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<document id="5E47A75BD3C10BB56B3021CACE551566" ID-CLB-Dataset="54626" ID-DOI="10.37828/em.2021.48.2" ID-GBIF-Dataset="7d008316-40f9-4624-b45c-4e5389a3b631" ID-ISSN="2336-9744" ID-Zenodo-Dep="8029263" ID-ZooBank="urn:lsid:zoobank.org:pub:1B209A4A-6CD7-4B69-9F03-06B613A9148D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1686315596766" checkinUser="felipe" docAuthor="Volynkin, Anton V." docDate="2021" docId="6C1D2B3C7F7CFFC8FF43FAFA9451AD2E" docLanguage="en" docName="EcolMontenegrina.48.9-17.pdf" docOrigin="Ecologica Montenegrina 48" docSource="http://dx.doi.org/10.37828/em.2021.48.2" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.1:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="1" docTitle="Manulea (Tortrilema) nubes Volynkin 2021, sp. n." docType="treatment" docVersion="3" lastPageNumber="14" masterDocId="902453447F7FFFCDFFD4FFC59723A943" masterDocTitle="Tortrilema, a new subgenus of the genus Manulea Wallengren with description of a new species from Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini)" masterLastPageNumber="17" masterPageNumber="9" pageNumber="12" updateTime="1686588900589" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="7A6DE319EE2A561F027F13F3BB078257" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D9765EED7FC936E58577DD188160AB91">
<mods:title id="177F7602155B02779F37A8E7F9F17733">Tortrilema, a new subgenus of the genus Manulea Wallengren with description of a new species from Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini)</mods:title>
</mods:titleInfo>
<mods:name id="4D8C81A5EBF24D641EB589B5F1AA26C6" type="personal">
<mods:role id="63FA6F61C65B8B1985CEF2C73F0AA892">
<mods:roleTerm id="A0242C3B090F400F39AD29D64B60F5B0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DA841CF0275DDA4104FB3C3EE2F1855B">Volynkin, Anton V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="62F52E6E5EA7E831787B4BAEF4F6976B">text</mods:typeOfResource>
<mods:relatedItem id="F8D984E0B6D473FBDB21F5644AC0B90A" type="host">
<mods:titleInfo id="653E73054162BD28C6B72E3623786FFF">
<mods:title id="35472EB69AAE6937429B2CFD104B6A30">Ecologica Montenegrina</mods:title>
</mods:titleInfo>
<mods:part id="CB83AAC5C35D88292A31E42362FAE829">
<mods:date id="7EE5199D8200776FF1131A69FF0C5ACA">2021</mods:date>
<mods:detail id="C68BDF579F60C30E631E6A1964375578" type="pubDate">
<mods:number id="819325237A09A1E2D2D74D5256A70763">2021-11-21</mods:number>
</mods:detail>
<mods:detail id="429CD32C9FFB8F3F44D8E50BA04567C3" type="volume">
<mods:number id="26BC717657A4D1698B375806041C221B">48</mods:number>
</mods:detail>
<mods:extent id="28045347438271A7F0187DD2A37D18EE" unit="page">
<mods:start id="0BDE7C0F961447D502E10987C296066D">9</mods:start>
<mods:end id="05DEE0069DF240C657B88EF510795C05">17</mods:end>
</mods:extent>
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<mods:location id="E7D2FDCDCC3EF3E78D488C5054E302DD">
<mods:url id="402351DCB9BAF1D2B8F919ACAC3D08AC">http://dx.doi.org/10.37828/em.2021.48.2</mods:url>
</mods:location>
<mods:classification id="17DECF8D193FF2E26E0BFECC4891956C">journal article</mods:classification>
<mods:identifier id="19F55EFDF86031ECF04EB70E0488BFDF" type="CLB-Dataset">54626</mods:identifier>
<mods:identifier id="CD845A2D296ACCA4C75F5A84F3A4F7E4" type="DOI">10.37828/em.2021.48.2</mods:identifier>
<mods:identifier id="353726FF3419F868D3A67A7A813CC70C" type="GBIF-Dataset">7d008316-40f9-4624-b45c-4e5389a3b631</mods:identifier>
<mods:identifier id="961A570CB4AB375D3A7EBEFE9E0098FC" type="ISSN">2336-9744</mods:identifier>
<mods:identifier id="6067296FA3327F2FFB417A2EF1FB167C" type="Zenodo-Dep">8029263</mods:identifier>
<mods:identifier id="3526E5F697466FD7C3207554076B5F4C" type="ZooBank">urn:lsid:zoobank.org:pub:1B209A4A-6CD7-4B69-9F03-06B613A9148D</mods:identifier>
</mods:mods>
<treatment id="6C1D2B3C7F7CFFC8FF43FAFA9451AD2E" LSID="urn:lsid:plazi:treatment:6C1D2B3C7F7CFFC8FF43FAFA9451AD2E" httpUri="http://treatment.plazi.org/id/6C1D2B3C7F7CFFC8FF43FAFA9451AD2E" lastPageId="5" lastPageNumber="14" pageId="3" pageNumber="12">
<subSubSection id="ACAEC9A17F7CFFCEFF43FAFA93A7AC34" pageId="3" pageNumber="12" type="nomenclature">
<paragraph id="E40B9A2A7F7CFFCEFF43FAFA9518AC1A" blockId="3.[151,1156,1343,1434]" box="[151,571,1343,1369]" pageId="3" pageNumber="12">
<heading id="BF432D467F7CFFCEFF43FAFA9518AC1A" box="[151,571,1343,1369]" fontSize="11" level="2" pageId="3" pageNumber="12" reason="2">
<emphasis id="D6C046387F7CFFCEFF43FAFA9518AC1A" bold="true" box="[151,571,1343,1369]" pageId="3" pageNumber="12">
<taxonomicName id="23B4E1A97F7CFFCEFF43FAFA96D3AC1A" authority="Volynkin, 2021" authorityName="Volynkin" authorityYear="2021" box="[151,496,1343,1369]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="12" phylum="Tracheophyta" rank="species" species="nubes" status="sp. n." subGenus="Tortrilema">
<emphasis id="D6C046387F7CFFCEFF43FAFA9627AC1A" bold="true" box="[151,260,1343,1369]" italics="true" pageId="3" pageNumber="12">Manulea</emphasis>
(
<emphasis id="D6C046387F7CFFCEFEC2FAFA96BBAC1A" bold="true" box="[278,408,1343,1369]" italics="true" pageId="3" pageNumber="12">Tortrilema</emphasis>
)
<emphasis id="D6C046387F7CFFCEFE7DFAFA96D3AC1A" bold="true" box="[425,496,1343,1369]" italics="true" pageId="3" pageNumber="12">nubes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDF3FB437F7CFFCEFE2CFAFA9518AC1A" box="[504,571,1343,1369]" pageId="3" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E40B9A2A7F7CFFCEFF43FA9A93A7AC34" blockId="3.[151,1156,1343,1434]" box="[151,1156,1375,1399]" pageId="3" pageNumber="12">
http://zoobank.org/
<uri id="902596287F7CFFCEFEB7FA9A93A7AC34" box="[355,1156,1375,1399]" pageId="3" pageNumber="12">
urn:lsid:zoobank.org:act:
<uuid id="9012A0FF7F7CFFCEFDBAFA9A93A7AC34" box="[622,1156,1375,1399]" pageId="3" pageNumber="12">EB8B031D-8FC7-43B5-A090-FCA7FCAB60BB</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="ACAEC9A17F7CFFCEFF43FABA96A5ACD9" box="[151,390,1407,1434]" pageId="3" pageNumber="12" type="description">
<paragraph id="E40B9A2A7F7CFFCEFF43FABA96A5ACD9" blockId="3.[151,1156,1343,1434]" box="[151,390,1407,1434]" pageId="3" pageNumber="12">
(
<figureCitation id="7C8F86AF7F7CFFCEFF4BFABA962AACD9" box="[159,265,1407,1434]" captionStart="Figures 1–8" captionStartId="2.[151,237,1975,1999]" captionTargetBox="[196,1409,317,1933]" captionTargetId="figure-51@2.[151,1436,315,1957]" captionTargetPageId="2" captionText="Figures 1–8. Manulea (Tortrilema subgen. n.): adults. Depositories of the specimens: 1, 2 and 5–8 in MWM/ZSM; 3 and 4 in AFM." figureDoi="http://doi.org/10.5281/zenodo.8029265" httpUri="https://zenodo.org/record/8029265/files/figure.png" pageId="3" pageNumber="12">Figs 1, 2</figureCitation>
,
<figureCitation id="7C8F86AF7F7CFFCEFEC2FABA9670ACD9" box="[278,339,1407,1434]" captionStart="Figures 9–12" captionStartId="4.[151,237,1989,2013]" captionTargetBox="[213,1394,475,1946]" captionTargetId="figure-152@4.[181,1407,471,1970]" captionTargetPageId="4" captionText="Figures 9–12. Manulea (Tortrilema subgen. n.): male genitalia. Depositories of the specimens dissected: 9 and 10 in MWM/ZSM; 11 and 12 in AFM." figureDoi="http://doi.org/10.5281/zenodo.8029267" httpUri="https://zenodo.org/record/8029267/files/figure.png" pageId="3" pageNumber="12">9, 10</figureCitation>
,
<figureCitation id="7C8F86AF7F7CFFCEFE8BFABA965EACD9" box="[351,381,1407,1434]" captionStart="Figures 13–17" captionStartId="6.[151,237,1968,1992]" captionTargetBox="[159,1416,191,1908]" captionTargetId="figure-17@6.[151,1436,189,1928]" captionTargetPageId="6" captionText="Figures 13–17. Manulea (Tortrilema subgen. n.): male (13, 14) and female (15–17) genitalia. The specimens dissected are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.8029269" httpUri="https://zenodo.org/record/8029269/files/figure.png" pageId="3" pageNumber="12">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="ACAEC9A17F7CFFCEFF43FA06950DAF8A" pageId="3" pageNumber="12" type="materials_examined">
<paragraph id="E40B9A2A7F7CFFCEFF43FA069442AF62" blockId="3.[151,1437,1474,2041]" pageId="3" pageNumber="12">
<emphasis id="D6C046387F7CFFCEFF43FA069671AC9E" bold="true" box="[151,338,1475,1501]" pageId="3" pageNumber="12">Type material</emphasis>
.
<materialsCitation id="54DC90777F7CFFCEFEBEFA069442AF62" collectingDate="1995-03-02" collectingDateMax="1995-03-04" collectingDateMin="1995-03-02" collectionCode="R, ZSM, MWM" collectorName="Dr. R. Brechlin" country="Vietnam" county="Nebelwald" elevation="2400" location="Nord" municipality="Cha-pa" pageId="3" pageNumber="12" specimenCount="1" typeStatus="holotype">
<emphasis id="D6C046387F7CFFCEFEBEFA0696FFAC9E" bold="true" box="[362,476,1475,1501]" pageId="3" pageNumber="12">
<typeStatus id="3B0F24887F7CFFCEFEBEFA0696FFAC9E" box="[362,476,1475,1501]" pageId="3" pageNumber="12">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="7C8F86AF7F7CFFCEFE22FA079569AC9E" box="[502,586,1474,1501]" captionStart="Figures 1–8" captionStartId="2.[151,237,1975,1999]" captionTargetBox="[196,1409,317,1933]" captionTargetId="figure-51@2.[151,1436,315,1957]" captionTargetPageId="2" captionText="Figures 1–8. Manulea (Tortrilema subgen. n.): adults. Depositories of the specimens: 1, 2 and 5–8 in MWM/ZSM; 3 and 4 in AFM." figureDoi="http://doi.org/10.5281/zenodo.8029265" httpUri="https://zenodo.org/record/8029265/files/figure.png" pageId="3" pageNumber="12">Figs 1</figureCitation>
,
<figureCitation id="7C8F86AF7F7CFFCEFDB6FA079557AC9E" box="[610,628,1474,1501]" captionStart="Figures 9–12" captionStartId="4.[151,237,1989,2013]" captionTargetBox="[213,1394,475,1946]" captionTargetId="figure-152@4.[181,1407,471,1970]" captionTargetPageId="4" captionText="Figures 9–12. Manulea (Tortrilema subgen. n.): male genitalia. Depositories of the specimens dissected: 9 and 10 in MWM/ZSM; 11 and 12 in AFM." figureDoi="http://doi.org/10.5281/zenodo.8029267" httpUri="https://zenodo.org/record/8029267/files/figure.png" pageId="3" pageNumber="12">9</figureCitation>
): male, “N-
<collectingCountry id="9CA3DABA7F7CFFCEFCC5FA079455AC9E" box="[785,886,1474,1501]" name="Vietnam" pageId="3" pageNumber="12">Vietnam</collectingCountry>
/ Tonkin, Mt. Fan-si-pan (
<location id="E16BCCF17F7CFFCEFB34FA079201AC9E" LSID="urn:lsid:plazi:treatment:6C1D2B3C7F7CFFC8FF43FAFA9451AD2E:E16BCCF17F7CFFCEFB34FA079201AC9E" box="[1248,1314,1474,1501]" country="Vietnam" county="Nebelwald" municipality="Cha-pa" name="Nord" pageId="3" pageNumber="12">Nord</location>
),
<collectingMunicipality id="046F00507F7CFFCEFA94FA0792BBAC9E" box="[1344,1432,1474,1501]" pageId="3" pageNumber="12">Cha-pa</collectingMunicipality>
,
<collectingCounty id="0D6AE2A67F7CFFCEFF43FA219635ACBC" box="[151,278,1508,1535]" pageId="3" pageNumber="12">Nebelwald</collectingCounty>
[cloud forest] (22.15ˊN 103.45ˊE)
<date id="900ABCEA7F7CFFCEFD11FA219441ACBC" box="[709,866,1508,1535]" pageId="3" pageNumber="12" value="1995-03-02" valueMax="1995-03-04" valueMin="1995-03-02">
<collectingDate id="804E45027F7CFFCEFD11FA219441ACBC" box="[709,866,1508,1535]" pageId="3" pageNumber="12" value="1995-03-02" valueMax="1995-03-04" valueMin="1995-03-02">2.–4.III.1995</collectingDate>
</date>
,
<quantity id="234C37CF7F7CFFCEFCA5FA2194E5ACBC" box="[881,966,1508,1535]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="12" unit="m" value="2400.0">
<elevation id="6F997D197F7CFFCEFCA5FA2194E5ACBC" box="[881,966,1508,1535]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="12" unit="m" value="2400.0">2400m</elevation>
</quantity>
, leg.
<collectorName id="4941FFFC7F7CFFCEFBDFFA209311ACBC" box="[1035,1074,1509,1535]" pageId="3" pageNumber="12">Dr.</collectorName>
<collectionCode id="82A502EF7F7CFFCEFBE8FA209370ACBC" box="[1084,1107,1509,1535]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="12">R</collectionCode>
. Brechlin”, gen. prep.
<collectorName id="4941FFFC7F7CFFCEFABDFA2092BBACBC" box="[1385,1432,1509,1535]" pageId="3" pageNumber="12">No.</collectorName>
:
<collectionCode id="82A502EF7F7CFFCEFF43F9C397F0AF62" box="[151,211,1542,1569]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="12" type="Museum">ZSM</collectionCode>
<collectorName id="4941FFFC7F7CFFCEFF0FF9C39634AF62" box="[219,279,1542,1569]" pageId="3" pageNumber="12">Arct.</collectorName>
2021-020 (prepared by
<collectorName id="4941FFFC7F7CFFCEFDE6F9C39580AF62" box="[562,675,1542,1569]" pageId="3" pageNumber="12">Volynkin</collectorName>
) (
<collectionCode id="82A502EF7F7CFFCEFD6EF9C3942DAF62" box="[698,782,1542,1569]" pageId="3" pageNumber="12">MWM</collectionCode>
/
<collectionCode id="82A502EF7F7CFFCEFCC3F9C39475AF62" box="[791,854,1542,1569]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="12" type="Museum">ZSM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="E40B9A2A7F7CFFCEFF22F9ED950DAF8A" blockId="3.[151,1437,1474,2041]" pageId="3" pageNumber="12">
<materialsCitation id="54DC90777F7CFFCEFF22F9ED92B8AF01" box="[246,1435,1575,1602]" collectionCode="ZSM" collectorName="Nos." country="Vietnam" pageId="3" pageNumber="12" specimenCount="17" specimenCount-female="9" specimenCount-male="8" typeStatus="paratype">
<emphasis id="D6C046387F7CFFCEFF22F9ED9657AF01" bold="true" box="[246,372,1576,1602]" pageId="3" pageNumber="12">
<typeStatus id="3B0F24887F7CFFCEFF22F9ED9657AF01" box="[246,372,1576,1602]" pageId="3" pageNumber="12">Paratypes</typeStatus>
</emphasis>
.
<emphasis id="D6C046387F7CFFCEFE57F9ED9532AF01" bold="true" box="[387,529,1576,1602]" pageId="3" pageNumber="12">
<collectingCountry id="9CA3DABA7F7CFFCEFE57F9ED9532AF01" box="[387,529,1576,1602]" name="Vietnam" pageId="3" pageNumber="12">VIETNAM</collectingCountry>
</emphasis>
:
<specimenCount id="F2B251A37F7CFFCEFDF5F9E2955CAF01" box="[545,639,1575,1602]" pageId="3" pageNumber="12" type="male">8 males</specimenCount>
,
<specimenCount id="F2B251A37F7CFFCEFD58F9E29422AF01" box="[652,769,1575,1602]" pageId="3" pageNumber="12" type="female">9 females</specimenCount>
, the same data as in the holotype, gen. prep.
<collectorName id="4941FFFC7F7CFFCEFACCF9ED9270AF01" box="[1304,1363,1576,1602]" pageId="3" pageNumber="12">Nos.</collectorName>
:
<collectionCode id="82A502EF7F7CFFCEFA8BF9E292B8AF01" box="[1375,1435,1575,1602]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="12" type="Museum">ZSM</collectionCode>
</materialsCitation>
<materialsCitation id="54DC90777F7CFFCEFF43F98C9369AF27" box="[151,1098,1609,1636]" collectionCode="ZSM" collectorName="Volynkin" country="Vietnam" location="Arct." pageId="3" pageNumber="12" specimenCount="1" typeStatus="paratype">
<location id="E16BCCF17F7CFFCEFF43F98C97F0AF27" LSID="urn:lsid:plazi:treatment:6C1D2B3C7F7CFFC8FF43FAFA9451AD2E:E16BCCF17F7CFFCEFF43F98C97F0AF27" box="[151,211,1609,1636]" country="Vietnam" name="Arct." pageId="3" pageNumber="12">Arct.</location>
2021-143 (male),
<collectionCode id="82A502EF7F7CFFCEFE62F98C96D1AF27" box="[438,498,1609,1636]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="12" type="Museum">ZSM</collectionCode>
<location id="E16BCCF17F7CFFCEFE29F98C951AAF27" LSID="urn:lsid:plazi:treatment:6C1D2B3C7F7CFFC8FF43FAFA9451AD2E:E16BCCF17F7CFFCEFE29F98C951AAF27" box="[509,569,1609,1636]" country="Vietnam" name="Arct." pageId="3" pageNumber="12">Arct.</location>
2021-144 (female) (prepared by
<collectorName id="4941FFFC7F7CFFCEFC1AF98C9361AF27" box="[974,1090,1609,1636]" pageId="3" pageNumber="12">Volynkin</collectorName>
)
</materialsCitation>
;
<materialsCitation id="54DC90777F7CFFCEFB8DF98C9654AFEB" collectingDate="1995-03-25" collectingDateMax="1995-03-30" collectingDateMin="1995-03-25" collectionCode="N" collectorName="Cha-pa" country="Vietnam" elevation="1600" location="Mt." pageId="3" pageNumber="12" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="F2B251A37F7CFFCEFB8DF98C939AAF27" box="[1113,1209,1609,1636]" pageId="3" pageNumber="12" type="male">3 males</specimenCount>
,
<specimenCount id="F2B251A37F7CFFCEFB1DF98C9216AF27" box="[1225,1333,1609,1636]" pageId="3" pageNumber="12" type="female">1 female</specimenCount>
,
<quantity id="234C37CF7F7CFFCEFA97F98C92B4AF27" box="[1347,1431,1609,1636]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="12" unit="m" value="1600.0">
<elevation id="6F997D197F7CFFCEFA97F98C92B4AF27" box="[1347,1431,1609,1636]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="12" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<location id="E16BCCF17F7CFFCEFF43F9A997E2AFC5" LSID="urn:lsid:plazi:treatment:6C1D2B3C7F7CFFC8FF43FAFA9451AD2E:E16BCCF17F7CFFCEFF43F9A997E2AFC5" box="[151,193,1644,1670]" country="Vietnam" name="Mt." pageId="3" pageNumber="12">Mt.</location>
Fan-si-pan (north),
<collectorName id="4941FFFC7F7CFFCEFE10F9AE953AAFC5" box="[452,537,1643,1670]" pageId="3" pageNumber="12">Cha-pa</collectorName>
, primary forest, 22°17''
<collectionCode id="82A502EF7F7CFFCEFC92F9AE947DAFC5" box="[838,862,1643,1670]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="12" type="Herbarium">N</collectionCode>
, 103°44''
<collectionCode id="82A502EF7F7CFFCEFC06F9AE94C5AFC5" box="[978,998,1643,1670]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="12" type="Herbarium">E</collectionCode>
,
<date id="900ABCEA7F7CFFCEFC2DF9AE938FAFC5" box="[1017,1196,1643,1670]" pageId="3" pageNumber="12" value="1995-03-25" valueMax="1995-03-30" valueMin="1995-03-25">
<collectingDate id="804E45027F7CFFCEFC2DF9AE938FAFC5" box="[1017,1196,1643,1670]" pageId="3" pageNumber="12" value="1995-03-25" valueMax="1995-03-30" valueMin="1995-03-25">25–30.III.1995</collectingDate>
</date>
,
<collectionCode id="82A502EF7F7CFFCEFB14F9A993FAAFC5" box="[1216,1241,1644,1670]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="12" type="Museum">V</collectionCode>
.
<location id="E16BCCF17F7CFFCEFB38F9AE9268AFC5" LSID="urn:lsid:plazi:treatment:6C1D2B3C7F7CFFC8FF43FAFA9451AD2E:E16BCCF17F7CFFCEFB38F9AE9268AFC5" box="[1260,1355,1643,1670]" country="Vietnam" name="Sinyaev" pageId="3" pageNumber="12">Sinyaev</location>
&amp;
<collectionCode id="82A502EF7F7CFFCEFA54F9AE92BAAFC5" box="[1408,1433,1643,1670]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="12" type="Herbarium">A</collectionCode>
.
<collectorName id="4941FFFC7F7CFFCEFF43F9489662AFEB" box="[151,321,1677,1704]" pageId="3" pageNumber="12">Schintlmeister</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="54DC90777F7CFFCEFE51F9489610AF8A" collectingDate="1993-05-08" collectingDateMax="1993-05-29" collectingDateMin="1993-05-08" collectionCode="N" collectorName="Sinyaev &amp; Simonov" country="Vietnam" elevation="2400" location="Mt." municipality="Cha" pageId="3" pageNumber="12" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="F2B251A37F7CFFCEFE51F94896CCAFEB" box="[389,495,1677,1704]" pageId="3" pageNumber="12" type="female">1 female</specimenCount>
,
<location id="E16BCCF17F7CFFCEFE2FF9489506AFE4" LSID="urn:lsid:plazi:treatment:6C1D2B3C7F7CFFC8FF43FAFA9451AD2E:E16BCCF17F7CFFCEFE2FF9489506AFE4" box="[507,549,1677,1703]" country="Vietnam" municipality="Cha" name="Mt." pageId="3" pageNumber="12">Mt.</location>
Fan-si-pan,
<collectingMunicipality id="046F00507F7CFFCEFD6EF94895CAAFEB" box="[698,745,1677,1704]" pageId="3" pageNumber="12">Cha</collectingMunicipality>
pa,
<quantity id="234C37CF7F7CFFCEFCC9F9489451AFEB" box="[797,882,1677,1704]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="12" unit="m" value="2400.0">
<elevation id="6F997D197F7CFFCEFCC9F9489451AFEB" box="[797,882,1677,1704]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="12" unit="m" value="2400.0">2400m</elevation>
</quantity>
, 22°15''
<collectionCode id="82A502EF7F7CFFCEFC1BF94894C4AFEB" box="[975,999,1677,1704]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="12" type="Herbarium">N</collectionCode>
, 103°46''
<collectionCode id="82A502EF7F7CFFCEFB81F948934AAFEB" box="[1109,1129,1677,1704]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="12" type="Herbarium">E</collectionCode>
,
<date id="900ABCEA7F7CFFCEFBA1F9489231AFEB" box="[1141,1298,1677,1704]" pageId="3" pageNumber="12" value="1993-05-08" valueMax="1993-05-29" valueMin="1993-05-08">
<collectingDate id="804E45027F7CFFCEFBA1F9489231AFEB" box="[1141,1298,1677,1704]" pageId="3" pageNumber="12" value="1993-05-08" valueMax="1993-05-29" valueMin="1993-05-08">
8–29.
<collectionCode id="82A502EF7F7CFFCEFB63F94893ECAFEB" box="[1207,1231,1677,1704]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="12" type="Museum">V</collectionCode>
.1993
</collectingDate>
</date>
,
<collectorName id="4941FFFC7F7CFFCEFACBF948925EAFEB" box="[1311,1405,1677,1704]" pageId="3" pageNumber="12">Sinyaev</collectorName>
&amp;
<collectorName id="4941FFFC7F7CFFCEFF43F96B9622AF8A" box="[151,257,1710,1737]" pageId="3" pageNumber="12">Simonov</collectorName>
leg.
</materialsCitation>
(all in MWM/
<collectionCode id="82A502EF7F7CFFCEFE30F96B9501AF8A" box="[484,546,1710,1737]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="12" type="Museum">ZSM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="ACAEC9A17F7CFFC9FF22F915946EA889" lastPageId="4" lastPageNumber="13" pageId="3" pageNumber="12" type="diagnosis">
<paragraph id="E40B9A2A7F7CFFC9FF22F915946EA889" blockId="3.[151,1437,1474,2041]" lastBlockId="4.[151,1436,195,458]" lastPageId="4" lastPageNumber="13" pageId="3" pageNumber="12">
<emphasis id="D6C046387F7CFFCEFF22F915964CAFA9" bold="true" box="[246,367,1744,1770]" pageId="3" pageNumber="12">Diagnosis</emphasis>
. The new species is externally very similar to
<taxonomicName id="23B4E1A97F7CFFCEFC02F9149376AFA8" authorityName="Volynkin &amp; Saldaitis" authorityYear="2021" box="[982,1109,1744,1771]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="12" phylum="Tracheophyta" rank="species" species="labahe">
<emphasis id="D6C046387F7CFFCEFC02F9149376AFA8" box="[982,1109,1744,1771]" italics="true" pageId="3" pageNumber="12">M. labahe</emphasis>
</taxonomicName>
known from the
<collectingRegion id="267054C87F7CFFCEFAEAF9159623AE4E" country="China" name="Sichuan" pageId="3" pageNumber="12">Sichuan Province</collectingRegion>
of
<collectingCountry id="9CA3DABA7F7CFFCEFEF8F9379651AE4E" box="[300,370,1778,1805]" name="China" pageId="3" pageNumber="12">China</collectingCountry>
but differs in the somewhat smaller size (the male forewing length is
<quantity id="234C37CF7F7CFFCEFB1CF937923FAE4E" box="[1224,1308,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="12" unit="mm" value="17.0">17 mm</quantity>
vs.
<quantity id="234C37CF7F7CFFCEFA85F93797E7AE6C" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="3" pageNumber="12" unit="mm" value="18.5" valueMax="19.0" valueMin="18.0">18–19 mm</quantity>
in
<taxonomicName id="23B4E1A97F7CFFCEFF3FF8D09640AE6C" authorityName="Volynkin &amp; Saldaitis" authorityYear="2021" box="[235,355,1812,1839]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="12" phylum="Tracheophyta" rank="species" species="labahe">
<emphasis id="D6C046387F7CFFCEFF3FF8D09640AE6C" box="[235,355,1812,1839]" italics="true" pageId="3" pageNumber="12">M. labahe</emphasis>
</taxonomicName>
). In the male genital capsule of
<taxonomicName id="23B4E1A97F7CFFCEFD34F8D09468AE6C" authority="Volynkin, 2021" authorityName="Volynkin" authorityYear="2021" box="[736,843,1812,1839]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="12" phylum="Tracheophyta" rank="species" species="nubes" status="sp. n.">
<emphasis id="D6C046387F7CFFCEFD34F8D09468AE6C" box="[736,843,1812,1839]" italics="true" pageId="3" pageNumber="12">M. nubes</emphasis>
</taxonomicName>
<emphasis id="D6C046387F7CFFCEFC87F8D194B5AE6D" bold="true" box="[851,918,1812,1838]" pageId="3" pageNumber="12">
<taxonomicNameLabel id="CDF3FB437F7CFFCEFC87F8D194B5AE6D" box="[851,918,1812,1838]" pageId="3" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, the uncus is broader than in
<taxonomicName id="23B4E1A97F7CFFCEFB26F8D09249AE6C" authorityName="Volynkin &amp; Saldaitis" authorityYear="2021" box="[1266,1386,1812,1839]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="12" phylum="Tracheophyta" rank="species" species="labahe">
<emphasis id="D6C046387F7CFFCEFB26F8D09249AE6C" box="[1266,1386,1812,1839]" italics="true" pageId="3" pageNumber="12">M. labahe</emphasis>
</taxonomicName>
, the valva is shorter and somewhat broader (the length to width ratio is 2.35–2.65 vs. 2.80–3.05 in
<taxonomicName id="23B4E1A97F7CFFCEFAC6F8F392AFAE13" authorityName="Volynkin &amp; Saldaitis" authorityYear="2021" box="[1298,1420,1845,1872]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="12" phylum="Tracheophyta" rank="species" species="labahe">
<emphasis id="D6C046387F7CFFCEFAC6F8F392AFAE13" box="[1298,1420,1845,1872]" italics="true" pageId="3" pageNumber="12">M. labahe</emphasis>
</taxonomicName>
), and the distal saccular process is shorter and has a more angular dorsal basal protrusion and a more curved distal section. The aedeagus of the new species is somewhat narrower distally than in
<taxonomicName id="23B4E1A97F7CFFCEFB5FF8BF9221AED7" authorityName="Volynkin &amp; Saldaitis" authorityYear="2021" box="[1163,1282,1913,1940]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="12" phylum="Tracheophyta" rank="species" species="labahe">
<emphasis id="D6C046387F7CFFCEFB5FF8BF9221AED7" box="[1163,1282,1913,1940]" italics="true" pageId="3" pageNumber="12">M. labahe</emphasis>
</taxonomicName>
. The vesicae of the two species display no remarkable differences. Since the female of
<taxonomicName id="23B4E1A97F7CFFCEFB9DF85993E4AEF6" authorityName="Volynkin &amp; Saldaitis" authorityYear="2021" box="[1097,1223,1946,1973]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="12" phylum="Tracheophyta" rank="species" species="labahe">
<emphasis id="D6C046387F7CFFCEFB9DF85993E4AEF6" box="[1097,1223,1946,1973]" italics="true" pageId="3" pageNumber="12">M. labahe</emphasis>
</taxonomicName>
is unknown, the copulatory organ of the new species is compared to
<taxonomicName id="23B4E1A97F7CFFCEFCDBF87894BCAE94" authorityName="Volynkin &amp; Saldaitis" authorityYear="2021" box="[783,927,1980,2007]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="12" phylum="Tracheophyta" rank="species" species="lienquan">
<emphasis id="D6C046387F7CFFCEFCDBF87894BCAE94" box="[783,927,1980,2007]" italics="true" pageId="3" pageNumber="12">M. lienquan</emphasis>
</taxonomicName>
and
<taxonomicName id="23B4E1A97F7CFFCEFC09F8789384AE94" authorityName="Matsumura" authorityYear="1927" box="[989,1191,1980,2007]" class="Insecta" family="Arctiidae" genus="Lithosia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="postmaculosa">
<emphasis id="D6C046387F7CFFCEFC09F8789384AE94" box="[989,1191,1980,2007]" italics="true" pageId="3" pageNumber="12">M. postmaculosa</emphasis>
</taxonomicName>
instead. The female genitalia of
<taxonomicName id="23B4E1A97F7CFFCEFEF9F81A96BDAEBA" authority="Volynkin, 2021" authorityName="Volynkin" authorityYear="2021" box="[301,414,2014,2041]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="12" phylum="Tracheophyta" rank="species" species="nubes" status="sp. n.">
<emphasis id="D6C046387F7CFFCEFEF9F81A96BDAEBA" box="[301,414,2014,2041]" italics="true" pageId="3" pageNumber="12">M. nubes</emphasis>
</taxonomicName>
<emphasis id="D6C046387F7CFFCEFE7FF81B96D1AEBB" bold="true" box="[427,498,2014,2040]" pageId="3" pageNumber="12">
<taxonomicNameLabel id="CDF3FB437F7CFFCEFE7FF81B96D1AEBB" box="[427,498,2014,2040]" pageId="3" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are characterised by the narrow, curved, well-sclerotised and longitudinally rugose ductus bursae whereas the ductus bursae of
<taxonomicName id="23B4E1A97F7BFFC9FD2BFF0094ACA99D" authorityName="Volynkin &amp; Saldaitis" authorityYear="2021" box="[767,911,195,222]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="13" phylum="Tracheophyta" rank="species" species="lienquan">
<emphasis id="D6C046387F7BFFC9FD2BFF0094ACA99D" box="[767,911,195,222]" italics="true" pageId="4" pageNumber="13">M. lienquan</emphasis>
</taxonomicName>
is broad, membranous and less curved, and that of
<taxonomicName id="23B4E1A97F7BFFC9FF3AFF23969AA843" authorityName="Matsumura" authorityYear="1927" box="[238,441,229,256]" class="Insecta" family="Arctiidae" genus="Lithosia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="13" phylum="Arthropoda" rank="species" species="postmaculosa">
<emphasis id="D6C046387F7BFFC9FF3AFF23969AA843" box="[238,441,229,256]" italics="true" pageId="4" pageNumber="13">M. postmaculosa</emphasis>
</taxonomicName>
is broader, almost straight, dorso-ventrally flattened and smoothly sclerotised. The postvaginal plate of the new species is markedly shorter and narrower than in the congeners. The corpus bursae of
<taxonomicName id="23B4E1A97F7BFFC9FEDAFEEF9658A800" authority="Volynkin, 2021" authorityName="Volynkin" authorityYear="2021" box="[270,379,296,323]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="13" phylum="Tracheophyta" rank="species" species="nubes" status="sp. n.">
<emphasis id="D6C046387F7BFFC9FEDAFEEF9658A800" box="[270,379,296,323]" italics="true" pageId="4" pageNumber="13">M. nubes</emphasis>
</taxonomicName>
<emphasis id="D6C046387F7BFFC9FE50FEEC96EBA800" bold="true" box="[388,456,297,323]" pageId="4" pageNumber="13">
<taxonomicNameLabel id="CDF3FB437F7BFFC9FE50FEEC96EBA800" box="[388,456,297,323]" pageId="4" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is teardrop-shaped, membranous and having a long, band-shaped signum whereas the corpus bursae of the congeners is sack-like, more or less gelatinous and lacks a signum. Additionally, the appendix bursae of the new species is very short, membranous and directed posteriorly whereas the appendix bursae of
<taxonomicName id="23B4E1A97F7BFFC9FED9FE4A96BEA8EA" authorityName="Volynkin &amp; Saldaitis" authorityYear="2021" box="[269,413,398,425]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="13" phylum="Tracheophyta" rank="species" species="lienquan">
<emphasis id="D6C046387F7BFFC9FED9FE4A96BEA8EA" box="[269,413,398,425]" italics="true" pageId="4" pageNumber="13">M. lienquan</emphasis>
</taxonomicName>
is gelatinous and directed postero-laterally, and that of
<taxonomicName id="23B4E1A97F7BFFC9FBE2FE4A93DCA8EA" authorityName="Matsumura" authorityYear="1927" box="[1078,1279,398,425]" class="Insecta" family="Arctiidae" genus="Lithosia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="13" phylum="Arthropoda" rank="species" species="postmaculosa">
<emphasis id="D6C046387F7BFFC9FBE2FE4A93DCA8EA" box="[1078,1279,398,425]" italics="true" pageId="4" pageNumber="13">M. postmaculosa</emphasis>
</taxonomicName>
is gelatinous with a heavily sclerotised inner plate and directed laterally.
</paragraph>
</subSubSection>
<caption id="B0CBCAA27F7BFFC9FF43F80096DBAEB8" ID-DOI="http://doi.org/10.5281/zenodo.8029267" ID-Zenodo-Dep="8029267" httpUri="https://zenodo.org/record/8029267/files/figure.png" pageId="4" pageNumber="13" startId="4.[151,237,1989,2013]" targetBox="[213,1394,475,1946]" targetPageId="4" targetType="figure">
<paragraph id="E40B9A2A7F7BFFC9FF43F80096DBAEB8" blockId="4.[151,1436,1989,2043]" pageId="4" pageNumber="13">
<emphasis id="D6C046387F7BFFC9FF43F8009608AE9E" bold="true" box="[151,299,1989,2013]" pageId="4" pageNumber="13">Figures 9–12</emphasis>
.
<taxonomicName id="23B4E1A97F7BFFC9FEEFF80096B9AE9E" box="[315,410,1989,2013]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="13" phylum="Tracheophyta" rank="genus">
<emphasis id="D6C046387F7BFFC9FEEFF80096B9AE9E" box="[315,410,1989,2013]" italics="true" pageId="4" pageNumber="13">Manulea</emphasis>
</taxonomicName>
(
<taxonomicName id="23B4E1A97F7BFFC9FE7FF800953DAE9E" authority="Volynkin, 2021" authorityName="Volynkin" authorityYear="2021" box="[427,542,1989,2013]" class="Magnoliopsida" family="Scrophulariaceae" genus="Manulea" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="13" phylum="Tracheophyta" rank="subGenus" status="subgen. n." subGenus="Tortrilema">
<emphasis id="D6C046387F7BFFC9FE7FF800953DAE9E" box="[427,542,1989,2013]" italics="true" pageId="4" pageNumber="13">Tortrilema</emphasis>
</taxonomicName>
<emphasis id="D6C046387F7BFFC9FDF3F80095BFAE9E" bold="true" box="[551,668,1989,2013]" pageId="4" pageNumber="13">
<taxonomicNameLabel id="CDF3FB437F7BFFC9FDF3F80095BFAE9E" box="[551,668,1989,2013]" pageId="4" pageNumber="13" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
): male genitalia. Depositories of the specimens dissected: 9 and 10 in MWM/ZSM; 11 and 12 in AFM.
</paragraph>
</caption>
<subSubSection id="ACAEC9A17F7AFFC8FF22FF0196A7AAA5" pageId="5" pageNumber="14" type="description">
<paragraph id="E40B9A2A7F7AFFC8FF22FF0196A7AAA5" blockId="5.[151,1437,195,1133]" pageId="5" pageNumber="14">
<emphasis id="D6C046387F7AFFC8FF22FF0196A4A99D" bold="true" box="[246,391,196,222]" pageId="5" pageNumber="14">Description</emphasis>
.
<emphasis id="D6C046387F7AFFC8FE42FF0196CAA99D" bold="true" box="[406,489,196,222]" pageId="5" pageNumber="14">Adults</emphasis>
(
<figureCitation id="7C8F86AF7F7AFFC8FE2DFF069547A99D" box="[505,612,195,222]" captionStart="Figures 1–8" captionStartId="2.[151,237,1975,1999]" captionTargetBox="[196,1409,317,1933]" captionTargetId="figure-51@2.[151,1436,315,1957]" captionTargetPageId="2" captionText="Figures 1–8. Manulea (Tortrilema subgen. n.): adults. Depositories of the specimens: 1, 2 and 5–8 in MWM/ZSM; 3 and 4 in AFM." figureDoi="http://doi.org/10.5281/zenodo.8029265" httpUri="https://zenodo.org/record/8029265/files/figure.png" pageId="5" pageNumber="14">Figs 1, 2</figureCitation>
). Forewing length 17.0 mm in males and
<quantity id="234C37CF7F7AFFC8FB84FF0693ECA99D" box="[1104,1231,195,222]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="5" pageNumber="14" unit="mm" value="18.5" valueMax="19.0" valueMin="18.0">18–19 mm</quantity>
in females. Male antenna ciliate, female antenna filiform. Head and thorax brown. Forewing elongate with postmedially strongly convex costal margin and elongate apex. Forewing ground colour brown. Antemedial and postmedial transverse line wide, dark greyish-brown, with sinuous margins, diffuse. Discal spot dark brown, small, round. Cilia brown with admixture of grey scales. Hindwing monotonous pale ochreous. Abdomen ochreous-brown with admixture of ochreous-yellow scales latero-anteriorly.
<emphasis id="D6C046387F7AFFC8FBEBFEA993D4A8C5" bold="true" box="[1087,1271,364,390]" pageId="5" pageNumber="14">Male genitalia</emphasis>
(
<figureCitation id="7C8F86AF7F7AFFC8FADFFEA992ACA8C4" box="[1291,1423,364,391]" captionStart="Figures 9–12" captionStartId="4.[151,237,1989,2013]" captionTargetBox="[213,1394,475,1946]" captionTargetId="figure-152@4.[181,1407,471,1970]" captionTargetPageId="4" captionText="Figures 9–12. Manulea (Tortrilema subgen. n.): male genitalia. Depositories of the specimens dissected: 9 and 10 in MWM/ZSM; 11 and 12 in AFM." figureDoi="http://doi.org/10.5281/zenodo.8029267" httpUri="https://zenodo.org/record/8029267/files/figure.png" pageId="5" pageNumber="14">Figs 9, 10</figureCitation>
). Uncus strongly dilated distally but tapered apically, with small claw-like tip and tiny triangular subapical protrusion ventrally. Tuba analis membranous, broad, equal in width to tegumen length; scaphium very thin, weakly sclerotised. Tegumen short but broad, with arms fused in distal two thirds. Vinculum elongate, equal in length to tegumen, with thin arms connected anteriorly by membranous commissure. Intravincular area with cluster of androconial hair-like scales. Valva with almost parallel margins and broadly lobular and apically rounded dorsal section. Costa thin. Sacculus relatively narrow basally and constricted medially. Distal saccular process with broad basal section forming triangular dorsal protrusion, and long and narrow distal section hook-like curved dorsally. Juxta weakly sclerotised, more or less trapezoid with short medial protrusion posteriorly. Aedeagus elongate and narrow, with short and rounded coecum, somewhat constricted subapically and weakly scobinated apically. Vesica short and ca. 2 times broader than aedeagus, with short bilobate diverticulum laterally and broad semiglobular ventral diverticulum bearing long, narrow and almost straight cornutus. Vesica ejaculatorius weakly scobinated basally, projected distally.
<emphasis id="D6C046387F7AFFC8FA96FCC49620AA7E" bold="true" pageId="5" pageNumber="14">Female genitalia</emphasis>
(
<figureCitation id="7C8F86AF7F7AFFC8FEC2FCE7964CAA7E" box="[278,367,802,829]" captionStart="Figures 13–17" captionStartId="6.[151,237,1968,1992]" captionTargetBox="[159,1416,191,1908]" captionTargetId="figure-17@6.[151,1436,189,1928]" captionTargetPageId="6" captionText="Figures 13–17. Manulea (Tortrilema subgen. n.): male (13, 14) and female (15–17) genitalia. The specimens dissected are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.8029269" httpUri="https://zenodo.org/record/8029269/files/figure.png" pageId="5" pageNumber="14">Fig. 15</figureCitation>
). Papilla analis broad, trapezoid with rounded corners, setose. Apophyses long and thin, apophysis anterioris somewhat longer than apophysis posterioris. Postvaginal plate short, more or less trapezoid. Ductus bursae narrow, C-like curved anteriorly, heavily sclerotised and longitudinally rugose. Corpus bursae broad, teardrop-shaped, membranous with long and narrow band-like longitudinal signum somewhat dilated anteriorly. Appendix bursae very short, conical, membranous, positioned postero-dorsally, directed posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="ACAEC9A17F7AFFC8FF22FC289632AD69" pageId="5" pageNumber="14" type="distribution">
<paragraph id="E40B9A2A7F7AFFC8FF22FC289632AD69" blockId="5.[151,1437,195,1133]" pageId="5" pageNumber="14">
<emphasis id="D6C046387F7AFFC8FF22FC2896B3AD44" bold="true" box="[246,400,1005,1031]" pageId="5" pageNumber="14">Distribution</emphasis>
. The new species is known only from its
<typeStatus id="3B0F24887F7AFFC8FC72FC2B94FBAD4B" box="[934,984,1006,1032]" pageId="5" pageNumber="14">type</typeStatus>
locality in North
<collectingCountry id="9CA3DABA7F7AFFC8FB69FC289201AD4B" box="[1213,1314,1005,1032]" name="Vietnam" pageId="5" pageNumber="14">Vietnam</collectingCountry>
(
<collectingRegion id="267054C87F7AFFC8FAE3FC289626AD69" country="Vietnam" name="Lao Cai" pageId="5" pageNumber="14">Lào Cai Province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="ACAEC9A17F7AFFC8FF22FBF49451AD2E" pageId="5" pageNumber="14" type="etymology">
<paragraph id="E40B9A2A7F7AFFC8FF22FBF49451AD2E" blockId="5.[151,1437,195,1133]" pageId="5" pageNumber="14">
<emphasis id="D6C046387F7AFFC8FF22FBF4965DAD08" bold="true" box="[246,382,1073,1099]" pageId="5" pageNumber="14">Etymology</emphasis>
. ‘Nubes’ means ‘a cloud’ in Latin. The specific epithet refers to the cloud forest habitats of the Mount Fan-si-pan where the new species was collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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