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<document id="CC97E8E8E021745A2BD768ADECEEED4A" ID-CLB-Dataset="36890" ID-DOI="10.11646/zootaxa.3772.1.1" ID-GBIF-Dataset="12787306-9151-40cd-8103-1ecac6bdb6b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="285653" ID-ZooBank="1A1578DD-4B10-4F70-8CB6-03B0ED07AB68" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460391597516" checkinUser="plazi" docAuthor="Sartori, André F., Gargominy, Olivier &amp; Fontaine, Benoît" docDate="2014" docId="222F879AFFF4FFEF578BFB48FC4AFDC3" docLanguage="en" docName="zt03772p068.pdf" docOrigin="Zootaxa 3772 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pseudolibera lillianae Solem 1976" docType="treatment" docVersion="7" lastPageNumber="41" masterDocId="DE16FFE2FFD2FFC7571CFFB4FFAEFFFE" masterDocTitle="Radiation and decline of endodontid land snails in Makatea, French Polynesia" masterLastPageNumber="68" masterPageNumber="1" pageNumber="39" updateTime="1698298295679" updateUser="ExternalLinkService">
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<mods:title id="EED948ACAB1ED691CA07AA0180886B03">Radiation and decline of endodontid land snails in Makatea, French Polynesia</mods:title>
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<mods:name id="77316F5817DCC7450DF6C118A9E9A035" type="personal">
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<mods:namePart id="39E22CEA39DFDCDF7B21DCBF3CE784FA">Sartori, André F.</mods:namePart>
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<mods:namePart id="27A7012AF13C6EF744F5C8FEDAA14C21">Gargominy, Olivier</mods:namePart>
</mods:name>
<mods:name id="403D30D6358C53EE73F8E0662DBB4C44" type="personal">
<mods:role id="55E7EE0972F9C333B94519A38E70C2BE">
<mods:roleTerm id="A3A1814294793F754E4429086ED1655B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EE7AD2EFE933913EE5D2EFFC792F38CA">Fontaine, Benoît</mods:namePart>
</mods:name>
<mods:typeOfResource id="99F443377C66F36ECEAABC11659AD7B8">text</mods:typeOfResource>
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<mods:title id="8E05FC37BFACDAAF9A4B4FF737BE669E">Zootaxa</mods:title>
</mods:titleInfo>
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<mods:date id="5F4AE4EDFEC55366C6A25D4832B2BFE8">2014</mods:date>
<mods:detail id="5A3B628209B547E00F2CCF33D9DC6CC5" type="volume">
<mods:number id="E0E6FE0AD4A0CEBE6E7215D8C9FE6A99">3772</mods:number>
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<mods:identifier id="0E6EC90F78372D241DEC619256914ABC" type="DOI">10.11646/zootaxa.3772.1.1</mods:identifier>
<mods:identifier id="D847F0FCA0E57AF2F2EFE8F462F2293D" type="GBIF-Dataset">12787306-9151-40cd-8103-1ecac6bdb6b8</mods:identifier>
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<treatment id="222F879AFFF4FFEF578BFB48FC4AFDC3" ID-DOI="http://doi.org/10.5281/zenodo.5612286" ID-GBIF-Taxon="127667190" ID-Zenodo-Dep="5612286" LSID="urn:lsid:plazi:treatment:222F879AFFF4FFEF578BFB48FC4AFDC3" httpUri="http://treatment.plazi.org/id/222F879AFFF4FFEF578BFB48FC4AFDC3" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
<subSubSection id="E29C6507FFF4FFE1578BFB48FE70FA82" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="AA39368CFFF4FFE1578BFB48FD93FAE8" blockId="38.[151,573,1276,1335]" box="[151,573,1276,1302]" pageId="38" pageNumber="39">
<heading id="F17181E0FFF4FFE1578BFB48FD93FAE8" bold="true" box="[151,573,1276,1302]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<taxonomicName id="6D864D0FFFF4FFE1578BFB48FD93FAE8" ID-CoL="4NZGV" authority="Solem, 1976" authorityName="Solem" authorityYear="1976" box="[151,573,1276,1302]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="lillianae">
<emphasis id="98F2EA9EFFF4FFE1578BFB48FD93FAE8" bold="true" box="[151,573,1276,1302]" pageId="38" pageNumber="39">
<emphasis id="98F2EA9EFFF4FFE1578BFB48FE0EFAE8" bold="true" box="[151,416,1276,1302]" italics="true" pageId="38" pageNumber="39">Pseudolibera lillianae</emphasis>
<bibRefCitation id="CE174B7DFFF4FFE156BBFB48FD93FAE8" author="Solem" box="[423,573,1276,1302]" pageId="38" pageNumber="39" refString="Solem, A. (1976) Endodontoid Land Snails from Pacific Islands (Mollusca: Pulmonata: Sigmurethra). Part 1. Family Endodontidae. Field Museum of Natural History, Chicago, Illinois, 508 pp." type="book" year="1976">Solem, 1976</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AA39368CFFF4FFE1578BFAAAFE4AFAC9" blockId="38.[151,573,1276,1335]" box="[151,484,1310,1335]" pageId="38" pageNumber="39">
<figureCitation id="32BD2A09FFF4FFE1578BFAAAFEBFFAC9" box="[151,273,1310,1335]" captionStart="FIGURE 23" captionStartId="39.[151,250,1687,1709]" captionTargetBox="[151,1436,877,1665]" captionTargetId="figure@39.[151,1436,877,1666]" captionTargetPageId="39" captionText="FIGURE 23. Pseudolibera lillianae. A. Holotype (BPBM 115805); B. Specimen 1 (MNHN 25589); Scale bars = 5 mm." httpUri="https://zenodo.org/record/285676/files/figure.png" pageId="38" pageNumber="39">Figures 23</figureCitation>
; 24; 36A; 37F; 39.
</paragraph>
<paragraph id="AA39368CFFF4FFE1578BFAD2FE70FA82" blockId="38.[151,1321,1382,1496]" box="[151,478,1382,1404]" pageId="38" pageNumber="39">
<taxonomicName id="6D864D0FFFF4FFE1578BFAD2FF75FA82" box="[151,219,1382,1404]" class="Gastropoda" family="Endodontidae" genus="Libera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">
<emphasis id="98F2EA9EFFF4FFE1578BFAD2FF75FA82" box="[151,219,1382,1404]" italics="true" pageId="38" pageNumber="39">Libera</emphasis>
</taxonomicName>
sp.
<bibRefCitation id="CE174B7DFFF4FFE15618FAD2FE2FFA82" author="Cooke" box="[260,385,1382,1404]" pageId="38" pageNumber="39" refString="Cooke, C. M. (1934) Land shells of Makatea. Occasional Papers of the Bernice Pauahi Bishop Museum of Polynesian Ethnology and Natural History, 10 (11), 1 - 11." type="journal article" year="1934">Cooke 1934</bibRefCitation>
, pp. 5–6.
</paragraph>
</subSubSection>
<subSubSection id="E29C6507FFF4FFE1578BFA31FCA0FA26" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="AA39368CFFF4FFE1578BFA31FCA0FA26" blockId="38.[151,1321,1382,1496]" pageId="38" pageNumber="39">
<taxonomicName id="6D864D0FFFF4FFE1578BFA31FE0DFA65" authority="Gould" authorityName="Gould" box="[151,419,1413,1435]" class="Insecta" family="Scarabaeidae" genus="Endodonta" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="obolus">
<emphasis id="98F2EA9EFFF4FFE1578BFA31FEE1FA65" box="[151,335,1413,1435]" italics="true" pageId="38" pageNumber="39">Endodonta obolus</emphasis>
(Gould)
</taxonomicName>
—Aubert de la
<bibRefCitation id="CE174B7DFFF4FFE1555FFA32FCACFA65" author="Rue" box="[579,770,1413,1435]" pageId="38" pageNumber="39" refString="Aubert de la Rue, E. &amp; Soyer, R. (1958) Faunule de mollusques terrestres recueillie dans l'ile de Makatea (Archipel des Tuamotu). Bulletin du Museum National d'Histoire Naturelle, 30 (4), 365 - 366." type="journal article" year="1958">Rüe &amp; Soyer 1958</bibRefCitation>
, p. 365,
<emphasis id="98F2EA9EFFF4FFE15445FA32FCD1FA65" box="[857,895,1414,1435]" italics="true" pageId="38" pageNumber="39">non</emphasis>
<taxonomicName id="6D864D0FFFF4FFE15499FA31FB3DFA65" authority="Gould 1846b" authorityName="Gould" authorityYear="1846" box="[901,1171,1413,1435]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="obolus">
<emphasis id="98F2EA9EFFF4FFE15499FA31FBA8FA65" box="[901,1030,1413,1435]" italics="true" pageId="38" pageNumber="39">Helix obolus</emphasis>
<bibRefCitation id="CE174B7DFFF4FFE15310FA31FB3DFA65" author="Gould" box="[1036,1171,1413,1435]" pageId="38" pageNumber="39" refString="Gould, A. A. (1846 b) Descriptions of the species Helix, from the shells of the United Stated Exploring Expedition. Proceedings of the Boston Society of Natural History, 2, 175 - 176." type="journal article" year="1846" yearSuffix="b">Gould 1846b</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="6D864D0FFFF4FFE1578BFA10FE76FA44" authority="Pease" authorityName="Pease" box="[151,472,1444,1466]" class="Gastropoda" family="Urocyclidae" genus="Trochonanina" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="obconica">
<emphasis id="98F2EA9EFFF4FFE1578BFA10FE29FA44" box="[151,391,1444,1466]" italics="true" pageId="38" pageNumber="39">Trochonanina obconica</emphasis>
(Pease)
</taxonomicName>
[in part]—Aubert de la
<bibRefCitation id="CE174B7DFFF4FFE155D1FA11FC23FA44" author="Rue" box="[717,909,1444,1466]" pageId="38" pageNumber="39" refString="Aubert de la Rue, E. &amp; Soyer, R. (1958) Faunule de mollusques terrestres recueillie dans l'ile de Makatea (Archipel des Tuamotu). Bulletin du Museum National d'Histoire Naturelle, 30 (4), 365 - 366." type="journal article" year="1958">Rüe &amp; Soyer 1958</bibRefCitation>
, p. 365,
<emphasis id="98F2EA9EFFF4FFE154FFFA11FBA7FA44" box="[995,1033,1445,1466]" italics="true" pageId="38" pageNumber="39">non</emphasis>
<taxonomicName id="6D864D0FFFF4FFE15313FA10FA8BFA44" authority="Pease 1865" authorityName="Pease" authorityYear="1865" box="[1039,1317,1444,1466]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="obconica">
<emphasis id="98F2EA9EFFF4FFE15313FA10FB07FA44" box="[1039,1193,1444,1466]" italics="true" pageId="38" pageNumber="39">Helix obconica</emphasis>
<bibRefCitation id="CE174B7DFFF4FFE153B3FA11FA8BFA44" author="Pease" box="[1199,1317,1444,1466]" pageId="38" pageNumber="39" refString="Pease, W. (1865) Descriptions of new species of land shells from the islands of the central Pacific. Proceedings of the Zoological Society of London, 1864, 668 - 676." type="proceedings paper" year="1865">Pease 1865</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="6D864D0FFFF4FFE1578BFA76FE55FA26" ID-CoL="4NZGV" authority="Solem 1976" authorityName="Solem" authorityYear="1976" box="[151,507,1474,1496]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="lillianae">
<emphasis id="98F2EA9EFFF4FFE1578BFA76FED6FA26" box="[151,376,1474,1496]" italics="true" pageId="38" pageNumber="39">Pseudolibera lillianae</emphasis>
<bibRefCitation id="CE174B7DFFF4FFE15663FA76FE55FA26" author="Solem" box="[383,507,1474,1496]" pageId="38" pageNumber="39" refString="Solem, A. (1976) Endodontoid Land Snails from Pacific Islands (Mollusca: Pulmonata: Sigmurethra). Part 1. Family Endodontidae. Field Museum of Natural History, Chicago, Illinois, 508 pp." type="book" year="1976">Solem 1976</bibRefCitation>
</taxonomicName>
, pp. 384–385, figs 168a–b.
</paragraph>
</subSubSection>
<subSubSection id="E29C6507FFF4FFE1578BF9B1FBBCF908" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="AA39368CFFF4FFE1578BF9B1FF4FF92C" blockId="38.[151,1436,1541,2034]" pageId="38" pageNumber="39">
<emphasis id="98F2EA9EFFF4FFE1578BF9B1FED2F9E0" bold="true" box="[151,380,1541,1566]" pageId="38" pageNumber="39">Examined material</emphasis>
(1140 specimens).
<emphasis id="98F2EA9EFFF4FFE15549F9B1FD6CF9E0" bold="true" box="[597,706,1541,1566]" pageId="38" pageNumber="39">
<typeStatus id="753D882EFFF4FFE15549F9B1FD6CF9E0" box="[597,706,1541,1566]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectionCode id="CC97AE49FFF4FFE155CDF9B2FCB1F9E3" LSID="urn:lsid:biocol.org:col:34697" box="[721,799,1542,1565]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="38" pageNumber="39">BPBM</collectionCode>
115805, Tuamotu Islands: Makatea,
<quantity id="6D7E9B69FFF4FFE153A1F9B2FAAAF9E0" box="[1213,1284,1541,1566]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="38" pageNumber="39" unit="mi" value="1.0">1 mile</quantity>
inland at
<quantity id="6D7E9B69FFF4FFE1526EF9B1FF02F9BC" metricMagnitude="1" metricUnit="m" metricValue="7.62" pageId="38" pageNumber="39" unit="ft" value="250.0">250 ft</quantity>
. elevation. Collected on a hillside around roots of a plant by Mrs. G.P. Wilder on
<date id="DE38104CFFF4FFE15354F99DFABFF9BC" box="[1096,1297,1577,1602]" pageId="38" pageNumber="39" value="1932-10-24">October 24, 1932</date>
.
<emphasis id="98F2EA9EFFF4FFE15200F99DFF52F998" bold="true" pageId="38" pageNumber="39">Additional material</emphasis>
:
<collectionCode id="CC97AE49FFF4FFE1560CF9FAFEC6F99B" LSID="urn:lsid:biocol.org:col:34988" box="[272,360,1614,1637]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="38" pageNumber="39">MNHN</collectionCode>
, unregistered, 13 shells, one of which gold coated, det. A. Solem [presumably collected by E. Aubert de la Rüe in
<quantity id="6D7E9B69FFF4FFE15680F9C5FE5FF974" box="[412,497,1649,1674]" metricMagnitude="1" metricUnit="m" metricValue="4.9657" pageId="38" pageNumber="39" unit="in" value="1955.0">1955 in</quantity>
Makatea; see remarks below];
<collectionCode id="CC97AE49FFF4FFE1544FF9C6FC06F977" LSID="urn:lsid:biocol.org:col:34988" box="[851,936,1650,1673]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="38" pageNumber="39">MNHN</collectionCode>
25589, 18 shells, Mk04; 4 shells, Mk09; 14 shells, Mk10; 361 shells, Mk12; 362 shells, Mk13; 226 shells, Mk16; 133 shells, Mk19; 6 shells, Mk20; 2 shells, Mk25.
</paragraph>
<paragraph id="AA39368CFFF4FFE157DDF969FBBCF908" blockId="38.[151,1436,1541,2034]" box="[193,1042,1757,1782]" pageId="38" pageNumber="39">
<emphasis id="98F2EA9EFFF4FFE157DDF969FECEF908" bold="true" box="[193,352,1757,1782]" pageId="38" pageNumber="39">
<typeStatus id="753D882EFFF4FFE157DDF969FF52F908" box="[193,252,1757,1782]" pageId="38" pageNumber="39">Type</typeStatus>
locality.
</emphasis>
Tuamotu Islands: Makatea,
<quantity id="6D7E9B69FFF4FFE15581F96AFD4BF908" box="[669,741,1757,1782]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="38" pageNumber="39" unit="mi" value="1.0">1 mile</quantity>
inland at
<quantity id="6D7E9B69FFF4FFE15448F969FC37F908" box="[852,921,1757,1782]" metricMagnitude="1" metricUnit="m" metricValue="7.62" pageId="38" pageNumber="39" unit="ft" value="250.0">250 ft</quantity>
. elevation.
</paragraph>
</subSubSection>
<subSubSection id="E29C6507FFF4FFE157DDF8B5FC0EF89C" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="AA39368CFFF4FFE157DDF8B5FC0EF89C" blockId="38.[151,1436,1541,2034]" pageId="38" pageNumber="39">
<emphasis id="98F2EA9EFFF4FFE157DDF8B5FE95F8E4" bold="true" box="[193,315,1793,1818]" pageId="38" pageNumber="39">Diagnosis.</emphasis>
Shell less than
<quantity id="6D7E9B69FFF4FFE156F1F8B6FD81F8E7" box="[493,559,1794,1818]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="39" unit="mm" value="9.0">9 mm</quantity>
in diameter, depressed, flammulated; apex and spire elevated; peripheral keel long and narrow; teleoconch sculptured by subequal axial and spiral ribs, reduced on shell base; palatal wall devoid of barriers; 1 parietal barrier extending 1/2 to 1 whorl, not bifurcated.
</paragraph>
</subSubSection>
<subSubSection id="E29C6507FFF4FFE057DDF8D9FEF0FCC6" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="description">
<paragraph id="AA39368CFFF4FFE057DDF8D9FEF0FCC6" blockId="38.[151,1436,1541,2034]" lastBlockId="39.[151,1436,151,824]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<emphasis id="98F2EA9EFFF4FFE157DDF8D9FEFDF878" bold="true" box="[193,339,1901,1926]" pageId="38" pageNumber="39">Description.</emphasis>
Shell depressed, dome-shaped, white, with regularly spaced, amber flammulations, frequently interrupted at the shell periphery and vicinity of the umbilicus; flammulations on the shell base larger and fewer, commonly absent on the last half whorl; shell base rarely tinted with an amber background coloration, in addition to flammulations. Shell wall thin, opaque to subpellucid; periostracum adherent, shiny. Apex and spire elevated, later whorls descending slightly more rapidly. Apical suture adpressed at apex, progressively deepening in subsequent whorls; umbilical suture adpressed. Whorls concave above and below long, narrow peripheral keel, transitioning into broadly convex toward the apical suture and shell base; junction of basal and columellar walls initially obtusely angled, developing a keel approximately from the fifth whorl onwards. Transition between protoconch and teleoconch indistinct. Primary axial sculpture of the protoconch composed of relatively broad ribs, with interspaces c. 3 times their width; 4–6 secondary axial riblets occupying the interspaces between primary ribs, each approximately 1/5 the width of the primary ribs and wavy in morphology. Primary ribs of the protoconch gradually transitioning into narrower and less conspicuous ribs, which comprise the primary axial sculpture of the teleoconch; secondary riblets of the protoconch persisting as the secondary axial sculpture of the teleoconch. Spiral sculpture developing approximately from the beginning of the third whorl onwards, composed of wavy ribs, similar in prominence and spacing to the primary axial ribs of the teleoconch; nodular projections present at intersections between spiral and axial ribs. Sculpture less prominent on shell base than above periphery. Umbilicus rapidly expanding in diameter for approximately the first 3 whorls, remaining constant in diameter or expanding slightly from approximately the third to the fifth whorl, subsequently constricted by inward growth of the lower columellar wall and lip. Peristome elongated crescent, with rostrate periphery; columellar lip reflected. Palatal wall devoid of barriers. Parietal wall with 1 barrier, extending 1/2 to 1 whorl, positioned slightly closer to the apical than the umbilical suture, descending gradually anteriorly and posteriorly, rarely flanked by 1 trace on each side. Parietal traces, when present, extending c. 1/4 whorl. Other shell features that can be expressed numerically are shown in
<tableCitation id="E7040337FFF5FFE05618FCABFEF6FCC6" box="[260,344,799,824]" captionStart="TABLE 3" captionStartId="56.[151,238,472,495]" captionText="TABLE 3. Dimensions (in mm) and ratios of dimensions of specimens of Pseudolibera. See Figure 3 for the placement" pageId="39" pageNumber="40">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="FEF96604FFF5FFE0578BF923FAF2F953" box="[151,1372,1687,1709]" httpUri="https://zenodo.org/record/285676/files/figure.png" pageId="39" pageNumber="40" targetBox="[151,1436,877,1665]" targetPageId="39">
<paragraph id="AA39368CFFF5FFE0578BF923FAF2F953" blockId="39.[151,1372,1687,1709]" box="[151,1372,1687,1709]" pageId="39" pageNumber="40">
<emphasis id="98F2EA9EFFF5FFE0578BF923FEB1F953" bold="true" box="[151,287,1687,1709]" pageId="39" pageNumber="40">FIGURE 23.</emphasis>
<taxonomicName id="6D864D0FFFF5FFE05639F923FDA9F953" box="[293,519,1687,1709]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="lillianae">
<emphasis id="98F2EA9EFFF5FFE05639F923FDA9F953" box="[293,519,1687,1709]" italics="true" pageId="39" pageNumber="40">Pseudolibera lillianae</emphasis>
</taxonomicName>
.
<emphasis id="98F2EA9EFFF5FFE05508F923FD88F953" bold="true" box="[532,550,1687,1709]" pageId="39" pageNumber="40">A</emphasis>
. Holotype (BPBM 115805);
<emphasis id="98F2EA9EFFF5FFE05451F923FCF0F953" bold="true" box="[845,862,1687,1709]" pageId="39" pageNumber="40">B</emphasis>
. Specimen 1 (MNHN 25589); Scale bars = 5 mm.
</paragraph>
</caption>
<subSubSection id="E29C6507FFF5FFEF57DDF96EFC4AFDC3" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="discussion">
<paragraph id="AA39368CFFF5FFE057DDF96EFBA8F87C" blockId="39.[151,1436,1754,2031]" pageId="39" pageNumber="40">
<emphasis id="98F2EA9EFFF5FFE057DDF96EFE9BF90D" bold="true" box="[193,309,1754,1779]" pageId="39" pageNumber="40">Remarks.</emphasis>
<bibRefCitation id="CE174B7DFFF5FFE05620F96EFDF8F90D" author="Solem" box="[316,598,1754,1779]" pageId="39" pageNumber="40" refString="Solem, A. (1976) Endodontoid Land Snails from Pacific Islands (Mollusca: Pulmonata: Sigmurethra). Part 1. Family Endodontidae. Field Museum of Natural History, Chicago, Illinois, 508 pp." type="book" year="1976">Solem (1976, pp. 5, 384)</bibRefCitation>
found a preliminary description of this species among Montague Cooke’s unpublished notes and, wishing to recognize Cooke’s contribution, attributed authorship of
<taxonomicName id="6D864D0FFFF5FFE053BBF94BFAB1F8E9" box="[1191,1311,1790,1815]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="lillianae">
<emphasis id="98F2EA9EFFF5FFE053BBF94BFAB1F8E9" box="[1191,1311,1790,1815]" italics="true" pageId="39" pageNumber="40">P. lillianae</emphasis>
</taxonomicName>
to “Cooke &amp; Solem”. However,
<bibRefCitation id="CE174B7DFFF5FFE05681F896FDD2F8C5" author="Solem" box="[413,636,1826,1851]" pageId="39" pageNumber="40" refString="Solem, A. (1976) Endodontoid Land Snails from Pacific Islands (Mollusca: Pulmonata: Sigmurethra). Part 1. Family Endodontidae. Field Museum of Natural History, Chicago, Illinois, 508 pp." type="book" year="1976" yearSuffix="p">Solem (1976, p. 5)</bibRefCitation>
clearly stated that all diagnoses and descriptions published in his monograph were prepared by himself. Hence, Solem was alone responsible for satisfying the criteria of availability and, following ICZN Article 50 (
<bibRefCitation id="CE174B7DFFF5FFE05515F8DFFD21F87D" author="ICZN" box="[521,655,1899,1923]" pageId="39" pageNumber="40" refString="ICZN (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. Fourth Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
), is the sole author of
<taxonomicName id="6D864D0FFFF5FFE0549AF8DFFC50F87D" box="[902,1022,1898,1923]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="lillianae">
<emphasis id="98F2EA9EFFF5FFE0549AF8DFFC50F87D" box="[902,1022,1898,1923]" italics="true" pageId="39" pageNumber="40">P. lillianae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="AA39368CFFF5FFEF57DDF83AFDD7FE79" blockId="39.[151,1436,1754,2031]" lastBlockId="40.[151,1436,151,573]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<bibRefCitation id="CE174B7DFFF5FFE057DDF83AFEF4F859" author="Solem" box="[193,346,1934,1959]" pageId="39" pageNumber="40" refString="Solem, A. (1976) Endodontoid Land Snails from Pacific Islands (Mollusca: Pulmonata: Sigmurethra). Part 1. Family Endodontidae. Field Museum of Natural History, Chicago, Illinois, 508 pp." type="book" year="1976">Solem (1976)</bibRefCitation>
established
<taxonomicName id="6D864D0FFFF5FFE056E2F83BFDD8F859" box="[510,630,1934,1959]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="lillianae">
<emphasis id="98F2EA9EFFF5FFE056E2F83BFDD8F859" box="[510,630,1934,1959]" italics="true" pageId="39" pageNumber="40">P. lillianae</emphasis>
</taxonomicName>
based on only two specimens—the considerably worn
<typeStatus id="753D882EFFF5FFE05226F83AFA32F859" box="[1338,1436,1934,1959]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
(
<figureCitation id="32BD2A09FFF5FFE05783F806FF5EF835" box="[159,240,1970,1995]" captionStart="FIGURE 23" captionStartId="39.[151,250,1687,1709]" captionTargetBox="[151,1436,877,1665]" captionTargetId="figure@39.[151,1436,877,1666]" captionTargetPageId="39" captionText="FIGURE 23. Pseudolibera lillianae. A. Holotype (BPBM 115805); B. Specimen 1 (MNHN 25589); Scale bars = 5 mm." httpUri="https://zenodo.org/record/285676/files/figure.png" pageId="39" pageNumber="40">Fig. 23</figureCitation>
A) and a very small juvenile specimen collected by Aubert de la
<bibRefCitation id="CE174B7DFFF5FFE054D0F807FB1FF835" author="Rue" box="[972,1201,1970,1995]" pageId="39" pageNumber="40" refString="Aubert de la Rue, E. &amp; Soyer, R. (1958) Faunule de mollusques terrestres recueillie dans l'ile de Makatea (Archipel des Tuamotu). Bulletin du Museum National d'Histoire Naturelle, 30 (4), 365 - 366." type="journal article" year="1958">Rüe &amp; Soyer (1958)</bibRefCitation>
and misidentified as
<taxonomicName id="6D864D0FFFF5FFE0578BF862FECAF811" box="[151,356,2006,2031]" class="Insecta" family="Scarabaeidae" genus="Endodonta" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="obolus">
<emphasis id="98F2EA9EFFF5FFE0578BF862FECAF811" box="[151,356,2006,2031]" italics="true" pageId="39" pageNumber="40">Endodonta obolus</emphasis>
</taxonomicName>
.
<bibRefCitation id="CE174B7DFFF5FFE0566DF862FD0BF811" author="Solem" box="[369,677,2006,2031]" pageId="39" pageNumber="40" refString="Solem, A. (1983) Endodontoid Land Snails from Pacific Islands (Mollusca: Pulmonata: Sigmurethra). Part 2. Families Punctidae and Charopidae, Zoogeography. Field Museum of Natural History, Chicago, Illinois, 336 pp." type="book" year="1983">Solem (1983, pp. 279–280)</bibRefCitation>
noted that, during a visit to the MNHN, he found several additional specimens of this species mixed in a lot that Aubert de la
<bibRefCitation id="CE174B7DFFFAFFEF5456FF2CFB98FF4E" author="Rue" box="[842,1078,151,176]" pageId="40" pageNumber="41" refString="Aubert de la Rue, E. &amp; Soyer, R. (1958) Faunule de mollusques terrestres recueillie dans l'ile de Makatea (Archipel des Tuamotu). Bulletin du Museum National d'Histoire Naturelle, 30 (4), 365 - 366." type="journal article" year="1958">Rüe &amp; Soyer (1958)</bibRefCitation>
had reported as
<taxonomicName id="6D864D0FFFFAFFEF521CFF23FF50FF2B" class="Gastropoda" family="Urocyclidae" genus="Trochonanina" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="obconica">
<emphasis id="98F2EA9EFFFAFFEF521CFF23FF50FF2B" italics="true" pageId="40" pageNumber="41">Trochonanina obconica</emphasis>
</taxonomicName>
, which also contained three new species of the genus. Solem’s premature death prevented him from establishing those three new species; they are described herein as
<taxonomicName id="6D864D0FFFFAFFEF5461FF55FC4DFF06" box="[893,995,223,248]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="solemi" status="sp. nov.">
<emphasis id="98F2EA9EFFFAFFEF5461FF55FC4DFF06" box="[893,995,223,248]" italics="true" pageId="40" pageNumber="41">P. solemi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C157E5FFFAFFEF54F5FF54FBECFF09" box="[1001,1090,224,247]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="6D864D0FFFFAFFEF5351FF55FAA1FF06" box="[1101,1295,223,248]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="aubertdelaruei" status="sp. nov.">
<emphasis id="98F2EA9EFFFAFFEF5351FF55FAA1FF06" box="[1101,1295,223,248]" italics="true" pageId="40" pageNumber="41">P. aubertdelaruei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C157E5FFFAFFEF520AFF54FAC3FF09" box="[1302,1389,224,247]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="6D864D0FFFFAFFEF578BFEB1FEA5FEE2" box="[151,267,261,284]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="extincta" status="sp. nov.">
<emphasis id="98F2EA9EFFFAFFEF578BFEB1FEA5FEE2" box="[151,267,261,284]" italics="true" pageId="40" pageNumber="41">P. extincta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C157E5FFFAFFEF560EFEB1FEC7FEE2" box="[274,361,261,284]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
<collectingCountry id="D291761CFFFAFFEF566BFEB0FE38FEE2" box="[375,406,260,284]" name="American Samoa" pageId="40" pageNumber="41">As</collectingCountry>
for the specimens of
<taxonomicName id="6D864D0FFFFAFFEF55ACFEB1FC87FEE3" box="[688,809,260,285]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="lillianae">
<emphasis id="98F2EA9EFFFAFFEF55ACFEB1FC87FEE3" box="[688,809,260,285]" italics="true" pageId="40" pageNumber="41">P. lillianae</emphasis>
</taxonomicName>
collected by Aubert de la
<bibRefCitation id="CE174B7DFFFAFFEF536DFEB1FACAFEE3" author="Rue" box="[1137,1380,260,285]" pageId="40" pageNumber="41" refString="Aubert de la Rue, E. &amp; Soyer, R. (1958) Faunule de mollusques terrestres recueillie dans l'ile de Makatea (Archipel des Tuamotu). Bulletin du Museum National d'Histoire Naturelle, 30 (4), 365 - 366." type="journal article" year="1958">Rüe &amp; Soyer (1958)</bibRefCitation>
and recognized by
<bibRefCitation id="CE174B7DFFFAFFEF5620FE93FE79FEBE" author="Solem" box="[316,471,295,320]" pageId="40" pageNumber="41" refString="Solem, A. (1983) Endodontoid Land Snails from Pacific Islands (Mollusca: Pulmonata: Sigmurethra). Part 2. Families Punctidae and Charopidae, Zoogeography. Field Museum of Natural History, Chicago, Illinois, 336 pp." type="book" year="1983">Solem (1983)</bibRefCitation>
, it seems clear that they are contained in the first lot we list in our examined material, even though the lot now lacks labels detailing its history and collection data. The lot contained
<specimenCount id="BC80FD05FFFAFFEF53CDFEF8FAC2FE9B" box="[1233,1388,332,357]" pageId="40" pageNumber="41" type="generic">14 specimens</specimenCount>
, but one of them proved to be
<taxonomicName id="6D864D0FFFFAFFEF56AAFEC5FDB5FE76" box="[438,539,367,392]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="solemi" status="sp. nov.">
<emphasis id="98F2EA9EFFFAFFEF56AAFEC5FDB5FE76" box="[438,539,367,392]" italics="true" pageId="40" pageNumber="41">P. solemi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C157E5FFFAFFEF553EFEC4FDD7FE79" box="[546,633,368,391]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="AA39368CFFFAFFEF57DDFE21FC4AFDC3" blockId="40.[151,1436,151,573]" pageId="40" pageNumber="41">
<taxonomicName id="6D864D0FFFFAFFEF57DDFE21FE97FE53" box="[193,313,404,429]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="lillianae">
<emphasis id="98F2EA9EFFFAFFEF57DDFE21FE97FE53" box="[193,313,404,429]" italics="true" pageId="40" pageNumber="41">P. lillianae</emphasis>
</taxonomicName>
was the most abundant
<taxonomicName id="6D864D0FFFFAFFEF5542FE20FD5DFE53" box="[606,755,404,429]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="genus">
<emphasis id="98F2EA9EFFFAFFEF5542FE20FD5DFE53" box="[606,755,404,429]" italics="true" pageId="40" pageNumber="41">Pseudolibera</emphasis>
</taxonomicName>
in the material recovered in 2005 (
<figureCitation id="32BD2A09FFFAFFEF5382FE20FB5CFE52" box="[1182,1266,404,429]" captionStart="FIGURE 39" captionStartId="64.[151,250,1340,1362]" captionTargetBox="[240,1346,337,1318]" captionTargetId="figure@64.[240,1347,337,1319]" captionTargetPageId="64" captionText="FIGURE 39. Diagrammatic representation of the abundance per station and per species of the endodontid specimens collected in Makatea. The thickness of the bars along the axes and the area of the circles in the grid is proportional to the number of collected specimens." httpUri="https://zenodo.org/record/285692/files/figure.png" pageId="40" pageNumber="41">Fig. 39</figureCitation>
). A few wellpreserved specimens were found, which reveal the color pattern of the species (
<figureCitation id="32BD2A09FFFAFFEF5344FE03FB07FE2E" box="[1112,1193,439,464]" captionStart="FIGURE 23" captionStartId="39.[151,250,1687,1709]" captionTargetBox="[151,1436,877,1665]" captionTargetId="figure@39.[151,1436,877,1666]" captionTargetPageId="39" captionText="FIGURE 23. Pseudolibera lillianae. A. Holotype (BPBM 115805); B. Specimen 1 (MNHN 25589); Scale bars = 5 mm." httpUri="https://zenodo.org/record/285676/files/figure.png" pageId="40" pageNumber="41">Fig. 23</figureCitation>
B) and details of its sculpture (
<figureCitation id="32BD2A09FFFAFFEF5611FE68FECEFE0A" box="[269,352,476,501]" captionStart="FIGURE 24" captionStartId="40.[151,250,1940,1962]" captionTargetBox="[151,1436,625,1918]" captionTargetId="figure@40.[151,1436,625,1919]" captionTargetPageId="40" captionText="FIGURE 24. Pseudolibera lillianae. A – C. Specimen 1 (MNHN 25589), apical views; A. General view; B. Sculpture of the protoconch and early teleoconch; C. Sculpture of the late teleoconch; D. Specimen 9 (station Mk 16), umbilical view, showing parietal traces flanking the barrier; Scale bars: A, D = 2 mm; B, C = 0.4 mm." httpUri="https://zenodo.org/record/285677/files/figure.png" pageId="40" pageNumber="41">Fig. 24</figureCitation>
A–C). Only one specimen among the hundreds recovered displayed parietal traces in addition to the single barrier (
<figureCitation id="32BD2A09FFFAFFEF5676FE4BFE10FDE6" box="[362,446,511,536]" captionStart="FIGURE 24" captionStartId="40.[151,250,1940,1962]" captionTargetBox="[151,1436,625,1918]" captionTargetId="figure@40.[151,1436,625,1919]" captionTargetPageId="40" captionText="FIGURE 24. Pseudolibera lillianae. A – C. Specimen 1 (MNHN 25589), apical views; A. General view; B. Sculpture of the protoconch and early teleoconch; C. Sculpture of the late teleoconch; D. Specimen 9 (station Mk 16), umbilical view, showing parietal traces flanking the barrier; Scale bars: A, D = 2 mm; B, C = 0.4 mm." httpUri="https://zenodo.org/record/285677/files/figure.png" pageId="40" pageNumber="41">Fig. 24</figureCitation>
D).
<taxonomicName id="6D864D0FFFFAFFEF56F4FDB5FDCFFDE6" box="[488,609,511,536]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="lillianae">
<emphasis id="98F2EA9EFFFAFFEF56F4FDB5FDCFFDE6" box="[488,609,511,536]" italics="true" pageId="40" pageNumber="41">P. lillianae</emphasis>
</taxonomicName>
is the largest
<taxonomicName id="6D864D0FFFFAFFEF541FFE4BFC36FDE6" box="[771,920,511,536]" class="Gastropoda" family="Endodontidae" genus="Pseudolibera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="genus">
<emphasis id="98F2EA9EFFFAFFEF541FFE4BFC36FDE6" box="[771,920,511,536]" italics="true" pageId="40" pageNumber="41">Pseudolibera</emphasis>
</taxonomicName>
in shell diameter and the only species of the genus with a long and narrow peripheral keel that is frequently chipped off.
</paragraph>
</subSubSection>
</treatment>
</document>
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