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<document id="1ED4CA90CE5FC051E9A51CAA8989E357" ID-CLB-Dataset="3079" ID-DOI="10.1649/072.068.0mo4.1" ID-GBIF-Dataset="62322993-ca80-4b6c-ac95-861cbb236abd" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907735" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623093043626" checkinUser="carolina" docAuthor="Park, J. - S. &amp; Carlton, C. E." docDate="2014" docId="03D82A3EBD0C112A3F65FE2EFD44FA8C" docLanguage="en" docName="TheColeopBull.68.mo13.1-156.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 13) 68" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Sagola misella Sharp Desig. Park and Carlton 2013" docType="key" docVersion="23" lastPageNumber="103" masterDocId="FFE15246BD69114C3C45FFF3FFD6FF9A" masterDocTitle="A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="102" updateTime="1699001444256" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="18FD986E4F884D8F76B9B079D7E27CAD">A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)</mods:title>
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<mods:namePart id="B38F7AA2BF15CACD75E0072B190BAFE9">Park, J. - S.</mods:namePart>
<mods:affiliation id="B0E744BBDE2071B0399EEBC84B5A7843">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:namePart id="44D3C75A36C54FABC558EEFAAC8E2D4F">Carlton, C. E.</mods:namePart>
<mods:affiliation id="2A2A8E564DCA1680A8586F4C0E5D2474">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:title id="C7484DF03E130EA8708B8FECCC56E09B">The Coleopterists Bulletin</mods:title>
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<mods:date id="2D557EAA8AFA18DFCA1DCA7A10532737">2014</mods:date>
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<mods:title id="84FED0192CBE6C7ABB46E43EB6981845">mo 13</mods:title>
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<treatment id="03D82A3EBD0C112A3F65FE2EFD44FA8C" ID-DOI="http://doi.org/10.5281/zenodo.5765207" ID-GBIF-Taxon="191245083" ID-Zenodo-Dep="5765207" LSID="urn:lsid:plazi:treatment:03D82A3EBD0C112A3F65FE2EFD44FA8C" httpUri="http://treatment.plazi.org/id/03D82A3EBD0C112A3F65FE2EFD44FA8C" lastPageId="102" lastPageNumber="103" pageId="101" pageNumber="102">
<subSubSection id="C36BC8A3BD0C11293F65FE2EFBF7FE68" box="[800,1057,477,498]" pageId="101" pageNumber="102" type="nomenclature">
<paragraph id="8BCE9B28BD0C11293F65FE2EFBF7FE68" blockId="101.[800,1057,477,525]" box="[800,1057,477,498]" pageId="101" pageNumber="102">
<heading id="D0862C44BD0C11293F65FE2EFBF7FE68" box="[800,1057,477,498]" centered="true" fontSize="10" level="2" pageId="101" pageNumber="102" reason="2">
<emphasis id="B905473ABD0C11293F65FE2EFBF7FE68" bold="true" box="[800,1057,477,498]" pageId="101" pageNumber="102">
25.
<taxonomicName id="4C71E0ABBD0C11293F0DFE2EFC5BFE68" ID-CoL="4TYZK" authorityName="Sharp Desig. Park and Carlton" authorityYear="2013" box="[840,909,477,498]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="misella">
<emphasis id="B905473ABD0C11293F0DFE2EFC5BFE68" bold="true" box="[840,909,477,498]" italics="true" pageId="101" pageNumber="102">misella</emphasis>
</taxonomicName>
species-group
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C36BC8A3BD0C11293F23FE0BFC0CFD97" box="[870,986,504,525]" pageId="101" pageNumber="102" type="description">
<paragraph id="8BCE9B28BD0C11293F23FE0BFC0CFD97" blockId="101.[800,1057,477,525]" box="[870,986,504,525]" pageId="101" pageNumber="102">(13 species)</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD0C11293E80FDD1FB47FCDB" pageId="101" pageNumber="102" type="diagnosis">
