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<mods:title id="6453783B532CDAE3BF29AA7914D7E647">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="EC99C89E5F7BE9CA49028F6B75B280C0">Bistroem, Olof</mods:namePart>
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<mods:namePart id="330CE209BA2988B83DAB1B92469ACFA6">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="CB7E66B4EB8B1AA69942DE089B8D1AF0">Bergsten, Johannes</mods:namePart>
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<treatment id="E7870D57C9F4380FEB78F010528E1B7D" ID-GBIF-Taxon="127862704" LSID="urn:lsid:plazi:treatment:E7870D57C9F4380FEB78F010528E1B7D" httpUri="http://treatment.plazi.org/id/E7870D57C9F4380FEB78F010528E1B7D" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<subSubSection id="7EB607D32566B2EDB379AB45BF7FDC53" pageId="26" pageNumber="27" type="multiple">
<paragraph id="5981D9C0A3B26C3532053D093E91AD73" pageId="26" pageNumber="27">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
</subSubSection>
<subSubSection id="0A82C9D0991A354EF8AF24BB3E1F8420" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="3F05210E9780CE6016948534D18254F0" pageId="26" pageNumber="27">
<taxonomicName id="062D9468DCC9D674490647D2F95E82B6" ID-CoL="6NRMF" authority="Omer-Cooper, 1970" authorityName="Omer-Cooper" authorityYear="1970" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mateui" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="mateui">Laccophilus mateui Omer-Cooper, 1970</taxonomicName>
Figs 29-30, 227, 389, 529
</paragraph>
</subSubSection>
<subSubSection id="1D9C3E0EBC62A98FC13406485926374D" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="02B0D35FE286A66C5BCEAC40C1F6D3B5" pageId="26" pageNumber="27">
<taxonomicName id="62D640178404D4B56342639410C1D779" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mateui" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="mateui">Laccophilus mateui</taxonomicName>
<bibRefCitation id="2E9A8936B9DB914CC2A1AFF706E306D9" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="285 - 294" title="New and little known species of African Laccophilus (Coleoptera: Dytiscidae)." volume="33" year="1970">Omer-Cooper 1970</bibRefCitation>
: 285, 287 (original description, faunistics);
<bibRefCitation id="1FCDD053A483E47AD05793FBDA7FFD9D" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 246 (catalogue, faunistics);
<bibRefCitation id="930AD4ECF2E23B6150D127F924866F14" author="Nilsson, AN" editor="Loebl, I" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" pagination="35 - 78" title="Dytiscidae." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol. 1. Archostemata-Myxophaga-Adephaga" year="2003">Nilsson 2003</bibRefCitation>
: 76 (catalogue, faunistics);
<bibRefCitation id="47FECCA8B0CD1109AA83633CCB5B4F37" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 214 (catalogue, faunistics).
</paragraph>
</subSubSection>
<subSubSection id="99A058EB2403123E7EBF3194142D0AC6" pageId="26" pageNumber="27" type="type locality">
<paragraph id="3E68B57982516C69A7A290C1876FB678" pageId="26" pageNumber="27">Type locality.</paragraph>
<paragraph id="CF2021DBC38CCE67BB090B01179DBAAA" pageId="26" pageNumber="27">Algeria: Sahara, Hoggar, Aguelm, Ymeleulauen.</paragraph>
</subSubSection>
<subSubSection id="E1489A672C6B804A7FA243E0C759007F" pageId="27" pageNumber="28" type="type material">
<paragraph id="C20F36086F213715010B8C334BD6E570" pageId="27" pageNumber="28">
<pageBreakToken id="1EDDBDB77EDD8967300BB24E689E7A57" pageId="27" pageNumber="28" start="start">Type</pageBreakToken>
material, studied
</paragraph>
<paragraph id="287552A773162A1F3BD8836501A6D2E8" pageId="27" pageNumber="28">
(1 ex.). Holotype: male: &quot;Type male / H. B. Leech Collection / det. J. Omer-Cooper
<taxonomicName id="FDA6382117A84DED1179B32B20BC6AA1" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mateui" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mateui">Laccophilus mateui</taxonomicName>
sp.n. / Aguelm, Ymeleulauen, Hoggar, Sahara J. Mateu coll. / 18-V- 1951&quot; (AMGS; according to original description, holotype to be deposited in CAS).
