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<document id="399C16C892ADABE0D85C25E4103C903C" ID-CLB-Dataset="3079" ID-DOI="10.1649/072.068.0mo4.1" ID-GBIF-Dataset="62322993-ca80-4b6c-ac95-861cbb236abd" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907735" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623093043626" checkinUser="carolina" docAuthor="Park, J. - S. &amp; Carlton, C. E." docDate="2014" docId="03D82A3EBD6F11443F7EF9BEFD44FDD2" docLanguage="en" docName="TheColeopBull.68.mo13.1-156.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 13) 68" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Sagola hirtalis Broun, 1893 b: 1050" docType="key" docVersion="23" lastPageNumber="9" masterDocId="FFE15246BD69114C3C45FFF3FFD6FF9A" masterDocTitle="A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="7" updateTime="1699001444256" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6C3FEA7A82480E7A926F814461961505">A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)</mods:title>
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<mods:namePart id="2E7B669AC0A49AD78B8AEC2E7B53025B">Park, J. - S.</mods:namePart>
<mods:affiliation id="272C1BD3D8EAF97B233CB2691A9B3336">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:namePart id="1EB81D390748B4B3578E3B119A2B64EE">Carlton, C. E.</mods:namePart>
<mods:affiliation id="0E6DB6AC45D1293D54493D04ACDC23BC">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:title id="E87DE1274585C8EC554492340F9DE349">The Coleopterists Bulletin</mods:title>
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<mods:date id="248487B1EC0753A90D41FF0E0E67552C">2014</mods:date>
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<mods:title id="FDD913D827031EB69EC8F2BC7CB1CB66">mo 13</mods:title>
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<treatment id="03D82A3EBD6F11443F7EF9BEFD44FDD2" ID-DOI="http://doi.org/10.5281/zenodo.5765044" ID-GBIF-Taxon="191245046" ID-Zenodo-Dep="5765044" LSID="urn:lsid:plazi:treatment:03D82A3EBD6F11443F7EF9BEFD44FDD2" httpUri="http://treatment.plazi.org/id/03D82A3EBD6F11443F7EF9BEFD44FDD2" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="C36BC8A3BD6F114A3F7EF9BEFC26F9E6" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BCE9B28BD6F114A3F7EF9BEFC26F9E6" blockId="6.[827,1072,1613,1660]" pageId="6" pageNumber="7">
<heading id="D0862C44BD6F114A3F7EF9BEFBE6F9F8" box="[827,1072,1613,1634]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="7" reason="2">
<emphasis id="B905473ABD6F114A3F7EF9BEFBE6F9F8" bold="true" box="[827,1072,1613,1634]" pageId="6" pageNumber="7">
1.
<taxonomicName id="4C71E0ABBD6F114A3F12F9BEFC4AF9F8" ID-CoL="6WZP3" authorityName="Broun, 1893 b:" authorityYear="1050" box="[855,924,1613,1634]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hirtalis">
<emphasis id="B905473ABD6F114A3F12F9BEFC4AF9F8" bold="true" box="[855,924,1613,1634]" italics="true" pageId="6" pageNumber="7">hirtalis</emphasis>
</taxonomicName>
species-group
</emphasis>
</heading>
(16 species)
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD6F114B3E9EF961FE99FD8C" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BCE9B28BD6F114B3E9EF961FE99FD8C" blockId="6.[707,1193,1682,1730]" lastBlockId="7.[151,638,194,534]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B905473ABD6F114A3E9EF961FC9CF93D" bold="true" box="[731,842,1682,1703]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
