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<document id="2193956FD55AD056C2D041F19A7207BE" ID-DOI="10.11646/zootaxa.3718.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="5266830" ID-ZooBank="4B34E42F-EA55-40F5-BF4A-BF40DE2AA91E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629908720460" checkinUser="felipe" docAuthor="Giachino, Pier Mauro, Latella, Leonardo &amp; Vailati, Dante" docDate="2013" docId="B872BD5BFFF8FFD6FF5BAA21FEDACA63" docLanguage="en" docName="zt03718p386.pdf" docOrigin="Zootaxa 3718 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anemadus lucarellii Giachino &amp; Latella &amp; Vailati 2013, sp. nov." docType="treatment" docVersion="4" lastPageNumber="381" masterDocId="444BC523FFF9FFD2FFCCAB45FFBACB23" masterDocTitle="Two new species of the genus Anemadus Reitter, 1885, from the Near East (Coleoptera: Cholevidae)" masterLastPageNumber="386" masterPageNumber="378" pageNumber="379" updateTime="1699127209946" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="F016AA2385D3808E9A201064C61670AC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="FE3957AAFD926412D8B7D80E05B58730">Two new species of the genus Anemadus Reitter, 1885, from the Near East (Coleoptera: Cholevidae)</mods:title>
</mods:titleInfo>
<mods:name id="12F543FD256CED17D42440CBA790822A" type="personal">
<mods:role id="BBC0D449A91865D1F02988D9FDCEA640">
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<mods:namePart id="4F528EE266F10538AB306FB65D28886C">Giachino, Pier Mauro</mods:namePart>
<mods:affiliation id="F6654A20A2C1D5D2AFBEAE3D72C8C935">Settore Fitosanitario Regionale, Environment Park, Palazzina A 2, Via Livorno 60, 10144 Torino (Italy).</mods:affiliation>
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<mods:roleTerm id="E290BFEDFD2098B3C11251E3AD30D49D">Author</mods:roleTerm>
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<mods:namePart id="94B79DC8174E06CB8936044918F25992">Latella, Leonardo</mods:namePart>
<mods:affiliation id="162425A8A3B29BB95E10D47ED69349A6">Museo Civico di Storia Naturale, Lungadige Porta Vittoria 9, 37129 Verona (Italy). E-mail: leonardo. latella @ comune. verona. it</mods:affiliation>
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</mods:role>
<mods:namePart id="A604139EF8CF78E78B71C54AC60BF64F">Vailati, Dante</mods:namePart>
<mods:affiliation id="1100ABE0DB6069E7DE37F76980626928">Via Interna 8, 25127 Brescia (Italy). E-mail: dante. vailati @ libero. it</mods:affiliation>
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<mods:title id="0F80B145D2694626C4C366680BF16812">Zootaxa</mods:title>
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<mods:part id="19CDF194F9567A9DA208DABE15487438">
<mods:date id="8F7F890E1F8F4D3EFDE41DAED3F6A398">2013</mods:date>
<mods:detail id="80E25818A221A3F100BFB88512077AC2" type="pubDate">
<mods:number id="94B33E59FDBBE87D5C5BF7478CFD8D77">2013-10-07</mods:number>
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<mods:detail id="0092CEB0031B5BEE5B5285BA60A55D3A" type="volume">
<mods:number id="1C9EB1C89A9BA2B11C0CF3108C121FBA">3718</mods:number>
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<mods:identifier id="5E485F512D3765E7FB2448AC2937C29B" type="DOI">10.11646/zootaxa.3718.4.7</mods:identifier>
<mods:identifier id="E67A55928DB7F4B363661F74A530CAFB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="00D3B1B692C05D1668B82CE432D974EC" type="Zenodo-Dep">5266830</mods:identifier>
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<treatment id="B872BD5BFFF8FFD6FF5BAA21FEDACA63" ID-DOI="http://doi.org/10.5281/zenodo.5267601" ID-Zenodo-Dep="5267601" LSID="urn:lsid:plazi:treatment:B872BD5BFFF8FFD6FF5BAA21FEDACA63" httpUri="http://treatment.plazi.org/id/B872BD5BFFF8FFD6FF5BAA21FEDACA63" lastPageId="4" lastPageNumber="381" pageId="1" pageNumber="379">
<subSubSection id="78C15FC6FFF8FFD3FF5BAA21FE5BCA5D" box="[151,481,356,382]" pageId="1" pageNumber="379" type="nomenclature">
<paragraph id="30640C4DFFF8FFD3FF5BAA21FE5BCA5D" blockId="1.[151,481,356,418]" box="[151,481,356,382]" pageId="1" pageNumber="379">
<heading id="6B2CBB21FFF8FFD3FF5BAA21FE5BCA5D" bold="true" box="[151,481,356,382]" fontSize="10" level="2" pageId="1" pageNumber="379" reason="2">
<emphasis id="02AFD05FFFF8FFD3FF5BAA21FE5BCA5D" bold="true" box="[151,481,356,382]" pageId="1" pageNumber="379">
<taxonomicName id="F7DB77CEFFF8FFD3FF5BAA21FE3BCA5E" ID-CoL="DWS4" authority="Giachino &amp; Latella &amp; Vailati, 2013" authorityName="Giachino &amp; Latella &amp; Vailati" authorityYear="2013" box="[151,385,356,381]" class="Insecta" family="Leiodidae" genus="Anemadus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="379" phylum="Arthropoda" rank="species" species="lucarellii" status="sp. nov.">Anemadus lucarellii</taxonomicName>
<taxonomicNameLabel id="199C6D24FFF8FFD3FE44AA20FE5BCA5D" box="[392,481,357,382]" pageId="1" pageNumber="379" rank="species">
<emphasis id="02AFD05FFFF8FFD3FE44AA20FE13CA5D" box="[392,425,357,382]" inLineHeading="true" pageId="1" pageNumber="379" reason="1">sp.</emphasis>
nov.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="78C15FC6FFF8FFD3FF5BAACCFEB6CA81" box="[151,268,393,418]" pageId="1" pageNumber="379" type="description">
<paragraph id="30640C4DFFF8FFD3FF5BAACCFEB6CA81" blockId="1.[151,481,356,418]" box="[151,268,393,418]" pageId="1" pageNumber="379">
(
<figureCitation id="A8E010C8FFF8FFD3FF53AACCFEB9CA81" box="[159,259,393,418]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2–4" captionStartId-0="2.[151,249,1674,1696]" captionStartId-1="3.[151,264,1546,1569]" captionTargetBox-1="[321,1264,195,1521]" captionTargetId-1="figure-226@3.[321,1266,195,1522]" captionTargetPageId-1="3" captionText-0="FIGURE 1. Anemadus lucarellii n. sp., habitus of the holotypus ♂. Scale bar: 1 mm." captionText-1="FIGURES 2–4. Anemadus lucarellii n. sp., holotypus ♂. 2: aedeagus in dorsal view; 3: aedeagus in lateral view; 4: apex of left paramere in dorsal view. Scale bars: 0.1 mm." figureDoi-1="http://doi.org/10.5281/zenodo.5266832" httpUri-1="https://zenodo.org/record/5266832/files/figure.png" pageId="1" pageNumber="379">figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="78C15FC6FFF8FFD3FF5BAA94FBD3C975" pageId="1" pageNumber="379" type="materials_examined">
<paragraph id="30640C4DFFF8FFD3FF5BAA94FC21CAC9" blockId="1.[151,1437,465,2002]" box="[151,923,465,490]" pageId="1" pageNumber="379">
<materialsCitation id="80B30610FFF8FFD3FF5BAA94FC2DCAC9" box="[151,919,465,490]" country="Turkey" county="Vilayet" location="Ferzene Magarasi" municipality="Seydisehir" pageId="1" pageNumber="379" specimenCount="1" stateProvince="Konya" typeStatus="holotype">
<emphasis id="02AFD05FFFF8FFD3FF5BAA94FE8FCAC9" bold="true" box="[151,309,465,490]" inLineHeading="true" pageId="1" pageNumber="379" reason="1">
<typeStatus id="EF60B2EFFFF8FFD3FF5BAA94FF68CAC9" box="[151,210,465,490]" pageId="1" pageNumber="379">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="48CC4CDDFFF8FFD3FEF0AA94FE37CAC9" box="[316,397,465,490]" name="Turkey" pageId="1" pageNumber="379">Turkey</collectingCountry>
,
<collectingRegion id="F21FC2AFFFF8FFD3FE54AA97FE59CAC9" box="[408,483,466,490]" country="Turkey" name="Konya" pageId="1" pageNumber="379">Konya</collectingRegion>
<collectingCounty id="D90574C1FFF8FFD3FE26AA94FD84CAC9" box="[490,574,465,490]" pageId="1" pageNumber="379">Vilayet</collectingCounty>
,
<collectingMunicipality id="D0009637FFF8FFD3FD84AA94FD7ACAC9" box="[584,704,465,490]" pageId="1" pageNumber="379">Seydişehir</collectingMunicipality>
,
<location id="35045A96FFF8FFD3FD00AA97FC2DCAC9" LSID="urn:lsid:plazi:treatment:B872BD5BFFF8FFD6FF5BAA21FEDACA63:35045A96FFF8FFD3FD00AA97FC2DCAC9" box="[716,919,465,490]" country="Turkey" county="Vilayet" municipality="Seydisehir" name="Ferzene Magarasi" pageId="1" pageNumber="379" stateProvince="Konya">Ferzene Mağarasi</location>
</materialsCitation>
.
</paragraph>
<paragraph id="30640C4DFFF8FFD3FF0BAAB0FBD3C975" blockId="1.[151,1437,465,2002]" pageId="1" pageNumber="379">
<materialsCitation id="80B30610FFF8FFD3FF0BAAB0FBDFC975" collectingDate="1998-09-27" collectorName="L. Latella" country="Turkey" county="Vilayet" elevation="1470" latitude="37.38912" location="Ferzene Magarasi" longLatPrecision="1" longitude="31.822845" municipality="Seydisehir" pageId="1" pageNumber="379" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Konya" typeStatus="holotype">
<emphasis id="02AFD05FFFF8FFD3FF0BAAB0FEE9C92D" bold="true" box="[199,339,501,526]" inLineHeading="true" pageId="1" pageNumber="379" reason="1">
<typeStatus id="EF60B2EFFFF8FFD3FF0BAAB0FEB8C92D" box="[199,258,501,526]" pageId="1" pageNumber="379">Type</typeStatus>
series.
</emphasis>
HT
<specimenCount id="26DDC7C4FFF8FFD3FE4AAAB3FE26C92E" box="[390,412,502,525]" pageId="1" pageNumber="379" type="male">♂</specimenCount>
,
<collectingCountry id="48CC4CDDFFF8FFD3FE65AAB0FE40C92D" box="[425,506,501,526]" name="Turkey" pageId="1" pageNumber="379">Turkey</collectingCountry>
,
<collectingRegion id="F21FC2AFFFF8FFD3FDCAAAB3FDEBC92D" box="[518,593,502,526]" country="Turkey" name="Konya" pageId="1" pageNumber="379">Konya</collectingRegion>
<collectingCounty id="D90574C1FFF8FFD3FD94AAB0FD16C92D" box="[600,684,501,526]" pageId="1" pageNumber="379">Vilayet</collectingCounty>
,
<collectingMunicipality id="D0009637FFF8FFD3FD7BAAB0FC95C92D" box="[695,815,501,526]" pageId="1" pageNumber="379">Seydişehir</collectingMunicipality>
,
<location id="35045A96FFF8FFD3FCF6AAB3FBBFC92D" LSID="urn:lsid:plazi:treatment:B872BD5BFFF8FFD6FF5BAA21FEDACA63:35045A96FFF8FFD3FCF6AAB3FBBFC92D" box="[826,1029,501,526]" country="Turkey" county="Vilayet" latitude="37.38912" longLatPrecision="1" longitude="31.822845" municipality="Seydisehir" name="Ferzene Magarasi" pageId="1" pageNumber="379" stateProvince="Konya">Ferzene Mağarasi</location>
,
<geoCoordinate id="55EF6A8AFFF8FFD3FBDCAAB0FB75C92D" box="[1040,1231,501,526]" degrees="37" direction="north" minutes="23" orientation="latitude" pageId="1" pageNumber="379" precision="1" seconds="20.82" value="37.38912">37° 23' 20.82&quot; N</geoCoordinate>
–
<geoCoordinate id="55EF6A8AFFF8FFD3FB12AAB0FA2DC92E" box="[1246,1431,501,526]" degrees="31" direction="east" minutes="49" orientation="longitude" pageId="1" pageNumber="379" precision="1" seconds="22.24" value="31.822845">31° 49' 22.24&quot; E</geoCoordinate>
,
<elevation id="BBF6EB7EFFF8FFD3FF5BA95CFEACC911" box="[151,278,537,562]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="379" unit="m" value="1470.0">
<quantity id="F723A1A8FFF8FFD3FF5BA95CFF56C912" box="[151,236,537,562]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="379" unit="m" value="1470.0">1470 m</quantity>
asl
</elevation>
,
<date id="44652A8DFFF8FFD3FEEDA95CFE1EC911" box="[289,420,537,562]" pageId="1" pageNumber="379" value="1998-09-27">
<collectingDate id="5421D365FFF8FFD3FEEDA95CFE1EC911" box="[289,420,537,562]" pageId="1" pageNumber="379" value="1998-09-27">27.IX.1998</collectingDate>
</date>
,
<collectorName id="9D2E699BFFF8FFD3FE63A95FFDA7C911" box="[431,541,537,562]" pageId="1" pageNumber="379">L. Latella</collectorName>
leg. (CLa). PTT:
<specimenCount id="26DDC7C4FFF8FFD3FD2AA95CFC97C912" box="[742,813,537,562]" pageId="1" pageNumber="379" type="female">2 ♀ ♀</specimenCount>
,
<collectingCountry id="48CC4CDDFFF8FFD3FCF7A95CFC36C911" box="[827,908,537,562]" name="Turkey" pageId="1" pageNumber="379">Turkey</collectingCountry>
,
<collectingRegion id="F21FC2AFFFF8FFD3FC54A95FFC59C911" box="[920,995,538,562]" country="Turkey" name="Konya" pageId="1" pageNumber="379">Konya</collectingRegion>
Vilayet, Seydişehir, Ferzene Mağarasi,
<geoCoordinate id="55EF6A8AFFF8FFD3FF5BA978FEEEC975" box="[151,340,573,598]" degrees="37" direction="north" minutes="23" orientation="latitude" pageId="1" pageNumber="379" precision="1" seconds="20.82" value="37.38912">37° 23' 20.82&quot; N</geoCoordinate>
–
<geoCoordinate id="55EF6A8AFFF8FFD3FEA8A978FDA1C976" box="[356,539,573,598]" degrees="31" direction="east" minutes="49" orientation="longitude" pageId="1" pageNumber="379" precision="1" seconds="22.24" value="31.822845">31° 49' 22.24&quot; E</geoCoordinate>
,
<elevation id="BBF6EB7EFFF8FFD3FDEBA978FD1FC975" box="[551,677,573,598]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="379" unit="m" value="1470.0">
<quantity id="F723A1A8FFF8FFD3FDEBA978FDC6C976" box="[551,636,573,598]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="379" unit="m" value="1470.0">1470 m</quantity>
asl
</elevation>
,
<date id="44652A8DFFF8FFD3FD7CA978FC89C975" box="[688,819,573,598]" pageId="1" pageNumber="379" value="1998-09-27">
<collectingDate id="5421D365FFF8FFD3FD7CA978FC89C975" box="[688,819,573,598]" pageId="1" pageNumber="379" value="1998-09-27">27.IX.1998</collectingDate>
</date>
,
<collectorName id="9D2E699BFFF8FFD3FCF2A97BFC11C975" box="[830,939,573,598]" pageId="1" pageNumber="379">L. Latella</collectorName>
leg. (CGi, CVa)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78C15FC6FFF8FFD3FF0BA924FEEDC89D" pageId="1" pageNumber="379" type="diagnosis">
<paragraph id="30640C4DFFF8FFD3FF0BA924FEEDC89D" blockId="1.[151,1437,465,2002]" pageId="1" pageNumber="379">
<emphasis id="02AFD05FFFF8FFD3FF0BA924FEFBC959" bold="true" box="[199,321,609,634]" inLineHeading="true" pageId="1" pageNumber="379" reason="1">Diagnosis.</emphasis>
An
<taxonomicName id="F7DB77CEFFF8FFD3FEB5A924FDF3C959" box="[377,585,609,634]" class="Insecta" family="Leiodidae" genus="Anemadus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="379" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="02AFD05FFFF8FFD3FEB5A924FE57C95A" box="[377,493,609,633]" italics="true" pageId="1" pageNumber="379">Anemadus</emphasis>
species
</taxonomicName>
wich, based on the characters of the aedeagus (apex of the median lobe regularly rounded, not abruptly expanded before the apex and internal sac with a median patch of spines) belongs to the
<taxonomicName id="F7DB77CEFFF8FFD3FF24A9ECFE80C9E2" authorityName="Reitter" authorityYear="1884" box="[232,314,681,705]" class="Insecta" family="Leiodidae" genus="Anemadus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="379" phylum="Arthropoda" rank="species" species="pellitus">
<emphasis id="02AFD05FFFF8FFD3FF24A9ECFE80C9E2" box="[232,314,681,705]" italics="true" pageId="1" pageNumber="379">pellitus</emphasis>
</taxonomicName>
species group (
<emphasis id="02AFD05FFFF8FFD3FE31A9EFFD81C9E2" box="[509,571,682,705]" italics="true" pageId="1" pageNumber="379">sensu</emphasis>
<bibRefCitation id="544A71BCFFF8FFD3FD84A9ECFCCCC9E1" author="Giachino, P. M. &amp; Vailati, D." box="[584,886,681,706]" pageId="1" pageNumber="379" pagination="1 - 314" refId="ref3824" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae Hatch, 1928 (Coleoptera, Cholevidae). Monografie di &quot; Natura Bresciana &quot;, 18, 1 - 314." type="journal article" year="1993">Giachino &amp; Vailati, 1993</bibRefCitation>
), which strongly resembles
<taxonomicName id="F7DB77CEFFF8FFD3FB0BA9EFFEDFC9C5" authority="Ganglbauer, 1900" authorityName="Ganglbauer" authorityYear="1900" class="Insecta" family="Leiodidae" genus="Anemadus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="379" phylum="Arthropoda" rank="species" species="cribratostriatus">
<emphasis id="02AFD05FFFF8FFD3FB0BA9EFFA26C9E2" box="[1223,1436,681,705]" italics="true" pageId="1" pageNumber="379">A. cribratostriatus</emphasis>
Ganglbauer,
<quantity id="F723A1A8FFF8FFD3FEE1A988FEDFC9C5" box="[301,357,717,742]" metricMagnitude="1" metricUnit="m" metricValue="4.826" pageId="1" pageNumber="379" unit="in" value="1900.0">1900</quantity>
</taxonomicName>
in body shape, elytral microsculpture of the
<typeStatus id="EF60B2EFFFF8FFD3FC5EA98BFC78C9C5" box="[914,962,718,742]" pageId="1" pageNumber="379">type</typeStatus>
“C” (with transverse regular rows and longitudinal striae deep and clearly visible) (sensu
<bibRefCitation id="544A71BCFFF8FFD3FD13A9B4FBBAC829" author="Giachino, P. M. &amp; Vailati, D." box="[735,1024,753,778]" pageId="1" pageNumber="379" pagination="1 - 314" refId="ref3824" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae Hatch, 1928 (Coleoptera, Cholevidae). Monografie di &quot; Natura Bresciana &quot;, 18, 1 - 314." type="journal article" year="1993">Giachino &amp; Vailati, 1993</bibRefCitation>
) and structure of the aedeagus. The new species differs from
<taxonomicName id="F7DB77CEFFF8FFD3FE08A853FD22C80E" authorityName="Ganglbauer" authorityYear="1900" box="[452,664,789,813]" class="Insecta" family="Leiodidae" genus="Anemadus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="379" phylum="Arthropoda" rank="species" species="cribratostriatus">
<emphasis id="02AFD05FFFF8FFD3FE08A853FD22C80E" box="[452,664,789,813]" italics="true" pageId="1" pageNumber="379">A. cribratostriatus</emphasis>
</taxonomicName>
in the longer elytra, with apex more strongly pointed and lateral margins less rounded; the pronotum with margins more rounded posteriorly (instead than sub-parallel sided as in
<taxonomicName id="F7DB77CEFFF8FFD3FA48A87FFEFDC856" authorityName="Ganglbauer" authorityYear="1900" class="Insecta" family="Leiodidae" genus="Anemadus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="379" phylum="Arthropoda" rank="species" species="cribratostriatus">
<emphasis id="02AFD05FFFF8FFD3FA48A87FFEFDC856" italics="true" pageId="1" pageNumber="379">A. cribratostriatus</emphasis>
</taxonomicName>
); the longer antennae; the stronger tibiae of all legs; the shape of the median lobe of the aedagus, which is apically more elongated; the shorter and thinner parameres, with the group of inner apical setae more apically inserted.
</paragraph>
</subSubSection>
<subSubSection id="78C15FC6FFF8FFD0FF0BA88CFB7FCC72" lastPageId="2" lastPageNumber="380" pageId="1" pageNumber="379" type="description">
<paragraph id="30640C4DFFF8FFD3FF0BA88CFF61CF0A" blockId="1.[151,1437,465,2002]" pageId="1" pageNumber="379">
<emphasis id="02AFD05FFFF8FFD3FF0BA88CFEE3C8C1" bold="true" box="[199,345,969,994]" inLineHeading="true" pageId="1" pageNumber="379" reason="1">Description.</emphasis>
Total length:
<specimenCount id="26DDC7C4FFF8FFD3FE35A88FFDAAC8C2" box="[505,528,970,993]" pageId="1" pageNumber="379" type="male">♂</specimenCount>
,
<quantity id="F723A1A8FFF8FFD3FDD1A88CFD3FC8C2" box="[541,645,969,994]" metricMagnitude="-3" metricUnit="m" metricValue="3.31" pageId="1" pageNumber="379" unit="mm" value="3.31">3.31 mm</quantity>
,
<specimenCount id="26DDC7C4FFF8FFD3FD5EA88FFD06C8C2" box="[658,700,970,993]" pageId="1" pageNumber="379" type="female">♀♀</specimenCount>
<quantity id="F723A1A8FFF8FFD3FD09A88CFCD6C8C2" box="[709,876,969,994]" metricMagnitude="-3" metricUnit="m" metricValue="3.385" metricValueMax="3.53" metricValueMin="3.24" pageId="1" pageNumber="379" unit="mm" value="3.385" valueMax="3.53" valueMin="3.24">3.24–3.53 mm</quantity>
. Body brown, with testaceous legs, antennae and palpi, antennomeres VII and IX brownish. Integument uniformly covered with yellow pubescence, short and semierect.
</paragraph>
<paragraph id="30640C4DFFF8FFD3FF0BAF70FE6ECFB5" blockId="1.[151,1437,465,2002]" pageId="1" pageNumber="379">Head retractile, eyes well-developed, pubescence short and recumbent on the frons, erect on the clypeus. Antennae short, thin, reaching the basal fifth of elytrae in males and females, VIII antennomere not transverse. Antennomeres length (mm):</paragraph>
<paragraph id="30640C4DFFF8FFD3FF0BAFE7FB78CF99" blockId="1.[151,1437,465,2002]" box="[199,1218,1185,1210]" pageId="1" pageNumber="379">
HT
<specimenCount id="26DDC7C4FFF8FFD3FF3FAFE7FEA7CF9A" box="[243,285,1186,1209]" pageId="1" pageNumber="379" type="male">♂♂</specimenCount>
0.150 - 0.136 - 0.157 - 0.102 - 0.109 - 0.095 - 0.123 - 0.068 - 0.109 - 0.095 - 0.191
</paragraph>
<paragraph id="30640C4DFFF8FFD3FF0BAF83FB1ACFFD" blockId="1.[151,1437,465,2002]" box="[199,1184,1221,1246]" pageId="1" pageNumber="379">
PT
<specimenCount id="26DDC7C4FFF8FFD3FF22AF83FEB9CFFE" box="[238,259,1222,1245]" pageId="1" pageNumber="379" type="female">♀</specimenCount>
0.164 - 0.157 - 0.164 - 0.116 - 0.123 - 0.116 - 0.130 - 0.061 - 0.102 -0.089 - 0.205
</paragraph>
<paragraph id="30640C4DFFF8FFD3FF0BAFAFFDB0CEB1" blockId="1.[151,1437,465,2002]" pageId="1" pageNumber="379">
Pronotum transverse (ratio maximum width/maximum length: 1.50
<specimenCount id="26DDC7C4FFF8FFD3FC02AFAFFC5FCE22" box="[974,997,1258,1281]" pageId="1" pageNumber="379" type="male">♂</specimenCount>
, 1.58–1.64
<specimenCount id="26DDC7C4FFF8FFD3FBA1AFACFB2FCE21" box="[1133,1173,1257,1282]" pageId="1" pageNumber="379" type="female">♀♀</specimenCount>
) widest at mid-length, without basal impression, with almost flat disk near the basal angles; lateral margin regularly curving anteriorly and near the basal corners; basal angles obtuse and slightly rounded. Base of pronotum slightly narrower than elytra, laterally subsinuate. Pubescence on disk yellow, short and recumbent. Pronotum disk roughly granulose with emphasized microsculpture.
</paragraph>
<paragraph id="30640C4DFFF8FFD3FF0BAED8FAB6CD01" blockId="1.[151,1437,465,2002]" pageId="1" pageNumber="379">
Elytra elliptical, very elongate (ratio maximum width/maximum length: 0.58
<specimenCount id="26DDC7C4FFF8FFD3FB92AEDBFBCFCE96" box="[1118,1141,1438,1461]" pageId="1" pageNumber="379" type="male">♂</specimenCount>
, 0.58
<specimenCount id="26DDC7C4FFF8FFD3FB09AED8FB54CE95" box="[1221,1262,1437,1462]" pageId="1" pageNumber="379" type="female">♀♀</specimenCount>
), each elytron rounded and clearly narrowed apically in both sexes. Elytra strongly separated apically, with disk convex, flattened along the suture in the central area. Sculpture of
<typeStatus id="EF60B2EFFFF8FFD3FD0DAEA3FD4BCEDD" box="[705,753,1510,1534]" pageId="1" pageNumber="379">type</typeStatus>
“C” (sensu
<bibRefCitation id="544A71BCFFF8FFD3FCB1AEA0FB24CEDD" author="Giachino, P. M. &amp; Vailati, D." box="[893,1182,1509,1534]" pageId="1" pageNumber="379" pagination="1 - 314" refId="ref3824" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae Hatch, 1928 (Coleoptera, Cholevidae). Monografie di &quot; Natura Bresciana &quot;, 18, 1 - 314." type="journal article" year="1993">Giachino &amp; Vailati, 1993</bibRefCitation>
). Sutural stria present, well-developed, sub-parallel with elytral suture for the basal 2/3, strongly convergent in the distal 1/3.
</paragraph>
<paragraph id="30640C4DFFF8FFD3FF0BAD68FDD8CD65" blockId="1.[151,1437,465,2002]" box="[199,610,1581,1606]" pageId="1" pageNumber="379">Metathoracic wings fully developed.</paragraph>
<paragraph id="30640C4DFFF8FFD3FF0BAD17FE21CD91" blockId="1.[151,1437,465,2002]" pageId="1" pageNumber="379">Legs relatively long, all tibiae strong, protibiae slightly dilated distally, with inner latero-ventral expansion in male. Mesotibiae curved, metatibiae slightly curved. Three basal segments of anterior tarsi dilated in male, as large as the apex of the tibia.</paragraph>
<paragraph id="30640C4DFFF8FFD3FF0BADF8FC91CC61" blockId="1.[151,1437,465,2002]" pageId="1" pageNumber="379">
Aedeagus (
<figureCitation id="A8E010C8FFF8FFD3FE88ADF8FE12CDF5" box="[324,424,1725,1750]" captionStart="FIGURES 2–4" captionStartId="3.[151,264,1546,1569]" captionTargetBox="[321,1264,195,1521]" captionTargetId="figure-226@3.[321,1266,195,1522]" captionTargetPageId="3" captionText="FIGURES 2–4. Anemadus lucarellii n. sp., holotypus ♂. 2: aedeagus in dorsal view; 3: aedeagus in lateral view; 4: apex of left paramere in dorsal view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5266832" httpUri="https://zenodo.org/record/5266832/files/figure.png" pageId="1" pageNumber="379">figs. 2–3</figureCitation>
) quite large, length
<quantity id="F723A1A8FFF8FFD3FD46ADF8FD54CDF6" box="[650,750,1725,1750]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="379" unit="mm" value="0.79">0.79 mm</quantity>
(0.86 if measured with parameres included). Median lobe, in dorsal view, elongate, with lateral edges convergent and subapically slightly concave. Apex acute but not sharp. Median lobe, in lateral view, straight, with slight double-sinuation in the distal half; with sinuation pointing first ventrally and after dorsally; apex slightly bent downwards.
</paragraph>
<paragraph id="30640C4DFFF8FFD3FF0BAC0BFD61CCF1" blockId="1.[151,1437,465,2002]" pageId="1" pageNumber="379">
Parameres (
<figureCitation id="A8E010C8FFF8FFD3FE87AC08FE15CC45" box="[331,431,1869,1894]" captionStart="FIGURES 2–4" captionStartId="3.[151,264,1546,1569]" captionTargetBox="[321,1264,195,1521]" captionTargetId="figure-226@3.[321,1266,195,1522]" captionTargetPageId="3" captionText="FIGURES 2–4. Anemadus lucarellii n. sp., holotypus ♂. 2: aedeagus in dorsal view; 3: aedeagus in lateral view; 4: apex of left paramere in dorsal view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5266832" httpUri="https://zenodo.org/record/5266832/files/figure.png" pageId="1" pageNumber="379">figs. 2–4</figureCitation>
) relatively weak, longer than the median lobe of about 1/7 of their length, slightly curved inwards and with small asymmetrical lateral expansions, larger in the proximal part and gradually tapering distally. Apex curved outwards and upwards, sickle-shaped, with pointed tip; chaetotaxy represented by a group of 4 long ventral setae and one more ventro-apically inserted.
</paragraph>
<caption id="64A45CC5FFFBFFD0FF5BADCFFC4DCDBC" box="[151,1015,1674,1696]" pageId="2" pageNumber="380" startId="2.[151,249,1674,1696]">
<paragraph id="30640C4DFFFBFFD0FF5BADCFFC4DCDBC" blockId="2.[151,1015,1674,1696]" box="[151,1015,1674,1696]" pageId="2" pageNumber="380">
<emphasis id="02AFD05FFFFBFFD0FF5BADCFFEA9CDBC" bold="true" box="[151,275,1674,1696]" pageId="2" pageNumber="380">FIGURE 1.</emphasis>
<taxonomicName id="F7DB77CEFFFBFFD0FED5ADCFFE5FCD83" authority="Giachino &amp; Latella &amp; Vailati, 2013" authorityName="Giachino &amp; Latella &amp; Vailati" authorityYear="2013" box="[281,485,1674,1696]" class="Insecta" family="Leiodidae" genus="Anemadus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="380" phylum="Arthropoda" rank="species" species="lucarellii" status="sp. nov.">
<emphasis id="02AFD05FFFFBFFD0FED5ADCFFE5FCD83" box="[281,485,1674,1696]" italics="true" pageId="2" pageNumber="380">Anemadus lucarellii</emphasis>
</taxonomicName>
<emphasis id="02AFD05FFFFBFFD0FE27ADCFFD9ECDBC" bold="true" box="[491,548,1674,1695]" pageId="2" pageNumber="380">
<taxonomicNameLabel id="199C6D24FFFBFFD0FE27ADCFFD9ECDBC" box="[491,548,1674,1695]" pageId="2" pageNumber="380" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus of the holotypus ♂. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="30640C4DFFFBFFD0FF0BAD89FCD1CC0E" blockId="2.[151,1436,1740,1981]" pageId="2" pageNumber="380">Internal sac of the aedeagus with an elongated patch of spines pointing apically, flanked on each side by a group of setae of different length, the most proximal setae forming towards the base two pairs of phanerae flattened in margins and crenellated, the first of which is not very evident.</paragraph>
<paragraph id="30640C4DFFFBFFD0FF0BAC7DFB7FCC72" blockId="2.[151,1436,1740,1981]" box="[199,1221,1848,1873]" pageId="2" pageNumber="380">
Female genitalia as typical for the genus (see
<bibRefCitation id="544A71BCFFFBFFD0FD0AAC7DFC5ACC72" author="Giachino, P. M. &amp; Vailati, D." box="[710,992,1848,1873]" pageId="2" pageNumber="380" pagination="1 - 314" refId="ref3824" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae Hatch, 1928 (Coleoptera, Cholevidae). Monografie di &quot; Natura Bresciana &quot;, 18, 1 - 314." type="journal article" year="1993">Giachino &amp; Vailati, 1993</bibRefCitation>
for the description).
</paragraph>
</subSubSection>
<subSubSection id="78C15FC6FFFBFFD0FF0BAC19FE0CCC9E" pageId="2" pageNumber="380" type="etymology">
<paragraph id="30640C4DFFFBFFD0FF0BAC19FE0CCC9E" blockId="2.[151,1436,1740,1981]" pageId="2" pageNumber="380">
<emphasis id="02AFD05FFFFBFFD0FF0BAC19FEF4CC56" bold="true" box="[199,334,1884,1909]" inLineHeading="true" pageId="2" pageNumber="380" reason="1">Etymology.</emphasis>
The new species is named after Marco Lucarelli, entomologist at the University of Rome “Tor Vergata”, who organized and took part to the biospeleological expedition in
<collectingCountry id="48CC4CDDFFFBFFD0FC26ACC5FBF0CCBA" box="[1002,1098,1920,1945]" name="Turkey" pageId="2" pageNumber="380">Anatolia</collectingCountry>
with the second author of this paper (Leonardo Latella).
</paragraph>
</subSubSection>
<caption id="64A45CC5FFFAFFD1FF5BAD4FFDE5CD1C" ID-DOI="http://doi.org/10.5281/zenodo.5266832" ID-Zenodo-Dep="5266832" httpUri="https://zenodo.org/record/5266832/files/figure.png" pageId="3" pageNumber="381" startId="3.[151,264,1546,1569]" targetBox="[321,1264,195,1521]" targetPageId="3">
<paragraph id="30640C4DFFFAFFD1FF5BAD4FFDE5CD1C" blockId="3.[151,1436,1546,1600]" pageId="3" pageNumber="381">
<emphasis id="02AFD05FFFFAFFD1FF5BAD4FFE80CD03" bold="true" box="[151,314,1546,1569]" pageId="3" pageNumber="381">FIGURES 2–4.</emphasis>
<taxonomicName id="F7DB77CEFFFAFFD1FEF3AD4EFDB1CD02" authority="Giachino &amp; Latella &amp; Vailati, 2013" authorityName="Giachino &amp; Latella &amp; Vailati" authorityYear="2013" box="[319,523,1547,1569]" class="Insecta" family="Leiodidae" genus="Anemadus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="381" phylum="Arthropoda" rank="species" species="lucarellii" status="sp. nov.">
<emphasis id="02AFD05FFFFAFFD1FEF3AD4EFDB1CD02" box="[319,523,1547,1569]" italics="true" pageId="3" pageNumber="381">Anemadus lucarellii</emphasis>
</taxonomicName>
<emphasis id="02AFD05FFFFAFFD1FDDDAD4EFDF3CD03" bold="true" box="[529,585,1547,1568]" pageId="3" pageNumber="381">
<taxonomicNameLabel id="199C6D24FFFAFFD1FDDDAD4EFDF3CD03" box="[529,585,1547,1568]" pageId="3" pageNumber="381" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotypus ♂. 2: aedeagus in dorsal view; 3: aedeagus in lateral view; 4: apex of left paramere in dorsal view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="78C15FC6FFFAFFD6FF0BAD29FEDACA63" lastPageId="4" lastPageNumber="382" pageId="3" pageNumber="381" type="distribution">
<paragraph id="30640C4DFFFAFFD1FF0BAD29FEAECC1A" blockId="3.[151,1436,1644,2029]" pageId="3" pageNumber="381">
<emphasis id="02AFD05FFFFAFFD1FF0BAD29FE45CDA6" bold="true" box="[199,511,1644,1669]" inLineHeading="true" pageId="3" pageNumber="381" reason="1">Distribution and ecology.</emphasis>
The species is so far known only from one locality, the Ferzene Cave near Seydişehir, in
<collectingRegion id="F21FC2AFFFFAFFD1FEF7ADD4FE3CCD8A" box="[315,390,1681,1705]" country="Turkey" name="Konya" pageId="3" pageNumber="381">Konya</collectingRegion>
Vilayet, South-Western
<collectingCountry id="48CC4CDDFFFAFFD1FD53ADD5FCB9CD8A" box="[671,771,1680,1705]" name="Turkey" pageId="3" pageNumber="381">Anatolia</collectingCountry>
. The cave is horizontal, around
<quantity id="F723A1A8FFFAFFD1FBBBADD5FB7ACD8B" box="[1143,1216,1680,1705]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="381" unit="m" value="400.0">400 m</quantity>
long, and opens at
<quantity id="F723A1A8FFFAFFD1FF5BADF1FF51CDEF" box="[151,235,1716,1741]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="3" pageNumber="381" unit="m" value="1470.0">1470 m</quantity>
a.s.l. It was formed within the contact zone between the Jura Cretaceous aged limestones and Triassic aged argillaceous thin-layered dolomite limestones (
<bibRefCitation id="544A71BCFFFAFFD1FD6FAD9DFC39CDD2" author="Tanindi, O. &amp; Tekin, E. &amp; Semiha, O." box="[675,899,1752,1777]" pageId="3" pageNumber="381" refId="ref4576" refString="Tanindi, O., Tekin, E., Semiha, O. (2003) Archaeological destruction in Turkey: year 2002 preliminary report: Central Anatolia regions: June-October 2002. Trans. E. Tekin, TASK Foundation, Istambul, 46 pp." type="book" year="2003">Tanindi et al., 2003</bibRefCitation>
). The internal temperature at the collecting time (h 12.00) measured 8.4°C. Specimens were found in the entrance gallery. The area outside the cave is a typical oak shrub land.
</paragraph>
<paragraph id="30640C4DFFFAFFD6FF0BAC01FEDACA63" blockId="3.[151,1436,1644,2029]" lastBlockId="4.[151,1436,151,320]" lastPageId="4" lastPageNumber="382" pageId="3" pageNumber="381">
The Ferzene cave, or “Ferzene Mağarasi”, or “Grotte de Fersine” is a cave well known to biospeleology scholars and of particular importance in the history of Turkish biospeleology (
<bibRefCitation id="544A71BCFFFAFFD1FC36AC2DFB75CCA2" author="Latella, L. &amp; Sbordoni, V. &amp; Vigna Taglianti, A." box="[1018,1231,1896,1921]" pageId="3" pageNumber="381" refId="ref4048" refString="Latella, L., Sbordoni, V. &amp; Vigna Taglianti, A. (1998) Biogeographical considerations on the Anatolian cave fauna. XXXII congresso della Societa Italiana di Biogeografia. Biogeografia dell'Anatolia. Roma 29 - 31 ottobre 1998. Book of abstract, pp. 8." type="book" year="1998">Latella et al., 1998</bibRefCitation>
). Known for more than a century, and subject of a monographic contribution by
<bibRefCitation id="544A71BCFFFAFFD1FC97ACC9FBB2CC86" author="Jeannel, R." box="[859,1032,1932,1957]" pageId="3" pageNumber="381" pagination="159 - 174" refId="ref3946" refString="Jeannel, R. (1934 a) Coleopteres cavernicoles de la grotte de Fersine, en Asie mineure. Annales de la Societe Entomologique de France, 103, 159 - 174." type="journal article" year="1934">Jeannel (1934a</bibRefCitation>
,
<bibRefCitation id="544A71BCFFFAFFD1FBD9ACC9FBE5CC86" author="Jeannel, R." box="[1045,1119,1932,1957]" pageId="3" pageNumber="381" pagination="345 - 346" refId="ref3980" refString="Jeannel, R. (1934 b) Coleopteres cavernicoles de la grotte de Fersine, en Asie mineure. Rectification. Annales de la Societe Entomologique de France, 103, 345 - 346." type="journal article" year="1934">1934b</bibRefCitation>
), it was visited by the most famous biospeleologists of the last century including, in 1933, Leo Weirather (
<bibRefCitation id="544A71BCFFFAFFD1FBDCACF5FB17CCEA" author="Pretner, E." box="[1040,1197,1968,1993]" pageId="3" pageNumber="381" pagination="85 - 234" refId="ref4492" refString="Pretner, E. (2011) Die Verdieste von Leo Weirather um die Biospelaologie, insbesondere Jugoslawiens, sein Hohlenkataster und seine Sammelplatze (edited by B. Hauser). Berichte des naturwissenschaftlich-medizinischen Vereins in Innsbruck, 97, 85 - 234." type="journal article" year="2011">Pretner, 2011</bibRefCitation>
). This cave is known to house an interesting subterranean fauna, including the following beetles: the carabids
<taxonomicName id="F7DB77CEFFFAFFD6FBBDAC90FF75CB93" authority="Jeannel, 1934" authorityName="Jeannel" authorityYear="1934" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" lastPageId="4" lastPageNumber="382" order="Coleoptera" pageId="3" pageNumber="381" phylum="Arthropoda" rank="species" species="huetheri">
<emphasis id="02AFD05FFFFAFFD1FBBDAC90FA80CCCE" box="[1137,1338,2005,2029]" italics="true" pageId="3" pageNumber="381">Duvalius huetheri</emphasis>
Jeannel, 1934
</taxonomicName>
and
<taxonomicName id="F7DB77CEFFFDFFD6FEC3ABDCFC6ECB93" authority="Vigna Taglianti, 1999" authorityName="Vigna Taglianti" authorityYear="1999" box="[271,980,151,176]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="382" phylum="Arthropoda" rank="species" species="agnolettii" subGenus="Antisphodrus">
<emphasis id="02AFD05FFFFDFFD6FEC3ABDCFE1BCB93" box="[271,417,153,176]" italics="true" pageId="4" pageNumber="382">Laemostenus</emphasis>
(
<emphasis id="02AFD05FFFFDFFD6FE7AABDDFDF1CB93" box="[438,587,152,176]" italics="true" pageId="4" pageNumber="382">Antisphodrus</emphasis>
)
<emphasis id="02AFD05FFFFDFFD6FDADABDDFD76CB93" box="[609,716,152,176]" italics="true" pageId="4" pageNumber="382">agnolettii</emphasis>
Vigna Taglianti, 1999
</taxonomicName>
(
<bibRefCitation id="544A71BCFFFDFFD6FC2BABD2FADBCB93" author="Casale, A. &amp; Vigna Taglianti, A." box="[999,1377,151,176]" pageId="4" pageNumber="382" pagination="277 - 406" refId="ref3704" refString="Casale, A. &amp; Vigna Taglianti, A. (1999) Caraboid beetles (excl. Cicindelidae) of Anatolia, and their biogeographical significance (Coleoptera, Caraboidea). Biogeographia, 20, 277 - 406." type="journal article" year="1999">Casale &amp; Vigna Taglianti, 1999</bibRefCitation>
), the colevid
<taxonomicName id="F7DB77CEFFFDFFD6FF3EABF9FD0CCBF6" authority="Jeannel, 1934" authorityName="Jeannel" authorityYear="1934" box="[242,694,188,213]" class="Insecta" family="Leiodidae" genus="Huetheriella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="382" phylum="Arthropoda" rank="species" species="maximiliani">
<emphasis id="02AFD05FFFFDFFD6FF3EABF9FDB4CBF7" box="[242,526,188,212]" italics="true" pageId="4" pageNumber="382">Huetheriella maximiliani</emphasis>
Jeannel, 1934
</taxonomicName>
, and the histerid
<taxonomicName id="F7DB77CEFFFDFFD6FCB0ABF9FADECBF6" authority="Jeannel, 1934" authorityName="Jeannel" authorityYear="1934" box="[892,1380,188,213]" class="Insecta" family="Histeridae" genus="Spelaeacritus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="382" phylum="Arthropoda" rank="species" species="anophthalmus">
<emphasis id="02AFD05FFFFDFFD6FCB0ABF9FB06CBF7" box="[892,1212,188,212]" italics="true" pageId="4" pageNumber="382">Spelaeacritus anophthalmus</emphasis>
Jeannel, 1934
</taxonomicName>
. The discovery of a new species of colevid, even if not a troglobite, in a cave considered faunistically well known is therefore of particular interest and underlines the importance of continuing to carry out research in areas considered to be well known.
</paragraph>
</subSubSection>
</treatment>
</document>
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