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<document id="C97C7DAE328D7AF37E5442AEB640C7A9" ID-CLB-Dataset="26733" ID-DOI="10.11646/zootaxa.4609.1.10" ID-GBIF-Dataset="4935cee9-8c79-4df4-b007-98cbf46b964e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3150305" ID-ZooBank="9F7084A1-00B7-46EE-BAD7-B0A0EB6F481D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1558533746644" checkinUser="plazi" docAuthor="Souma, Jun" docDate="2019" docId="C951D3010333FFA5FF45FADDFDC11CDC" docLanguage="en" docName="zootaxa.4609.1.10.pdf" docOrigin="Zootaxa 4609 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acalypta pallidicoronata Souma 2019, sp. nov." docType="treatment" docUuid="8B7F5F56-2EE0-4D38-BB62-12AD0C936BDE" docUuidSource="ZooBank" docVersion="8" lastPageNumber="183" masterDocId="3568AB790331FFA0FFD2FFD6FF95181B" masterDocTitle="A new species and a new record of the genus Acalypta (Hemiptera: Heteroptera: Tingidae) from Japan" masterLastPageNumber="184" masterPageNumber="178" pageNumber="180" updateTime="1698731483076" updateUser="ExternalLinkService">
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<mods:title id="B6FDE931B37315CD6532F81F22FCA21A">A new species and a new record of the genus Acalypta (Hemiptera: Heteroptera: Tingidae) from Japan</mods:title>
</mods:titleInfo>
<mods:name id="7F3955FE9C911BC61533FB38FB9865EF" type="personal">
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<mods:namePart id="4BF4EFE5C661DBC671884640A45D4A55">Souma, Jun</mods:namePart>
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<mods:title id="A1FCEDC7852F35DA7734ACDC697A2404">Zootaxa</mods:title>
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<mods:part id="7A690122D153A1DFBDEF57309CE068DD">
<mods:date id="D36134EE45D2A1195E1B74A746C7D694">2019</mods:date>
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<mods:number id="26002196CCC6B1FA4FBD656DFEF82CCE">2019-05-22</mods:number>
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<mods:classification id="CF594542F0F7AD3BE9A5BBE07E540D59">journal article</mods:classification>
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<mods:identifier id="ABBF35FE2348278482F87244D03B5A0E" type="DOI">10.11646/zootaxa.4609.1.10</mods:identifier>
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<treatment id="C951D3010333FFA5FF45FADDFDC11CDC" ID-GBIF-Taxon="157239483" LSID="urn:lsid:zoobank.org:act:8B7F5F56-2EE0-4D38-BB62-12AD0C936BDE" httpUri="http://treatment.plazi.org/id/C951D3010333FFA5FF45FADDFDC11CDC" lastPageId="5" lastPageNumber="183" pageId="2" pageNumber="180">
<subSubSection id="09E2319C0333FFA2FF45FADDFDB81D3E" box="[151,557,1291,1318]" pageId="2" pageNumber="180" type="nomenclature">
<paragraph id="414762170333FFA2FF45FADDFDB81D3E" blockId="2.[151,557,1291,1318]" box="[151,557,1291,1318]" pageId="2" pageNumber="180">
<heading id="1A0FD57B0333FFA2FF45FADDFDB81D3E" bold="true" box="[151,557,1291,1318]" fontSize="11" level="1" pageId="2" pageNumber="180" reason="1">
<emphasis id="738CBE050333FFA2FF45FADDFDB81D3E" bold="true" box="[151,557,1291,1318]" pageId="2" pageNumber="180">
<taxonomicName id="86F819940333FFA2FF45FADDFE551D3D" authority="Souma, 2019" authorityName="Souma" authorityYear="2019" box="[151,448,1291,1318]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="180" phylum="Arthropoda" rank="species" species="pallidicoronata" status="sp. nov.">
<emphasis id="738CBE050333FFA2FF45FADDFE551D3D" bold="true" box="[151,448,1291,1318]" italics="true" pageId="2" pageNumber="180">Acalypta pallidicoronata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="68BF037E0333FFA2FE1DFADDFDB81D3E" box="[463,557,1291,1317]" pageId="2" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="09E2319C0333FFA2FF45FA85FDAE1D76" box="[151,571,1363,1389]" pageId="2" pageNumber="180" type="description">
<paragraph id="414762170333FFA2FF45FA85FDAE1D76" blockId="2.[151,571,1363,1389]" box="[151,571,1363,1389]" pageId="2" pageNumber="180">
(
<figureCitation id="D9C37E920333FFA2FF4DFA85FF7D1D76" box="[159,232,1363,1389]" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="2" pageNumber="180">Figs. 1</figureCitation>
CD, 1GH, 2B, 2D, 3B, 3D)
</paragraph>
</subSubSection>
<subSubSection id="09E2319C0333FFA2FF45FA4DFE311E39" pageId="2" pageNumber="180" type="materials_examined">
