,710,732]" pageId="96" pageNumber="97" refId="ref63125" refString="Imadate, G. (1974 b) Fauna Japonica. Protura (Insecta). Keigaku Publishing Company, Ltd., Tokyo, 351 pp." type="book" year="1974">Imadaté, 1974b: 328</bibRefCitation>
</treatmentCitation>
, figs. 178179.
<emphasis id="B96730719B42FFEB469A39CEF28E7B8A" bold="true" box="[151,408,151,176]" inLineHeading="true" pageId="97" pageNumber="98" reason="1">Specimens examined.</emphasis>
Specimens deposited in NSMT:
<specimenCount id="9D1527EA9B42FFEB451E39C1F0B67B95" box="[787,928,151,175]" pageId="97" pageNumber="98" type="female">two females</specimenCount>
, NSMTAp 77, Kamishishiori, Kesennumashi,
<collectingRegion id="49D722819B42FFEB46C839E5F2847BEE" box="[197,402,188,212]" country="Japan" name="Miyagi" pageId="97" pageNumber="98">Miyagi Prefecture</collectingRegion>
,
<date id="FFADCAA39B42FFEB479139E5F1357BEE" box="[412,547,188,212]" pageId="97" pageNumber="98" value="1971-11-27">27-XI-1971</date>
, T. Watanabe leg. (same data as for the
<typeStatus id="54A852C19B42FFEB45D739E5F7577BEE" box="[986,1089,188,212]" pageId="97" pageNumber="98" type="holotype">holotype</typeStatus>
);
<specimenCount id="9D1527EA9B42FFEB425E39E5F7C87BEE" box="[1107,1246,188,212]" pageId="97" pageNumber="98" type="female">two females</specimenCount>
, same data as for the
<typeStatus id="54A852C19B42FFEB46CD3986F2347BCD" box="[192,290,223,247]" pageId="97" pageNumber="98" type="holotype">holotype</typeStatus>
(No.0086
<quantity id="4CEB41869B42FFEB47963986F2D37BCD" box="[411,453,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="97" pageNumber="98" unit="in" value="2.0">2 in</quantity>
Imadatés Proturan Collection).
</treatmentCitationGroup>
</paragraph>
<caption id="DF6CBCEB9B43FFEA469A3E3AF1C77CE0" pageId="96" pageNumber="97" startId="96.[151,255,1891,1915]" targetBox="[195,1373,775,1876]" targetPageId="96">
<paragraph id="8BACEC639B43FFEA469A3E3AF1C77CE0" blockId="96.[151,1436,1891,2011]" pageId="96" pageNumber="97">
<emphasis id="B96730719B43FFEA469A3E3AF23E7C41" bold="true" box="[151,296,1891,1915]" pageId="96" pageNumber="97">FIGURE 54.</emphasis>
<taxonomicName id="4C1397E09B43FFEA47223E3AF1587C40" authority="Imadate" authorityName="Imadate" box="[303,590,1891,1914]" class="Entognatha" family="Eosentomidae" genus="Eosentomon" kingdom="Animalia" order="Protura" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="tokui">
<emphasis id="B96730719B43FFEA47223E3AF2E67C40" box="[303,496,1891,1914]" italics="true" pageId="96" pageNumber="97">Eosentomon tokui</emphasis>
Imadaté
</taxonomicName>
from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm.
</paragraph>
</caption>
<paragraph id="8BACEC639B42FFEB46CA385AF2AC792D" blockId="97.[151,1437,151,608]" pageId="97" pageNumber="98">
<emphasis id="B96730719B42FFEB46CA385AF2F97A26" bold="true" box="[199,495,259,284]" inLineHeading="true" pageId="97" pageNumber="98" reason="1">Additions to description.</emphasis>
On head, setae
<emphasis id="B96730719B42FFEB44AE385DF1D37A26" box="[675,709,260,284]" italics="true" pageId="97" pageNumber="98">m4</emphasis>
present,
<emphasis id="B96730719B42FFEB4521385CF05E7A26" box="[812,840,261,284]" italics="true" pageId="97" pageNumber="98">aa</emphasis>
and
<emphasis id="B96730719B42FFEB4572385CF08D7A26" box="[895,923,261,284]" italics="true" pageId="97" pageNumber="98">pa</emphasis>
absent, sensilla
<emphasis id="B96730719B42FFEB4259385CF77B7A26" box="[1108,1133,261,284]" italics="true" pageId="97" pageNumber="98">as</emphasis>
and
<emphasis id="B96730719B42FFEB42A9385CF7AB7A26" box="[1188,1213,261,284]" italics="true" pageId="97" pageNumber="98">ps</emphasis>
present (
<figureCitation id="1328F0E69B42FFEB432B385DF6997A26" box="[1318,1423,260,284]" captionStart="FIGURE 54" captionStartId="96.[151,255,1891,1915]" captionTargetBox="[195,1373,775,1876]" captionTargetId="figure-263@96.[190,1379,770,1885]" captionTargetPageId="96" captionText="FIGURE 54. Eosentomon tokui Imadaté from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm." pageId="97" pageNumber="98">Fig. 54A</figureCitation>