<paragraph id="8BCE9B28BD0C11293E80FDD1FB47FCDB" blockId="101.[685,1172,546,833]" pageId="101" pageNumber="102">
<emphasis id="B905473ABD0C11293E80FDD1FCF8FDAD" bold="true" box="[709,814,546,567]" pageId="101" pageNumber="102">Diagnosis.</emphasis>
Members of the
<taxonomicName id="4C71E0ABBD0C11293FA8FDD1FBE0FDAD" authorityName="Sharp Desig. Park and Carlton" authorityYear="2013" box="[1005,1078,546,567]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="misella">
<emphasis id="B905473ABD0C11293FA8FDD1FBE0FDAD" box="[1005,1078,546,567]" italics="true" pageId="101" pageNumber="102">misella</emphasis>
</taxonomicName>
speciesgroup can be distinguished from other
<taxonomicName id="4C71E0ABBD0C1129380AFDCEFB42FDC8" box="[1103,1172,573,594]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B905473ABD0C1129380AFDCEFB42FDC8" box="[1103,1172,573,594]" italics="true" pageId="101" pageNumber="102">Sagola</emphasis>
</taxonomicName>
species-groups by the following combination of characters: body length
<quantity id="4C8936CDBD0C11293FE8FD81FBFBFD1D" box="[941,1069,625,647]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.3" metricValueMin="2.1" pageId="101" pageNumber="102" unit="mm" value="2.7" valueMax="3.3" valueMin="2.1">2.13.3 mm</quantity>
; antennomere 1 approximately 1.52.0 times longer than wide, male antennomeres enlarged; frontal rostrum rectangular, lobes continuous, reaching posterior frontal fovea (
<figureCitation id="134A87ADBD0C11293F7AFD2FFC4EFD6B" box="[831,920,732,753]" captionStart="Fig" captionStartId="104.[197,230,972,991]" captionTargetBox="[183,1189,190,937]" captionTargetId="figure-477@104.[173,1193,183,948]" captionTargetPageId="104" captionText="Fig. 59. Sagola species in the misella species-group. Aedeagi, dorsal views: a) S. misella, b) S. bituberata, c) S. prisca, d) S. murchisonensis, e) S. fiordlandensis, f) S. hunterensis, g) S. humpridgensis, h) S. clunieae, i) S. turretensis, j) S. otagoensis, k) S. otagoensis, apical lobe, lateral view, l) S. southlandensis, m) S. huvali, n) S. tennysonensis. S. misella: o) Dorsal surface of head, p) Prosternum, ventral view, q) Meso- and metaventrites, ventral view, r) Abdomen, ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907896" httpUri="https://zenodo.org/record/4907896/files/figure.png" pageId="101" pageNumber="102">Fig. 59o</figureCitation>
); frontal sulcus reaching midpoint of eye, posterior frontal fovea present (
<figureCitation id="134A87ADBD0C11293EF0FCE2FCD9FCBC" box="[693,783,785,806]" captionStart="Fig" captionStartId="104.[197,230,972,991]" captionTargetBox="[183,1189,190,937]" captionTargetId="figure-477@104.[173,1193,183,948]" captionTargetPageId="104" captionText="Fig. 59. Sagola species in the misella species-group. Aedeagi, dorsal views: a) S. misella, b) S. bituberata, c) S. prisca, d) S. murchisonensis, e) S. fiordlandensis, f) S. hunterensis, g) S. humpridgensis, h) S. clunieae, i) S. turretensis, j) S. otagoensis, k) S. otagoensis, apical lobe, lateral view, l) S. southlandensis, m) S. huvali, n) S. tennysonensis. S. misella: o) Dorsal surface of head, p) Prosternum, ventral view, q) Meso- and metaventrites, ventral view, r) Abdomen, ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907896" httpUri="https://zenodo.org/record/4907896/files/figure.png" pageId="101" pageNumber="102">Fig. 59o</figureCitation>
); female hind wings reduced to small pads; abdominal tergites IVVI with discal carinae.
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD0C112A3EB0FC80FD44FA8C" lastPageId="102" lastPageNumber="103" pageId="101" pageNumber="102" type="key">
<paragraph id="8BCE9B28BD0C11293EB0FC80FC25FC3C" blockId="101.[757,1100,883,935]" pageId="101" pageNumber="102">
<emphasis id="B905473ABD0C11293EB0FC80FC25FC3C" bold="true" pageId="101" pageNumber="102">
KEY TO SPECIES OF THE
<emphasis id="B905473ABD0C11293FBEFC8BFB9AFC12" bold="true" box="[1019,1100,888,904]" italics="true" pageId="101" pageNumber="102">MISELLA</emphasis>
SPECIES- GROUP
</emphasis>
</paragraph>
<paragraph id="8BCE9B28BD0C11293E80FC4EFC4DFB9D" blockId="101.[685,1172,957,1031]" pageId="101" pageNumber="102">The key is based on male specimens because most female specimens are indistinguishable based on external morphology.</paragraph>
<key id="FFE0CC19BD0C112A3EE8FBEFFD44FA8C" lastPageId="102" lastPageNumber="103" pageId="101" pageNumber="102">
<keyStep id="3085838DBD0C11293EE8FBEFFB42FB13" pageId="101" pageNumber="102">
<paragraph id="8BCE9B28BD0C11293EE8FBEFFB42FBD6" blockId="101.[685,1172,1052,1729]" pageId="101" pageNumber="102">
<keyLead id="3080381DBD0C11293EE8FBEFFB42FBD6" pageId="101" pageNumber="102">1. Eye small, approximately one-third length of temple .................................. 2</keyLead>
</paragraph>
<paragraph id="8BCE9B28BD0C11293EE8FBAAFB42FB13" blockId="101.[685,1172,1052,1729]" pageId="101" pageNumber="102">
<keyLead id="3080381DBD0C11293EE8FBAAFB42FB13" pageId="101" pageNumber="102">1. Eye large, approximately one-half length of temple................................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="3085838DBD0C11293EE8FB65FB42FA84" pageId="101" pageNumber="102">
<paragraph id="8BCE9B28BD0C11293EE8FB65FB42FB7A" blockId="101.[685,1172,1052,1729]" pageId="101" pageNumber="102">
<keyLead id="3080381DBD0C11293EE8FB65FB42FB7A" pageId="101" pageNumber="102">
2(1). Apical lobe of genitalia arrowheadshaped (
<figureCitation id="134A87ADBD0C11293F22FB42FC6FFB5C" box="[871,953,1201,1222]" captionStart="Fig" captionStartId="104.[197,230,972,991]" captionTargetBox="[183,1189,190,937]" captionTargetId="figure-477@104.[173,1193,183,948]" captionTargetPageId="104" captionText="Fig. 59. Sagola species in the misella species-group. Aedeagi, dorsal views: a) S. misella, b) S. bituberata, c) S. prisca, d) S. murchisonensis, e) S. fiordlandensis, f) S. hunterensis, g) S. humpridgensis, h) S. clunieae, i) S. turretensis, j) S. otagoensis, k) S. otagoensis, apical lobe, lateral view, l) S. southlandensis, m) S. huvali, n) S. tennysonensis. S. misella: o) Dorsal surface of head, p) Prosternum, ventral view, q) Meso- and metaventrites, ventral view, r) Abdomen, ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907896" httpUri="https://zenodo.org/record/4907896/files/figure.png" pageId="101" pageNumber="102">Fig. 59b</figureCitation>
) .................................. ..............................
<taxonomicName id="4C71E0ABBD0C11293F89FB38FB42FB7A" ID-CoL="6WZNZ" authority="Broun" authorityName="Broun" authorityYear="1914" box="[972,1172,1227,1248]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="bituberata">
<emphasis id="B905473ABD0C11293F89FB38FB42FB7A" bold="true" box="[972,1172,1227,1248]" pageId="101" pageNumber="102">
<emphasis id="B905473ABD0C11293F89FB38FB9DFB7A" bold="true" box="[972,1099,1227,1248]" italics="true" pageId="101" pageNumber="102">S. bituberata</emphasis>
Broun
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCE9B28BD0C11293EE8FB1DFB42FA84" blockId="101.[685,1172,1052,1729]" pageId="101" pageNumber="102">
<keyLead id="3080381DBD0C11293EE8FB1DFB42FA84" pageId="101" pageNumber="102">2. Apical lobe of genitalia not arrowheadshaped ..................................................3</keyLead>
</paragraph>
</keyStep>
<keyStep id="3085838DBD0C11293EE8FAD8FB42FA02" pageId="101" pageNumber="102">
<paragraph id="8BCE9B28BD0C11293EE8FAD8FB42FAEF" blockId="101.[685,1172,1052,1729]" pageId="101" pageNumber="102">
<keyLead id="3080381DBD0C11293EE8FAD8FB42FAEF" pageId="101" pageNumber="102">
3(2). Major process of median lobe of genitalia S-shaped (
<figureCitation id="134A87ADBD0C11293FE9FAB5FC28FAC1" box="[940,1022,1350,1371]" captionStart="Fig" captionStartId="104.[197,230,972,991]" captionTargetBox="[183,1189,190,937]" captionTargetId="figure-477@104.[173,1193,183,948]" captionTargetPageId="104" captionText="Fig. 59. Sagola species in the misella species-group. Aedeagi, dorsal views: a) S. misella, b) S. bituberata, c) S. prisca, d) S. murchisonensis, e) S. fiordlandensis, f) S. hunterensis, g) S. humpridgensis, h) S. clunieae, i) S. turretensis, j) S. otagoensis, k) S. otagoensis, apical lobe, lateral view, l) S. southlandensis, m) S. huvali, n) S. tennysonensis. S. misella: o) Dorsal surface of head, p) Prosternum, ventral view, q) Meso- and metaventrites, ventral view, r) Abdomen, ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907896" httpUri="https://zenodo.org/record/4907896/files/figure.png" pageId="101" pageNumber="102">Fig. 59a</figureCitation>
) ....................... ...................................
<taxonomicName id="4C71E0ABBD0C11293FA8FA93FB42FAEF" ID-CoL="4TYZK" authority="Sharp" authorityName="Sharp Desig. Park and Carlton" authorityYear="2013" box="[1005,1172,1376,1397]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="misella">
<emphasis id="B905473ABD0C11293FA8FA93FB42FAEF" bold="true" box="[1005,1172,1376,1397]" pageId="101" pageNumber="102">
<emphasis id="B905473ABD0C11293FA8FA93FB9BFAEF" bold="true" box="[1005,1101,1376,1397]" italics="true" pageId="101" pageNumber="102">S. misella</emphasis>
Sharp
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCE9B28BD0C11293EE8FA70FB42FA02" blockId="101.[685,1172,1052,1729]" box="[685,1172,1410,1432]" pageId="101" pageNumber="102">
<keyLead id="3080381DBD0C11293EE8FA70FB42FA02" box="[685,1172,1410,1432]" pageId="101" pageNumber="102">3. Median lobe of genitalia not curved.... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="3085838DBD0C11293EE8FA56FB45F9B7" pageId="101" pageNumber="102">
<paragraph id="8BCE9B28BD0C11293EE8FA56FB42FA6A" blockId="101.[685,1172,1052,1729]" pageId="101" pageNumber="102">
<keyLead id="3080381DBD0C11293EE8FA56FB42FA6A" pageId="101" pageNumber="102">
4(3). Median lobe of genitalia triangular with membranous tube (
<figureCitation id="134A87ADBD0C11293F95FA33FBF7FA4F" box="[976,1057,1472,1493]" captionStart="Fig" captionStartId="104.[197,230,972,991]" captionTargetBox="[183,1189,190,937]" captionTargetId="figure-477@104.[173,1193,183,948]" captionTargetPageId="104" captionText="Fig. 59. Sagola species in the misella species-group. Aedeagi, dorsal views: a) S. misella, b) S. bituberata, c) S. prisca, d) S. murchisonensis, e) S. fiordlandensis, f) S. hunterensis, g) S. humpridgensis, h) S. clunieae, i) S. turretensis, j) S. otagoensis, k) S. otagoensis, apical lobe, lateral view, l) S. southlandensis, m) S. huvali, n) S. tennysonensis. S. misella: o) Dorsal surface of head, p) Prosternum, ventral view, q) Meso- and metaventrites, ventral view, r) Abdomen, ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907896" httpUri="https://zenodo.org/record/4907896/files/figure.png" pageId="101" pageNumber="102">Fig. 59c</figureCitation>
) ................. .....................................
<taxonomicName id="4C71E0ABBD0C11293FB2FA29FB42FA6A" ID-CoL="4TZ2F" authority="Sharp" authorityName="Sharp Desig. Park and Carlton" authorityYear="2013" box="[1015,1172,1498,1520]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="prisca">
<emphasis id="B905473ABD0C11293FB2FA29FB42FA6A" bold="true" box="[1015,1172,1498,1520]" pageId="101" pageNumber="102">
<emphasis id="B905473ABD0C11293FB2FA29FB9BFA75" bold="true" box="[1015,1101,1498,1519]" italics="true" pageId="101" pageNumber="102">S. prisca</emphasis>
Sharp
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCE9B28BD0C11293EE8FA0EFB45F9B7" blockId="101.[685,1172,1052,1729]" pageId="101" pageNumber="102">
<keyLead id="3080381DBD0C11293EE8FA0EFB45F9B7" pageId="101" pageNumber="102">4. Medina lobe of genitalia not triangular.............................................................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="3085838DBD0C11293EE8F9C9FB42F95B" pageId="101" pageNumber="102">
<paragraph id="8BCE9B28BD0C11293EE8F9C9FB42F9F0" blockId="101.[685,1172,1052,1729]" pageId="101" pageNumber="102">
<keyLead id="3080381DBD0C11293EE8F9C9FB42F9F0" pageId="101" pageNumber="102">5(4). Median lobe of genitalia depressed, left apex and as long as paramere............. 6</keyLead>
</paragraph>
<paragraph id="8BCE9B28BD0C11293EE8F984FB42F95B" blockId="101.[685,1172,1052,1729]" pageId="101" pageNumber="102">
<keyLead id="3080381DBD0C11293EE8F984FB42F95B" pageId="101" pageNumber="102">5. Median lobe of genitalia divided, minor lobe longer than major lobe and paramere .................................................................. 7</keyLead>
</paragraph>
</keyStep>
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6(5). Paramere depressed apically (
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); only known from
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(
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: triangle) ............................................. .......
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6. Paramere blunt; known from Fiordland (
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: star)......................................... .......................
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7(5). Abdominal ventrite VII with small and acute process; major lobe of median lobe of genitalia slender and simple (
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Fig.
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) ..............
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<emphasis id="B905473ABD0F112A3DCCFE28FDC1FE6A" bold="true" box="[393,535,475,496]" italics="true" pageId="102" pageNumber="103">humpridgensis</emphasis>
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7. Abdominal ventrite VII without process; major lobe of median lobe of genitalia diamond-shaped and broad (
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)............................................... .................
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8(1). Elytra subquadrate, hind wings reduced to small pads .....
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9(8). Antennomeres 48 longer than wide .... ..............
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10(9). Median lobe of genitalia twisted, with membranous tube (
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11(10). Apical lobe of genitalia rectangular vertically (
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12(11). Median lobe of genitalia C-shaped vertically (
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12. Median lobe of genitalia S-shaped vertically (
<figureCitation id="134A87ADBD0F112A3D17FB15FE4BFB61" box="[338,413,1254,1275]" captionStart="Fig" captionStartId="104.[197,230,972,991]" captionTargetBox="[183,1189,190,937]" captionTargetId="figure-477@104.[173,1193,183,948]" captionTargetPageId="104" captionText="Fig. 59. Sagola species in the misella species-group. Aedeagi, dorsal views: a) S. misella, b) S. bituberata, c) S. prisca, d) S. murchisonensis, e) S. fiordlandensis, f) S. hunterensis, g) S. humpridgensis, h) S. clunieae, i) S. turretensis, j) S. otagoensis, k) S. otagoensis, apical lobe, lateral view, l) S. southlandensis, m) S. huvali, n) S. tennysonensis. S. misella: o) Dorsal surface of head, p) Prosternum, ventral view, q) Meso- and metaventrites, ventral view, r) Abdomen, ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907896" httpUri="https://zenodo.org/record/4907896/files/figure.png" pageId="102" pageNumber="103">Fig. 59l</figureCitation>
)....................................... ............
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<taxonomicName id="4C71E0ABBD0F112A3D23FAF2FDC2FA8C" ID-CoL="4TZ3C" box="[358,532,1281,1302]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="southlandensis" status="sp. nov.">
<emphasis id="B905473ABD0F112A3D23FAF2FEA5FA8C" bold="true" box="[358,371,1281,1302]" italics="true" pageId="102" pageNumber="103">S</emphasis>
.
<emphasis id="B905473ABD0F112A3DC7FAF2FDC2FA8C" bold="true" box="[386,532,1281,1302]" italics="true" pageId="102" pageNumber="103">southlandensis</emphasis>
</taxonomicName>
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