treatments-xml/data/1B/21/87/1B2187E01F1C6658FC032121BE172DB5.xml

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<document id="3935C97185CCCAE39861655A7D82AE41" ID-DOI="10.6620/ZS.2017.56-32" ID-ISSN="1810-522X" ID-PMC="PMC6517758" ID-PubMed="31966231" ID-Zenodo-Dep="12825856" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721914379609" checkinUser="felipe" docAuthor="Salazar-Vallejo, Sergio I." docDate="2017" docId="1B2187E01F1C6658FC032121BE172DB5" docLanguage="en" docName="ZoolStud.56.32.1-16.pdf.imf" docOrigin="Zoological Studies (Zool. Stud.) 56 (32)" docSource="http://dx.doi.org/10.5281/zenodo.12825164" docStyle="DocumentStyle:CB122E4603BA284FA287D3847868CAF6.4:ZoolStud.2016-2018.journal_article" docStyleId="CB122E4603BA284FA287D3847868CAF6" docStyleName="ZoolStud.2016-2018.journal_article" docStyleVersion="4" docTitle="Caulleryaspis villamari Salazar-Vallejo, 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="4" masterDocId="E718FF981F1D665BFF90235DBC642B7E" masterDocTitle="Cymonomus curvirostris Sakai 1965" masterLastPageNumber="16" masterPageNumber="16" pageNumber="2" updateTime="1721932462749" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C0E3521E2E05518CC6C2FBF78EAD487B">Cymonomus curvirostris Sakai 1965</mods:title>
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<mods:namePart id="8D2893F0B01977C33B81F5852A124A78">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:title id="36E00657032DF018E0EB442E66AC63F4">Zoological Studies</mods:title>
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<mods:date id="1BFD6390ADE5C212215A55AA64B1A68C">2017</mods:date>
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<mods:title id="95E133C9CEE3B123FC6E019C4C16F796">Zool. Stud.</mods:title>
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<mods:number id="F5EAEE5D8F1F1CA0A3FDC22C9F2A528E">2017-10-31</mods:number>
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<mods:identifier id="2DCAD349E79F5BCD6D697E6F05108E55" type="DOI">10.6620/ZS.2017.56-32</mods:identifier>
<mods:identifier id="0F0408EC11E508E3E50AE5329A66C325" type="ISSN">1810-522X</mods:identifier>
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<mods:identifier id="607FB656776D206C6916AA1BC4ED40CB" type="PubMed">31966231</mods:identifier>
<mods:identifier id="C2FCF9FCAA2727B0BADED90E0770D926" type="Zenodo-Dep">12825856</mods:identifier>
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<treatment id="1B2187E01F1C6658FC032121BE172DB5" LSID="urn:lsid:plazi:treatment:1B2187E01F1C6658FC032121BE172DB5" httpUri="http://treatment.plazi.org/id/1B2187E01F1C6658FC032121BE172DB5" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<subSubSection id="DB92657D1F1C665AFC032121B97A29EB" box="[915,1310,636,662]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="933736F61F1C665AFC032121B97A29EB" blockId="1.[818,1408,636,746]" box="[915,1310,636,662]" pageId="1" pageNumber="2">
<heading id="C87F819A1F1C665AFC032121B97A29EB" bold="true" box="[915,1310,636,662]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
<emphasis id="A1FCEAE41F1C665AFC032121B97A29EB" bold="true" box="[915,1310,636,662]" pageId="1" pageNumber="2">
<taxonomicName id="54884D751F1C665AFC032121B8D029EB" ID-CoL="85RBJ" authorityName="Salazar-Vallejo" authorityYear="2017" box="[915,1204,636,662]" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="villamari" status="sp. nov.">
<emphasis id="A1FCEAE41F1C665AFC032121B8D029EB" bold="true" box="[915,1204,636,662]" italics="true" pageId="1" pageNumber="2">Caulleryaspis villamari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BACF579F1F1C665AFB2C2121B97A29EB" box="[1212,1310,636,661]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DB92657D1F1C665AFBBF21C1B8D42994" pageId="1" pageNumber="2" type="description">
<paragraph id="933736F61F1C665AFBBF21C1B8E729CB" blockId="1.[818,1408,636,746]" box="[1071,1155,668,693]" pageId="1" pageNumber="2">
(
<figureCitation id="0BB32A731F1C665AFBA621C1B81D29CB" box="[1078,1145,668,693]" captionStart="Fig" captionStartId="2.[175,209,1870,1890]" captionTargetBox="[192,1435,936,1844]" captionTargetId="figure-347@2.[189,1437,934,1844]" captionTargetPageId="2" captionText="Fig. 1. Caulleryaspis villamari sp. nov. (A) Holotype (USNM 1437645), ventral view. (B) Same, close-up of prostomium, mouth and first series of hooks. (C) Same, shield. (D) Same, branchial plate after removal of branchiae and interbranchial filaments. (E) Smaller specimens (USNM 1437646), ventral view. (F) Close-up of shield of smallest one. (G) Close-up of shield of largest one. Scale bars: A = 2.7 mm, B = 0.6 mm, C, D = 1.3 mm, E = 2.8 mm, F = 0.4 mm, G = 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.12825858" httpUri="https://zenodo.org/record/12825858/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="933736F61F1C665AFCA221EAB8D42994" blockId="1.[818,1408,636,746]" pageId="1" pageNumber="2">
<uri id="E7193AF41F1C665AFCA221EAB8D42994" pageId="1" pageNumber="2">
urn:lsid:zoobank.org:act:
<uuid id="E72E0C231F1C665AFBA121EAB8D42994" pageId="1" pageNumber="2">38C77F7E-EAC6-477D-A4DF-265F6B52CDB0</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="DB92657D1F1C665AFCBD2052BF81280D" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="933736F61F1C665AFCBD2052BFCE2840" blockId="1.[813,1415,783,883]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="B39811D81F1C665AFCBD2052BFCE2840" pageId="1" pageNumber="2">