); seta
<emphasis id="B96730719B42FFEB46C63870F3F27A7A" box="[203,228,297,320]" italics="true" pageId="97" pageNumber="98">sp</emphasis>
1.41.5 times longer than
<emphasis id="B96730719B42FFEB44173870F13E7A7A" box="[538,552,297,320]" italics="true" pageId="97" pageNumber="98">p</emphasis>
; sensilla
<emphasis id="B96730719B42FFEB44983870F1A77A7A" box="[661,689,297,320]" italics="true" pageId="97" pageNumber="98">pp</emphasis>
rudimentary. Labral setae present. Seta
<emphasis id="B96730719B42FFEB42733870F7827A7A" box="[1150,1172,297,320]" italics="true" pageId="97" pageNumber="98">rs</emphasis>
inflated, equal to
<emphasis id="B96730719B42FFEB436A3870F66B7A7A" box="[1383,1405,297,320]" italics="true" pageId="97" pageNumber="98">sr</emphasis>
in length (
<figureCitation id="1328F0E69B42FFEB46E03815F2427A5E" box="[237,340,332,356]" captionStart="FIGURE 54" captionStartId="96.[151,255,1891,1915]" captionTargetBox="[195,1373,775,1876]" captionTargetId="figure-263@96.[190,1379,770,1885]" captionTargetPageId="96" captionText="FIGURE 54. Eosentomon tokui Imadaté from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm." pageId="97" pageNumber="98">Fig. 54B</figureCitation>
). On maxillary palpus (
<figureCitation id="1328F0E69B42FFEB44533815F1D27A5E" box="[606,708,332,356]" captionStart="FIGURE 54" captionStartId="96.[151,255,1891,1915]" captionTargetBox="[195,1373,775,1876]" captionTargetId="figure-263@96.[190,1379,770,1885]" captionTargetPageId="96" captionText="FIGURE 54. Eosentomon tokui Imadaté from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm." pageId="97" pageNumber="98">Fig. 54C</figureCitation>
) sensillum
<emphasis id="B96730719B42FFEB454B3815F07E7A5E" box="[838,872,332,356]" italics="true" pageId="97" pageNumber="98">md</emphasis>
slightly longer then
<emphasis id="B96730719B42FFEB425E3815F7797A5E" box="[1107,1135,332,356]" italics="true" pageId="97" pageNumber="98">ml</emphasis>
. On galea (
<figureCitation id="1328F0E69B42FFEB42FF3815F64F7A5E" box="[1266,1369,332,356]" captionStart="FIGURE 54" captionStartId="96.[151,255,1891,1915]" captionTargetBox="[195,1373,775,1876]" captionTargetId="figure-263@96.[190,1379,770,1885]" captionTargetPageId="96" captionText="FIGURE 54. Eosentomon tokui Imadaté from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm." pageId="97" pageNumber="98">Fig. 54D</figureCitation>
) digit O longer than M and I; M slightly shorter than or as long as I. Mandible with two teeth (
<figureCitation id="1328F0E69B42FFEB42733836F7F27ABD" box="[1150,1252,367,391]" captionStart="FIGURE 54" captionStartId="96.[151,255,1891,1915]" captionTargetBox="[195,1373,775,1876]" captionTargetId="figure-263@96.[190,1379,770,1885]" captionTargetPageId="96" captionText="FIGURE 54. Eosentomon tokui Imadaté from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm." pageId="97" pageNumber="98">Fig. 54E</figureCitation>
). On hind tarsus (
<figureCitation id="1328F0E69B42FFEB469238CDF2157A96" box="[159,259,404,428]" captionStart="FIGURE 54" captionStartId="96.[151,255,1891,1915]" captionTargetBox="[195,1373,775,1876]" captionTargetId="figure-263@96.[190,1379,770,1885]" captionTargetPageId="96" captionText="FIGURE 54. Eosentomon tokui Imadaté from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm." pageId="97" pageNumber="98">Fig. 54F</figureCitation>
)
<emphasis id="B96730719B42FFEB471B38CDF22E7A96" box="[278,312,404,428]" italics="true" pageId="97" pageNumber="98">D2</emphasis>
and
<emphasis id="B96730719B42FFEB477838CDF2817A96" box="[373,407,404,428]" italics="true" pageId="97" pageNumber="98">D4</emphasis>
spine-like,
<emphasis id="B96730719B42FFEB442C38CDF1557A96" box="[545,579,404,428]" italics="true" pageId="97" pageNumber="98">D4</emphasis>
slightly more slender than
<emphasis id="B96730719B42FFEB458838CDF0B17A96" box="[901,935,404,428]" italics="true" pageId="97" pageNumber="98">D2</emphasis>