<paragraph id="414762170333FFA2FF45FA4DFE311E39" blockId="2.[151,1437,1434,2039]" pageId="2" pageNumber="180">
<emphasis id="738CBE050333FFA2FF45FA4DFEBE1DAE" bold="true" box="[151,299,1434,1461]" pageId="2" pageNumber="180">Type series.</emphasis>
<materialsCitation id="F190684A0333FFA2FEE8FA4DFD491DC2" ID-GBIF-Occurrence="2243135987" collectingDate="2017-09-06" collectorName="Tsushima Is. Kamiagata-machi" country="Japan" latitude="34.5902" location="Mt. Konoki-yama" longLatPrecision="1" longitude="129.38092" pageId="2" pageNumber="180" specimenCount="1" specimenCount-male="1" stateProvince="Ishikawa" typeStatus="holotype">
<typeStatus id="9E43DCB50333FFA2FEE8FA4DFE321DAE" box="[314,423,1435,1461]" pageId="2" pageNumber="180" type="holotype">Holotype</typeStatus>
: brachypterous
<specimenCount id="57FEA99E0333FFA2FDBAFA4AFDEA1DAD" box="[616,639,1436,1462]" pageId="2" pageNumber="180" type="male"></specimenCount>
, “
<collectingCountry id="39EF22870333FFA2FD4EFA4DFD611DAE" box="[668,756,1435,1461]" name="Japan" pageId="2" pageNumber="180">JAPAN</collectingCountry>
<collectorName id="EC0D07C10333FFA2FCD1FA4DFBE41DAE" box="[771,1137,1435,1462]" pageId="2" pageNumber="180">Tsushima Is. Kamiagata-machi</collectorName>
, near
<location id="442734CC0333FFA2FB16FA4AFA0D1DAE" LSID="urn:lsid:plazi:treatment:C951D3010333FFA5FF45FADDFDC11CDC:442734CC0333FFA2FB16FA4AFA0D1DAE" box="[1220,1432,1435,1462]" country="Japan" latitude="34.5902" longLatPrecision="1" longitude="129.38092" name="Mt. Konoki-yama" pageId="2" pageNumber="180" stateProvince="Ishikawa">Mt. Kônoki-yama</location>
,
<geoCoordinate id="24CC04D00333FFA2FF45FA68FE801DC2" box="[151,277,1470,1497]" direction="north" orientation="latitude" pageId="2" pageNumber="180" precision="1" value="34.5902">34.59020N</geoCoordinate>
<geoCoordinate id="24CC04D00333FFA2FECFFA69FE3D1DC2" box="[285,424,1471,1497]" direction="east" orientation="longitude" pageId="2" pageNumber="180" precision="1" value="129.38092">129.38092E</geoCoordinate>
,
<collectingDate id="2502BD3F0333FFA2FE67FA69FDAE1DC2" box="[437,571,1471,1498]" pageId="2" pageNumber="180" value="2017-09-06">
<date id="354644D70333FFA2FE67FA69FDAE1DC2" box="[437,571,1471,1498]" pageId="2" pageNumber="180" value="2017-09-06">6. IX. 2017</date>
</collectingDate>
, T.
<collectingRegion id="833CACF50333FFA2FDB4FA69FD441DC2" box="[614,721,1471,1497]" country="Japan" name="Ishikawa" pageId="2" pageNumber="180">Ishikawa</collectingRegion>
</materialsCitation>
.
<typeStatus id="9E43DCB50333FFA2FD3AFA16FCC01DC1" box="[744,853,1472,1498]" pageId="2" pageNumber="180" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="57FEA99E0333FFA2FCB4FA68FC201DC1" box="[870,949,1470,1498]" pageId="2" pageNumber="180" type="male">15 ♂♂</specimenCount>
<specimenCount id="57FEA99E0333FFA2FC6CFA69FB9B1DC1" box="[958,1038,1471,1498]" pageId="2" pageNumber="180" type="female">13 ♀♀</specimenCount>
, all brachypterous): as
<typeStatus id="9E43DCB50333FFA2FAC8FA69FAE91DC2" box="[1306,1404,1471,1497]" pageId="2" pageNumber="180" type="holotype">holotype</typeStatus>
(
<specimenCount id="57FEA99E0333FFA2FA5FFA69FF511DE5" pageId="2" pageNumber="180" type="male">4 ♂♂</specimenCount>
); as
<typeStatus id="9E43DCB50333FFA2FF2BFA35FECE1DE6" box="[249,347,1507,1533]" pageId="2" pageNumber="180" type="holotype">holotype</typeStatus>
but
<date id="354644D70333FFA2FE5FFA35FE621DE6" box="[397,503,1507,1533]" pageId="2" pageNumber="180" value="2017-09-08">8.ix.2017</date>
(
<specimenCount id="57FEA99E0333FFA2FDD8FA35FDA71DE5" box="[522,562,1507,1534]" pageId="2" pageNumber="180" type="female">1 ♀</specimenCount>
); as
<typeStatus id="9E43DCB50333FFA2FDB8FA35FD591DE6" box="[618,716,1507,1533]" pageId="2" pageNumber="180" type="holotype">holotype</typeStatus>
but leg. J. Souma (
<specimenCount id="57FEA99E0333FFA2FC7AFA35FC721DE5" box="[936,999,1507,1534]" pageId="2" pageNumber="180" type="male">3 ♂♂</specimenCount>
<specimenCount id="57FEA99E0333FFA2FC3DFA35FBB81DE5" box="[1007,1069,1507,1534]" pageId="2" pageNumber="180" type="female">3 ♀♀</specimenCount>
); as
<typeStatus id="9E43DCB50333FFA2FBB4FA35FB5D1DE6" box="[1126,1224,1507,1533]" pageId="2" pageNumber="180" type="holotype">holotype</typeStatus>
but