Members of the
<taxonomicName id="4C71E0ABBD6F114A3841F961FB9BF93D" authorityName="Broun, 1893 b:" authorityYear="1050" box="[1028,1101,1682,1703]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hirtalis">
<emphasis id="B905473ABD6F114A3841F961FB9BF93D" box="[1028,1101,1682,1703]" italics="true" pageId="6" pageNumber="7">hirtalis</emphasis>
</taxonomicName>
speciesgroup can be distinguished from other
<taxonomicName id="4C71E0ABBD6F114A3821F95EFB7FF958" box="[1124,1193,1709,1730]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B905473ABD6F114A3821F95EFB7FF958" box="[1124,1193,1709,1730]" italics="true" pageId="6" pageNumber="7">Sagola</emphasis>
</taxonomicName>
species-groups by the following combination of characters: body length 2.04.0 mm; gular surface of male head with deep, round depression; male antennomeres longer than those of female, antennomere 1 at least 2 times longer than wide, with dull surface; prosternum with lateral procoxal foveae; male fore femur with semicircular depression (
<figureCitation id="134A87ADBD6E114B3C94FE8FFEC8FE0B" box="[209,286,380,401]" captionStart="Fig" captionStartId="9.[175,207,1144,1163]" captionTargetBox="[166,1166,194,1123]" captionTargetId="figure-247@9.[151,1171,183,1135]" captionTargetPageId="9" captionText="Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907749" httpUri="https://zenodo.org/record/4907749/files/figure.png" pageId="7" pageNumber="8">Fig. 4u</figureCitation>
: arrow); abdominal tergites IVVI with discal carinae; abdominal ventrites IVVI with basolateral foveae; parameres distinctly asymmetrical and finely punctured (
<figureCitation id="134A87ADBD6E114B3E48FE3FFDA7FE7B" box="[525,625,459,481]" captionStart="Fig" captionStartId="9.[175,207,1144,1163]" captionTargetBox="[166,1166,194,1123]" captionTargetId="figure-247@9.[151,1171,183,1135]" captionTargetPageId="9" captionText="Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907749" httpUri="https://zenodo.org/record/4907749/files/figure.png" pageId="7" pageNumber="8">Fig. 4ap</figureCitation>
); present on North Island, not known from South Island (
<figureCitation id="134A87ADBD6E114B3CA6FDF2FE95FD8C" box="[227,323,512,534]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="10.[731,763,905,924]" captionStartId-1="14.[731,763,912,931]" captionStartId-2="17.[709,741,904,923]" captionTargetBox-0="[710,1193,188,881]" captionTargetBox-1="[710,1192,195,891]" captionTargetBox-2="[688,1170,188,881]" captionTargetId-0="figure-608@10.[707,1194,183,884]" captionTargetId-1="figure-613@14.[707,1193,193,894]" captionTargetId-2="figure-602@17.[685,1171,183,884]" captionTargetPageId-0="10" captionTargetPageId-1="14" captionTargetPageId-2="17" captionText-0="Fig. 5. Known collection localities of New Zealand Sagola species. S. hirtalis: black circles; S. terricola: triangles; S. convexa: black squares; S. coromandelensis: star; S. northlandensis: white circle; S. taranakiensis: white square." captionText-1="Fig. 6. Known collection localities of New Zealand Sagola species. S. kuscheli: black circle; S. newtoni: triangles; S. thorpei: black squares; S. dugdalei: star; S. thayerae: white circle; S. hunuaensis: white square." captionText-2="Fig. 7. Known collection localities of New Zealand Sagola species. S. waikatoensis: black circle; S. gisbornensis: triangles; S. weiri: black square; S. brookesi: star." figureDoi-0="http://doi.org/10.5281/zenodo.4907753" figureDoi-1="http://doi.org/10.5281/zenodo.4907755" figureDoi-2="http://doi.org/10.5281/zenodo.4907759" httpUri-0="https://zenodo.org/record/4907753/files/figure.png" httpUri-1="https://zenodo.org/record/4907755/files/figure.png" httpUri-2="https://zenodo.org/record/4907759/files/figure.png" pageId="7" pageNumber="8">Figs. 57</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD6E11443C99FDBAFD44FDD2" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="key">
<paragraph id="8BCE9B28BD6E114B3C99FDBAFE0BFDE6" blockId="7.[220,568,585,638]" pageId="7" pageNumber="8">
<emphasis id="B905473ABD6E114B3C99FDBAFE0BFDE6" bold="true" pageId="7" pageNumber="8">
KEY TO SPECIES OF THE
<emphasis id="B905473ABD6E114B3DA6FDBCFDEEFDC5" bold="true" box="[483,568,591,607]" italics="true" pageId="7" pageNumber="8">HIRTALIS</emphasis>
SPECIES- GROUP
</emphasis>
</paragraph>
<paragraph id="8BCE9B28BD6E114B3CEAFD60FE53FD47" blockId="7.[151,638,659,733]" pageId="7" pageNumber="8">The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</paragraph>
<key id="FFE0CC19BD6E11443CD2FD0BFD44FDD2" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<keyStep id="3085838DBD6E114B3CD2FD0BFDA8FC04" pageId="7" pageNumber="8">
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<keyLead id="3080381DBD6E114B3CD2FD0BFDA8FCD8" pageId="7" pageNumber="8">1. Hind wings reduced to small pads; abdominal tergite IV without patches of microtrichia.....................................2</keyLead>
</paragraph>
<paragraph id="8BCE9B28BD6E114B3CD2FCA7FDA8FC04" blockId="7.[151,638,851,926]" pageId="7" pageNumber="8">
<keyLead id="3080381DBD6E114B3CD2FCA7FDA8FC04" pageId="7" pageNumber="8">1. Hind wings well-developed; abdominal tergite IV with pair of transverse patches of microtrichia reaching middle............3</keyLead>
</paragraph>
</keyStep>
<keyStep id="3085838DBD6E114B3CD2FC43FDABFB5A" pageId="7" pageNumber="8">
<paragraph id="8BCE9B28BD6E114B3CD2FC43FDA8FBB5" blockId="7.[151,638,944,1071]" pageId="7" pageNumber="8">
<keyLead id="3080381DBD6E114B3CD2FC43FDA8FBB5" pageId="7" pageNumber="8">
2(1). Smaller body size, length
<quantity id="4C8936CDBD6E114B3E61FC43FDAFFC5F" box="[548,633,944,965]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="8" unit="mm" value="2.4">2.4 mm</quantity>
; antennomere 1 approximately 2 times longer than wide, 45 subquadrate; mid-tibia bent (
<figureCitation id="134A87ADBD6E114B3DDAFBF3FE2EFB8F" box="[415,504,1024,1045]" captionStart="Fig" captionStartId="26.[197,229,1062,1081]" captionTargetBox="[178,1187,186,1039]" captionTargetId="figure-283@26.[173,1193,183,1044]" captionTargetPageId="26" captionText="Fig. 12. Sagola species in the bifida species-group. Aedeagi, dorsal views: a) S. bifida, b) S. latistriata, c) S. crawi, d) S. earlyi, e) S. embersoni, f) S. mayae, g) S. ranatungae, h) S. baylessae, i) S. tekoauensis, j) S. ferroi, k) S. fairburni. S. latistriata, male: l) Dorsal surface of head, m) Ventral surface of head (arrow indicates large depression), n) Mid-femur, lateral view (arrow indicates shallow depression), o) Mid-tibia, lateral view (arrow indicates bend). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907775" httpUri="https://zenodo.org/record/4907775/files/figure.png" pageId="7" pageNumber="8">Fig. 12o</figureCitation>
: arrow) ........ .....................