. Tracheal camerae contracted distally (
<figureCitation id="1328F0E69B42FFEB436338CDF3DD7AF5" captionStart="FIGURE 54" captionStartId="96.[151,255,1891,1915]" captionTargetBox="[195,1373,775,1876]" captionTargetId="figure-263@96.[190,1379,770,1885]" captionTargetPageId="96" captionText="FIGURE 54. Eosentomon tokui Imadaté from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm." pageId="97" pageNumber="98">Fig. 54G</figureCitation>
). Central lobe slightly sinuate (
<figureCitation id="1328F0E69B42FFEB442038EEF1807AF5" box="[557,662,439,463]" captionStart="FIGURE 54" captionStartId="96.[151,255,1891,1915]" captionTargetBox="[195,1373,775,1876]" captionTargetId="figure-263@96.[190,1379,770,1885]" captionTargetPageId="96" captionText="FIGURE 54. Eosentomon tokui Imadaté from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm." pageId="97" pageNumber="98">Fig. 54H</figureCitation>
). Laterostigmata IIIV large (
<figureCitation id="1328F0E69B42FFEB45E538EEF7507AF5" box="[1000,1094,439,463]" captionStart="FIGURE 54" captionStartId="96.[151,255,1891,1915]" captionTargetBox="[195,1373,775,1876]" captionTargetId="figure-263@96.[190,1379,770,1885]" captionTargetPageId="96" captionText="FIGURE 54. Eosentomon tokui Imadaté from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm." pageId="97" pageNumber="98">Fig. 54I</figureCitation>
), II with a central spot, IIIIV with no inner structure; that on V small. On abdominal tergite VIII (
<figureCitation id="1328F0E69B42FFEB45BC3885F7067ACE" box="[945,1040,476,500]" captionStart="FIGURE 54" captionStartId="96.[151,255,1891,1915]" captionTargetBox="[195,1373,775,1876]" captionTargetId="figure-263@96.[190,1379,770,1885]" captionTargetPageId="96" captionText="FIGURE 54. Eosentomon tokui Imadaté from Kamishishiori, Miyagi Prefecture (type locality), female. A, dorsal view of head, right side; B, labrum and rostral region; C, maxillary palpus; D, galea; E, mandible; F, dorsal view of hind tarsus; G, tracheal camerae on thoracic tergite III; H, central lobe on abdominal tergite VI; I, laterostigmata IIIV; J, posterior margin on abdominal tergite VIII. Scales: 20 µm." pageId="97" pageNumber="98">Fig. 54J</figureCitation>
)
<emphasis id="B96730719B42FFEB422C3885F7427ACE" box="[1057,1108,476,500]" italics="true" pageId="97" pageNumber="98">P1a'</emphasis>
without basal dilatation and anterior to
<emphasis id="B96730719B42FFEB471F3B59F2277922" box="[274,305,512,536]" italics="true" pageId="97" pageNumber="98">P2</emphasis>
;
<emphasis id="B96730719B42FFEB47323B59F27B7922" box="[319,365,512,536]" italics="true" pageId="97" pageNumber="98">P2a</emphasis>
linear.
</paragraph>
</subSubSection>
<subSubSection id="C309BFE89B42FFEB46CA3B7DF68B7906" box="[199,1437,548,572]" pageId="97" pageNumber="98" type="discussion">
<paragraph id="8BACEC639B42FFEB46CA3B7DF68B7906" blockId="97.[151,1437,151,608]" box="[199,1437,548,572]" pageId="97" pageNumber="98">
<emphasis id="B96730719B42FFEB46CA3B7DF22D7906" bold="true" box="[199,315,548,572]" inLineHeading="true" pageId="97" pageNumber="98" reason="1">Remarks.</emphasis>
The
<typeStatus id="54A852C19B42FFEB47793B7DF2C07906" box="[372,470,548,572]" pageId="97" pageNumber="98" type="holotype">holotype</typeStatus>
was not found in the collection of NSMT; hence, topotype specimens were examined.
</paragraph>
</subSubSection>
<subSubSection id="C309BFE89B42FFEB46CA3B1EF10E7965" box="[199,536,583,608]" pageId="97" pageNumber="98" type="distribution">
<paragraph id="8BACEC639B42FFEB46CA3B1EF10E7965" blockId="97.[151,1437,151,608]" box="[199,536,583,608]" pageId="97" pageNumber="98">
<emphasis id="B96730719B42FFEB46CA3B1EF276795A" bold="true" box="[199,352,583,608]" inLineHeading="true" pageId="97" pageNumber="98" reason="1">Distribution.</emphasis>
<collectingCountry id="F304ACF39B42FFEB476A3B11F2B17965" box="[359,423,584,607]" name="Japan" pageId="97" pageNumber="98">Japan</collectingCountry>
(Honshu)
</paragraph>
</subSubSection>
</treatment>
</document>