430 lines
54 KiB
XML
430 lines
54 KiB
XML
<document id="ED3120271BE1622B53BD53D24F30EF74" ID-CLB-Dataset="264974" ID-DOI="10.5852/ejt.2023.890.2261" ID-GBIF-Dataset="ea7ad0da-111f-4709-819b-6a84832ceec5" ID-ISSN="2118-9773" ID-Zenodo-Dep="8305633" ID-ZooBank="5BA69DED-87A2-4041-886F-E11A6D38D52D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1693494219877" checkinUser="felipe" docAuthor="Calcetas, Orlando A." docDate="2023" docId="8B4587A1FFFF59768F67245B9E032673" docLanguage="en" docName="ejt-890_184-203.pdf" docOrigin="European Journal of Taxonomy 890" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucopholis semperi Brenske 1896" docType="treatment" docVersion="6" lastPageNumber="196" masterDocId="777CFFD9FFF7597B8D2323739F57247C" masterDocTitle="Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896" masterLastPageNumber="203" masterPageNumber="184" pageNumber="192" updateTime="1699623951618" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="760AFAA5BA22CA6C5A0E334D23D99EDC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="35AA21004F583148AFE75FA38BA8C782">
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<mods:title id="F99AC158D879DD4BFCA403B9C4A59EA2">Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896</mods:title>
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<mods:roleTerm id="9B662425DD97E8C3CF85020861B0DF12">Author</mods:roleTerm>
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<mods:namePart id="0E0FAC7D26FF0F2FF336D366109CC82D">Calcetas, Orlando A.</mods:namePart>
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<mods:affiliation id="72BEB624C8094269750549B17E895A0D">Department of Agriculture, IVA-CALABARZON, Regional Crop Protection Center (DA-RCPC) Lipa Agricultural Research and Experiment Station (LARES) Brgy. Maraouy, Lipa City, Batangas 4217, Philippines</mods:affiliation>
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<mods:nameIdentifier id="087EA9966EA2DF331D6697F37571D541" type="email">orlando.calcetas@calabarzon.da.gov.ph & orly.calcetas@yahoo.com.ph</mods:nameIdentifier>
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<mods:typeOfResource id="CFD04B6B0C03E0361A395AC1D8768A58">text</mods:typeOfResource>
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<mods:title id="9A6B90D98FADC4B61826C562ED8D30DC">European Journal of Taxonomy</mods:title>
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<mods:part id="9F54E589EAE15E10C31C1C6F30C0017C">
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<mods:date id="77372568461EA695FC5C3A9D8E91A771">2023</mods:date>
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<mods:number id="38E38710B6A1567333304E51B28EF270">2023-08-31</mods:number>
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<mods:detail id="F2C8F52CB91841A6EE9596C573A30295" type="volume">
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<mods:number id="A26C9E63E73A933DD93550EAF70879F0">890</mods:number>
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<mods:identifier id="A60D74A262CB1F5454F7DBE92312A965" type="DOI">10.5852/ejt.2023.890.2261</mods:identifier>
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<treatment id="8B4587A1FFFF59768F67245B9E032673" ID-DOI="http://doi.org/10.5281/zenodo.8306344" ID-GBIF-Taxon="212742581" ID-Zenodo-Dep="8306344" LSID="urn:lsid:plazi:treatment:8B4587A1FFFF59768F67245B9E032673" httpUri="http://treatment.plazi.org/id/8B4587A1FFFF59768F67245B9E032673" lastPageId="13" lastPageNumber="196" pageId="8" pageNumber="192">
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<subSubSection id="4BF6653CFFFF59738F67245B9CA7233F" box="[580,1008,1832,1859]" pageId="8" pageNumber="192" type="nomenclature">
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<paragraph id="035336B7FFFF59738F67245B9CA7233F" blockId="8.[580,1008,1832,1894]" box="[580,1008,1832,1859]" pageId="8" pageNumber="192">
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<heading id="581B81DBFFFF59738F67245B9CA7233F" box="[580,1008,1832,1859]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="192" reason="2">
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<taxonomicName id="C4EC4D34FFFF59738F67245B9CA7233F" ID-CoL="6Q6HW" authority="Brenske, 1896" authorityName="Brenske" authorityYear="1896" box="[580,1008,1832,1859]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="192" phylum="Arthropoda" rank="species" species="semperi">
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<emphasis id="3198EAA5FFFF59738F67245B9C68233F" bold="true" box="[580,831,1832,1859]" italics="true" pageId="8" pageNumber="192">Leucopholis semperi</emphasis>
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<bibRefCitation id="677D4B46FFFF59738E65245A9CA7233F" author="Brenske E." box="[838,1008,1833,1859]" pageId="8" pageNumber="192" pagination="178 - 205" refId="ref7881" refString="Brenske E. 1896. Neue Melolonthiden aus Africa und Asien. Stettiner entomologische Zeitschrift 57: 178 - 205." type="journal article" year="1896">Brenske, 1896</bibRefCitation>
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</taxonomicName>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4BF6653CFFFF59738FFA243F9C0C231A" box="[729,859,1868,1894]" pageId="8" pageNumber="192" type="description">
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||
<paragraph id="035336B7FFFF59738FFA243F9C0C231A" blockId="8.[580,1008,1832,1894]" box="[729,859,1868,1894]" pageId="8" pageNumber="192">
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<figureCitation id="9BD72A32FFFF59738FFA243F9C0C231A" box="[729,859,1868,1894]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="10.[189,233,1962,1988]" captionStartId-1="11.[189,232,1927,1953]" captionTargetBox-0="[202,1398,500,1918]" captionTargetBox-1="[189,1398,480,1860]" captionTargetId-1="figure-86@11.[416,1172,1122,1594]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Fig. 11–14. Leucopholis semperi Brenske, 1896, neotype, ♂ (MNHN). 11. Habitus, dorsal aspect. 12. Metaventral process, dorsal aspect. 13. Male genitalia, dorsal aspect. 14. Male genitalia, lateral aspect." captionText-1="Fig. 15–16. Leucopholis semperi Brenske, 1896. 15. Neotype, ♂ (MNHN), metaventral process, dorsal aspect. 16. Nontypes, ♀♀, dorsal aspect (photograph of Brenske’s non-type series from ZMHB courtesy of Dr Joachim Willers)." figureDoi-0="http://doi.org/10.5281/zenodo.8306342" figureDoi-1="http://doi.org/10.5281/zenodo.8306346" httpUri-0="https://zenodo.org/record/8306342/files/figure.png" httpUri-1="https://zenodo.org/record/8306346/files/figure.png" pageId="8" pageNumber="192">Figs 11–16</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4BF6653CFFFF59738D9E24FE9A522395" pageId="8" pageNumber="192" type="reference_group">
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||
<paragraph id="035336B7FFFF59738D9E24FE9B6923D4" blockId="8.[189,1086,1932,1960]" box="[189,1086,1932,1960]" pageId="8" pageNumber="192">
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||
<treatmentCitationGroup id="23FC1199FFFF59738D9E24FE9B6923D4" box="[189,1086,1932,1960]" pageId="8" pageNumber="192">
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||
<treatmentCitation id="824D10A6FFFF59738D9E24FE9DBB23DB" author="Brenske E." box="[189,748,1932,1959]" page="194" pageId="8" pageNumber="192" year="1896">
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<taxonomicName id="C4EC4D34FFFF59738D9E24FE9DBB23DB" authority="Brenske, 1896: 194" authorityName="Brenske" authorityPageNumber="194" authorityYear="1896" box="[189,748,1932,1959]" class="Insecta" family="Scarabaeidae" genus="Leucopholus" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="192" phylum="Arthropoda" rank="species" species="semperi">
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<emphasis id="3198EAA5FFFF59738D9E24FE9E0423DB" box="[189,339,1933,1959]" italics="true" pageId="8" pageNumber="192">Leucopholus</emphasis>
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[sic!]
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<emphasis id="3198EAA5FFFF59738CBC24FE9EA823DB" box="[415,511,1933,1959]" italics="true" pageId="8" pageNumber="192">Semperi</emphasis>
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<bibRefCitation id="677D4B46FFFF59738F2424FE9DBB23DB" author="Brenske E." box="[519,748,1933,1959]" pageId="8" pageNumber="192" pagination="178 - 205" refId="ref7881" refString="Brenske E. 1896. Neue Melolonthiden aus Africa und Asien. Stettiner entomologische Zeitschrift 57: 178 - 205." type="journal article" year="1896">Brenske, 1896: 194</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(incorrect original spelling).
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="035336B7FFFF59738D9E24BD9A522395" blockId="8.[189,1285,1998,2025]" box="[189,1285,1998,2025]" pageId="8" pageNumber="192">
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<treatmentCitationGroup id="23FC1199FFFF59738D9E24BD9A522395" box="[189,1285,1998,2025]" pageId="8" pageNumber="192">
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<taxonomicName id="C4EC4D34FFFF59738D9E24BD9EE62394" ID-CoL="6Q6HW" authorityName="Brenske" authorityYear="1896" box="[189,433,1998,2024]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="192" phylum="Arthropoda" rank="species" species="semperi">
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<emphasis id="3198EAA5FFFF59738D9E24BD9EE62394" box="[189,433,1998,2024]" italics="true" pageId="8" pageNumber="192">Leucopholis semperi</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="824D10A6FFFF59738CED24BD9D862394" author="Dalla Torre K. W." box="[462,721,1998,2025]" page="178" pageId="8" pageNumber="192" year="1912">
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||
<bibRefCitation id="677D4B46FFFF59738CED24BD9D862394" author="Dalla Torre K. W." box="[462,721,1998,2025]" pageId="8" pageNumber="192" pagination="135 - 209" refId="ref8028" refString="Dalla Torre K. W. 1912. Scarabaeidae: Melolonthinae III. In: Junk W. & Schenkling S. (eds) Coleopterorum Catalogus Vol. 20, Pars 49: 135 - 209. W. Junk, Berlin." type="journal article" year="1912">Dalla Torre 1912: 178</bibRefCitation>
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||
</treatmentCitation>
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(catalogue). —
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<treatmentCitation id="824D10A6FFFF59738EAC24BD9B252394" author="Schultze W. A." box="[911,1138,1998,2024]" page="179" pageId="8" pageNumber="192" year="1916">
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||
<bibRefCitation id="677D4B46FFFF59738EAC24BD9B252394" author="Schultze W. A." box="[911,1138,1998,2024]" pageId="8" pageNumber="192" pagination="1 - 194" refId="ref8197" refString="Schultze W. A. 1916. Catalogue of Philippine Coleoptera. The Philippine Journal of Science 11: 1 - 194." type="journal article" year="1916">Schultze 1916: 179</bibRefCitation>
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</treatmentCitation>
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(catalogue).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="4BF6653CFFFE59728D9E227B9A7C2684" pageId="9" pageNumber="193" type="diagnosis">
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<paragraph id="035336B7FFFE59728D9E227B9E91255E" blockId="9.[189,454,264,290]" box="[189,454,264,290]" pageId="9" pageNumber="193">
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<heading id="581B81DBFFFE59728D9E227B9E91255E" bold="true" box="[189,454,264,290]" fontSize="11" level="3" pageId="9" pageNumber="193" reason="3">
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<emphasis id="3198EAA5FFFE59728D9E227B9E91255E" bold="true" box="[189,454,264,290]" pageId="9" pageNumber="193">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="035336B7FFFE59728D9E22449A7C2684" blockId="9.[187,1400,311,761]" pageId="9" pageNumber="193">
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The metaventral process is barely or not constricted subapically. The pronotum has the medio-subanterior portion with very few to no scales (easily detached) and is mostly covered with ovoid scales. The elytron is covered mostly with narrow spindle-shaped to lanceolate scales. The species can be distinguished from all other species of
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<taxonomicName id="C4EC4D34FFFE59728C8A22D29D6E25C7" authorityName="Dejean" authorityYear="1833" box="[425,569,417,443]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="genus">
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<emphasis id="3198EAA5FFFE59728C8A22D29D6E25C7" box="[425,569,417,443]" italics="true" pageId="9" pageNumber="193">Leucopholis</emphasis>
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</taxonomicName>
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in the
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<collectingCountry id="7BFB7627FFFE59728FAE22D29C4625C7" box="[653,785,417,443]" name="Philippines" pageId="9" pageNumber="193">Philippines</collectingCountry>
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by the paramere with the apical process being bird bill-like and rounded apically. Both
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<taxonomicName id="C4EC4D34FFFE59728F4822B69DBF25A2" authorityName="Brenske" authorityYear="1896" box="[619,744,453,478]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="semperi">
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<emphasis id="3198EAA5FFFE59728F4822B69DBF25A2" box="[619,744,453,478]" italics="true" pageId="9" pageNumber="193">L. semperi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="C4EC4D34FFFE59728E0122B69CCB25A2" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[802,924,453,478]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="stainesi" status="sp. nov.">
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<emphasis id="3198EAA5FFFE59728E0122B69CCB25A2" box="[802,924,453,478]" italics="true" pageId="9" pageNumber="193">L. stainesi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2AAB57DEFFFE59728E8522B69B5625A3" box="[934,1025,453,479]" pageId="9" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
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have a shorter apical processes and distinctly shorter phallobase while they are long and spatulate apically in
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<taxonomicName id="C4EC4D34FFFE59728975229B9B8C267D" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[1110,1243,487,513]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="ratcliffei" status="sp. nov.">
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<emphasis id="3198EAA5FFFE59728975229B9B8C267D" box="[1110,1243,487,513]" italics="true" pageId="9" pageNumber="193">L. ratcliffei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2AAB57DEFFFE597289C1229B9A17267E" box="[1250,1344,488,514]" pageId="9" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
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, and long and tapered apically in
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<taxonomicName id="C4EC4D34FFFE59728F28217F9DD52659" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[523,642,523,549]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
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<emphasis id="3198EAA5FFFE59728F28217F9DD52659" box="[523,642,523,549]" italics="true" pageId="9" pageNumber="193">L. bezdeki</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="2AAB57DEFFFE59728FAA21789DB32659" box="[649,740,523,549]" pageId="9" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
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The depression on each side of the lateral margin of the paramere is shallow and with a rugose surface in
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<taxonomicName id="C4EC4D34FFFE59728E2A215C9CD22634" authorityName="Brenske" authorityYear="1896" box="[777,901,559,584]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="semperi">
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||
<emphasis id="3198EAA5FFFE59728E2A215C9CD22634" box="[777,901,559,584]" italics="true" pageId="9" pageNumber="193">L. semperi</emphasis>
|
||
</taxonomicName>
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,
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<taxonomicName id="C4EC4D34FFFE59728EB6215C9B5A2634" authorityName="Calcetas" authorityYear="2023" box="[917,1037,558,584]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="bezdeki">
|
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<emphasis id="3198EAA5FFFE59728EB6215C9B5A2634" box="[917,1037,558,584]" italics="true" pageId="9" pageNumber="193">L. bezdeki</emphasis>
|
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</taxonomicName>
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,
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<taxonomicName id="C4EC4D34FFFE5972893F215C9BF62634" authorityName="Calcetas" authorityYear="2023" box="[1052,1185,558,584]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="ratcliffei">
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<emphasis id="3198EAA5FFFE5972893F215C9BF62634" box="[1052,1185,558,584]" italics="true" pageId="9" pageNumber="193">L. ratcliffei</emphasis>
|
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</taxonomicName>
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and
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||
<taxonomicName id="C4EC4D34FFFE597289FF215C9A012634" authorityName="Calcetas" authorityYear="2023" box="[1244,1366,559,584]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="stainesi">
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<emphasis id="3198EAA5FFFE597289FF215C9A012634" box="[1244,1366,559,584]" italics="true" pageId="9" pageNumber="193">L. stainesi</emphasis>
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</taxonomicName>
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or with a distinct deep depression in
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<taxonomicName id="C4EC4D34FFFE59728F6D21219D842617" authorityName="Calcetas" authorityYear="2023" box="[590,723,593,619]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="ratcliffei">
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<emphasis id="3198EAA5FFFE59728F6D21219D842617" box="[590,723,593,619]" italics="true" pageId="9" pageNumber="193">L. ratcliffei</emphasis>
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</taxonomicName>
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and
|
||
<taxonomicName id="C4EC4D34FFFE59728E2F21219CD22617" authorityName="Calcetas" authorityYear="2023" box="[780,901,594,619]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="stainesi">
|
||
<emphasis id="3198EAA5FFFE59728E2F21219CD22617" box="[780,901,594,619]" italics="true" pageId="9" pageNumber="193">L. stainesi</emphasis>
|
||
</taxonomicName>
|
||
. The metaventral process is also extended in front of the prosternal process in all four above mentioned species while it is slightly extended in all other species of Philippine
|
||
<taxonomicName id="C4EC4D34FFFE59728F1221EB9D9526CE" authorityName="Dejean" authorityYear="1833" box="[561,706,664,690]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3198EAA5FFFE59728F1221EB9D9526CE" box="[561,706,664,690]" italics="true" pageId="9" pageNumber="193">Leucopholis</emphasis>
|
||
</taxonomicName>
|
||
. The abdominal scales of
|
||
<taxonomicName id="C4EC4D34FFFE5972892621EA9BD426CE" authorityName="Brenske" authorityYear="1896" box="[1029,1155,665,690]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFE5972892621EA9BD426CE" box="[1029,1155,665,690]" italics="true" pageId="9" pageNumber="193">L. semperi</emphasis>
|
||
</taxonomicName>
|
||
are short ovoid and with longitudinal furrows while these furrows are absent in
|
||
<taxonomicName id="C4EC4D34FFFE59728E5D21CF9CA226A9" authorityName="Calcetas" authorityYear="2023" box="[894,1013,699,725]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="bezdeki">
|
||
<emphasis id="3198EAA5FFFE59728E5D21CF9CA226A9" box="[894,1013,699,725]" italics="true" pageId="9" pageNumber="193">L. bezdeki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C4EC4D34FFFE5972892021CF9BDF26A9" authorityName="Calcetas" authorityYear="2023" box="[1027,1160,699,725]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="ratcliffei">
|
||
<emphasis id="3198EAA5FFFE5972892021CF9BDF26A9" box="[1027,1160,699,725]" italics="true" pageId="9" pageNumber="193">L. ratcliffei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C4EC4D34FFFE597289E221CF9A6C26A9" authorityName="Calcetas" authorityYear="2023" box="[1217,1339,700,725]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="stainesi">
|
||
<emphasis id="3198EAA5FFFE597289E221CF9A6C26A9" box="[1217,1339,700,725]" italics="true" pageId="9" pageNumber="193">L. stainesi</emphasis>
|
||
</taxonomicName>
|
||
. The posterolateral margin of the elytron of
|
||
<taxonomicName id="C4EC4D34FFFE59728FA021AC9DA82684" authorityName="Brenske" authorityYear="1896" box="[643,767,735,760]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFE59728FA021AC9DA82684" box="[643,767,735,760]" italics="true" pageId="9" pageNumber="193">L. semperi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C4EC4D34FFFE59728E1B21AC9CE62684" authorityName="Calcetas" authorityYear="2023" box="[824,945,735,760]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="stainesi">
|
||
<emphasis id="3198EAA5FFFE59728E1B21AC9CE62684" box="[824,945,735,760]" italics="true" pageId="9" pageNumber="193">L. stainesi</emphasis>
|
||
</taxonomicName>
|
||
are vertically flattened laterally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BF6653CFFFE59728D9E20539C162044" pageId="9" pageNumber="193" type="materials_examined">
|
||
<paragraph id="035336B7FFFE59728D9E20539EFD2746" blockId="9.[189,426,800,826]" box="[189,426,800,826]" pageId="9" pageNumber="193">
|
||
<heading id="581B81DBFFFE59728D9E20539EFD2746" bold="true" box="[189,426,800,826]" fontSize="11" level="3" pageId="9" pageNumber="193" reason="3">
|
||
<emphasis id="3198EAA5FFFE59728D9E20539EFD2746" bold="true" box="[189,426,800,826]" pageId="9" pageNumber="193">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFE59728DC9203C9B4C27CC" blockId="9.[189,1398,847,944]" pageId="9" pageNumber="193">
|
||
<materialsCitation id="B3843CEAFFFE59728DC9203C9B4C27CC" ID-GBIF-Occurrence="4402534306" collectionCode="MNHN" country="Philippines" determinerName="O. A. Calcetas" location="Mindanao" municipality="Surigao" pageId="9" pageNumber="193" specimenCode="MNHN-16275" specimenCount="2" specimenCount-male="2" typeStatus="neotype">
|
||
<emphasis id="3198EAA5FFFE59728DC9203C9E062715" bold="true" box="[234,337,847,873]" pageId="9" pageNumber="193">
|
||
<typeStatus id="DC578815FFFE59728DC9203C9E062715" box="[234,337,847,873]" pageId="9" pageNumber="193" type="neotype">Neotype</typeStatus>
|
||
</emphasis>
|
||
(here designated)
|
||
<collectingCountry id="7BFB7627FFFE59728D9E20019E3927F0" box="[189,366,882,908]" name="Philippines" pageId="9" pageNumber="193">PHILIPPINES</collectingCountry>
|
||
•
|
||
<specimenCount id="15EAFD3EFFFE59728CAF20019EF227F0" box="[396,421,882,908]" pageId="9" pageNumber="193" type="male">♂</specimenCount>
|
||
; “
|
||
<collectingMunicipality id="E337ACCDFFFE59728C9D20019D7527F0" box="[446,546,882,908]" pageId="9" pageNumber="193">Surigao</collectingMunicipality>
|
||
,
|
||
<location id="0633606CFFFE59728F1220019DF927F0" LSID="urn:lsid:plazi:treatment:8B4587A1FFFF59768F67245B9E032673:0633606CFFFE59728F1220019DF927F0" box="[561,686,882,908]" country="Philippines" municipality="Surigao" name="Mindanao" pageId="9" pageNumber="193">Mindanao</location>
|
||
,
|
||
<location id="0633606CFFFE59728F9F20019C5D27F0" LSID="urn:lsid:plazi:treatment:8B4587A1FFFF59768F67245B9E032673:0633606CFFFE59728F9F20019C5D27F0" box="[700,778,882,908]" country="Philippines" municipality="Surigao" name="Baker" pageId="9" pageNumber="193">Baker</location>
|
||
”; “
|
||
<taxonomicName id="C4EC4D34FFFE59728E1220019BF727F0" authority="Brenske" authorityName="Brenske" authorityYear="1896" box="[817,1184,882,908]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFE59728E1220019B7A27F0" box="[817,1069,882,908]" italics="true" pageId="9" pageNumber="193">Leucopholis semperi</emphasis>
|
||
Brenske
|
||
</taxonomicName>
|
||
”; “NEOTYPE
|
||
<specimenCount id="15EAFD3EFFFE5972887B20019A2727F0" box="[1368,1392,882,908]" pageId="9" pageNumber="193" type="male">♂</specimenCount>
|
||
,
|
||
<taxonomicName id="C4EC4D34FFFE59728D9E20E69D4C27D3" authority="Brenske" authorityName="Brenske" authorityYear="1896" box="[189,539,917,943]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="193" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFE59728D9E20E69EE627D3" box="[189,433,917,943]" italics="true" pageId="9" pageNumber="193">Leucopholis semperi</emphasis>
|
||
Brenske
|
||
</taxonomicName>
|
||
, des.
|
||
<determinerName id="F68FDA73FFFE59728F7F20E69C5427D3" box="[604,771,917,943]" pageId="9" pageNumber="193">O.A. Calcetas</determinerName>
|
||
, 2022”;
|
||
<specimenCode id="534A9ECCFFFE59728E4520E69B4027CC" box="[870,1047,917,944]" collectionCode="MNHN-" pageId="9" pageNumber="193">MNHN-16275</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFE59728DC920A59EB6278C" blockId="9.[189,1398,982,1080]" box="[234,481,982,1008]" pageId="9" pageNumber="193">
|
||
<emphasis id="3198EAA5FFFE59728DC920A59EB6278C" bold="true" box="[234,481,982,1008]" pageId="9" pageNumber="193">Additional material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFE59728D9E20899C162044" blockId="9.[189,1398,982,1080]" pageId="9" pageNumber="193">
|
||
<materialsCitation id="B3843CEAFFFE59728D9E20899BB72069" ID-GBIF-Occurrence="4402534304" box="[189,1248,1018,1045]" collectingDate="1915-04-16" collectionCode="ZMHB" collectorName="S. Antln" country="Philippines" location="Burauen" pageId="9" pageNumber="193" specimenCount="2" specimenCount-female="2" stateProvince="Leyte">
|
||
<collectingCountry id="7BFB7627FFFE59728D9E20899E3D2068" box="[189,362,1018,1044]" name="Philippines" pageId="9" pageNumber="193">PHILIPPINES</collectingCountry>
|
||
•
|
||
<specimenCount id="15EAFD3EFFFE59728CA520899E9D2069" box="[390,458,1018,1045]" pageId="9" pageNumber="193" type="female">2 ♀♀</specimenCount>
|
||
; Philippinen,
|
||
<collectingRegion id="C128F855FFFE59728F5220889DE02069" box="[625,695,1019,1045]" country="Philippines" name="Southern Leyte" pageId="9" pageNumber="193">Leyte</collectingRegion>
|
||
,
|
||
<location id="0633606CFFFE59728FE720889C7C2069" LSID="urn:lsid:plazi:treatment:8B4587A1FFFF59768F67245B9E032673:0633606CFFFE59728FE720889C7C2069" box="[708,811,1019,1045]" country="Philippines" municipality="Philippinen" name="Burauen" pageId="9" pageNumber="193" stateProvince="Leyte">Burauen</location>
|
||
;
|
||
<date id="77521077FFFE59728E1A20899C802069" box="[825,983,1018,1045]" pageId="9" pageNumber="193" value="1915-04-16">
|
||
<collectingDate id="6716E99FFFFE59728E1A20899C802069" box="[825,983,1018,1045]" pageId="9" pageNumber="193" value="1915-04-16">16 Apr. 1915</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="AE195361FFFE59728EC620899B102068" box="[997,1095,1018,1044]" pageId="9" pageNumber="193">S. Antln</collectorName>
|
||
leg.;
|
||
<collectionCode id="65FDAE72FFFE597289A820889BB72069" box="[1163,1248,1019,1045]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="9" pageNumber="193">ZMHB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="B3843CEAFFFE597289DF20899C6A2044" ID-GBIF-Occurrence="4402534301" collectingDate="1915-04-20" collectionCode="ZMHB" collectorName="S. Antln" country="Philippines" location="Burauen" pageId="9" pageNumber="193" specimenCount="1" specimenCount-female="1" stateProvince="Leyte">
|
||
<specimenCount id="15EAFD3EFFFE597289DF20899A7D2069" box="[1276,1322,1018,1045]" pageId="9" pageNumber="193" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<date id="77521077FFFE59728F1E276E9D8F2044" box="[573,728,1053,1080]" pageId="9" pageNumber="193" value="1915-04-20">
|
||
<collectingDate id="6716E99FFFFE59728F1E276E9D8F2044" box="[573,728,1053,1080]" pageId="9" pageNumber="193" value="1915-04-20">20 Apr. 1915</collectingDate>
|
||
</date>
|
||
;
|
||
<collectionCode id="65FDAE72FFFE59728FC7276D9C6A2044" box="[740,829,1054,1080]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="9" pageNumber="193">ZMHB</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BF6653CFFFE59778D9E272C9B2421AC" lastPageId="12" lastPageNumber="196" pageId="9" pageNumber="193" type="description">
|
||
<paragraph id="035336B7FFFE59728D9E272C9E0A2005" blockId="9.[189,349,1119,1145]" box="[189,349,1119,1145]" pageId="9" pageNumber="193">
|
||
<heading id="581B81DBFFFE59728D9E272C9E0A2005" bold="true" box="[189,349,1119,1145]" fontSize="11" level="3" pageId="9" pageNumber="193" reason="3">
|
||
<emphasis id="3198EAA5FFFE59728D9E272C9E0A2005" bold="true" box="[189,349,1119,1145]" pageId="9" pageNumber="193">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFE59728D9E27FE9EFC20DB" blockId="9.[189,427,1165,1191]" box="[189,427,1165,1191]" pageId="9" pageNumber="193">
|
||
<collectingCountry id="7BFB7627FFFE59728D9E27FE9E1120DB" box="[189,326,1165,1191]" name="Philippines" pageId="9" pageNumber="193">Philippines</collectingCountry>
|
||
: Surigao
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFE59728D9E27BC9D742095" blockId="9.[189,547,1231,1257]" box="[189,547,1231,1257]" pageId="9" pageNumber="193">
|
||
<heading id="581B81DBFFFE59728D9E27BC9D742095" bold="true" box="[189,547,1231,1257]" fontSize="11" level="3" pageId="9" pageNumber="193" reason="3">
|
||
<emphasis id="3198EAA5FFFE59728D9E27BC9D742095" bold="true" box="[189,547,1231,1257]" pageId="9" pageNumber="193">
|
||
Redescription of the
|
||
<typeStatus id="DC578815FFFE59728CE227BC9D742095" box="[449,547,1231,1257]" pageId="9" pageNumber="193" type="neotype">neotype</typeStatus>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFE59728D9E278D9EB32164" blockId="9.[189,484,1277,1304]" box="[189,484,1277,1304]" pageId="9" pageNumber="193">BODY LENGTH. 36.0 mm.</paragraph>
|
||
<paragraph id="035336B7FFFE59728D9E264C9E812126" blockId="9.[189,470,1343,1370]" box="[189,470,1343,1370]" pageId="9" pageNumber="193">BODY WIDTH. 17.0 mm.</paragraph>
|
||
<paragraph id="035336B7FFFE59728D9E26F29BAE21C2" blockId="9.[189,1399,1409,1470]" pageId="9" pageNumber="193">
|
||
COLOUR. Dorsum monochromatic blackish. Head, pronotum, scutellum, elytra, legs, blackish; covered with yellowish white scales (
|
||
<figureCitation id="9BD72A32FFFE59728F3226D79D3021C2" box="[529,615,1444,1470]" captionStart="Fig" captionStartId="10.[189,233,1962,1988]" captionTargetBox="[202,1398,500,1918]" captionTargetPageId="10" captionText="Fig. 11–14. Leucopholis semperi Brenske, 1896, neotype, ♂ (MNHN). 11. Habitus, dorsal aspect. 12. Metaventral process, dorsal aspect. 13. Male genitalia, dorsal aspect. 14. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306342" httpUri="https://zenodo.org/record/8306342/files/figure.png" pageId="9" pageNumber="193">Fig. 11</figureCitation>
|
||
). Venter blackish, covered with whitish scales (
|
||
<figureCitation id="9BD72A32FFFE597289B726D79BBB21C2" box="[1172,1260,1444,1470]" captionStart="Fig" captionStartId="10.[189,233,1962,1988]" captionTargetBox="[202,1398,500,1918]" captionTargetPageId="10" captionText="Fig. 11–14. Leucopholis semperi Brenske, 1896, neotype, ♂ (MNHN). 11. Habitus, dorsal aspect. 12. Metaventral process, dorsal aspect. 13. Male genitalia, dorsal aspect. 14. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306342" httpUri="https://zenodo.org/record/8306342/files/figure.png" pageId="9" pageNumber="193">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFE59728D9E26959C202366" blockId="9.[187,1400,1509,1818]" pageId="9" pageNumber="193">
|
||
HEAD. Clypeus with medial anterior margin nearly straight anteriorly; slightly cleft, slanted at 60° angle laterally; anterior surface uneven, lustrous, impunctate subanteriorly; above clypeo-labral suture with row of moderate-sized, rounded to rugose punctures adjacent to posterior margin; each puncture with stiff brownish hair or seta; anterior and lateral margins carinate dorsally, slightly concave medially; anterior angle widely rounded; dorsal surface covered with short or long, elongate, lanceolate, rounded to tapered apically scales. Apical maXillary palpomere rice grain-like, with spindle-shaped to oval flattened area. Mentum pot-shaped, lustrous, glabrous; with very few hairbrushes on each side of medial cleft subanteriorly; with slightly sunken surface on each four corners. Antennal lamellae length
|
||
<quantity id="C4149B52FFFE5972883725AE9A23228B" box="[1300,1396,1757,1783]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="193" unit="mm" value="3.2">3.2 mm</quantity>
|
||
, distinctly longer than entire length of antennomeres II–VII.
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFE59718D9E24329C5025F0" blockId="9.[187,1400,1857,2025]" lastBlockId="10.[187,1399,265,397]" lastPageId="10" lastPageNumber="194" pageId="9" pageNumber="193">PRONOTUM. Anterior margin evenly, widely concave, nearly straight medially; anterior angle obtuse, with lustrous, glabrous, impunctate callosity subapically; rounded apically; lateral margin widely convex; anterolateral at 45° angle, crenulate; medial margin crenulate; posterolateral margin wedge-shaped, slightly crenulate; posterior angle at 90° angle, strongly sinuate, rounded apically; posterior margin widely concave, deeply rounded medially, distinctly extended posteriorly; surface covered with variable scales; covered mostly with short, ovoid, elongate, rounded apically scales medio-subanteriorly; covered mostly with elongate, spindle-shaped, tapered apically scales medio-subposteriorly; covered mostly with elongate, lanceolate, tapered apically scales submedially; covered mostly with short, ovoid, stout medially, rounded apically scales adjacent to lateral margin.</paragraph>
|
||
<caption id="5793663FFFFD59718D9E24D99A202394" ID-DOI="http://doi.org/10.5281/zenodo.8306342" ID-Zenodo-Dep="8306342" httpUri="https://zenodo.org/record/8306342/files/figure.png" pageId="10" pageNumber="194" startId="10.[189,233,1962,1988]" targetBox="[202,1398,500,1918]" targetPageId="10">
|
||
<paragraph id="035336B7FFFD59718D9E24D99A202394" blockId="10.[189,1399,1962,2024]" pageId="10" pageNumber="194">
|
||
<emphasis id="3198EAA5FFFD59718D9E24D99E1D23B8" bold="true" box="[189,330,1962,1988]" pageId="10" pageNumber="194">Fig. 11–14.</emphasis>
|
||
<taxonomicName id="C4EC4D34FFFD59718C7724D99C5823B8" authority="Brenske, 1896" authorityName="Brenske" authorityYear="1896" box="[340,783,1962,1988]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="194" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFD59718C7724D99D0623B8" box="[340,593,1962,1988]" italics="true" pageId="10" pageNumber="194">Leucopholis semperi</emphasis>
|
||
<bibRefCitation id="677D4B46FFFD59718F7824D99C5823B8" author="Brenske E." box="[603,783,1962,1988]" pageId="10" pageNumber="194" pagination="178 - 205" refId="ref7881" refString="Brenske E. 1896. Neue Melolonthiden aus Africa und Asien. Stettiner entomologische Zeitschrift 57: 178 - 205." type="journal article" year="1896">Brenske, 1896</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="DC578815FFFD59718E3D24D99CD723B8" box="[798,896,1962,1988]" pageId="10" pageNumber="194" type="neotype">neotype</typeStatus>
|
||
,
|
||
<specimenCount id="15EAFD3EFFFD59718EAC24D99CF223B8" box="[911,933,1962,1988]" pageId="10" pageNumber="194" type="male">♂</specimenCount>
|
||
(MNHN).
|
||
<emphasis id="3198EAA5FFFD5971891124D99B1923B8" bold="true" box="[1074,1102,1962,1988]" pageId="10" pageNumber="194">11</emphasis>
|
||
. Habitus, dorsal aspect.
|
||
<emphasis id="3198EAA5FFFD59718D9E24BE9F8D239B" bold="true" box="[189,218,1997,2023]" pageId="10" pageNumber="194">12</emphasis>
|
||
. Metaventral process, dorsal aspect.
|
||
<emphasis id="3198EAA5FFFD59718F5E24BE9DCD239B" bold="true" box="[637,666,1997,2023]" pageId="10" pageNumber="194">13</emphasis>
|
||
. Male genitalia, dorsal aspect.
|
||
<emphasis id="3198EAA5FFFD59718ED824BE9B4F239B" bold="true" box="[1019,1048,1997,2023]" pageId="10" pageNumber="194">14</emphasis>
|
||
. Male genitalia, lateral aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="035336B7FFFC59778D9E227A9E9925CC" blockId="11.[187,1399,265,397]" lastBlockId="12.[189,1400,265,432]" lastPageId="12" lastPageNumber="196" pageId="11" pageNumber="195">
|
||
VENTRAL SIDE OF THORAX. Prosternal process short, nearly isosceles triangulate; widely rounded apically; with flattened impunctate area medially and apically; covered with very few hairs or seta, with very few elongate, thin, narrow, parallel-sided, tapered apically scales. Metaventral process length
|
||
<quantity id="C4149B52FFFC5970898A22239A542516" box="[1193,1283,336,362]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="11" pageNumber="195" unit="mm" value="4.8">4.8 mm</quantity>
|
||
; distinctly wide, triangulate basally, elongate towards apex; not constricted subapically; subapical lateral margin distinctly long, nearly parallel-sided; widely rounded apically; surface lustrous, glabrous, impunctate dorsally; with faint, longitudinal thin dark line medially; dark line much distinct in metasternum; each side and ventrally covered with scales of variable shapes and sizes; covered with very long stiff hairs (
|
||
<figureCitation id="9BD72A32FFFB59778DE522019E4825F1" box="[198,287,370,397]" captionStart="Fig" captionStartId="11.[189,232,1927,1953]" captionTargetBox="[189,1398,480,1860]" captionTargetId="figure-86@11.[416,1172,1122,1594]" captionTargetPageId="11" captionText="Fig. 15–16. Leucopholis semperi Brenske, 1896. 15. Neotype, ♂ (MNHN), metaventral process, dorsal aspect. 16. Nontypes, ♀♀, dorsal aspect (photograph of Brenske’s non-type series from ZMHB courtesy of Dr Joachim Willers)." figureDoi="http://doi.org/10.5281/zenodo.8306346" httpUri="https://zenodo.org/record/8306346/files/figure.png" pageId="12" pageNumber="196">Fig. 15</figureCitation>
|
||
). Metasternum with medial depression, covered mostly with elongate, ovoid, stout medially, tapered apically scales.
|
||
</paragraph>
|
||
<caption id="5793663FFFFC59708D9E24F49E852395" ID-DOI="http://doi.org/10.5281/zenodo.8306346" ID-Zenodo-Dep="8306346" httpUri="https://zenodo.org/record/8306346/files/figure.png" pageId="11" pageNumber="195" startId="11.[189,232,1927,1953]" targetBox="[189,1398,480,1860]" targetPageId="11">
|
||
<paragraph id="035336B7FFFC59708D9E24F49E852395" blockId="11.[189,1399,1927,2025]" pageId="11" pageNumber="195">
|
||
<emphasis id="3198EAA5FFFC59708D9E24F49E1223DD" bold="true" box="[189,325,1927,1953]" pageId="11" pageNumber="195">Fig. 15–16.</emphasis>
|
||
<taxonomicName id="C4EC4D34FFFC59708C6F24F49DA323DD" authority="Brenske, 1896" authorityName="Brenske" authorityYear="1896" box="[332,756,1927,1953]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="195" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFC59708C6F24F49D1723DD" box="[332,576,1927,1953]" italics="true" pageId="11" pageNumber="195">Leucopholis semperi</emphasis>
|
||
<bibRefCitation id="677D4B46FFFC59708F6424F49DA323DD" author="Brenske E." box="[583,756,1927,1953]" pageId="11" pageNumber="195" pagination="178 - 205" refId="ref7881" refString="Brenske E. 1896. Neue Melolonthiden aus Africa und Asien. Stettiner entomologische Zeitschrift 57: 178 - 205." type="journal article" year="1896">Brenske, 1896</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="3198EAA5FFFC59708FDC24F49C4B23DD" bold="true" box="[767,796,1927,1953]" pageId="11" pageNumber="195">15</emphasis>
|
||
.
|
||
<typeStatus id="DC578815FFFC59708E0824FB9CC623DE" box="[811,913,1928,1954]" pageId="11" pageNumber="195" type="neotype">Neotype</typeStatus>
|
||
,
|
||
<specimenCount id="15EAFD3EFFFC59708EBE24FB9CE423DE" box="[925,947,1928,1954]" pageId="11" pageNumber="195" type="male">♂</specimenCount>
|
||
(MNHN), metaventral process, dorsal aspect.
|
||
<emphasis id="3198EAA5FFFC59708C3724D99E6623B8" bold="true" box="[276,305,1962,1988]" pageId="11" pageNumber="195">16</emphasis>
|
||
. Nontypes,
|
||
<specimenCount id="15EAFD3EFFFC59708C9E24D89EBD23B9" box="[445,490,1963,1989]" pageId="11" pageNumber="195" type="female">♀♀</specimenCount>
|
||
, dorsal aspect (photograph of Brenske’s non-type series from ZMHB courtesy of Dr Joachim Willers).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="035336B7FFFB59778D9E22A49BEA258D" blockId="12.[189,1213,471,497]" box="[189,1213,471,497]" pageId="12" pageNumber="196">LEG. Foretibia bidentate, widely rounded apically; posterior metatibia with 22 spicules.</paragraph>
|
||
<paragraph id="035336B7FFFB59778D9E216A9BB5262A" blockId="12.[187,1399,537,598]" pageId="12" pageNumber="196">SCUTELLUM. Covered with small, moderate-sized to long, elongate, ovoid, slightly stout medially, rounded apically scales; lustrous, impunctate, without scales medially and around posterior margin.</paragraph>
|
||
<paragraph id="035336B7FFFB59778D9E210E9C4C2716" blockId="12.[187,1400,637,875]" pageId="12" pageNumber="196">ELYTRON. Anterior margin slightly wedge-shaped; anterior angle widely rounded; thickly carinate, explanate; with narrow suture towards anterior angle; margin, evanescent medially; posterolateral margin vertically flattened towards posterior angle; posterior angle obtuse, widely rounded, moderately eXplanate, evanescent towards sutural angle; posterior margin smooth, slightly concave towards sutural angle; sutural angle slightly obtuse; sutural margin medially and towards sutural angle distinctly carinate, towards scutellum infleXed, not carinate; surface mostly covered with elongate, narrow, spindle-shaped to lanceolate, tapered to round, bent apically scales.</paragraph>
|
||
<paragraph id="035336B7FFFB59778D9E20E19BBB278E" blockId="12.[187,1398,914,1011]" pageId="12" pageNumber="196">ABDOMEN.Abdominal sternites I–V covered mostly with short, ovoid, inverted-lanceolate, tapered basally, widely rounded to truncate apically scales; scales with distinct longitudinal grooves or furrows starting basally and terminating medially; sternite VI covered with variable shapes and sizes scales.</paragraph>
|
||
<paragraph id="035336B7FFFB59778D9E27699BD720E1" blockId="12.[189,1399,1050,1182]" pageId="12" pageNumber="196">PYGIDIUM. Anterolateral margin thickly carinate towards anterior angle, disappearing medially; posterolateral margin widely explanate medially towards posterior margin; posterior margin slightly concave, narrowly explanate, depressed medially; subposterior margin sloping downward laterally; surface covered mostly with large, elongate spindle-shaped, stout medially scales.</paragraph>
|
||
<paragraph id="035336B7FFFB59778D9E27B69A3E2110" blockId="12.[187,1400,1221,1388]" pageId="12" pageNumber="196">
|
||
MALE GENITALIA. Genitalia length
|
||
<quantity id="C4149B52FFFB59778F6627B69DF920A3" box="[581,686,1221,1247]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="12" pageNumber="196" unit="mm" value="12.8">12.8 mm</quantity>
|
||
; phallobase with dorsal anterior margin W-shaped, moderately convex medially (
|
||
<figureCitation id="9BD72A32FFFB59778CAD279B9EB1217E" box="[398,486,1256,1282]" captionStart="Fig" captionStartId="10.[189,233,1962,1988]" captionTargetBox="[202,1398,500,1918]" captionTargetPageId="10" captionText="Fig. 11–14. Leucopholis semperi Brenske, 1896, neotype, ♂ (MNHN). 11. Habitus, dorsal aspect. 12. Metaventral process, dorsal aspect. 13. Male genitalia, dorsal aspect. 14. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306342" httpUri="https://zenodo.org/record/8306342/files/figure.png" pageId="12" pageNumber="196">Fig. 13</figureCitation>
|
||
). Paramere with dorsal basal margin bisinuate, bowl-shaped, slightly concave medially. Apical process length 3.0 mm, distinctly shorter than phallobase; bird bill-like, with widely rounded apex; apical margin bisinuate; posterior margin slightly extended, rounded apically. Lateral margin of paramere with wide, shallow, rugose surface longitudinal depression on each side (
|
||
<figureCitation id="9BD72A32FFFB5977882726219A0B2110" box="[1284,1372,1362,1388]" captionStart="Fig" captionStartId="10.[189,233,1962,1988]" captionTargetBox="[202,1398,500,1918]" captionTargetPageId="10" captionText="Fig. 11–14. Leucopholis semperi Brenske, 1896, neotype, ♂ (MNHN). 11. Habitus, dorsal aspect. 12. Metaventral process, dorsal aspect. 13. Male genitalia, dorsal aspect. 14. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306342" httpUri="https://zenodo.org/record/8306342/files/figure.png" pageId="12" pageNumber="196">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFB59778D9E26E09B2421AC" blockId="12.[189,1398,1427,1489]" pageId="12" pageNumber="196">FEMALE. Anterior margin of clypeus slightly slanted medially at 45° angle laterally. Antennal lamellae nearly as long as antennomeres II–VII. Posterior metatibiae with 24–32 spicules.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BF6653CFFFB59778D9E268B9D13223D" box="[189,580,1528,1601]" pageId="12" pageNumber="196" type="distribution">
|
||
<paragraph id="035336B7FFFB59778D9E268B9E01226E" blockId="12.[189,342,1528,1554]" box="[189,342,1528,1554]" pageId="12" pageNumber="196">
|
||
<heading id="581B81DBFFFB59778D9E268B9E01226E" bold="true" box="[189,342,1528,1554]" fontSize="11" level="3" pageId="12" pageNumber="196" reason="3">
|
||
<emphasis id="3198EAA5FFFB59778D9E268B9E01226E" bold="true" box="[189,342,1528,1554]" pageId="12" pageNumber="196">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFB59778D9E25559D13223D" blockId="12.[189,580,1574,1601]" box="[189,580,1574,1601]" pageId="12" pageNumber="196">
|
||
<collectingCountry id="7BFB7627FFFB59778D9E25559E16223C" box="[189,321,1574,1600]" name="Philippines" pageId="12" pageNumber="196">Philippines</collectingCountry>
|
||
(Visayas, Mindanao).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BF6653CFFFB59768D9E251B9E032673" lastPageId="13" lastPageNumber="197" pageId="12" pageNumber="196" type="discussion">
|
||
<paragraph id="035336B7FFFB59778D9E251B9E7822FE" blockId="12.[189,303,1640,1666]" box="[189,303,1640,1666]" pageId="12" pageNumber="196">
|
||
<heading id="581B81DBFFFB59778D9E251B9E7822FE" bold="true" box="[189,303,1640,1666]" fontSize="11" level="3" pageId="12" pageNumber="196" reason="3">
|
||
<emphasis id="3198EAA5FFFB59778D9E251B9E7822FE" bold="true" box="[189,303,1640,1666]" pageId="12" pageNumber="196">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFB59778D9E25E59CCD2341" blockId="12.[188,1399,1686,1854]" pageId="12" pageNumber="196">
|
||
<bibRefCitation id="677D4B46FFFB59778D9E25E59E2622CD" author="Brenske E." box="[189,369,1686,1713]" pageId="12" pageNumber="196" pagination="178 - 205" refId="ref7881" refString="Brenske E. 1896. Neue Melolonthiden aus Africa und Asien. Stettiner entomologische Zeitschrift 57: 178 - 205." type="journal article" year="1896">Brenske (1896)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="C4EC4D34FFFB59778CCE25E49D3F22CC" authorityName="Brenske" authorityYear="1896" box="[493,616,1687,1712]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="196" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFB59778CCE25E49D3F22CC" box="[493,616,1687,1712]" italics="true" pageId="12" pageNumber="196">L. semperi</emphasis>
|
||
</taxonomicName>
|
||
probably from a single male specimen from: “
|
||
<collectingCountry id="7BFB7627FFFB597789A125E59A5822CC" box="[1154,1295,1686,1712]" name="Philippines" pageId="12" pageNumber="196">Philippinen</collectingCountry>
|
||
, Dugang (Semper,
|
||
<date id="77521077FFFB59778C0825C99EBF22A8" box="[299,488,1721,1748]" pageId="12" pageNumber="196" value="1864-07-04" valueMax="1864-07-10" valueMin="1864-07-04">4.-10. Juli 1864</date>
|
||
)”. However, there is no place in the
|
||
<collectingCountry id="7BFB7627FFFB59778EA525C99B5D22A8" box="[902,1034,1722,1748]" name="Philippines" pageId="12" pageNumber="196">Philippines</collectingCountry>
|
||
with the name ʻDugangʼ, it is a Waray word for ʻadditionalʼ, probably it is a case of miscommunication about the locale during that time. Moreover, it is highly probable that the
|
||
<typeStatus id="DC578815FFFB59778FA624729DEF2367" box="[645,696,1793,1819]" pageId="12" pageNumber="196">type</typeStatus>
|
||
locality of the specimen is in the Visayas region particularly, in the
|
||
<collectingRegion id="C128F855FFFB59778C2624579E1F2342" box="[261,328,1828,1854]" country="Philippines" name="Southern Leyte" pageId="12" pageNumber="196">Leyte</collectingRegion>
|
||
area since Waray is the local dialect of the people.