<emphasis id="B905473ABD6E114B3DC1FBE9FDA8FBB5" bold="true" box="[388,638,1050,1071]" pageId="7" pageNumber="8">
<taxonomicName id="4C71E0ABBD6E114B3DC1FBE9FE21FBB5" ID-CoL="4TYVR" box="[388,503,1050,1071]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="brookesi" status="sp. nov.">
<emphasis id="B905473ABD6E114B3DC1FBE9FE21FBB5" bold="true" box="[388,503,1050,1071]" italics="true" pageId="7" pageNumber="8">S. brookesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236FA41BD6E114B3E47FBE9FDA8FBB5" box="[514,638,1050,1071]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCE9B28BD6E114B3CD2FBB2FDABFB5A" blockId="7.[151,638,1088,1216]" pageId="7" pageNumber="8">
<keyLead id="3080381DBD6E114B3CD2FBB2FDABFB5A" pageId="7" pageNumber="8">
2. Larger body size, length
<quantity id="4C8936CDBD6E114B3DBAFBB2FDACFBCC" box="[511,634,1088,1110]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="7" pageNumber="8" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.52.9 mm</quantity>
; antennomere 1 approximately 2.5 times longer than wide, 45 longer than wide; mid-tibia straight ................................... ..................
<emphasis id="B905473ABD6E114B3D36FB58FDABFB5A" bold="true" box="[371,637,1195,1216]" pageId="7" pageNumber="8">
<taxonomicName id="4C71E0ABBD6E114B3D36FB58FE29FB5A" ID-CoL="78ZP3" box="[371,511,1195,1216]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hunuaensis" status="sp. nov.">
<emphasis id="B905473ABD6E114B3D36FB58FE29FB5A" bold="true" box="[371,511,1195,1216]" italics="true" pageId="7" pageNumber="8">S. hunuaensis</emphasis>
</taxonomicName>
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</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3085838DBD6E114B3CD2FB21FDA8FAE2" pageId="7" pageNumber="8">
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<keyLead id="3080381DBD6E114B3CD2FB21FDA8FA86" pageId="7" pageNumber="8">3(1). Larger body, length 3.24.0 mm; antennomere 5 elongate, at least 1.5 times longer than wide.................................... 4</keyLead>
</paragraph>
<paragraph id="8BCE9B28BD6E114B3CD2FADDFDA8FAE2" blockId="7.[151,638,1326,1400]" pageId="7" pageNumber="8">
<keyLead id="3080381DBD6E114B3CD2FADDFDA8FAE2" pageId="7" pageNumber="8">
3. Smaller body, length 2.0
<quantity id="4C8936CDBD6E114B3E67FADDFDACFAD9" box="[546,634,1326,1347]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="8" unit="mm" value="3.3">3.3 mm</quantity>
; antennomere 5 elongate, not exceeding 1.5 times longer than wide ................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3085838DBD6E114B3CD2FA79FDA8F9FF" pageId="7" pageNumber="8">
<paragraph id="8BCE9B28BD6E114B3CD2FA79FDABFA75" blockId="7.[151,638,1418,1519]" pageId="7" pageNumber="8">
<keyLead id="3080381DBD6E114B3CD2FA79FDABFA75" pageId="7" pageNumber="8">
4(3). Apical lobe of genitalia bird-head-shaped; left paramere with two minor slender lobes, originating from base (
<figureCitation id="134A87ADBD6E114B3E69FA4CFDA0FA4E" box="[556,630,1471,1492]" captionStart="Fig" captionStartId="9.[175,207,1144,1163]" captionTargetBox="[166,1166,194,1123]" captionTargetId="figure-247@9.[151,1171,183,1135]" captionTargetPageId="9" captionText="Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907749" httpUri="https://zenodo.org/record/4907749/files/figure.png" pageId="7" pageNumber="8">Fig. 4b</figureCitation>
) ..................................
<taxonomicName id="4C71E0ABBD6E114B3D8EFA29FDABFA75" ID-CoL="4TZ3X" authority="Broun" authorityName="Broun Desig. Park and Carlton" authorityYear="2013" box="[459,637,1498,1519]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="terricola">
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<emphasis id="B905473ABD6E114B3D8EFA29FDE1FA75" bold="true" box="[459,567,1498,1519]" italics="true" pageId="7" pageNumber="8">S. terricola</emphasis>
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4. Apical lobe of genitalia shaped differently; left paramere with one lobe originating anteriorly (
<figureCitation id="134A87ADBD6E114B3D9FF9C5FDC8F9D1" box="[474,542,1590,1611]" captionStart="Fig" captionStartId="9.[175,207,1144,1163]" captionTargetBox="[166,1166,194,1123]" captionTargetId="figure-247@9.[151,1171,183,1135]" captionTargetPageId="9" captionText="Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907749" httpUri="https://zenodo.org/record/4907749/files/figure.png" pageId="7" pageNumber="8">Fig. 4a</figureCitation>
).............. ...................................
