176 lines
20 KiB
XML
176 lines
20 KiB
XML
<document id="63A83347CACFFB3043B0EE2D909DC0EE" ID-CLB-Dataset="2636" ID-DOI="10.11646/zootaxa.5091.1.1" ID-GBIF-Dataset="51b42715-ddb1-4b4a-aad4-8be0f600a287" ID-ISSN="1175-5326" ID-Zenodo-Dep="5840434" ID-ZooBank="ABC8AF70-F691-4D07-8F20-70934642C8BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1641986550580" checkinUser="plazi" docAuthor="Bergh, Eugene W. & Compton, John S." docDate="2022" docId="197787BAFFDF93227FC99B9BFAA1FC2D" docLanguage="en" docName="zootaxa.5091.1.1.pdf" docOrigin="Zootaxa 5091 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Plectofrondicularia undefined-b" docType="treatment" docVersion="7" lastPageNumber="13" masterDocId="E54EFFC2FFD3932E7F5E9B0CFFC4FF88" masterDocTitle="Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf" masterLastPageNumber="55" masterPageNumber="1" pageNumber="13" updateTime="1699301522325" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="7D13B12698B401F475AB6A31C30540DF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="873127F467C73F85FE25F11DB07485B6">
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<mods:title id="D250A11C964649F4299CD7012B3FB985">Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf</mods:title>
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</mods:titleInfo>
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<mods:name id="45F0EC678BBAA66E077D96F757F3FE9E" type="personal">
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<mods:role id="AA7719120276E1A3ACB1B7852C83AE41">
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<mods:roleTerm id="B380AC345CBB19725EE7F55282DB64FA">Author</mods:roleTerm>
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<mods:namePart id="9A7237A665616734B8D2CD89779B07C7">Bergh, Eugene W.</mods:namePart>
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<mods:nameIdentifier id="6B3AFD09E88690D70333F32278D620D8" type="ORCID">0000-0002-0765-4141</mods:nameIdentifier>
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<mods:affiliation id="DB0D61D0D1284226D8D0A7876CA441F0">Marine Research Institute and Department of Geological Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa. & Department of Environmental and Geographical Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa; & Invertebrate palaeontology and Geology, Iziko South African Museum, P. O. Box 61, Cape Town 8000, South Africa. john. compton @ uct. ac. za; https: // orcid. org / 0000 - 0002 - 0765 - 4141</mods:affiliation>
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<mods:nameIdentifier id="BF8EFA5780A912F00DBEC45AA3D111AE" type="email">john.compton@uct.ac.za</mods:nameIdentifier>
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<mods:roleTerm id="01ADE12F5CB8C05A4DF6E73BB6C3E98B">Author</mods:roleTerm>
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<mods:namePart id="1A296E710A2AEB6CD2579CD591DB3658">Compton, John S.</mods:namePart>
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<mods:affiliation id="308540B8D8C7E3B06649E9248D06F647">Marine Research Institute and Department of Geological Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa.</mods:affiliation>
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<mods:typeOfResource id="61786780F98449974FE79E5DB56BB519">text</mods:typeOfResource>
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<mods:title id="34AE46709C721D8D364B5733D048F312">Zootaxa</mods:title>
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<mods:part id="BC95737A50D8085CB53505E00596047E">
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<mods:date id="6E58237FBF184288DD30DB9248F36D81">2022</mods:date>
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<mods:number id="D4BDBCD2159151C64F3CCF1001C82823">2022-01-12</mods:number>
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<mods:number id="F1CB93CEF4F14677D56AFC32B1D38674">5091</mods:number>
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<mods:identifier id="2AE18B52EF5CD4352EAAFA290034201C" type="DOI">10.11646/zootaxa.5091.1.1</mods:identifier>