<date id="354644D70333FFA2FB29FA35FAF21DE6" box="[1275,1383,1507,1533]" pageId="2" pageNumber="180" value="2017-09-08">8.ix.2017</date>
, leg. J. Souma (
<specimenCount id="57FEA99E0333FFA2FEDCF9D1FED81E39" box="[270,333,1543,1570]" pageId="2" pageNumber="180" type="male">8 ♂♂</specimenCount>
<specimenCount id="57FEA99E0333FFA2FE86F9D1FE061E39" box="[340,403,1543,1570]" pageId="2" pageNumber="180" type="female">9 ♀♀</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="09E2319C0333FFA2FF15F9FCFDC21FEC" pageId="2" pageNumber="180" type="diagnosis">
<paragraph id="414762170333FFA2FF15F9FCFDC21FEC" blockId="2.[151,1437,1434,2039]" pageId="2" pageNumber="180">
<emphasis id="738CBE050333FFA2FF15F9FCFED71E5E" bold="true" box="[199,322,1578,1605]" pageId="2" pageNumber="180">Diagnosis.</emphasis>
Recognized among other species of
<taxonomicName id="86F819940333FFA2FCC2F9FCFC171E5C" authorityName="Westwood" authorityYear="1840" box="[784,898,1578,1607]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="738CBE050333FFA2FCC2F9FCFC171E5C" box="[784,898,1578,1607]" italics="true" pageId="2" pageNumber="180">Acalypta</emphasis>
</taxonomicName>
by a combination of the following characters: body pale fuscous, elliptical in male and ovate in female, approximately
<quantity id="8600CFF20333FFA2FBBEF998FB211E71" box="[1132,1204,1614,1643]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="180" unit="mm" value="2.0">2 mm</quantity>
long from head to apices of hemelytra (
<figureCitation id="D9C37E920333FFA2FE71F9A4FE631E94" box="[419,502,1650,1679]" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="2" pageNumber="180">Figs. 1</figureCitation>
CD, 1GH); antenniferous tubercles obtuse and slightly curved inward; pronotum unicarinate, 0.8 times as long as maximum width across paranota; calli with a faint median transverse furrow; paranotum with 2 rows of areolae throughout its length; anterolateral angle of paranotum rounded, not protruding anteriad, posterolateral angle not protruding posteriad; posterior process of pronotum short, 4 times as wide as its length; posterior margin of hemelytron straight, but slightly curved outward at apex; costal area with 2 rows of areolae in basal part and a single row in remaining parts; subcostal area with 7 rows of areolae at widest part; discoidal area considerably expanded beyond apical fourth of hemelytron, distinctly wider than subcostal area; discoidal-sutural boundary vein gradually obscured in its basal third; pygophore roundly inflated and semicircular, bulged evenly in venter, smooth on surface (
<figureCitation id="D9C37E920333FFA2FF0FF860FEDE1FC8" box="[221,331,1974,2003]" captionStart="FIGURE 2" captionStartId="3.[151,250,971,996]" captionTargetBox="[151,1436,170,960]" captionTargetId="figure@3.[151,1436,170,960]" captionTargetPageId="3" captionText="FIGURE 2. AD. Apical part of abdomen, ventral view. A. cooleyi, male (A) and female (C); A. pallidicoronata sp. nov., male (B) and female (D). Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/3150313/files/figure.png" pageId="2" pageNumber="180">Figs. 2B</figureCitation>
, 3B); apical margin of female abdomen rounded (
<figureCitation id="D9C37E920333FFA2FC6DF860FBB01FC8" box="[959,1061,1974,2003]" captionStart="FIGURE 2" captionStartId="3.[151,250,971,996]" captionTargetBox="[151,1436,170,960]" captionTargetId="figure@3.[151,1436,170,960]" captionTargetPageId="3" captionText="FIGURE 2. AD. Apical part of abdomen, ventral view. A. cooleyi, male (A) and female (C); A. pallidicoronata sp. nov., male (B) and female (D). Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/3150313/files/figure.png" pageId="2" pageNumber="180">Fig. 2D</figureCitation>
); and paramere flat and short, angularly curved inward (Fig. 3D).
</paragraph>
</subSubSection>
<caption id="1587329F0332FFA3FF45FC1DFDBA1C1F" httpUri="https://zenodo.org/record/3150313/files/figure.png" pageId="3" pageNumber="181" startId="3.[151,250,971,996]" targetBox="[151,1436,170,960]" targetPageId="3">
<paragraph id="414762170332FFA3FF45FC1DFDBA1C1F" blockId="3.[151,1436,971,1028]" pageId="3" pageNumber="181">
<emphasis id="738CBE050332FFA3FF45FC1DFEC51BFF" bold="true" box="[151,336,971,996]" pageId="3" pageNumber="181">FIGURE 2. AD.</emphasis>
Apical part of abdomen, ventral view.