<taxonomicName id="4C71E0ABBD6E114B3D90F9A3FDA8F9FF" ID-CoL="6WZP3" authority="Broun" authorityName="Broun, 1893 b:" authorityYear="1050" box="[469,638,1616,1637]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hirtalis">
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<emphasis id="B905473ABD6E114B3D90F9A3FDE3F9FF" bold="true" box="[469,565,1616,1637]" italics="true" pageId="7" pageNumber="8">S. hirtalis</emphasis>
Broun
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5(3). Smaller, body length, 2.0
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; antennomere 1 not exceeding 2.5 times longer than wide...............................6
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5. Larger, body length,
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; antennomere 1 approximately 3 times longer than wide.................................... 8
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<paragraph id="8BCE9B28BD6E114B3EE8FEEFFB42FEFC" blockId="7.[685,1172,284,706]" pageId="7" pageNumber="8">
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6(5). Phallobase of median lobe symmetrical and elongate (
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) ........................... .................................
<taxonomicName id="4C71E0ABBD6E114B3FA5FEA2FB42FEFC" ID-CoL="4TYWF" authority="Broun" authorityName="Broun" authorityYear="1886" box="[992,1172,337,358]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="convexa">
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<emphasis id="B905473ABD6E114B3FA5FEA2FB9DFEFC" bold="true" box="[992,1099,337,358]" italics="true" pageId="7" pageNumber="8">S. convexa</emphasis>
Broun
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<paragraph id="8BCE9B28BD6E114B3EE8FE86FB42FE3F" blockId="7.[685,1172,284,706]" pageId="7" pageNumber="8">
<keyLead id="3080381DBD6E114B3EE8FE86FB42FE3F" pageId="7" pageNumber="8">6. Phallobase of median lobe asymmetrical and triangular.................................7</keyLead>
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<paragraph id="8BCE9B28BD6E114B3EE8FE47FB45FDAE" blockId="7.[685,1172,284,706]" pageId="7" pageNumber="8">
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7(6). Eye small, one-third length of temple; mid-femur and tibia normal; apical lobe of genitalia curved; left paramere rectangular (
<figureCitation id="134A87ADBD6E114B3F29FDF7FC67FD83" box="[876,945,516,537]" captionStart="Fig" captionStartId="9.[175,207,1144,1163]" captionTargetBox="[166,1166,194,1123]" captionTargetId="figure-247@9.[151,1171,183,1135]" captionTargetPageId="9" captionText="Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907749" httpUri="https://zenodo.org/record/4907749/files/figure.png" pageId="7" pageNumber="8">Fig. 4k</figureCitation>
).................................... .......................
<emphasis id="B905473ABD6E114B3FE3FDECFB45FDAE" bold="true" box="[934,1171,543,564]" pageId="7" pageNumber="8">
<taxonomicName id="4C71E0ABBD6E114B3FE3FDECFBC3FDAE" ID-CoL="4TZ3Y" box="[934,1045,543,564]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="thayerae" status="sp. nov.">
<emphasis id="B905473ABD6E114B3FE3FDECFBC3FDAE" bold="true" box="[934,1045,543,564]" italics="true" pageId="7" pageNumber="8">S. thayerae</emphasis>
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7. Eye large, as long as temple; mid-femur with semicircular depression; mid-tibia bent; apical lobe of genitalia straight; left paramere L-shaped (
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) .......... ................
<emphasis id="B905473ABD6E114B3F3BFD5EFB45FD58" bold="true" box="[894,1171,685,706]" pageId="7" pageNumber="8">
<taxonomicName id="4C71E0ABBD6E114B3F3BFD5EFBC3FD58" ID-CoL="4TYXX" box="[894,1045,685,706]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gisbornensis" status="sp. nov.">
<emphasis id="B905473ABD6E114B3F3BFD5EFC5DFD58" bold="true" box="[894,907,685,706]" italics="true" pageId="7" pageNumber="8">S</emphasis>
.
<emphasis id="B905473ABD6E114B3FDEFD5EFBC3FD58" bold="true" box="[923,1045,685,706]" italics="true" pageId="7" pageNumber="8">gisbornensis</emphasis>
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</keyStep>
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<paragraph id="8BCE9B28BD6E114B3EE8FD21FB45FC98" blockId="7.[685,1172,722,985]" pageId="7" pageNumber="8">
<keyLead id="3080381DBD6E114B3EE8FD21FB45FC98" pageId="7" pageNumber="8">8(5). Apical lobe of genitalia bulbous and semicircular..........................................9</keyLead>
</paragraph>
<paragraph id="8BCE9B28BD6E114B3EE8FCE2FB42FCDB" blockId="7.[685,1172,722,985]" pageId="7" pageNumber="8">
<keyLead id="3080381DBD6E114B3EE8FCE2FB42FCDB" pageId="7" pageNumber="8">8. Apical lobe not bulbous and not semicircular ...............................................12</keyLead>
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<paragraph id="8BCE9B28BD6E114B3EE8FCA3FB42FC00" blockId="7.[685,1172,722,985]" pageId="7" pageNumber="8">
<keyLead id="3080381DBD6E114B3EE8FCA3FB42FC00" pageId="7" pageNumber="8">9(8). Process, either slender or triangular, extending from middle of apical lobe of genitalia .........................................10</keyLead>
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<keyLead id="3080381DBD6E114B3EE8FC59FB45FC43" pageId="7" pageNumber="8">9. Middle of apical lobe of genitalia lacking ...............................................11</keyLead>
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<paragraph id="8BCE9B28BD6E114B3EE8FC1AFB45FBA9" blockId="7.[685,1172,1001,1729]" pageId="7" pageNumber="8">
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10(9). Slender process extending from middle of apical lobe of genitalia (
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) ...... ............
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<taxonomicName id="4C71E0ABBD6E114B3F22FBEDFBC3FBA9" ID-CoL="4TYZT" box="[871,1045,1054,1075]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="northlandensis" status="sp. nov.">
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.
<emphasis id="B905473ABD6E114B3FC6FBEDFBC3FBA9" bold="true" box="[899,1045,1054,1075]" italics="true" pageId="7" pageNumber="8">northlandensis</emphasis>
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10. Broad, triangular process extending from middle of apical lobe of genitalia (
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)..........
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.
<emphasis id="B905473ABD6E114B3F8DFB8BFBC0FB17" bold="true" box="[968,1046,1144,1165]" italics="true" pageId="7" pageNumber="8">newtoni</emphasis>
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11(9). Slender process extending from base of apical lobe of genitalia; minor lobes of left paramere folded (
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)................ .......................
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.
<emphasis id="B905473ABD6E114B3F84FB1FFBC3FA9B" bold="true" box="[961,1045,1260,1281]" italics="true" pageId="7" pageNumber="8">dugdalei</emphasis>
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11. Apical lobe of genitalia lacking process; minor lobes of left paramere weakly bent (
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12(8). Slender and short process extending from apical lobe of genitalia; left paramere lacking minor lobe (
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13(12). Apical lobe of genitalia deeply divided into two lobes (
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Fig.
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14(13). Apical lobe of genitalia faucet-shaped; minor lobe of left paramere crescentshaped (
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15(14). Minor lobes of left paramere divided and U-shaped; right paramere rectangular (
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15. Minor lobes of left paramere divided and V-shaped; right paramere L-shaped (
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