</paragraph>
</subSubSection>
<subSubSection id="12A5175EBE0DF22EDF8DE2401EED28CC" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="6658D703399DC2A17515FCFEA912CDEE" pageId="27" pageNumber="28">Additional material, studied</paragraph>
<paragraph id="A1896A0FAF5D912F8DF7AC8EEE7AD0F3" pageId="27" pageNumber="28">(5 exs.). Algeria: &quot;55 km N Tamanrasset 16-17 March 1971 J.A. Gruwell&quot; (4 exs. USNM, 1 ex. MZH; habitus in Fig. 389).</paragraph>
</subSubSection>
<subSubSection id="C679952E0CA90E20F7277C9D8E248B16" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="488F0FF7CF5B89562524463B9400FC98" pageId="27" pageNumber="28">Diagnosis.</paragraph>
<paragraph id="87B1606D1088E37F37FDBD15ACFB914C" pageId="27" pageNumber="28">
<taxonomicName id="0826D734011B77433A5FFFECB36067A1" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mateui" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mateui">Laccophilus mateui</taxonomicName>
is a close relative to
<taxonomicName id="A8599F373D15EF4C3F78F05187682775" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus minutus" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="minutus">Laccophilus minutus</taxonomicName>
and
<taxonomicName id="92EE23A5FBD6BAF86AC5637E7487EAC4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus sordidus" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sordidus">Laccophilus sordidus</taxonomicName>
. All three species have similar general appearance and same ground plan regarding penis-shape. Absence of stridulatory apparatus separates it from
<taxonomicName id="18EA2FAB2B5DE5B63380B1D87A7EA07F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus hyalinus" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="hyalinus">Laccophilus hyalinus</taxonomicName>
and
<taxonomicName id="19EA71E2684F891D429714B257A3483F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus demoflysi" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="demoflysi">Laccophilus demoflysi</taxonomicName>
. Shape of penis distinguishes it from
<taxonomicName id="1C06208DDB8EAF415A6A3899963DD57F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus minutus" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="minutus">Laccophilus minutus</taxonomicName>
(penis apex is broader in
<taxonomicName id="A1265A6E388BA6CE36860ABA9D0F6B30" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mateui" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mateui">Laccophilus mateui</taxonomicName>
) and
<taxonomicName id="48CDEB2F5B41E007D24548532212E69F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus sordidus" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sordidus">Laccophilus sordidus</taxonomicName>
(penis is stouter in
<taxonomicName id="8D595A5D44F1A98CC572973E9E70F0A9" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mateui" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mateui">Laccophilus mateui</taxonomicName>
and longer in
<taxonomicName id="2BB36BF6768763FAA5D22E92A2A60C4D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus sordidus" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sordidus">Laccophilus sordidus</taxonomicName>
).
<taxonomicName id="84777991E7B8035F9CB0565229E1598A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mateui" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mateui">Laccophilus mateui</taxonomicName>
(&gt;5 mm) is also larger sized than
<taxonomicName id="858D4B1219D7F5819F5145354B8F0396" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus minutus" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="minutus">Laccophilus minutus</taxonomicName>
, a species which
<normalizedToken id="2936EBB3DB77B3498FBD8DEEE32D5C45" originalValue="dont">don't</normalizedToken>
exceed 5 mm in length.