<taxonomicName id="86F819940332FFA3FD08FC1BFCAB1BFF" authorityName=": Drake &amp; Lattin" authorityYear="1963" box="[730,830,972,996]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="181" phylum="Arthropoda" rank="species" species="cooleyi">
<emphasis id="738CBE050332FFA3FD08FC1BFCAB1BFF" box="[730,830,972,996]" italics="true" pageId="3" pageNumber="181">A. cooleyi</emphasis>
</taxonomicName>
, male (A) and female (C);
<taxonomicName id="86F819940332FFA3FB9CFC1BFA9D1BFF" authority="Souma, 2019" authorityName="Souma" authorityYear="2019" box="[1102,1288,972,996]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="181" phylum="Arthropoda" rank="species" species="pallidicoronata" status="sp. nov.">
<emphasis id="738CBE050332FFA3FB9CFC1BFA9D1BFF" box="[1102,1288,972,996]" italics="true" pageId="3" pageNumber="181">A. pallidicoronata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="68BF037E0332FFA3FADFFC1AFACA1BFF" box="[1293,1375,972,996]" pageId="3" pageNumber="181" rank="species">
<emphasis id="738CBE050332FFA3FADFFC1AFACA1BFF" bold="true" box="[1293,1375,972,996]" pageId="3" pageNumber="181">sp. nov.</emphasis>
</taxonomicNameLabel>
, male (B) and female (D). Scale bars: 0.2 mm.
</paragraph>
</caption>
<subSubSection id="09E2319C0332FFA3FF15FB92FA081FF8" pageId="3" pageNumber="181" type="description">
<paragraph id="414762170332FFA3FF15FB92FBF31C98" blockId="3.[151,1437,1092,2020]" pageId="3" pageNumber="181">
<emphasis id="738CBE050332FFA3FF15FB92FECC1C44" bold="true" box="[199,345,1092,1119]" pageId="3" pageNumber="181">Description.</emphasis>
Brachypterous male. General color pale fuscous; eyes red; antennae except segment III, calli, abdomen beneath and femora dark brown; body with golden pubescence (
<figureCitation id="D9C37E920332FFA3FC01FBBFFBCC1C98" box="[979,1113,1129,1155]" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="3" pageNumber="181">Figs. 1C, G</figureCitation>
).
</paragraph>
<paragraph id="414762170332FFA3FF15FB5BFC921DDC" blockId="3.[151,1437,1092,2020]" pageId="3" pageNumber="181">
Body ellipsoid, 1.8 times as long as maximum width across hemelytra (
<figureCitation id="D9C37E920332FFA3FC3DFB5BFBD21CBC" box="[1007,1095,1165,1191]" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="3" pageNumber="181">Fig. 1C</figureCitation>
). Head with several punctures on vertex; with a pair of very short frontal spines, their apices far remote from tip of tylus; antenniferous tubercles short and obtuse, slightly curved inward; juga deeply and coarsely punctate. Compound eyes prominent laterally, ellipsoidal in dorsal view. Antennae smooth, covered with pubescence; segments I thickest, not close to each other at their base, approximately 1.5 times as long as its width; segment II as long as its width; segment III tapering apicad, narrower than segment IV, slightly longer than head width across eyes; segment IV fusiform, widest a little beyond middle; relative lengths from segments I to IV as 1.6: 1.0: 6.0: 2.9. Bucculae widely open in front, approximately 1/3 as high as its length, with 3 rows of areolae at highest part. Rostrum almost same in length as antennae, slightly extending beyond posterior end of metasternal laminae.
</paragraph>
<paragraph id="414762170332FFA3FF15FA04FDEF1EFC" blockId="3.[151,1437,1092,2020]" pageId="3" pageNumber="181">
Pronotum (
<figureCitation id="D9C37E920332FFA3FE9AFA07FE0A1DF0" box="[328,415,1489,1515]" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="3" pageNumber="181">Fig. 1C</figureCitation>
) unicarinate, 0.8 times as long as maximum width across paranota. Hood low and roofshaped (
<figureCitation id="D9C37E920332FFA3FF27FA23FEDB1E14" box="[245,334,1525,1551]" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="3" pageNumber="181">Fig. 1G</figureCitation>
); anterior margin roundly protruding forward, only concealing basal part of vertex. Median carina low but slightly concave at level of calli in lateral view, with a single row of minute areolae arranged throughout its length, but one of these areolae at level of calli conspicuously large. Calli weakly depressed, with faint median transverse furrow. Paranotum transversely expanding laterad, approximately 3.1 times as long as its width, with 2 rows of areolae throughout its length which are of subequal size (
<figureCitation id="D9C37E920332FFA3FCBEF953FC571E84" box="[876,962,1669,1695]" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="3" pageNumber="181">Fig. 1C</figureCitation>
); anterolateral angle rounded, not protruding anteriad; lateral margin gently curved outward; posterolateral angle rounded, not protruding posteriad. Posterior process short, 4 times as wide as its length.
</paragraph>
<paragraph id="414762170332FFA3FF15F927FA081FF8" blockId="3.[151,1437,1092,2020]" pageId="3" pageNumber="181">
Hemelytron narrow, 2.5 times as long as its width, strongly elevated in discoidal area, slightly extended beyond apex of abdomen (
<figureCitation id="D9C37E920332FFA3FEBBF8C3FE641F34" box="[361,497,1813,1839]" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="3" pageNumber="181">Figs. 1C, G</figureCitation>
); posterior margin straight, but slightly curved outward at apex; costal area narrow, slightly reflexed upward, with 2 rows of areolae in basal part and in a single row in remaining parts; subcostal area 3.3 times as wide as costal area at middle of hemelytron, with 6 rows of areolae at widest part; discoidal area very long and wide, considerably expanded beyond apical fourth of hemelytron, distinctly wider than subcostal area, with 6 rows of areolae at widest part; sutural area indistinctly delimited from discoidal area in basal part, with a single row of areolae in middle part and 2 rows in apical part; boundary veins strongly carinate, but discoidal-sutural
</paragraph>
</subSubSection>
<subSubSection id="09E2319C0335FFA5FF15F8A7FC621C2C" lastPageId="5" lastPageNumber="183" pageId="4" pageNumber="182" type="discussion">
<paragraph id="414762170335FFA4FF15F8A7FB0F1FEC" blockId="4.[151,1437,1905,2040]" pageId="4" pageNumber="182">
<emphasis id="738CBE050335FFA4FF15F8A7FEAE1F90" bold="true" box="[199,315,1905,1931]" pageId="4" pageNumber="182">Remarks.</emphasis>
In keys to the Japanese species of
<taxonomicName id="86F819940335FFA4FD02F8A7FCA11F90" authorityName="Westwood" authorityYear="1840" box="[720,820,1905,1931]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="738CBE050335FFA4FD02F8A7FCA11F90" box="[720,820,1905,1931]" italics="true" pageId="4" pageNumber="182">Acalypta</emphasis>
</taxonomicName>
(
<bibRefCitation id="25691FE60335FFA4FC91F8A7FC4F1F90" author="Takeya, C." box="[835,986,1905,1931]" pageId="4" pageNumber="182" pagination="41 - 75" refId="ref4316" refString="Takeya, C. (1962) Taxonomic revision of the Tingidae of Japan, Korea, the Ryukyus and Formosa Part 1 (Hemiptera). Mushi, 36 (5), 41 - 75." type="journal article" year="1962">Takeya 1962</bibRefCitation>
,
<bibRefCitation id="25691FE60335FFA4FC34F8A7FB091F90" author="Tomokuni, M." box="[998,1180,1905,1931]" pageId="4" pageNumber="182" pagination="87 - 91" refId="ref4354" refString="Tomokuni, M. (1972) Japanese species of the genus Acalypta (Hemiptera: Tingidae). Transactions of the Shikoku Entomological Society, 11, 87 - 91." type="journal article" year="1972">Tomokuni 1972</bibRefCitation>
,
<bibRefCitation id="25691FE60335FFA4FB7AF8A7FF441FB4" author="Yamada, K. &amp; Tomokuni, M." pageId="4" pageNumber="182" pagination="2 - 13" refId="ref4492" refString="Yamada, K. &amp; Tomokuni, M. (2012) Family Tingidae Laporte, 1832. In: Ishikawa, T., Takai, M. &amp; Yasunaga, T. (Eds.), A field guide to Japanese bugs. Terrestrial heteropterans III. Zenkoku Noson Kyoiku Kyokai, Tokyo, pp. 180 - 213, pls. 2 - 13. [in Japanese]" type="book chapter" year="2012">Yamada &amp; Tomokuni 2012</bibRefCitation>
) this new species does not run to any species due to having a combination of the following characters: body ellipsoidal or ovate; pronotum 0.8 times as long as maximum width across paranota; anterolateral angle of paranotum not protruding forward; and discoidal area distinctly wider than subcostal area (
<figureCitation id="D9C37E920335FFA4FBC0F80BFBC51FEC" box="[1042,1104,2013,2039]" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="4" pageNumber="182">Fig. 1</figureCitation>
CD).
</paragraph>
<paragraph id="414762170334FFA5FF15FF4FFBBC1A00" blockId="5.[151,1437,153,1223]" pageId="5" pageNumber="183">
<taxonomicName id="86F819940334FFA5FF15FF4FFE4B18A8" authority="Souma, 2019" authorityName="Souma" authorityYear="2019" box="[199,478,153,179]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="pallidicoronata" status="sp. nov.">
<emphasis id="738CBE050334FFA5FF15FF4FFE4B18A8" box="[199,478,153,179]" italics="true" pageId="5" pageNumber="183">Acalypta pallidicoronata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="68BF037E0334FFA5FE36FF4FFDA818A8" box="[484,573,153,179]" pageId="5" pageNumber="183" rank="species">
<emphasis id="738CBE050334FFA5FE36FF4FFDA818A8" bold="true" box="[484,573,153,179]" pageId="5" pageNumber="183">sp. nov.</emphasis>
</taxonomicNameLabel>
closely matches the original description of
<taxonomicName id="86F819940334FFA5FBF2FF4FFABF18A8" authority="(Drake 1942)" authorityName="Drake" authorityYear="1942" baseAuthorityName="Drake" baseAuthorityYear="1942" box="[1056,1322,153,179]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="738CBE050334FFA5FBF2FF4FFBA418A8" box="[1056,1073,153,179]" italics="true" pageId="5" pageNumber="183">A</emphasis>
.