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<treatment id="197787BAFFDF93227FC99B9BFAA1FC2D" ID-DOI="http://doi.org/10.5281/zenodo.5842160" ID-GBIF-Taxon="192044087" ID-Zenodo-Dep="5842160" LSID="urn:lsid:plazi:treatment:197787BAFFDF93227FC99B9BFAA1FC2D" httpUri="http://treatment.plazi.org/id/197787BAFFDF93227FC99B9BFAA1FC2D" lastPageNumber="13" pageId="12" pageNumber="13">
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<subSubSection id="D9C46527FFDF93227FC99B9BFEF8FF5D" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="916136ACFFDF93227FC99B9BFE0DFF39" blockId="12.[151,457,151,213]" box="[151,457,151,178]" pageId="12" pageNumber="13">
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||
<heading id="CA2981C0FFDF93227FC99B9BFE0DFF39" bold="true" box="[151,457,151,178]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
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<taxonomicName id="56DE4D2FFFDF93227FC99B9BFE0DFF39" box="[151,457,151,178]" class="Nodosariata" family="Plectofrondiculariidae" genus="Plectofrondicularia" kingdom="Chromista" order="Polymorphinida" pageId="12" pageNumber="13" phylum="Foraminifera" rank="species" species="undefined-b">
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<emphasis id="A3AAEABEFFDF93227FC99B9BFE0DFF39" bold="true" box="[151,457,151,178]" pageId="12" pageNumber="13">
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<emphasis id="A3AAEABEFFDF93227FC99B9BFE47FF3A" bold="true" box="[151,387,151,178]" italics="true" pageId="12" pageNumber="13">Plectofrondicularia</emphasis>
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sp. B
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="916136ACFFDF93227FC99BB7FEF8FF5D" blockId="12.[151,457,151,213]" box="[151,316,187,213]" pageId="12" pageNumber="13">
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<heading id="CA2981C0FFDF93227FC99BB7FEF8FF5D" bold="true" box="[151,316,187,213]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
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<emphasis id="A3AAEABEFFDF93227FC99BB7FEF8FF5D" bold="true" box="[151,316,187,213]" pageId="12" pageNumber="13">Pl. 2, figs. 1–2</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="D9C46527FFDF93227FC99A0FFB13FE01" pageId="12" pageNumber="13" type="description">
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<paragraph id="916136ACFFDF93227FC99A0FFB13FE01" blockId="12.[151,1437,259,934]" pageId="12" pageNumber="13">
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<emphasis id="A3AAEABEFFDF93227FC99A0FFEEFFE95" bold="true" box="[151,299,259,285]" pageId="12" pageNumber="13">Description:</emphasis>
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The test wall is calcareous. The test is flat with a keeled periphery and biserial in the early stage, becoming uniserial as chambers are added. The later chambers are chevron-shaped. The side abruptly widens and curves where it narrows towards the apertural end. The initial stage protrudes more than the later stage chambers. The sutures are limbate and the aperture terminal, radial with projecting laminae fusing centrally.
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</paragraph>
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</subSubSection>
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<subSubSection id="D9C46527FFDF93227F999A9FFAA1FC2D" pageId="12" pageNumber="13" type="discussion">
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<paragraph id="916136ACFFDF93227F999A9FFEF8FE5A" blockId="12.[151,1437,259,934]" pageId="12" pageNumber="13">
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<emphasis id="A3AAEABEFFDF93227F999A9FFEF9FE25" bold="true" box="[199,317,403,429]" pageId="12" pageNumber="13">Remarks:</emphasis>
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The tests are large in size (up to
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<quantity id="56269B49FFDF93227DED9A9FFD30FE26" box="[691,756,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="mm" value="1.0">1 mm</quantity>
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in width and
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<quantity id="56269B49FFDF93227CD19A9FFC15FE26" box="[911,977,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="13" unit="mm" value="3.0">3 mm</quantity>
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in length), in trace abundances (<1%) in all three cores.
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</paragraph>
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<paragraph id="916136ACFFDF93227F999AD7FED1FD21" blockId="12.[151,1437,259,934]" pageId="12" pageNumber="13">
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The tests range between narrow (pl. 2, fig. 1) and broad (pl. 2, fig. 2). The tests are mostly broken and resemble
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<taxonomicName id="56DE4D2FFFDF93227FC99AF3FE5AFD91" box="[151,414,511,537]" class="Nodosariata" family="Nodosariidae" genus="Frondicularia" kingdom="Chromista" order="Nodosariida" pageId="12" pageNumber="13" phylum="Foraminifera" rank="species" species="sagittula">
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<emphasis id="A3AAEABEFFDF93227FC99AF3FE5AFD91" box="[151,414,511,537]" italics="true" pageId="12" pageNumber="13">Frondicularia sagittula</emphasis>
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</taxonomicName>
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in
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<bibRefCitation id="F54F4B5DFFDF93227E9D990CFD89FD91" author="Jones, R. W." box="[451,589,511,538]" pageId="12" pageNumber="13" refId="ref33343" refString="Jones, R. W. (1994) The Challenger Foraminifera. Oxford University Press, Oxford, 149 pp." type="book" year="1994">Jones (1994</bibRefCitation>
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; pl. 65, fig. 23) and
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<taxonomicName id="56DE4D2FFFDF93227C6F9AF3FBB9FD91" authorityName="Cushman" authorityYear="1927" box="[817,1149,511,537]" class="Nodosariata" family="Plectofrondiculariidae" genus="Plectofrondicularia" kingdom="Chromista" order="Polymorphinida" pageId="12" pageNumber="13" phylum="Foraminifera" rank="species" species="vaughani">
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<emphasis id="A3AAEABEFFDF93227C6F9AF3FBB9FD91" box="[817,1149,511,537]" italics="true" pageId="12" pageNumber="13">Plectofrondicularia vaughani</emphasis>
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</taxonomicName>
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in
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<bibRefCitation id="F54F4B5DFFDF93227BFF9AF3FA5CFD91" author="Holbourn, A. & Henderson, A. S. & MacLeod, N." box="[1185,1432,511,537]" pageId="12" pageNumber="13" refId="ref32986" refString="Holbourn, A., Henderson, A. S. & MacLeod, N. (2013) Atlas of Benthic Foraminifera. Wiley-Blackwell, Oxford, 654 pp. https: // doi. org / 10.1002 / 9781118452493" type="book" year="2013">
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Holbourn
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<emphasis id="A3AAEABEFFDF93227A4A990CFA89FD91" box="[1300,1357,511,537]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
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||
(2013
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</bibRefCitation>
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, p. 422), but with a peripheral keel and the absence of a basal spine. In this study, the specimens have been classified as
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<taxonomicName id="56DE4D2FFFDF93227FE6994BFE50FDE9" authorityName="Liebus" authorityYear="1902" box="[184,404,583,609]" class="Nodosariata" family="Plectofrondiculariidae" genus="Plectofrondicularia" kingdom="Chromista" order="Polymorphinida" pageId="12" pageNumber="13" phylum="Foraminifera" rank="genus">
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<emphasis id="A3AAEABEFFDF93227FE6994BFE50FDE9" box="[184,404,583,609]" italics="true" pageId="12" pageNumber="13">Plectofrondicularia</emphasis>
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</taxonomicName>
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based on the presence of a keeled margin (
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<bibRefCitation id="F54F4B5DFFDF93227CCE994BFB79FDE9" author="Loeblich, A. R. & Tappan, H." box="[912,1213,583,610]" pageId="12" pageNumber="13" refId="ref34837" refString="Loeblich, A. R. & Tappan, H. (1988) Foraminiferal Genera and their Classification. Van Nostrand Reinhold, New York, 970 pp. https: // doi. org / 10.1007 / 978 - 1 - 4899 - 5760 - 3" type="book" year="1988">Loeblich & Tappan, 1988</bibRefCitation>
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). The specimens in this study were not assigned to
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<emphasis id="A3AAEABEFFDF93227D5E9960FD48FD0D" box="[512,652,619,645]" italics="true" pageId="12" pageNumber="13">
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<taxonomicName id="56DE4D2FFFDF93227D5E9960FD4CFD0D" authorityName="Cushman" authorityYear="1927" box="[512,648,619,645]" class="Nodosariata" family="Plectofrondiculariidae" genus="Plectofrondicularia" kingdom="Chromista" order="Polymorphinida" pageId="12" pageNumber="13" phylum="Foraminifera" rank="species" species="vaughani">P. vaughani</taxonomicName>
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,
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</emphasis>
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because of its broader peripheral keel and thicker, more pronounced proloculus.
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</paragraph>
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<paragraph id="916136ACFFDF93227F9999BFFDBDFD7A" blockId="12.[151,1437,259,934]" pageId="12" pageNumber="13">
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<emphasis id="A3AAEABEFFDF93227F9999BFFE98FD45" bold="true" box="[199,348,691,717]" pageId="12" pageNumber="13">Life strategy</emphasis>
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:
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<taxonomicName id="56DE4D2FFFDF93227E3799BFFDBDFD46" box="[361,633,691,718]" class="Nodosariata" family="Plectofrondiculariidae" genus="Plectofrondicularia" kingdom="Chromista" order="Polymorphinida" pageId="12" pageNumber="13" phylum="Foraminifera" rank="species" species="undetermined">
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<emphasis id="A3AAEABEFFDF93227E3799BFFD81FD45" box="[361,581,691,717]" italics="true" pageId="12" pageNumber="13">Plectofrondicularia</emphasis>
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spp.
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</taxonomicName>
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have been documented to occur as infaunal and under suboxic conditions (
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<bibRefCitation id="F54F4B5DFFDF93227FC199DBFEAEFD79" author="Pezelj, D & Mandic, O. & Coric, S." box="[159,362,727,754]" pageId="12" pageNumber="13" pagination="81 - 100" refId="ref36415" refString="Pezelj, D, Mandic, O. & Coric, S. (2013) Paleoenvironmental dynamics in the southern Pannonian Basin during initial Middle Miocene marine flooding. Geologica Carpathica, 64, 81 - 100." type="journal article" year="2013">
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Pezelj
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<emphasis id="A3AAEABEFFDF93227FB299D4FED9FD79" box="[236,285,727,753]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
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||
., 2013
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</bibRefCitation>
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and references therein).
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</paragraph>
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<paragraph id="916136ACFFDF93227F9999F7FAA1FC2D" blockId="12.[151,1437,259,934]" pageId="12" pageNumber="13">
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<emphasis id="A3AAEABEFFDF93227F9999F7FE0CFC9D" bold="true" box="[199,456,763,789]" pageId="12" pageNumber="13">Regional occurrence:</emphasis>
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This species occurs in middle Miocene sediments from the northern Namibian outer continental shelf, south of the
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<collectingRegion id="531AF84EFFDF93227EB5982CFD86FCB2" box="[491,578,800,826]" country="Namibia" name="Kunene" pageId="12" pageNumber="13">Kunene</collectingRegion>
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<collectingRegion id="531AF84EFFDF93227D179813FD43FCB1" box="[585,647,799,825]" country="Nigeria" name="Rivers" pageId="12" pageNumber="13">River</collectingRegion>
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mouth (this study).
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<bibRefCitation id="F54F4B5DFFDF93227C379813FBFAFCB1" author="Wefer, G. & Berger, W. H. & Richter, C. & Adams, D. D. & Anderon, L. D. & Andreasen, D. J. & Bruchert, V. & Cambray, H. & Christensen, B. A. & Frost, G. M. & Giraudeau, J. & Gorgas, T. J. & Hermelin, O. & Lange, C. B. & Laser, B. & Lin, H-L. & Maslin, M. & Meyers, P. A. & Motoyama, I. & Murray, R. W. & Pato, D. & Perez, M. E. & Pufahl, P. K. & Spiess, V. & Wigley, R. & Yamazaki, T." box="[873,1086,799,826]" pageId="12" pageNumber="13" refId="ref38965" refString="Wefer, G., Berger, W. H., Richter, C., Adams, D. D., Anderon, L. D., Andreasen, D. J., Bruchert, V., Cambray, H., Christensen, B. A., Frost, G. M., Giraudeau, J., Gorgas, T. J., Hermelin, O., Lange, C. B., Laser, B., Lin, H-L., Maslin, M., Meyers, P. A., Motoyama, I., Murray, R. W., Pato, D., Perez, M. E., Pufahl, P. K., Spiess, V., Vidal. L., Wigley, R. & Yamazaki, T. (1998) Proceedings of the Ocean Drilling Program, Initial Reports. Vol. 175. Ocean Drilling Program, College Station, Texas, 577 pp." type="book" year="1998">
|
||
Wefer
|
||
<emphasis id="A3AAEABEFFDF93227CEA982CFC21FCB1" box="[948,997,799,825]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
|
||
. (1998)
|
||
</bibRefCitation>
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||
recorded minor occurrences of
|
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<taxonomicName id="56DE4D2FFFDF93227FC9984FFE60FCD6" box="[151,420,835,862]" class="Nodosariata" family="Plectofrondiculariidae" genus="Plectofrondicularia" kingdom="Chromista" order="Polymorphinida" pageId="12" pageNumber="13" phylum="Foraminifera" rank="species" species="undetermined">
|
||
<emphasis id="A3AAEABEFFDF93227FC9984FFEB7FCD5" box="[151,371,835,861]" italics="true" pageId="12" pageNumber="13">Plectofrondicularia</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
(
|
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<taxonomicName id="56DE4D2FFFDF93227EEE984FFCE7FCD5" baseAuthorityName="Costa" baseAuthorityYear="1855" box="[432,803,835,861]" class="Nodosariata" family="Plectofrondiculariidae" genus="Plectofrondicularia" kingdom="Chromista" order="Polymorphinida" pageId="12" pageNumber="13" phylum="Foraminifera" rank="species" species="inaequalis">
|
||
<emphasis id="A3AAEABEFFDF93227EEE984FFD48FCD5" box="[432,652,835,861]" italics="true" pageId="12" pageNumber="13">Plectofrondicularia</emphasis>
|
||
cf.
|
||
<emphasis id="A3AAEABEFFDF93227DF0984FFCE7FCD5" box="[686,803,835,861]" italics="true" pageId="12" pageNumber="13">inaequalis</emphasis>
|
||
</taxonomicName>
|
||
,
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||
<taxonomicName id="56DE4D2FFFDF93227C73984FFB56FCD5" box="[813,1170,835,861]" class="Nodosariata" family="Plectofrondiculariidae" genus="Plectofrondicularia" kingdom="Chromista" order="Polymorphinida" pageId="12" pageNumber="13" phylum="Foraminifera" rank="species" species="raricosta">
|
||
<emphasis id="A3AAEABEFFDF93227C73984FFBCDFCD5" box="[813,1033,835,861]" italics="true" pageId="12" pageNumber="13">Plectofrondicularia</emphasis>
|
||
cf.
|
||
<emphasis id="A3AAEABEFFDF93227B759848FB56FCD5" box="[1067,1170,836,861]" italics="true" pageId="12" pageNumber="13">raricosta</emphasis>
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</taxonomicName>
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and
|
||
<taxonomicName id="56DE4D2FFFDF93227B9E984FFEEEFC09" class="Nodosariata" family="Plectofrondiculariidae" genus="Plectofrondicularia" kingdom="Chromista" order="Polymorphinida" pageId="12" pageNumber="13" phylum="Foraminifera" rank="species" species="semicosta">
|
||
<emphasis id="A3AAEABEFFDF93227B9E984FFA58FCD5" box="[1216,1436,835,861]" italics="true" pageId="12" pageNumber="13">Plectofrondicularia</emphasis>
|
||
cf.
|
||
<emphasis id="A3AAEABEFFDF93227FE29864FEEEFC09" box="[188,298,872,897]" italics="true" pageId="12" pageNumber="13">semicosta</emphasis>
|
||
</taxonomicName>
|
||
) in
|
||
<taxonomicName id="56DE4D2FFFDF93227E07986BFE2FFC09" authority="Miocene" authorityName="Miocene" box="[345,491,871,897]" class="Insecta" family="Cetoniidae" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="late">late Miocene</taxonomicName>
|
||
to Pleistocene-aged sediments along the Namibian and southwestern South African slope. The highest abundances (<10%) were recorded along the northern Namibian slope (
|
||
<bibRefCitation id="F54F4B5DFFDF93227BD49887FA9DFC2D" author="Wefer, G. & Berger, W. H. & Richter, C. & Adams, D. D. & Anderon, L. D. & Andreasen, D. J. & Bruchert, V. & Cambray, H. & Christensen, B. A. & Frost, G. M. & Giraudeau, J. & Gorgas, T. J. & Hermelin, O. & Lange, C. B. & Laser, B. & Lin, H-L. & Maslin, M. & Meyers, P. A. & Motoyama, I. & Murray, R. W. & Pato, D. & Perez, M. E. & Pufahl, P. K. & Spiess, V. & Wigley, R. & Yamazaki, T." box="[1162,1369,907,934]" pageId="12" pageNumber="13" refId="ref38965" refString="Wefer, G., Berger, W. H., Richter, C., Adams, D. D., Anderon, L. D., Andreasen, D. J., Bruchert, V., Cambray, H., Christensen, B. A., Frost, G. M., Giraudeau, J., Gorgas, T. J., Hermelin, O., Lange, C. B., Laser, B., Lin, H-L., Maslin, M., Meyers, P. A., Motoyama, I., Murray, R. W., Pato, D., Perez, M. E., Pufahl, P. K., Spiess, V., Vidal. L., Wigley, R. & Yamazaki, T. (1998) Proceedings of the Ocean Drilling Program, Initial Reports. Vol. 175. Ocean Drilling Program, College Station, Texas, 577 pp." type="book" year="1998">
|
||
Wefer
|
||
<emphasis id="A3AAEABEFFDF93227B899880FACCFC2D" box="[1239,1288,907,933]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
|
||
., 1998
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |