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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="C485D1A5CDFA4F04DEB39FE96F721D68" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus remotus Caterino &amp; Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="118" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="117" updateTime="1668155382655" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>271</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>401</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152040911" LSID="urn:lsid:zoobank.org:act:0AF82D92-CAB5-4263-BA19-43AD7E6E3DED" httpUri="http://treatment.plazi.org/id/C485D1A5CDFA4F04DEB39FE96F721D68" lastPageId="117" lastPageNumber="118" pageId="116" pageNumber="117">
<subSubSection pageId="116" pageNumber="117" type="nomenclature">
<paragraph pageId="116" pageNumber="117">
<taxonomicName LSID="urn:lsid:zoobank.org:act:0AF82D92-CAB5-4263-BA19-43AD7E6E3DED" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus remotus" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="remotus">Operclipygus remotus</taxonomicName>
<taxonomicNameLabel pageId="116" pageNumber="117">sp. n.</taxonomicNameLabel>
Fig. 33AMap 11
</paragraph>
</subSubSection>
<subSubSection pageId="116" pageNumber="117" type="type locality">
<paragraph pageId="116" pageNumber="117">Type locality.</paragraph>
<paragraph pageId="116" pageNumber="117">
PANAMA:
<normalizedToken originalValue="Darién">Darien</normalizedToken>
: Cana, Pirre Camp [
<geoCoordinate direction="north" orientation="latitude" precision="1" value="7.76375">7°45.825'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-77.722084">77°43.325'W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection pageId="116" pageNumber="117" type="type material">
<paragraph pageId="116" pageNumber="117">Type material.</paragraph>
<paragraph pageId="116" pageNumber="117">
Holotype male: &quot;PANAMA:
<normalizedToken originalValue="Darién">Darien</normalizedToken>
Prov. Cana, Pirre Camp, 1320,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="7.76375">7°45.825'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-77.722084">77°43.325'W</geoCoordinate>
, 6-MAY-2008, A. R. Gillogly&quot;/ &quot;Caterino/Tishechkin
<taxonomicName lsidName="" pageId="116" pageNumber="117" rank="tribe" tribe="Exosternini">Exosternini</taxonomicName>
Voucher EXO-00281&quot;(FMNH).
</paragraph>
</subSubSection>
<subSubSection lastPageId="117" lastPageNumber="118" pageId="116" pageNumber="117" type="description">
<paragraph pageId="116" pageNumber="117">Diagnostic description.</paragraph>
<paragraph lastPageId="117" lastPageNumber="118" pageId="116" pageNumber="117">
This species is extremely similar to
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus lunulus" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="lunulus">Operclipygus lunulus</taxonomicName>
, differing mainly in the following characters: length: 1.65 mm, width: 1.34 mm; anterior submarginal pronotal stria fine, recurved obliquely to about middle of pronotum; lateral region of pronotal disk with few coarse punctures; mesometaventral stria more broadly arched forward on mesoventral disk; propygidial punctures small, round, sparse, separated by at least 2
<normalizedToken originalValue="×">x</normalizedToken>
their diameters throughout; pygidium with coarse punctures restricted to
<pageBreakToken pageId="117" pageNumber="118" start="start">narrow</pageBreakToken>
basal marginal band. Male genitalia very similar to those of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus lunulus" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="lunulus">Operclipygus lunulus</taxonomicName>
(see Figs 32
<normalizedToken originalValue="I–K">I-K</normalizedToken>
), the aedeagus indistinguishable; S9 only weakly widened toward base (Fig. 33A).
</paragraph>
<caption pageId="117" pageNumber="118">
<paragraph pageId="117" pageNumber="118">
Figure 33. Male genitalia of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus dubius" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="dubius">Operclipygus dubius</taxonomicName>
group. A S9 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus remotus" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="remotus">Operclipygus remotus</taxonomicName>
B S9 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus occultus" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="occultus">Operclipygus occultus</taxonomicName>
C S9 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus perplexus" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="perplexus">Operclipygus perplexus</taxonomicName>
D Aedeagus, dorsal and lateral views, of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus occultus" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="occultus">Operclipygus occultus</taxonomicName>
E S8 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus occultus" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="occultus">Operclipygus occultus</taxonomicName>
F T9 &amp; T10 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus occultus" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="occultus">Operclipygus occultus</taxonomicName>
G Aedeagus, dorsal and lateral views, of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus perplexus" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="perplexus">Operclipygus perplexus</taxonomicName>
H S8 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus perplexus" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="perplexus">Operclipygus perplexus</taxonomicName>
I T9 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus perplexus" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="perplexus">Operclipygus perplexus</taxonomicName>
.
</paragraph>
</caption>
<caption pageId="117" pageNumber="118">
<paragraph pageId="117" pageNumber="118">
Map 11. Records of the
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus dubius" order="Coleoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="dubius">Operclipygus dubius</taxonomicName>
group.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="117" pageNumber="118" type="remarks">
<paragraph pageId="117" pageNumber="118">Remarks.</paragraph>
<paragraph pageId="117" pageNumber="118">This species can only be recognized by the combination of characters given above.</paragraph>
</subSubSection>
<subSubSection pageId="117" pageNumber="118" type="etymology">
<paragraph pageId="117" pageNumber="118">Etymology.</paragraph>
<paragraph pageId="117" pageNumber="118">
This
<normalizedToken originalValue="species’">species'</normalizedToken>
name refers to the remoteness of its type locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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