<emphasis id="738CBE050334FFA5FBECFF4CFB1918A8" box="[1086,1164,154,179]" italics="true" pageId="5" pageNumber="183">sauteri</emphasis>
(Drake 1942)
</taxonomicName>
. Although
<bibRefCitation id="25691FE60334FFA5FF45FF6BFEC818CC" author="Tomokuni, M." box="[151,349,189,215]" pageId="5" pageNumber="183" pagination="87 - 91" refId="ref4354" refString="Tomokuni, M. (1972) Japanese species of the genus Acalypta (Hemiptera: Tingidae). Transactions of the Shikoku Entomological Society, 11, 87 - 91." type="journal article" year="1972">Tomokuni (1972)</bibRefCitation>
pointed out that
<taxonomicName id="86F819940334FFA5FDF0FF6BFD0418CC" baseAuthorityName="Drake" baseAuthorityYear="1942" box="[546,657,189,215]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="738CBE050334FFA5FDF0FF6BFDA618CC" box="[546,563,189,215]" italics="true" pageId="5" pageNumber="183">A</emphasis>
.
<emphasis id="738CBE050334FFA5FD91FF68FD0418CC" box="[579,657,190,215]" italics="true" pageId="5" pageNumber="183">sauteri</emphasis>
</taxonomicName>
exhibits considerable intraspecific variation, based on a direct comparison of the
<typeStatus id="9E43DCB50334FFA5FEE5FF37FEF218E0" box="[311,359,225,251]" pageId="5" pageNumber="183">type</typeStatus>
material of the new species with hundreds of specimens of
<taxonomicName id="86F819940334FFA5FBD0FF37FBFA18E0" baseAuthorityName="Drake" baseAuthorityYear="1942" box="[1026,1135,225,251]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="738CBE050334FFA5FBD0FF37FBFA18E0" box="[1026,1135,225,251]" italics="true" pageId="5" pageNumber="183">A. sauteri</emphasis>
</taxonomicName>
from different localities of
<collectingCountry id="39EF22870334FFA5FF45FED3FF4F1904" box="[151,218,261,287]" name="Japan" pageId="5" pageNumber="183">Japan</collectingCountry>
, the following characters are recognized to readily differentiate
<taxonomicName id="86F819940334FFA5FC62FED3FBE81904" authority="Souma, 2019" authorityName="Souma" authorityYear="2019" box="[944,1149,261,287]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="pallidicoronata" status="sp. nov.">
<emphasis id="738CBE050334FFA5FC62FED3FBE81904" box="[944,1149,261,287]" italics="true" pageId="5" pageNumber="183">A. pallidicoronata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="68BF037E0334FFA5FB57FED3FB751904" box="[1157,1248,261,287]" pageId="5" pageNumber="183" rank="species">
<emphasis id="738CBE050334FFA5FB57FED3FB751904" bold="true" box="[1157,1248,261,287]" pageId="5" pageNumber="183">sp. nov.</emphasis>
</taxonomicNameLabel>
from
<taxonomicName id="86F819940334FFA5FAF4FED3FA001904" baseAuthorityName="Drake" baseAuthorityYear="1942" box="[1318,1429,261,287]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="738CBE050334FFA5FAF4FED3FA001904" box="[1318,1429,261,287]" italics="true" pageId="5" pageNumber="183">A. sauteri</emphasis>
</taxonomicName>
: general color pale fuscous; pronotum 0.8 times as long as maximum width across paranota; anterolateral angle of paranotum not protruding anteriad; posterolateral angle of paranotum not protruding posteriad; and discoidal area considerably expanded beyond apical fourth of hemelytron, distinctly wider than subcostal area (
<figureCitation id="D9C37E920334FFA5FB0DFEA7FABC1990" box="[1247,1321,369,395]" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="5" pageNumber="183">Figs. 1</figureCitation>
CD). On the other hand,
<taxonomicName id="86F819940334FFA5FE9FFE43FE2919B4" baseAuthorityName="Drake" baseAuthorityYear="1942" box="[333,444,405,431]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="738CBE050334FFA5FE9FFE43FECB19B4" box="[333,350,405,431]" italics="true" pageId="5" pageNumber="183">A</emphasis>
.
<emphasis id="738CBE050334FFA5FEBCFE40FE2919B4" box="[366,444,406,431]" italics="true" pageId="5" pageNumber="183">sauteri</emphasis>
</taxonomicName>
has the following features: general color dark fuscous; pronotum 0.6 times as long as maximum width across paranota; anterolateral angle of paranotum weakly protruding anteriad, not reaching midlevel of compound eye; posterolateral angle of paranotum protruding posteriad; and discoidal area not expanded beyond apical fourth of hemelytron, as wide as subcostal area at each widest part.
</paragraph>
<paragraph id="414762170334FFA5FF15FDF3FDAD1AD4" blockId="5.[151,1437,153,1223]" pageId="5" pageNumber="183">
The new species resembles
<taxonomicName id="86F819940334FFA5FDD5FDF3FD371A24" authorityName="Takeya" authorityYear="1962" box="[519,674,549,575]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="hirashimai">
<emphasis id="738CBE050334FFA5FDD5FDF3FD8D1A24" box="[519,536,549,575]" italics="true" pageId="5" pageNumber="183">A</emphasis>
.
<emphasis id="738CBE050334FFA5FDFBFDF3FD371A24" box="[553,674,549,575]" italics="true" pageId="5" pageNumber="183">hirashimai</emphasis>
</taxonomicName>
and
<taxonomicName id="86F819940334FFA5FD0FFDF3FB8C1A24" authority="Downes, 1927" authorityName="Downes" authorityYear="1927" box="[733,1049,549,575]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="saundersi">
<emphasis id="738CBE050334FFA5FD0FFDF3FD7B1A24" box="[733,750,549,575]" italics="true" pageId="5" pageNumber="183">A</emphasis>
.
<emphasis id="738CBE050334FFA5FD2DFDF3FCF91A24" box="[767,876,549,575]" italics="true" pageId="5" pageNumber="183">saundersi</emphasis>
Downes,
<quantity id="8600CFF20334FFA5FC33FDF3FB8C1A24" box="[993,1049,549,575]" metricMagnitude="1" metricUnit="m" metricValue="4.8945799999999995" pageId="5" pageNumber="183" unit="in" value="1927.0">1927</quantity>
</taxonomicName>
in the width of the costal area of hemelytron relative to the subcostal area, but is distinguished from them by the following characters: pronotum 0.8 times as long as maximum width across paranota; posterolateral angle of paranotum not protruding backward; posterior process short, 4 times as wide as its length; and costal area with 2 rows of areolae in basal part and a single row in remaining parts (
<figureCitation id="D9C37E920334FFA5FE77FD63FE7B1AD4" box="[421,494,693,719]" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="5" pageNumber="183">Figs. 1</figureCitation>
CD).
</paragraph>
<paragraph id="414762170334FFA5FF15FD0FFF7E1BBC" blockId="5.[151,1437,153,1223]" pageId="5" pageNumber="183">
In general appearance, this new species slightly resembles
<taxonomicName id="86F819940334FFA5FCB5FD0FFBCE1AE8" authority="Drake, 1930" authorityName="Drake" authorityYear="1930" box="[871,1115,729,755]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="duryi">
<emphasis id="738CBE050334FFA5FCB5FD0FFCED1AE8" box="[871,888,729,755]" italics="true" pageId="5" pageNumber="183">A</emphasis>
.
<emphasis id="738CBE050334FFA5FC5AFD0FFC561AE8" box="[904,963,729,755]" italics="true" pageId="5" pageNumber="183">duryi</emphasis>
Drake, 1930
</taxonomicName>
,
<taxonomicName id="86F819940334FFA5FBBAFD0FFB6A1AE8" authorityName="Takeya" authorityYear="1962" box="[1128,1279,729,755]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="miyamotoi">
<emphasis id="738CBE050334FFA5FBBAFD0FFBEC1AE8" box="[1128,1145,729,755]" italics="true" pageId="5" pageNumber="183">A</emphasis>
.
<emphasis id="738CBE050334FFA5FB5BFD0CFB6A1AE8" box="[1161,1279,730,755]" italics="true" pageId="5" pageNumber="183">miyamotoi</emphasis>
</taxonomicName>
and
<taxonomicName id="86F819940334FFA5FAEBFD0FFF771B0C" authorityName="Tomokuni" authorityYear="1972" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="tsurugisana">
<emphasis id="738CBE050334FFA5FAEBFD0FFADF1AE8" box="[1337,1354,729,755]" italics="true" pageId="5" pageNumber="183">A</emphasis>
.
<emphasis id="738CBE050334FFA5FA8BFD0CFF771B0C" italics="true" pageId="5" pageNumber="183">tsurugisana</emphasis>
</taxonomicName>
, but it is easily distinguished from them by the following characters: pronotum 0.8 times as long as maximum width across paranota; paranotum with 2 rows of areolae throughout its length; anterolateral angle of paranotum not protruding anteriad; discoidal area considerably expanded beyond apical fourth of hemelytron, distinctly wider than subcostal area; and costal area with 2 rows of areolae in basal part and a single row in remaining parts (
<figureCitation id="D9C37E920334FFA5FAB6FCBFFF341BBC" captionStart="FIGURE 1" captionStartId="2.[151,250,1184,1209]" captionTargetBox="[156,1427,176,1166]" captionTargetId="figure@2.[151,1436,174,1172]" captionTargetPageId="2" captionText="FIGURE 1. AH. Habitus of Acalypta species from Japan, dorsal and lateral view. A. cooleyi, brachypterous male (A, E) and macropterous female (B, F); A. pallidicoronata sp. nov., brachypterous male (C, G) and female (D, H). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3150307/files/figure.png" pageId="5" pageNumber="183">Figs. 1</figureCitation>
CD).
</paragraph>
<paragraph id="414762170334FFA5FF15FC67FC621C2C" blockId="5.[151,1437,153,1223]" pageId="5" pageNumber="183">
Members of
<taxonomicName id="86F819940334FFA5FE86FC67FE2D1BD0" authorityName="Westwood" authorityYear="1840" box="[340,440,945,971]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="738CBE050334FFA5FE86FC67FE2D1BD0" box="[340,440,945,971]" italics="true" pageId="5" pageNumber="183">Acalypta</emphasis>
</taxonomicName>
are divided into two species groups: those of which the pronotum is unicarinate and those of which it is tricarinate. All members of the former group are only known from brachypterous morphs (
<bibRefCitation id="25691FE60334FFA5FAFAFC03FF461C08" author="Tomokuni, M." pageId="5" pageNumber="183" pagination="145 - 154" refId="ref4386" refString="Tomokuni, M. (1983) Notes on the Japanese species of Acalypta (Hemiptera, Tingidae). Memoirs of the National Science Museum, 16, 145 - 154. [in Japanese, English summary]" type="journal article" year="1983">Tomokuni 1983</bibRefCitation>
). Having an unicarinate pronotum,
<taxonomicName id="86F819940334FFA5FDB8FC2FFCAD1C08" authority="Souma, 2019" authorityName="Souma" authorityYear="2019" box="[618,824,1017,1043]" class="Insecta" family="Tingidae" genus="Acalypta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="183" phylum="Arthropoda" rank="species" species="pallidicoronata" status="sp. nov.">
<emphasis id="738CBE050334FFA5FDB8FC2FFDEE1C08" box="[618,635,1017,1043]" italics="true" pageId="5" pageNumber="183">A</emphasis>
.
<emphasis id="738CBE050334FFA5FD59FC2FFCAD1C08" box="[651,824,1017,1043]" italics="true" pageId="5" pageNumber="183">pallidicoronata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="68BF037E0334FFA5FC93FC2FFC091C08" box="[833,924,1017,1043]" pageId="5" pageNumber="183" rank="species">
<emphasis id="738CBE050334FFA5FC93FC2FFC091C08" bold="true" box="[833,924,1017,1043]" pageId="5" pageNumber="183">sp. nov.</emphasis>
</taxonomicNameLabel>
belongs to the first group; similarly to other members of the group, no macropterous morph is know in this species either.
</paragraph>
</subSubSection>
<subSubSection id="09E2319C0334FFA5FF15FB96FDEA1C47" box="[199,639,1088,1116]" pageId="5" pageNumber="183" type="distribution">
<paragraph id="414762170334FFA5FF15FB96FDEA1C47" blockId="5.[151,1437,153,1223]" box="[199,639,1088,1116]" pageId="5" pageNumber="183">
<emphasis id="738CBE050334FFA5FF15FB96FEF41C40" bold="true" box="[199,353,1088,1115]" pageId="5" pageNumber="183">Distribution.</emphasis>
<collectingCountry id="39EF22870334FFA5FEB5FB97FE321C40" box="[359,423,1089,1115]" name="Japan" pageId="5" pageNumber="183">Japan</collectingCountry>
(Tsushima Island).
</paragraph>
</subSubSection>
<subSubSection id="09E2319C0334FFA5FF15FBB3FDC11CDC" pageId="5" pageNumber="183" type="etymology">
<paragraph id="414762170334FFA5FF15FBB3FC011CB8" blockId="5.[151,1437,153,1223]" pageId="5" pageNumber="183">
<emphasis id="738CBE050334FFA5FF15FBB3FEDB1C64" bold="true" box="[199,334,1125,1151]" pageId="5" pageNumber="183">Etymology.</emphasis>
The species epithet is the Latin adjective
<emphasis id="738CBE050334FFA5FCCAFBB3FCE71C64" box="[792,882,1125,1151]" italics="true" pageId="5" pageNumber="183">pallidus</emphasis>
(meaning pale) and the Latin noun
<emphasis id="738CBE050334FFA5FB2BFBB0FAD21C64" box="[1273,1351,1126,1151]" italics="true" pageId="5" pageNumber="183">corona</emphasis>
(meaning crown), referring to features of the pale and rounded paranotum.
</paragraph>
<paragraph id="414762170334FFA5FF15FB7BFDC11CDC" blockId="5.[151,1437,153,1223]" box="[199,596,1197,1223]" pageId="5" pageNumber="183">
<emphasis id="738CBE050334FFA5FF15FB7BFEDB1CDC" bold="true" box="[199,334,1197,1223]" pageId="5" pageNumber="183">Host-plant.</emphasis>
Undetermined mosses.
</paragraph>
</subSubSection>
</treatment>
</document>