<taxonomicName id="54884D751F1C665AFCBD2052BFB1285D" ID-CoL="52F4Q" authorityName="Stimpson" authorityYear="1853" box="[813,981,783,803]" class="Polychaeta" family="Sternaspidae" genus="Sternaspis" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="fossor">
<emphasis id="A1FCEAE41F1C665AFCBD2052BFB1285D" box="[813,981,783,803]" italics="true" pageId="1" pageNumber="2">Sternaspis fossor</emphasis>
</taxonomicName>
:
<treatmentCitation id="122910E71F1C665AFC732052B8EE285D" author="Villamar F." box="[995,1162,783,803]" page="36" pageId="1" pageNumber="2" year="1989">
<bibRefCitation id="F7194B071F1C665AFC732052B8EE285D" author="Villamar F." box="[995,1162,783,803]" pageId="1" pageNumber="2" pagination="34 - 40" refId="ref11793" refString="Villamar F. 1989. Estudio de los poliquetos bentonicos en el Golfo de Guayaquil, Exterior (Canal del Morro y Jambeli). Acta Oceanogr Pac INOCAR, Ecuador 5: 34 - 40." type="journal article" year="1989">Villamar 1989:36</bibRefCitation>
</treatmentCitation>
, 40 (
<emphasis id="A1FCEAE41F1C665AFB2C2052B884285D" box="[1212,1248,783,803]" italics="true" pageId="1" pageNumber="2">non</emphasis>
<bibRefCitation id="F7194B071F1C665AFB782052B9E7285D" author="Stimpson W." box="[1256,1411,783,803]" pageId="1" pageNumber="2" pagination="5 - 67" refId="ref11656" refString="Stimpson W. 1853. Synopsis of the marine Invertebrata of Grand Manan: or the region about the mouth of the Bay of Fundy. Smithson Contr Knowledge 6: 5 - 67. [printed and distributed in March, 1853]." type="journal article" year="1853">Stimpson, 1853</bibRefCitation>
,
<emphasis id="A1FCEAE41F1C665AFCF22077BFF92840" box="[866,925,810,830]" italics="true" pageId="1" pageNumber="2">partim</emphasis>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="933736F61F1C665AFCBD2019BF81280D" blockId="1.[813,1415,783,883]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="B39811D81F1C665AFCBD2019BF81280D" pageId="1" pageNumber="2">
<taxonomicName id="54884D751F1C665AFCBD2019BF802826" ID-CoL="52F56" baseAuthorityName="Ranzani" baseAuthorityYear="1817" box="[813,996,836,856]" class="Polychaeta" family="Sternaspidae" genus="Sternaspis" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="2" phylum="Annelida" rank="species" species="scutata">
<emphasis id="A1FCEAE41F1C665AFCBD2019BF802826" box="[813,996,836,856]" italics="true" pageId="1" pageNumber="2">Sternaspis scutata</emphasis>
</taxonomicName>
:
<treatmentCitation id="122910E71F1C665AFC622019B8962826" author="Villamar F &amp; Cruz M." box="[1010,1266,836,856]" page="151" pageId="1" pageNumber="2" year="2007">
<bibRefCitation id="F7194B071F1C665AFC622019B8962826" author="Villamar F &amp; Cruz M." box="[1010,1266,836,856]" pageId="1" pageNumber="2" pagination="147 - 153" refId="ref11830" refString="Villamar F, Cruz M. 2007. Poliquetos y moluscos macrobentonicos de la zona intermareal y submareal en la provincia del Guayas (Monteverde, Ecuador). Acta Oceanogr Pac 14: 147 - 153." type="journal article" year="2007">Villamar &amp; Cruz 2007:151</bibRefCitation>
</treatmentCitation>
(
<emphasis id="A1FCEAE41F1C665AFA922019B9422826" box="[1282,1318,836,856]" italics="true" pageId="1" pageNumber="2">non</emphasis>
<bibRefCitation id="F7194B071F1C665AFABF2019BFF0280D" author="Ranzani C." pageId="1" pageNumber="2" pagination="1457 - 1461" refId="ref11358" refString="Ranzani C. 1817. Descrizione di una nuova specie del genere Thalassema. Opus scient 2: 112 [Oken's Isis 12 - 13 (183): 1457 - 1461. (transl. German with additional comments in 1817)]." type="journal article" year="1817">Ranzani, 1817</bibRefCitation>
,
<emphasis id="A1FCEAE41F1C665AFC0D2002BFBC280D" box="[925,984,863,883]" italics="true" pageId="1" pageNumber="2">partim</emphasis>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DB92657D1F1C6658FCF920C7BCB72808" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="933736F61F1C665AFCF920C7B8C22FCD" blockId="1.[813,1414,922,1971]" pageId="1" pageNumber="2">
<emphasis id="A1FCEAE41F1C665AFCF920C7B86F28CD" box="[873,1035,922,947]" italics="true" pageId="1" pageNumber="2">Type material</emphasis>
:
<materialsCitation id="23E03CAB1F1C665AFB8B20C7B8C22FCD" collectingDate="1966-06-05" collectionCode="USNM" collectorName="R. V. Anton Bruun &amp; Cruise &amp; Sta. &amp; de Tierra" country="Peru" elevation="160" latitude="-6.383333" location="Eastern Tropical Pacific" longLatPrecision="1304" longitude="-80.916664" municipality="Lobos de Tierra" pageId="1" pageNumber="2" specimenCode="USNM 1437645, USNM 1437646" specimenCount="10" typeStatus="holotype">
<location id="9657602D1F1C665AFB8B20C7B95E28CD" LSID="urn:lsid:plazi:treatment:1B2187E01F1C6658FC032121BE172DB5:9657602D1F1C665AFB8B20C7B95E28CD" box="[1051,1338,922,947]" country="Peru" latitude="-6.383333" longLatPrecision="1304" longitude="-80.916664" municipality="Lobos de Tierra" name="Eastern Tropical Pacific" pageId="1" pageNumber="2">Eastern Tropical Pacific</location>
,
<collectingCountry id="EB9F76661F1C665AFAD620C7B9E628CD" box="[1350,1410,922,947]" name="Peru" pageId="1" pageNumber="2">Peru</collectingCountry>
.
<typeStatus id="4C3388541F1C665AFCBD20E7BFFB28AD" box="[813,927,954,979]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
(
<specimenCode id="C32E9E8D1F1C665AFC2120E7B8EE28AD" box="[945,1162,954,980]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="2" type="Museum">USNM 1437645</specimenCode>
), and
<specimenCount id="858EFD7F1F1C665AFB4C20E7B9E028AD" box="[1244,1412,954,979]" count="10" pageId="1" pageNumber="2" type="generic" typeStatus="paratype">10 paratypes</specimenCount>
(
<specimenCode id="C32E9E8D1F1C665AFCA32087B860288D" box="[819,1028,986,1012]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="2" type="Museum">USNM 1437646</specimenCode>
),
<collectorName id="3E7D53201F1C665AFB892087B892288D" box="[1049,1270,986,1011]" pageId="1" pageNumber="2">R.V. Anton Bruun</collectorName>
,
<collectorName id="3E7D53201F1C665AFA942087B930288A" box="[1284,1364,986,1012]" pageId="1" pageNumber="2">Cruise</collectorName>
16,
<collectorName id="3E7D53201F1C665AFCBD20A7BF392F6A" box="[813,861,1018,1044]" pageId="1" pageNumber="2">Sta.</collectorName>
635A (
<geoCoordinate id="F6BC50311F1C665AFC3F20A7B8762F6A" box="[943,1042,1018,1044]" degrees="06" direction="south" minutes="27" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="-6.45">06°27'S</geoCoordinate>
,
<geoCoordinate id="F6BC50311F1C665AFB8C20A7B8E42F6D" box="[1052,1152,1018,1043]" degrees="80" direction="west" minutes="56" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="-80.933334">80°56'W</geoCoordinate>
to
<geoCoordinate id="F6BC50311F1C665AFB3620A7B9622F6A" box="[1190,1286,1018,1044]" degrees="06" direction="south" minutes="23" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="-6.383333">06°23'S</geoCoordinate>
,
<geoCoordinate id="F6BC50311F1C665AFA8120A7B9182F6D" box="[1297,1404,1018,1043]" degrees="80" direction="west" minutes="55" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="-80.916664">80°55'W</geoCoordinate>
), off Isla
<collectingMunicipality id="7353AC8C1F1C665AFC122747B8242F4D" box="[898,1088,1050,1075]" pageId="1" pageNumber="2">
Lobos
<collectorName id="3E7D53201F1C665AFC422747B8242F4D" box="[978,1088,1050,1075]" pageId="1" pageNumber="2">de Tierra</collectorName>
</collectingMunicipality>
,
<quantity id="54709B131F1C665AFBDB2747B8FD2F4D" box="[1099,1177,1050,1075]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="2" unit="m" value="160.0">
<elevation id="18A5D1C51F1C665AFBDB2747B8FD2F4D" box="[1099,1177,1050,1075]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="2" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<date id="E73610361F1C665AFB342747B94B2F4D" box="[1188,1327,1050,1075]" pageId="1" pageNumber="2" value="1966-06-05">
<collectingDate id="F772E9DE1F1C665AFB342747B94B2F4D" box="[1188,1327,1050,1075]" pageId="1" pageNumber="2" value="1966-06-05">5 Jun. 1966</collectingDate>
</date>
(shield dirty orange in smaller specimens, becoming grayish; largest
<typeStatus id="4C3388541F1C665AFC6A2707B8032F0D" box="[1018,1127,1114,1139]" pageId="1" pageNumber="2" type="paratype">paratype</typeStatus>
with abdomen
<quantity id="54709B131F1C665AFABF2707B9E02F0D" box="[1327,1412,1114,1139]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="2" unit="mm" value="17.0">17 mm</quantity>
long,
<quantity id="54709B131F1C665AFCE52727BFA82FED" box="[885,972,1146,1171]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="2" unit="mm" value="12.0">12 mm</quantity>
wide; smaller
<typeStatus id="4C3388541F1C665AFB1B2727B9622FED" box="[1163,1286,1146,1171]" pageId="1" pageNumber="2" type="paratype">paratypes</typeStatus>
abdomen 3.5-5.0 mm long,
<quantity id="54709B131F1C665AFC6C27C7B8322FCD" box="[1020,1110,1178,1203]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="2" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3-5 mm</quantity>
wide).
</materialsCitation>
</paragraph>
<paragraph id="933736F61F1C665AFCF927E7B9362E2D" blockId="1.[813,1414,922,1971]" pageId="1" pageNumber="2">
<emphasis id="A1FCEAE41F1C665AFCF927E7B8282FAD" box="[873,1100,1210,1235]" italics="true" pageId="1" pageNumber="2">Additional material</emphasis>
: Eastern Tropical Pacific,
<collectingCountry id="EB9F76661F1C665AFCBD2787BFFF2F8D" box="[813,923,1242,1267]" name="Ecuador" pageId="1" pageNumber="2">Ecuador</collectingCountry>
.
<specimenCount id="858EFD7F1F1C665AFC3A2787B83A2F8D" box="[938,1118,1242,1267]" count="23" pageId="1" pageNumber="2" type="generic">23 specimens</specimenCount>
(
<collectionCode id="F599AE331F1C665AFBE12787B8A22F8A" box="[1137,1222,1242,1268]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="2" type="Museum">USNM</collectionCode>
1437647), RV Anton Bruun, Cruise 18B, Sta. 777D (
<geoCoordinate id="F6BC50311F1C665AFA8727A7B9E52E6D" box="[1303,1409,1274,1299]" degrees="00" direction="north" minutes="48" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="0.8">00°48'N</geoCoordinate>
,
<geoCoordinate id="F6BC50311F1C665AFCBD2647BFF52E4D" box="[813,913,1306,1331]" degrees="80" direction="west" minutes="37" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="-80.61667">80°37'W</geoCoordinate>
to
<geoCoordinate id="F6BC50311F1C665AFC292647B87E2E4A" box="[953,1050,1306,1332]" degrees="00" direction="south" minutes="47" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="-0.78333336">00°47'S</geoCoordinate>
,
<geoCoordinate id="F6BC50311F1C665AFBB72647B8F52E4D" box="[1063,1169,1306,1331]" degrees="80" direction="west" minutes="37" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="-80.61667">80°37'W</geoCoordinate>
),
<date id="E73610361F1C665AFB342647B92F2E4D" box="[1188,1355,1306,1332]" pageId="1" pageNumber="2" value="1966-09-12">12 Sep. 1966</date>
, T.R. Menzies, coll. (small to very small specimens).
</paragraph>
<paragraph id="933736F61F1C665AFCF92607BFCD2D4D" blockId="1.[813,1414,922,1971]" pageId="1" pageNumber="2">
<emphasis id="A1FCEAE41F1C665AFCF92607B8652E0D" box="[873,1025,1370,1395]" italics="true" pageId="1" pageNumber="2">Description</emphasis>
:
<typeStatus id="4C3388541F1C665AFB882607B8EA2E0D" box="[1048,1166,1370,1395]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
(
<collectionCode id="F599AE331F1C665AFB322607B89E2E0A" box="[1186,1274,1370,1396]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="2" type="Museum">USNM</collectionCode>
1437645) complete, flaccid. Body grayish, with introvert exposed, shield grayish (
<figureCitation id="0BB32A731F1C665AFBE726C7B8BF2ECD" box="[1143,1243,1434,1459]" captionStart="Fig" captionStartId="2.[175,209,1870,1890]" captionTargetBox="[192,1435,936,1844]" captionTargetId="figure-347@2.[189,1437,934,1844]" captionTargetPageId="2" captionText="Fig. 1. Caulleryaspis villamari sp. nov. (A) Holotype (USNM 1437645), ventral view. (B) Same, close-up of prostomium, mouth and first series of hooks. (C) Same, shield. (D) Same, branchial plate after removal of branchiae and interbranchial filaments. (E) Smaller specimens (USNM 1437646), ventral view. (F) Close-up of shield of smallest one. (G) Close-up of shield of largest one. Scale bars: A = 2.7 mm, B = 0.6 mm, C, D = 1.3 mm, E = 2.8 mm, F = 0.4 mm, G = 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.12825858" httpUri="https://zenodo.org/record/12825858/files/figure.png" pageId="1" pageNumber="2">Fig. 1A</figureCitation>
). Integument papillae mostly eroded; remaining ones short, filiform, with fine sediment particles. Body
<quantity id="54709B131F1C665AFAA12687B9E12E8D" box="[1329,1413,1498,1523]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="2" unit="mm" value="32.0">32 mm</quantity>
long,
<quantity id="54709B131F1C665AFCFE26A7BFDB2D6D" box="[878,959,1530,1555]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="2" unit="mm" value="10.0">10 mm</quantity>
wide, abdomen
<quantity id="54709B131F1C665AFB1326A7B8B02D6D" box="[1155,1236,1530,1555]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="2" unit="mm" value="20.0">20 mm</quantity>
long, about 28 segments.
</paragraph>
<paragraph id="933736F61F1C665AFCF92567B85A2D8D" blockId="1.[813,1414,922,1971]" pageId="1" pageNumber="2">
Prostomium hemispherical, opaque, distorted after being depressed (
<figureCitation id="0BB32A731F1C665AFBD12507B8FE2D0D" box="[1089,1178,1626,1651]" captionStart="Fig" captionStartId="2.[175,209,1870,1890]" captionTargetBox="[192,1435,936,1844]" captionTargetId="figure-347@2.[189,1437,934,1844]" captionTargetPageId="2" captionText="Fig. 1. Caulleryaspis villamari sp. nov. (A) Holotype (USNM 1437645), ventral view. (B) Same, close-up of prostomium, mouth and first series of hooks. (C) Same, shield. (D) Same, branchial plate after removal of branchiae and interbranchial filaments. (E) Smaller specimens (USNM 1437646), ventral view. (F) Close-up of shield of smallest one. (G) Close-up of shield of largest one. Scale bars: A = 2.7 mm, B = 0.6 mm, C, D = 1.3 mm, E = 2.8 mm, F = 0.4 mm, G = 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.12825858" httpUri="https://zenodo.org/record/12825858/files/figure.png" pageId="1" pageNumber="2">Fig. 1B</figureCitation>
). Peristomium oval, with abundant papillae (mostly eroded), extended as a wide band over prostomium. Mouth oval, as large as prostomium, completely covered by papillae (most eroded).
</paragraph>
<paragraph id="933736F61F1C6659FCF925A7BEBC2A28" blockId="1.[813,1414,922,1971]" lastBlockId="2.[174,775,253,886]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">First three chaetigers with 16-18 falcate, thin introvert hooks per bundle, each with a narrow darker band subdistally, tips transparent, straight in smaller hooks, curved in larger ones. Genital papillae whitish, thick, corrugated, digitate protruding from intersegmental groove between segments 7 and 8. Pre-shield region with 7 segments; two capillary chaetae in segment 9, not visible in other segments (probably broken off).</paragraph>
<footnote id="F0932AF81F1C665AFF182480BDFC2C91" box="[136,408,2013,2031]" pageId="1" pageNumber="2">
<paragraph id="933736F61F1C665AFF182480BDFC2C91" blockId="1.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="1" pageNumber="2">© 2017 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="933736F61F1F6659FF7A2200BE2F2908" blockId="2.[174,775,253,886]" pageId="2" pageNumber="3">
Shield grayish, soft, with a thick integument layer, without sediment particles (
<figureCitation id="0BB32A731F1F6659FDC52220BED22AE9" box="[597,694,381,407]" captionStart="Fig" captionStartId="2.[175,209,1870,1890]" captionTargetBox="[192,1435,936,1844]" captionTargetId="figure-347@2.[189,1437,934,1844]" captionTargetPageId="2" captionText="Fig. 1. Caulleryaspis villamari sp. nov. (A) Holotype (USNM 1437645), ventral view. (B) Same, close-up of prostomium, mouth and first series of hooks. (C) Same, shield. (D) Same, branchial plate after removal of branchiae and interbranchial filaments. (E) Smaller specimens (USNM 1437646), ventral view. (F) Close-up of shield of smallest one. (G) Close-up of shield of largest one. Scale bars: A = 2.7 mm, B = 0.6 mm, C, D = 1.3 mm, E = 2.8 mm, F = 0.4 mm, G = 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.12825858" httpUri="https://zenodo.org/record/12825858/files/figure.png" pageId="2" pageNumber="3">Fig. 1C</figureCitation>
). Ribs faintly defined, no concentric lines. Anterior margins rounded; anterior depression shallow; anterior keels not visible. Lateral margins projected medially, reduced posteriorly. Fan truncate, not extended beyond posterior corners; median notch shallow, lateral notches shallow, displaced laterally; posterior margin barely crenulated.
</paragraph>
<paragraph id="933736F61F1F6659FF7A2120BDFB2848" blockId="2.[174,775,253,886]" pageId="2" pageNumber="3">Marginal chaetal fascicles include 10 lateral ones, chaetae in oval arrangement, and 8 posterior fascicles, chaetae in oblique series. Peg chaetae forming small, thick spines. Additional capillary chaetae broken, present in two fascicles on each side of peg chaetae.</paragraph>
<paragraph id="933736F61F1F6659FF7A2060B8482A48" blockId="2.[174,775,253,886]" lastBlockId="2.[850,1452,253,887]" pageId="2" pageNumber="3">
Branchiae and interbranchial filaments lost. Branchial plates parallel, anteriorly expanded, rounded (
<figureCitation id="0BB32A731F1F6659FC5423A0B8792A68" box="[964,1053,253,278]" captionStart="Fig" captionStartId="2.[175,209,1870,1890]" captionTargetBox="[192,1435,936,1844]" captionTargetId="figure-347@2.[189,1437,934,1844]" captionTargetPageId="2" captionText="Fig. 1. Caulleryaspis villamari sp. nov. (A) Holotype (USNM 1437645), ventral view. (B) Same, close-up of prostomium, mouth and first series of hooks. (C) Same, shield. (D) Same, branchial plate after removal of branchiae and interbranchial filaments. (E) Smaller specimens (USNM 1437646), ventral view. (F) Close-up of shield of smallest one. (G) Close-up of shield of largest one. Scale bars: A = 2.7 mm, B = 0.6 mm, C, D = 1.3 mm, E = 2.8 mm, F = 0.4 mm, G = 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.12825858" httpUri="https://zenodo.org/record/12825858/files/figure.png" pageId="2" pageNumber="3">Fig. 1D</figureCitation>
), with up to 14 longitudinal series of branchial scars.
</paragraph>
<paragraph id="933736F61F1F6659FC1F2260B82329E9" blockId="2.[850,1452,253,887]" pageId="2" pageNumber="3">
<emphasis id="A1FCEAE41F1F6659FC1F2260B8682A28" box="[911,1036,317,342]" italics="true" pageId="2" pageNumber="3">Juveniles</emphasis>
: Small
<typeStatus id="4C3388541F1F6659FBEA2260B8992A28" box="[1146,1277,317,342]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
with shields orange, soft (
<figureCitation id="0BB32A731F1F6659FC7E2200B8222A08" box="[1006,1094,349,374]" captionStart="Fig" captionStartId="2.[175,209,1870,1890]" captionTargetBox="[192,1435,936,1844]" captionTargetId="figure-347@2.[189,1437,934,1844]" captionTargetPageId="2" captionText="Fig. 1. Caulleryaspis villamari sp. nov. (A) Holotype (USNM 1437645), ventral view. (B) Same, close-up of prostomium, mouth and first series of hooks. (C) Same, shield. (D) Same, branchial plate after removal of branchiae and interbranchial filaments. (E) Smaller specimens (USNM 1437646), ventral view. (F) Close-up of shield of smallest one. (G) Close-up of shield of largest one. Scale bars: A = 2.7 mm, B = 0.6 mm, C, D = 1.3 mm, E = 2.8 mm, F = 0.4 mm, G = 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.12825858" httpUri="https://zenodo.org/record/12825858/files/figure.png" pageId="2" pageNumber="3">Fig. 1E</figureCitation>
), damaged. Smallest one with shield orange, with shallow anterior depression, anterior margins barely projected anteriorly, lateral margins straight, progressively thinner, posterior margin with shallow medial notch (
<figureCitation id="0BB32A731F1F6659FB772280B9592A88" box="[1255,1341,477,502]" captionStart="Fig" captionStartId="2.[175,209,1870,1890]" captionTargetBox="[192,1435,936,1844]" captionTargetId="figure-347@2.[189,1437,934,1844]" captionTargetPageId="2" captionText="Fig. 1. Caulleryaspis villamari sp. nov. (A) Holotype (USNM 1437645), ventral view. (B) Same, close-up of prostomium, mouth and first series of hooks. (C) Same, shield. (D) Same, branchial plate after removal of branchiae and interbranchial filaments. (E) Smaller specimens (USNM 1437646), ventral view. (F) Close-up of shield of smallest one. (G) Close-up of shield of largest one. Scale bars: A = 2.7 mm, B = 0.6 mm, C, D = 1.3 mm, E = 2.8 mm, F = 0.4 mm, G = 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.12825858" httpUri="https://zenodo.org/record/12825858/files/figure.png" pageId="2" pageNumber="3">Fig. 1F</figureCitation>
). Largest
<typeStatus id="4C3388541F1F6659FCC222A0BFD92968" box="[850,957,509,534]" pageId="2" pageNumber="3" type="paratype">paratype</typeStatus>
with shield orange, anterior depression and posterior notch shallower than in smaller
<typeStatus id="4C3388541F1F6659FCC22160BFBD2928" box="[850,985,573,598]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
; lateral margins rounded, medially projected, posterior margin straight, barely crenulated (
<figureCitation id="0BB32A731F1F6659FC4E2120B85D29E9" box="[990,1081,637,663]" captionStart="Fig" captionStartId="2.[175,209,1870,1890]" captionTargetBox="[192,1435,936,1844]" captionTargetId="figure-347@2.[189,1437,934,1844]" captionTargetPageId="2" captionText="Fig. 1. Caulleryaspis villamari sp. nov. (A) Holotype (USNM 1437645), ventral view. (B) Same, close-up of prostomium, mouth and first series of hooks. (C) Same, shield. (D) Same, branchial plate after removal of branchiae and interbranchial filaments. (E) Smaller specimens (USNM 1437646), ventral view. (F) Close-up of shield of smallest one. (G) Close-up of shield of largest one. Scale bars: A = 2.7 mm, B = 0.6 mm, C, D = 1.3 mm, E = 2.8 mm, F = 0.4 mm, G = 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.12825858" httpUri="https://zenodo.org/record/12825858/files/figure.png" pageId="2" pageNumber="3">Fig. 1G</figureCitation>
).
</paragraph>
<paragraph id="933736F61F1F6659FC1F21C0B96E2868" blockId="2.[850,1452,253,887]" pageId="2" pageNumber="3">
<emphasis id="A1FCEAE41F1F6659FC1F21C0B87429C8" box="[911,1040,669,694]" italics="true" pageId="2" pageNumber="3">Etymology</emphasis>
: This species is named after Dr. Francisco Villamar, in recognition of his many publications on polychaetes from
<collectingCountry id="EB9F76661F1F6659FA922180B90A2988" box="[1282,1390,733,758]" name="Ecuador" pageId="2" pageNumber="3">Ecuador</collectingCountry>
. The epithet is a noun in the genitive case.
</paragraph>
<paragraph id="933736F61F1F6658FC1F2040BEB32848" blockId="2.[850,1452,253,887]" lastBlockId="3.[136,738,253,886]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="A1FCEAE41F1F6659FC1F2040B8642848" box="[911,1024,797,822]" italics="true" pageId="2" pageNumber="3">Remarks</emphasis>
:
<taxonomicName id="54884D751F1F6659FB852040B95E2848" authorityName="Salazar-Vallejo" authorityYear="2017" box="[1045,1338,797,823]" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="3" phylum="Annelida" rank="species" species="villamari" status="sp. nov.">
<emphasis id="A1FCEAE41F1F6659FB852040B95E2848" box="[1045,1338,797,823]" italics="true" pageId="2" pageNumber="3">Caulleryaspis villamari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BACF579F1F1F6659FAD62040B9CF2848" box="[1350,1451,797,822]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
has a soft shield, not stiff or brittle as in
<taxonomicName id="54884D751F1F6659FABD2060BF852808" authority="Otto, 1821" authorityName="Otto" authorityYear="1821" class="Polychaeta" family="Sternaspidae" genus="Sternaspis" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="3" phylum="Annelida" rank="genus">
<emphasis id="A1FCEAE41F1F6659FABD2060B9CF2828" box="[1325,1451,829,854]" italics="true" pageId="2" pageNumber="3">Sternaspis</emphasis>
<bibRefCitation id="F7194B071F1F6659FCC22000BF852808" author="Otto AG" box="[850,993,861,887]" pageId="2" pageNumber="3" pagination="618 - 634" refId="ref11326" refString="Otto AG. 1821. Animalium maritimorum nondum editorum genera duo. Verh K Leopold-Carol Akad Naturf 10 (2): 618 - 634, Pls. 50 - 51." type="journal article" year="1821">Otto, 1821</bibRefCitation>
</taxonomicName>
. Because the shield is deprived of sediment particles, it resembles
<taxonomicName id="54884D751F1E6658FD9D23A0BE462A48" authority="(Zhadan, Tzetlin and Salazar-Vallejo 2017)" authorityName="Salazar-Vallejo" authorityYear="2017" baseAuthorityName="Zhadan, Tzetlin and Salazar-Vallejo" baseAuthorityYear="2017" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="nana" status="n. comb.">
<emphasis id="A1FCEAE41F1E6658FD9D23A0BE082A68" box="[525,620,253,278]" italics="true" pageId="3" pageNumber="4">C. nana</emphasis>
(
<bibRefCitation id="F7194B071F1E6658FDED23A0BE7F2A48" author="Zhadan AE &amp; Tzetlin AB &amp; Salazar-Vallejo SI" pageId="3" pageNumber="4" pagination="75 - 92" refId="ref12084" refString="Zhadan AE, Tzetlin AB, Salazar-Vallejo SI. 2017. Sternaspidae (Annelida, Sedentaria) from Vietnam with description of three new species and clarification of some morphological features. Zootaxa 4226: 75 - 92. doi: 10.11646 / zootaxa. 4226.1.3." type="journal article" year="2017">Zhadan, Tzetlin and Salazar-Vallejo 2017</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BACF579F1F1E6658FDBD2240BEF92A48" box="[557,669,285,310]" pageId="3" pageNumber="4" rank="species">n. comb.</taxonomicNameLabel>
, from
<collectingCountry id="EB9F76661F1E6658FF182260BC952A28" box="[136,241,317,342]" name="Vietnam" pageId="3" pageNumber="4">Vietnam</collectingCountry>
. Their main differences are indicated in the key below and include the color and presence of ribs in the shield; in
<taxonomicName id="54884D751F1E6658FE282220BE2C2AE8" authorityName="Salazar-Vallejo" authorityYear="2017" box="[440,584,381,406]" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="villamari">
<emphasis id="A1FCEAE41F1E6658FE282220BE2C2AE8" box="[440,584,381,406]" italics="true" pageId="3" pageNumber="4">C. villamari</emphasis>
</taxonomicName>
shields are yellowish to pale brown, with distinct ribs, whereas in
<taxonomicName id="54884D751F1E6658FF3522E0BD662AA8" authorityName="Salazar-Vallejo" authorityYear="2017" baseAuthorityName="Zhadan, Tzetlin and Salazar-Vallejo" baseAuthorityYear="2017" box="[165,258,445,470]" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="nana">
<emphasis id="A1FCEAE41F1E6658FF3522E0BD662AA8" box="[165,258,445,470]" italics="true" pageId="3" pageNumber="4">C. nana</emphasis>
</taxonomicName>
shields are red to brownish without ribs. The juveniles are included here but with some hesitation because they are from abyssal depths. They differ from the juveniles another regional species,
<taxonomicName id="54884D751F1E6658FF6E2160BD072908" authority="Salazar-Vallejo &amp; Buzhinskaja, 2013" authorityName="Salazar-Vallejo &amp; Buzhinskaja" authorityYear="2013" class="Polychaeta" family="Sternaspidae" genus="Sternaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="maureri">
<emphasis id="A1FCEAE41F1E6658FF6E2160BD902928" box="[254,500,573,598]" italics="true" pageId="3" pageNumber="4">Sternaspis maureri</emphasis>
<bibRefCitation id="F7194B071F1E6658FD912160BD072908" author="Salazar-Vallejo SI &amp; Buzhinskaja G." pageId="3" pageNumber="4" pagination="1 - 27" refId="ref11501" refString="Salazar-Vallejo SI, Buzhinskaja G. 2013. Six new deepwater sternaspid species (Annelida, Sternaspidae) from the Pacific Ocean. ZooKeys 348: 1 - 27. doi: 10.3897 / zookeys. 348.5449." type="journal article" year="2013">Salazar-Vallejo &amp; Buzhinskaja, 2013</bibRefCitation>
</taxonomicName>
because those herein regarded as related to
<taxonomicName id="54884D751F1E6658FEB72120BDC929E8" authorityName="Salazar-Vallejo" authorityYear="2017" box="[295,429,637,662]" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="villamari">
<emphasis id="A1FCEAE41F1E6658FEB72120BDC929E8" box="[295,429,637,662]" italics="true" pageId="3" pageNumber="4">C. villamari</emphasis>
</taxonomicName>
have orange shields with fans with small median notch, and poorly defined ribs, whereas in
<taxonomicName id="54884D751F1E6658FECF21E0BD8729A8" authorityName="Salazar-Vallejo &amp; Buzhinskaja" authorityYear="2013" box="[351,483,701,726]" class="Polychaeta" family="Sternaspidae" genus="Sternaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="maureri">
<emphasis id="A1FCEAE41F1E6658FECF21E0BD8729A8" box="[351,483,701,726]" italics="true" pageId="3" pageNumber="4">S. maureri</emphasis>
</taxonomicName>
shields are reddish with a distinct median notch, and ribs are better defined. Clarifying their affinities would need fresh material in order to assess their genetic attributes.
</paragraph>
<caption id="C7F7667E1F1F6659FF3F2413BF1E2CC8" ID-DOI="http://doi.org/10.5281/zenodo.12825858" ID-Zenodo-Dep="12825858" httpUri="https://zenodo.org/record/12825858/files/figure.png" pageId="2" pageNumber="3" startId="2.[175,209,1870,1890]" targetBox="[192,1435,936,1844]" targetPageId="2" targetType="figure">
<paragraph id="933736F61F1F6659FF3F2413BF1E2CC8" blockId="2.[174,1452,1869,1974]" pageId="2" pageNumber="3">
<emphasis id="A1FCEAE41F1F6659FF3F2413BC8B2C1C" bold="true" box="[175,239,1870,1890]" pageId="2" pageNumber="3">Fig. 1.</emphasis>
<taxonomicName id="54884D751F1F6659FF6E2410BDB22C1C" authorityName="Salazar-Vallejo" authorityYear="2017" box="[254,470,1869,1890]" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="3" phylum="Annelida" rank="species" species="villamari" status="sp. nov.">
<emphasis id="A1FCEAE41F1F6659FF6E2410BDB22C1C" box="[254,470,1869,1890]" italics="true" pageId="2" pageNumber="3">Caulleryaspis villamari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BACF579F1F1F6659FE4D2413BE4E2C1C" box="[477,554,1870,1890]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
(A) Holotype (USNM 1437645), ventral view. (B) Same, close-up of prostomium, mouth and first series of hooks. (C) Same, shield. (D) Same, branchial plate after removal of branchiae and interbranchial filaments. (E) Smaller specimens (USNM 1437646), ventral view. (F) Close-up of shield of smallest one. (G) Close-up of shield of largest one. Scale bars: A = 2.7 mm, B = 0.6 mm, C, D = 1.3 mm, E = 2.8 mm, F = 0.4 mm, G = 0.7 mm.
</paragraph>
</caption>
<footnote id="F0932AF81F1F6659FB0C2480B9CF2C91" box="[1180,1451,2013,2031]" pageId="2" pageNumber="3">
<paragraph id="933736F61F1F6659FB0C2480B9CF2C91" blockId="2.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="2" pageNumber="3">
© 2017 Academia Sinica,
<collectingCountry id="EB9F76661F1F6659FAE12480B9CF2C91" box="[1393,1451,2013,2031]" name="Taiwan" pageId="2" pageNumber="3">Taiwan</collectingCountry>
</paragraph>
</footnote>
<paragraph id="933736F61F1E6658FF552060BCB72808" blockId="3.[136,738,253,886]" pageId="3" pageNumber="4">
<emphasis id="A1FCEAE41F1E6658FF552060BD3E2828" box="[197,346,829,854]" italics="true" pageId="3" pageNumber="4">Distribution</emphasis>
: Off northern
<collectingCountry id="EB9F76661F1E6658FD8A2060BE382828" box="[538,604,829,854]" name="Peru" pageId="3" pageNumber="4">Peru</collectingCountry>
, in
<quantity id="54709B131F1E6658FD1F2060BE842828" box="[655,736,829,854]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="4" unit="m" value="160.0">160 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="DB92657D1F1E6658FF1820C0BE172DB5" pageId="3" pageNumber="4" type="key">
<paragraph id="933736F61F1E6658FF1820C0BDE928A8" blockId="3.[136,665,924,1014]" pageId="3" pageNumber="4">
<emphasis id="A1FCEAE41F1E6658FF1820C0BDE928A8" bold="true" pageId="3" pageNumber="4">
Key to species of
<taxonomicName id="54884D751F1E6658FEFB20C1BDE928A8" ID-CoL="84N6X" authority="Sendall &amp; Salazar-Vallejo, 2013" authorityName="Sendall &amp; Salazar-Vallejo" authorityYear="2013" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="A1FCEAE41F1E6658FEFB20C1BE7C28C8" bold="true" box="[363,536,924,950]" italics="true" pageId="3" pageNumber="4">Caulleryaspis</emphasis>
<bibRefCitation id="F7194B071F1E6658FDB020C1BDE928A8" author="Sendall K &amp; Salazar-Vallejo SI" pageId="3" pageNumber="4" pagination="1 - 74" refId="ref11584" refString="Sendall K, Salazar-Vallejo SI. 2013. Revision of Sternaspis Otto, 1821 (Polychaeta: Sternaspidae). ZooKeys 286: 1 - 74. doi: https: // doi. org / 10.3897 / zookeys. 286.4438." type="journal article" year="2013">Sendall &amp; Salazar-Vallejo, 2013</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="933736F61F1E6658FF182080BEEA2888" blockId="3.[136,665,924,1014]" box="[136,654,988,1014]" pageId="3" pageNumber="4">
(modif.
<bibRefCitation id="F7194B071F1E6658FF712081BEEC2888" author="Salazar-Vallejo SI &amp; Buzhinskaja G." box="[225,648,988,1014]" pageId="3" pageNumber="4" pagination="1 - 27" refId="ref11501" refString="Salazar-Vallejo SI, Buzhinskaja G. 2013. Six new deepwater sternaspid species (Annelida, Sternaspidae) from the Pacific Ocean. ZooKeys 348: 1 - 27. doi: 10.3897 / zookeys. 348.5449." type="journal article" year="2013">Salazar-Vallejo &amp; Buzhinskaja 2013</bibRefCitation>
)
</paragraph>
<paragraph id="933736F61F1E6658FF182741BE852F35" blockId="3.[136,739,1052,1739]" pageId="3" pageNumber="4">1. Shield with sediment particles firmly adhered; shield surface not visible ........................................................... 2</paragraph>
<paragraph id="933736F61F1E6658FF18270CBE852FFE" blockId="3.[136,739,1052,1739]" pageId="3" pageNumber="4">- Shield without firmly adhered sediment particles; shield surface visible ................................................................. 4</paragraph>
<paragraph id="933736F61F1E6658FF1827DABE852FCB" blockId="3.[136,739,1052,1739]" pageId="3" pageNumber="4">2(1). Shield with anterior depression deep; peg chaetae robust ......................................................................................... 3</paragraph>
<paragraph id="933736F61F1E6658FF1827E1BE852F95" blockId="3.[136,739,1052,1739]" pageId="3" pageNumber="4">
- Shield with anterior depression shallow; peg chaetae indistinct .................
<taxonomicName id="54884D751F1E6658FE12278BBE142F95" ID-CoL="RX8N" authority="(Caullery, 1944)" baseAuthorityName="Caullery" baseAuthorityYear="1944" box="[386,624,1238,1259]" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="laevis">
<emphasis id="A1FCEAE41F1E6658FE12278BBDB12F95" box="[386,469,1238,1259]" italics="true" pageId="3" pageNumber="4">C. laevis</emphasis>
(
<bibRefCitation id="F7194B071F1E6658FE4F278BBE0F2F95" author="Caullery M." box="[479,619,1238,1259]" pageId="3" pageNumber="4" pagination="1 - 204" refId="ref10835" refString="Caullery M. 1944. Polychetes sedentaires de l'Expedition du Siboga: Ariciidae, Spionidae, Chaetopteridae, Chlorhaemidae, Opheliidae, Oweniidae, Sabellaridae, Sternaspidae, Amphictenidae, Ampharetidae, Terebellidae. Siboga-Expeditie, Leiden 24 (2 bis): 1 - 204." type="journal article" year="1944">Caullery, 1944</bibRefCitation>
)
</taxonomicName>
(
<collectingCountry id="EB9F76661F1E6658FDEB278ABEBF2F95" box="[635,731,1239,1259]" name="Indonesia" pageId="3" pageNumber="4">Indonesia</collectingCountry>
)
</paragraph>
<paragraph id="933736F61F1E6658FF1827ACBD3F2E2B" blockId="3.[136,739,1052,1739]" pageId="3" pageNumber="4">
3(2). Shield with anterior margins angular; peg chaetae forming thick, large spines ..............................................................
<taxonomicName id="54884D751F1E6658FF2D267BBEFA2E45" authority="Sendall &amp; Salazar-Vallejo, 2013" authorityName="Sendall &amp; Salazar-Vallejo" authorityYear="2013" box="[189,670,1318,1339]" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspsis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="gudmundssoni">
C. gudmundssoni
<bibRefCitation id="F7194B071F1E6658FEFD267BBEFA2E45" author="Sendall K &amp; Salazar-Vallejo SI" box="[365,670,1318,1339]" pageId="3" pageNumber="4" pagination="1 - 74" refId="ref11584" refString="Sendall K, Salazar-Vallejo SI. 2013. Revision of Sternaspis Otto, 1821 (Polychaeta: Sternaspidae). ZooKeys 286: 1 - 74. doi: https: // doi. org / 10.3897 / zookeys. 286.4438." type="journal article" year="2013">Sendall &amp; Salazar-Vallejo, 2013</bibRefCitation>
</taxonomicName>
(North Atlantic,
<collectingCountry id="EB9F76661F1E6658FE9F261CBD322E2B" box="[271,342,1345,1365]" name="Iceland" pageId="3" pageNumber="4">Iceland</collectingCountry>
)
</paragraph>
<paragraph id="933736F61F1E6658FF182601BDE72EBE" blockId="3.[136,739,1052,1739]" pageId="3" pageNumber="4">
- Shield with anterior margins rounded; peg chaetae forming thin, small spines ..............................
<taxonomicName id="54884D751F1E6658FDE0262BBE692EDB" ID-CoL="RX8L" authority="Salazar-Vallejo &amp; Buzhinskaja 2013" authorityName="Salazar-Vallejo &amp; Buzhinskaja" authorityYear="2013" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="fauchaldi">
<emphasis id="A1FCEAE41F1E6658FDE0262BBE852EF5" box="[624,737,1398,1419]" italics="true" pageId="3" pageNumber="4">C. fauchaldi</emphasis>
<bibRefCitation id="F7194B071F1E6658FF2D26CCBE692EDB" author="Salazar-Vallejo SI &amp; Buzhinskaja G." box="[189,525,1425,1445]" pageId="3" pageNumber="4" pagination="1 - 27" refId="ref11501" refString="Salazar-Vallejo SI, Buzhinskaja G. 2013. Six new deepwater sternaspid species (Annelida, Sternaspidae) from the Pacific Ocean. ZooKeys 348: 1 - 27. doi: 10.3897 / zookeys. 348.5449." type="journal article" year="2013">Salazar-Vallejo &amp; Buzhinskaja 2013</bibRefCitation>
</taxonomicName>
(Northeastern Pacific,
<collectingRegion id="514CF8141F1E6658FF2D26F1BD602EBE" box="[189,260,1452,1472]" country="United States of America" name="Oregon" pageId="3" pageNumber="4">Oregon</collectingRegion>
to
<collectingRegion id="514CF8141F1E6658FEB226F1BD1A2EBE" box="[290,382,1452,1472]" country="United States of America" name="California" pageId="3" pageNumber="4">California</collectingRegion>
)
</paragraph>
<paragraph id="933736F61F1E6658FF18269ABD172D54" blockId="3.[136,739,1052,1739]" pageId="3" pageNumber="4">
4(1). Shield with lateral margins rounded, medially expanded, ribs barely visible ...............................................................
<taxonomicName id="54884D751F1E6658FF2D26A1BE3E2D6E" ID-CoL="RX8P" authority="Salazar-Vallejo &amp; Buzhinskaja 2013" authorityName="Salazar-Vallejo &amp; Buzhinskaja" authorityYear="2013" box="[189,602,1532,1552]" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="nuda">
<emphasis id="A1FCEAE41F1E6658FF2D26A1BD632D6E" box="[189,263,1532,1552]" italics="true" pageId="3" pageNumber="4">C. nuda</emphasis>
<bibRefCitation id="F7194B071F1E6658FE9D26A1BE3E2D6E" author="Salazar-Vallejo SI &amp; Buzhinskaja G." box="[269,602,1532,1552]" pageId="3" pageNumber="4" pagination="1 - 27" refId="ref11501" refString="Salazar-Vallejo SI, Buzhinskaja G. 2013. Six new deepwater sternaspid species (Annelida, Sternaspidae) from the Pacific Ocean. ZooKeys 348: 1 - 27. doi: 10.3897 / zookeys. 348.5449." type="journal article" year="2013">Salazar-Vallejo &amp; Buzhinskaja 2013</bibRefCitation>
</taxonomicName>
(Northeastern Pacific, off
<collectingRegion id="514CF8141F1E6658FEB5254BBD0A2D54" box="[293,366,1558,1578]" country="United States of America" name="Oregon" pageId="3" pageNumber="4">Oregon</collectingRegion>
)
</paragraph>
<paragraph id="933736F61F1E6658FF18256CBE852D3B" blockId="3.[136,739,1052,1739]" box="[136,737,1585,1605]" pageId="3" pageNumber="4">- Shield with lateral margins straight to barely curved ...... 5</paragraph>
<paragraph id="933736F61F1E6658FF182511BD3D2DEB" blockId="3.[136,739,1052,1739]" pageId="3" pageNumber="4">
5(4). Ribs indistinct, shield dark red to brownish .......................
<taxonomicName id="54884D751F1E6658FF2D253BBEA12D05" ID-CoL="8W8HK" authority="(Zhadan, Tzetlin &amp; Salazar-Vallejo, 2017)" authorityName="Salazar-Vallejo" authorityYear="2017" baseAuthorityName="Zhadan, Tzetlin &amp; Salazar-Vallejo" baseAuthorityYear="2017" box="[189,709,1638,1659]" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="nana" status="comb. nov.">
C. nana (
<bibRefCitation id="F7194B071F1E6658FE8E253ABEA42D05" author="Zhadan AE &amp; Tzetlin AB &amp; Salazar-Vallejo SI" box="[286,704,1638,1659]" pageId="3" pageNumber="4" pagination="75 - 92" refId="ref12084" refString="Zhadan AE, Tzetlin AB, Salazar-Vallejo SI. 2017. Sternaspidae (Annelida, Sedentaria) from Vietnam with description of three new species and clarification of some morphological features. Zootaxa 4226: 75 - 92. doi: 10.11646 / zootaxa. 4226.1.3." type="journal article" year="2017">Zhadan, Tzetlin &amp; Salazar-Vallejo, 2017</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BACF579F1F1E6658FD5E253ABC9D2DEB" pageId="3" pageNumber="4" rank="species">n. comb.</taxonomicNameLabel>
(
<collectingCountry id="EB9F76661F1E6658FE9225DCBD302DEB" box="[258,340,1665,1685]" name="Vietnam" pageId="3" pageNumber="4">Vietnam</collectingCountry>
)
</paragraph>
<paragraph id="933736F61F1E6658FF1825C1BE172DB5" blockId="3.[136,739,1052,1739]" pageId="3" pageNumber="4">
- Ribs distinct, shield yellowish to pale brown ......................
<taxonomicName id="54884D751F1E6658FF2D25EBBD422DB5" ID-CoL="85RBJ" authorityName="Salazar-Vallejo" authorityYear="2017" box="[189,294,1718,1739]" class="Polychaeta" family="Sternaspidae" genus="Caulleryaspis" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="villamari" status="sp. nov.">
<emphasis id="A1FCEAE41F1E6658FF2D25EBBD422DB5" box="[189,294,1718,1739]" italics="true" pageId="3" pageNumber="4">C. villamari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BACF579F1F1E6658FEBC25EABD132DB5" box="[300,375,1719,1739]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
(Eastern Pacific,
<collectingCountry id="EB9F76661F1E6658FD8D25EABE0B2DB5" box="[541,623,1719,1739]" name="Ecuador" pageId="3" pageNumber="4">Ecuador</collectingCountry>
)
</paragraph>
</subSubSection>
</treatment>
</document>