</paragraph>
</subSubSection>
<subSubSection id="EB3A55F90556C3F5F960237EEA219164" pageId="27" pageNumber="28" type="description">
<paragraph id="A1C2CA25054E2255EC840A29C4F30149" pageId="27" pageNumber="28">Description.</paragraph>
<paragraph id="825C23A5061B2C3010D3C689C6335803" pageId="27" pageNumber="28">Body length 5.1-5.3 mm, width 2.8-3.0 mm. Body dorsally pale ferrugineous to ferrugineous, elytral colour pattern vague to fairly distinct (Fig. 389).</paragraph>
<paragraph id="E7A08A40E1000914185139EA3E018F85" pageId="27" pageNumber="28">Head: Pale ferrugineous, at pronotum darker, ferrugineous to dark ferrugineous (delimitation of colours often vague). Almost impunctate, at eyes with a few, scattered, somewhat indistinct punctures. Submat, finely microsculptured; reticulation double. In central part of head fine reticulation indistinct, in part obliterated; in lateral parts of head fine reticulation clearly discernible; large meshes contain 3-6 fine meshes.</paragraph>
<paragraph id="8ED351199160BA8C2E77BC757DB7006F" pageId="27" pageNumber="28">Pronotum: Ferrugineous, laterally pale ferrugineous (gradual change; no distinct delimitation of colours). Frontally, sometimes with a quite distinct dark ferrugineous area. Impunctate, at margin with a few, indistinct, coarser punctures discernible. Submat, rather densely microsculptured; reticulation double, fine meshes in part almost absent or indistinct.</paragraph>
<paragraph id="FC8250BB7550EC5EC67D71F032C1DEA3" pageId="27" pageNumber="28">Elytra: Ferrugineous, with vague pale ferrugineous markings (Fig. 389). Elytral colour pattern sometimes quite distinct. Submat, rather finely microsculptured; reticulation double, large meshes contain between 2-6 fine meshes. Each elytron with a discal, dorsolateral and lateral row of punctures, which are sparse and somewhat irregular, in part indistinct. Lateral, pre-apical furrow long, shallow, finely punctate and pubescent.</paragraph>
<paragraph id="7FB413D001E98EBAC6B493B45BC14A54" pageId="27" pageNumber="28">Ventral aspect: Ferrugineous, metacoxal plate in part dark ferrugineous but no distinct colour pattern formed (delimitation vague). Almost impunctate. Submat, finely microsculptured. No stridulatory apparatus. Metacoxal plates in anterior half with some vague, transversely located, slightly irregular furrows. Abdomen in basal half with rather distinct curved striae. Apex of prosternal process broken in holotype; it is keeled, short and apex pointed. Apical ventrite as in Fig. 29.</paragraph>
<paragraph id="A0A9EB9488819F5F7DEF64D5150BFA81" pageId="27" pageNumber="28">Legs: Pro- and mesotarsus somewhat enlarged, provided with suckers.</paragraph>
<paragraph id="083540FBE9AAC6B6100AC587B975C5EC" pageId="27" pageNumber="28">Male genitalia: Penis comparatively robust; in lateral aspect apical half evenly curved (Fig. 227).</paragraph>
<paragraph id="2CADB787530FADDE467D9AA914778F7F" pageId="27" pageNumber="28">Female: Pro- and mesotarsus slender. Apical ventrite as in Fig. 30.</paragraph>
</subSubSection>
<subSubSection id="5163F6D3652032B1F2077E8FA2A15B29" pageId="27" pageNumber="28" type="distribution">
<paragraph id="A5BD2FD65D90DEB24A6B6FFD1D6C902C" pageId="27" pageNumber="28">Distribution.</paragraph>
<paragraph id="90DCB2D723AD45FEF9427453CF52ADDF" pageId="27" pageNumber="28">Algeria (Hoggar, Sahara) (Fig. 529).</paragraph>
</subSubSection>
<subSubSection id="76D72B1553883323BCE261F06D45F7EB" pageId="27" pageNumber="28" type="collecting circumstances">
<paragraph id="848D5522A4F1C4F7D05547CA7273DEEF" pageId="27" pageNumber="28">Collecting circumstances.</paragraph>
<paragraph id="4F0344F54B47E35A856A27353B642AA4" pageId="27" pageNumber="28">Unknown.</paragraph>
</subSubSection>
</treatment>
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