|
||
</paragraph>
|
||
<paragraph id="035336B7FFFB59768D9E24169E032673" blockId="12.[189,1400,1893,2025]" lastBlockId="13.[187,1400,265,527]" lastPageId="13" lastPageNumber="197" pageId="12" pageNumber="196">
|
||
According to primary description, the type series was deposited in the collection of René Oberthür. During and after the Covid-19 pandemic of 2020–2021 several request attempts were made to MNHN for the whereabouts of the type material of
|
||
<taxonomicName id="C4EC4D34FFFB59778FE724DF9C1723B9" authorityName="Brenske" authorityYear="1896" box="[708,832,1964,1989]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="196" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFB59778FE724DF9C1723B9" box="[708,832,1964,1989]" italics="true" pageId="12" pageNumber="196">L. semperi</emphasis>
|
||
</taxonomicName>
|
||
, but curators at MNHN did not find it. No type specimen was found in the collection of Ernst Brenske (now housed at ZMHB,
|
||
<collectingRegion id="C128F855FFFB5977897A24BD9BF22394" box="[1113,1189,1998,2024]" country="Germany" name="Berlin" pageId="12" pageNumber="196">Berlin</collectingRegion>
|
||
,
|
||
<collectingCountry id="7BFB7627FFFB5977899324BD9A742394" box="[1200,1315,1998,2024]" name="Germany" pageId="12" pageNumber="196">Germany</collectingCountry>
|
||
). Thus, it is probable, that original type material of
|
||
<taxonomicName id="C4EC4D34FFFA59768FE222799C6A255F" authorityName="Brenske" authorityYear="1896" box="[705,829,266,291]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFA59768FE222799C6A255F" box="[705,829,266,291]" italics="true" pageId="13" pageNumber="197">L. semperi</emphasis>
|
||
</taxonomicName>
|
||
is lost. To avoid ambiguity about the identity of
|
||
<taxonomicName id="C4EC4D34FFFA59768D9D22589E6E2538" authorityName="Brenske" authorityYear="1896" box="[190,313,299,324]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFA59768D9D22589E6E2538" box="[190,313,299,324]" italics="true" pageId="13" pageNumber="197">L. semperi</emphasis>
|
||
</taxonomicName>
|
||
, a
|
||
<typeStatus id="DC578815FFFA59768C7E22589EED2539" box="[349,442,299,325]" pageId="13" pageNumber="197" type="neotype">neotype</typeStatus>
|
||
specimen is designated in the present paper. Fortunately, in Brenske’s collection,
|
||
<specimenCount id="15EAFD3EFFFA59768D9E223E9E7A251A" box="[189,301,332,359]" pageId="13" pageNumber="197" type="male">one male</specimenCount>
|
||
specimen collected by Charles Baker in Surigao, Mindanao and
|
||
<specimenCount id="15EAFD3EFFFA59768975223F9BB8251A" box="[1110,1263,332,358]" pageId="13" pageNumber="197" type="female">three female</specimenCount>
|
||
specimens from Burauen, Leyte identified as
|
||
<taxonomicName id="C4EC4D34FFFA59768F70221C9D8725F4" authorityName="Brenske" authorityYear="1896" box="[595,720,367,392]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFA59768F70221C9D8725F4" box="[595,720,367,392]" italics="true" pageId="13" pageNumber="197">L. semperi</emphasis>
|
||
</taxonomicName>
|
||
were found. The male (
|
||
<figureCitation id="9BD72A32FFFA59768EC8221D9B1525F5" box="[1003,1090,366,393]" captionStart="Fig" captionStartId="11.[189,232,1927,1953]" captionTargetBox="[189,1398,480,1860]" captionTargetId="figure-86@11.[416,1172,1122,1594]" captionTargetPageId="11" captionText="Fig. 15–16. Leucopholis semperi Brenske, 1896. 15. Neotype, ♂ (MNHN), metaventral process, dorsal aspect. 16. Nontypes, ♀♀, dorsal aspect (photograph of Brenske’s non-type series from ZMHB courtesy of Dr Joachim Willers)." figureDoi="http://doi.org/10.5281/zenodo.8306346" httpUri="https://zenodo.org/record/8306346/files/figure.png" pageId="13" pageNumber="197">Fig. 16</figureCitation>
|
||
) perfectly fits the original description of
|
||
<taxonomicName id="C4EC4D34FFFA59768C4822E39EBF25D5" authorityName="Brenske" authorityYear="1896" box="[363,488,400,425]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="197" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFFA59768C4822E39EBF25D5" box="[363,488,400,425]" italics="true" pageId="13" pageNumber="197">L. semperi</emphasis>
|
||
</taxonomicName>
|
||
and was collected near the original type locality. That is why, this specimen was selected as the
|
||
<typeStatus id="DC578815FFFA59768C8722C19D5225B0" box="[420,517,434,460]" pageId="13" pageNumber="197" type="neotype">neotype</typeStatus>
|
||
. The
|
||
<typeStatus id="DC578815FFFA59768F6722C19DF625B0" box="[580,673,434,460]" pageId="13" pageNumber="197" type="neotype">neotype</typeStatus>
|
||
specimen is currently on loan from ZMHB,
|
||
<collectingRegion id="C128F855FFFA5976898E22C29BAE25B7" box="[1197,1273,433,459]" country="Germany" name="Berlin" pageId="13" pageNumber="197">Berlin</collectingRegion>
|
||
,
|
||
<collectingCountry id="7BFB7627FFFA5976882622C29A2425B7" box="[1285,1395,433,459]" name="Germany" pageId="13" pageNumber="197">Germany</collectingCountry>
|
||
, and will be deposited at Oberthür’s collection at MNHN, Paris,
|
||
<collectingCountry id="7BFB7627FFFA59768EB422A09CB12591" box="[919,998,467,493]" name="France" pageId="13" pageNumber="197">France</collectingCountry>
|
||
for stability and upon consultation with experts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |