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<document id="62F71A81C368AF82651AD2E6C7B44AD6" ID-CLB-Dataset="2636" ID-DOI="10.11646/zootaxa.5091.1.1" ID-GBIF-Dataset="51b42715-ddb1-4b4a-aad4-8be0f600a287" ID-ISSN="1175-5326" ID-Zenodo-Dep="5840434" ID-ZooBank="ABC8AF70-F691-4D07-8F20-70934642C8BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1641986550580" checkinUser="plazi" docAuthor="Bergh, Eugene W. &amp; Compton, John S." docDate="2022" docId="197787BAFFDC93217FC99BD3FCA6F9DE" docLanguage="en" docName="zootaxa.5091.1.1.pdf" docOrigin="Zootaxa 5091 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marginulinopsis costata" docType="treatment" docVersion="6" lastPageNumber="16" masterDocId="E54EFFC2FFD3932E7F5E9B0CFFC4FF88" masterDocTitle="Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf" masterLastPageNumber="55" masterPageNumber="1" pageNumber="16" updateTime="1699301522325" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2FD6AF916AC7FE7A4A7CA5AEE1A81B33">Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf</mods:title>
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<mods:namePart id="D8B00AB6C99C4A696C6FB6E6F0BA89E9">Bergh, Eugene W.</mods:namePart>
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<mods:affiliation id="8E0278B05FAC01F78449AD380EC5DC68">Marine Research Institute and Department of Geological Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa. &amp; Department of Environmental and Geographical Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa; &amp; Invertebrate palaeontology and Geology, Iziko South African Museum, P. O. Box 61, Cape Town 8000, South Africa. john. compton @ uct. ac. za; https: // orcid. org / 0000 - 0002 - 0765 - 4141</mods:affiliation>
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<paragraph id="916136ACFFDC93217FC99BD3FDB0FF71" blockId="15.[151,628,223,285]" box="[151,628,223,250]" pageId="15" pageNumber="16">
<heading id="CA2981C0FFDC93217FC99BD3FDB0FF71" bold="true" box="[151,628,223,250]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName id="56DE4D2FFFDC93217FC99BD3FDB0FF71" ID-CoL="3Y5ZZ" authority="(Batsch, 1791)" baseAuthorityName="Batsch" baseAuthorityYear="1791" box="[151,628,223,250]" class="Nodosariata" family="Vaginulinidae" genus="Marginulinopsis" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="costata">
<emphasis id="A3AAEABEFFDC93217FC99BD3FDB0FF71" bold="true" box="[151,628,223,250]" pageId="15" pageNumber="16">
<emphasis id="A3AAEABEFFDC93217FC99BD3FE78FF71" bold="true" box="[151,444,223,250]" italics="true" pageId="15" pageNumber="16">Marginulinopsis costata</emphasis>
(
<bibRefCitation id="F54F4B5DFFDC93217E959BD3FDA8FF71" author="Batsch, A. I. G. C." box="[459,620,223,249]" pageId="15" pageNumber="16" refId="ref27825" refString="Batsch, A. I. G. C. (1791) Sechs Kupfertafeln mit Conchylien des Seesandes, gezeichnet und gestochen von A. J. G. K. Batsch, Jena, 6 pls." type="book" year="1791">Batsch, 1791</bibRefCitation>
)
</emphasis>
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</heading>
</paragraph>
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<heading id="CA2981C0FFDC93217FC99A0FFE9DFE95" bold="true" box="[151,345,259,285]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="A3AAEABEFFDC93217FC99A0FFE9DFE95" bold="true" box="[151,345,259,285]" pageId="15" pageNumber="16">Pl. 2, figs. 1012</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFDC93217FC99A44FD39FE71" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="916136ACFFDC93217FC99A44FCF2FEE8" blockId="15.[151,1436,328,505]" box="[151,822,328,352]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="B1CE1182FFDC93217FC99A44FCF2FEE8" box="[151,822,328,352]" pageId="15" pageNumber="16">
<treatmentCitation id="107F10BDFFDC93217FC99A44FD54FEE8" author="Batsch, A. I. G. C." box="[151,656,328,352]" page="2" pageId="15" pageNumber="16" year="1791">
<taxonomicName id="56DE4D2FFFDC93217FC99A44FD54FEE8" authority="Batsch, 1791, p. 2" authorityName="Batsch" authorityPageNumber="2" authorityYear="1791" box="[151,656,328,352]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" kingdom="Animalia" order="Nautilida" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="costatus" subGenus="Orthoceras">
<emphasis id="A3AAEABEFFDC93217FC99A44FE15FED7" box="[151,465,328,352]" italics="true" pageId="15" pageNumber="16">Nautilus (Orthoceras) costatus</emphasis>
<bibRefCitation id="F54F4B5DFFDC93217E899A44FD9AFEE8" author="Batsch, A. I. G. C." box="[471,606,328,352]" pageId="15" pageNumber="16" refId="ref27825" refString="Batsch, A. I. G. C. (1791) Sechs Kupfertafeln mit Conchylien des Seesandes, gezeichnet und gestochen von A. J. G. K. Batsch, Jena, 6 pls." type="book" year="1791">Batsch, 1791</bibRefCitation>
, p. 2
</taxonomicName>
</treatmentCitation>
, pl. 1, fig. 1ag.
</treatmentCitationGroup>
</paragraph>
<paragraph id="916136ACFFDC93217FC99A6AFCD7FEF6" blockId="15.[151,1436,328,505]" box="[151,787,358,382]" pageId="15" pageNumber="16">
<treatmentCitation id="107F10BDFFDC93217FC99A6AFDB7FEF6" author="d'Orbigny, A. D." box="[151,627,358,382]" page="258" pageId="15" pageNumber="16" year="1826">
<taxonomicName id="56DE4D2FFFDC93217FC99A6AFDB7FEF6" ID-CoL="3Y5ZG" authority="d'Orbigny 1826, p. 258" authorityName="d'Orbigny" authorityPageNumber="258" authorityYear="1826" box="[151,627,358,382]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="raphanus">
<emphasis id="A3AAEABEFFDC93217FC99A6AFEB9FEF6" box="[151,381,358,382]" italics="true" pageId="15" pageNumber="16">Marginulina raphanus</emphasis>
<bibRefCitation id="F54F4B5DFFDC93217EDD9A6AFDECFEF6" author="d'Orbigny, A. D." box="[387,552,358,382]" pageId="15" pageNumber="16" pagination="245 - 314" refId="ref30526" refString="d'Orbigny, A. D. (1826) Tableau methodique de la classe des Cephalopodes. Annales des Sciences Naturelles, 7, 245 - 314." type="journal article" year="1826">dOrbigny 1826</bibRefCitation>
, p. 258
</taxonomicName>
</treatmentCitation>
, pl. X, fig. 78.
</paragraph>
<paragraph id="916136ACFFDC93217FC99A89FBDAFE52" blockId="15.[151,1436,328,505]" pageId="15" pageNumber="16">
<treatmentCitation id="107F10BDFFDC93217FC99A89FDF0FE15" author="Brady, H. B." box="[151,564,389,413]" page="528" pageId="15" pageNumber="16" year="1884">
<taxonomicName id="56DE4D2FFFDC93217FC99A89FDF0FE15" ID-CoL="89BX5" authority="Brady, 1884, p. 528" authorityName="Brady" authorityPageNumber="528" authorityYear="1884" box="[151,564,389,413]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="costata">
<emphasis id="A3AAEABEFFDC93217FC99A89FEA1FE15" box="[151,357,389,413]" italics="true" pageId="15" pageNumber="16">Marginulina costata</emphasis>
<bibRefCitation id="F54F4B5DFFDC93217E329A89FE2DFE15" author="Brady, H. B." box="[364,489,389,413]" pageId="15" pageNumber="16" pagination="1 - 814" refId="ref28770" refString="Brady, H. B. (1884) Report of the foraminifera dredged by H. M. S. Challenger during the years 1873 - 1876. Zoology, 9, 1 - 814." type="journal article" year="1884">Brady, 1884</bibRefCitation>
, p. 528
</taxonomicName>
</treatmentCitation>
, pl. 65, fig. 1013;
<treatmentCitation id="107F10BDFFDC93217DA49A89FC74FE15" author="Bagg, R. M." box="[762,944,389,413]" page="62" pageId="15" pageNumber="16" year="1912">
<bibRefCitation id="F54F4B5DFFDC93217DA49A89FCB4FE15" author="Bagg, R. M." box="[762,880,389,413]" pageId="15" pageNumber="16" pagination="1 - 153" refId="ref27521" refString="Bagg, R. M. (1912) Pliocene and Pleistocene foraminifera from Southern California. United States Geological Survey Bulletin, 513, 1 - 153. https: // doi. org / 10.3133 / b 513" type="journal article" year="1912">Bagg, 1912</bibRefCitation>
, p. 62
</treatmentCitation>
, pl. XVIII, fig. 4;
<treatmentCitation id="107F10BDFFDC93217B389A89FA96FE15" author="Cushman, J. A." box="[1126,1362,389,413]" page="256" pageId="15" pageNumber="16" year="1921">
<bibRefCitation id="F54F4B5DFFDC93217B389A89FAC3FE15" author="Cushman, J. A." box="[1126,1287,389,413]" pageId="15" pageNumber="16" pagination="1 - 608" refId="ref29695" refString="Cushman, J. A. (1921) Foraminifera of the Philippine and adjacent seas. Bulletin of the United States National Museum, 100, 1 - 608." type="journal article" year="1921">Cushman, 1921</bibRefCitation>
, p. 256
</treatmentCitation>
, pl. 41, fig. 58;
<bibRefCitation id="F54F4B5DFFDC93217E7B9AA8FD9CFE34" author="Heron-Allen, E. &amp; Earland, A." box="[293,600,420,444]" pageId="15" pageNumber="16" pagination="25 - 268" refId="ref32862" refString="Heron-Allen, E. &amp; Earland, A. (1922) Protozoa Part II - Foraminifera. British Antarctic (&quot; Terra Nova &quot;) Expedition 1910. Zoology, 6, 25 - 268." type="journal article" year="1922">Heron-Allen &amp; Earland, 1922</bibRefCitation>
, p. 176;
<bibRefCitation id="F54F4B5DFFDC93217DEF9AA8FC9BFE34" author="Cushman, J. A." box="[689,863,420,444]" pageId="15" pageNumber="16" pagination="1 - 228" refId="ref29789" refString="Cushman, J. A. (1923 a) The foraminifera of the Atlantic Ocean: part 4 - Lagenidae. Bulletin of the United States National Museum, 104 (4), 1 - 228. https: // doi. org / 10.5479 / si. 03629236.104.3" type="journal article" year="1923">Cushman, 1923a</bibRefCitation>
, p. 132, pl. 37, fig. 2;
<bibRefCitation id="F54F4B5DFFDC93217B1A9AA8FB7AFE34" author="Jones, R. W." box="[1092,1214,420,444]" pageId="15" pageNumber="16" refId="ref33343" refString="Jones, R. W. (1994) The Challenger Foraminifera. Oxford University Press, Oxford, 149 pp." type="book" year="1994">Jones, 1994</bibRefCitation>
, p. 77, pl. 65, fig. 13;
<bibRefCitation id="F54F4B5DFFDC93217F999ACEFE0AFE52" author="Milker, Y. &amp; Schmiedl, G." box="[199,462,450,474]" pageId="15" pageNumber="16" pagination="1 - 134" refId="ref35283" refString="Milker, Y. &amp; Schmiedl, G. (2012) A taxonomic guide to modern benthic shelf foraminifera of the western Mediterranean Sea. Palaeontologica Electronica, 15, 1 - 134. https: // doi. org / 10.26879 / 271" type="journal article" year="2012">Milker &amp; Schmiedl, 2012</bibRefCitation>
, p. 74, fig. 18.26;
<bibRefCitation id="F54F4B5DFFDC93217DD99ACEFCB0FE52" author="Obaje, S. O. &amp; Okosun, E. A." box="[647,884,450,474]" pageId="15" pageNumber="16" pagination="357 - 364" refId="ref35709" refString="Obaje, S. O. &amp; Okosun, E. A. (2013) Taxonomic notes on marker benthic foraminifera of Tomboy Field, offshore western Niger Delta, Nigeria. International Journal of Science and Technology, 3, 357 - 364." type="journal article" year="2013">Obaje &amp; Okosun, 2013</bibRefCitation>
, p. 360, pl. 1.18.
</paragraph>
<paragraph id="916136ACFFDC93217FC99AEDFD39FE71" blockId="15.[151,1436,328,505]" box="[151,765,481,505]" pageId="15" pageNumber="16">
<taxonomicName id="56DE4D2FFFDC93217FC99AEDFDF1FE70" ID-CoL="7F89B" authority="Jones, 1994, p. 77" authorityName="Jones" authorityPageNumber="77" authorityYear="1994" box="[151,565,481,505]" class="Nodosariata" family="Vaginulinidae" genus="Vaginulinopsis" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="bradyi">
<emphasis id="A3AAEABEFFDC93217FC99AEDFEB1FE71" box="[151,373,481,505]" italics="true" pageId="15" pageNumber="16">Vaginulinopsis bradyi</emphasis>
<bibRefCitation id="F54F4B5DFFDC93217E259AEDFE31FE71" author="Jones, R. W." box="[379,501,481,505]" pageId="15" pageNumber="16" refId="ref33343" refString="Jones, R. W. (1994) The Challenger Foraminifera. Oxford University Press, Oxford, 149 pp." type="book" year="1994">Jones, 1994</bibRefCitation>
, p. 77
</taxonomicName>
, pl. 65, figs. 1112.
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFDC93217FC9992BFDA0FD7D" pageId="15" pageNumber="16">
<paragraph id="916136ACFFDC93217FC9992BFDA0FD7D" blockId="15.[151,1437,551,1622]" pageId="15" pageNumber="16">
<emphasis id="A3AAEABEFFDC93217FC9992BFEEFFDC9" bold="true" box="[151,299,551,577]" pageId="15" pageNumber="16">Description:</emphasis>
The test wall is calcareous and finely perforate. The test is elongate, uniserial and circular in crosssection. The initial portion is coiled and round. Up to nine chambers are visible along the exterior. The chambers are rectilinearly arranged, separated by depressed sutures. The test is covered with thick longitudinal costae. The costae may extend from the initial to the terminal chamber and are slightly curved. Many of the specimens do not have the costae covering the terminal chambers. The final chamber is globular in shape, with a terminal and radiate aperture at the dorsal angle of a pronounced neck.
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFDC93217F9999F3FCA6F9DE" pageId="15" pageNumber="16" type="discussion">
<paragraph id="916136ACFFDC93217F9999F3FD0DFC0D" blockId="15.[151,1437,551,1622]" pageId="15" pageNumber="16">
<emphasis id="A3AAEABEFFDC93217F9999F3FEF9FC91" bold="true" box="[199,317,767,793]" pageId="15" pageNumber="16">Remarks:</emphasis>
The relative abundance of
<taxonomicName id="56DE4D2FFFDC93217D27980CFD35FC91" authorityName="Brady" authorityYear="1884" box="[633,753,768,793]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="costata">
<emphasis id="A3AAEABEFFDC93217D27980CFD35FC91" box="[633,753,768,793]" italics="true" pageId="15" pageNumber="16">M. costata</emphasis>
</taxonomicName>
is generally low, forming trace components (&lt;1%) in some of the samples of core 2670. There is variation in the size of the tests. Smaller specimens measure up to
<quantity id="56269B49FFDC93217A77982FFA44FCB6" box="[1321,1408,803,830]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="mm" value="0.2">0.2 mm</quantity>
in cross section diameter and
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in length, while some tests that are ornamented throughout, are large, measuring up to
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in cross section and
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in length.
</paragraph>
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The tests show a variable degree of size and ornamentation. Similarly, there appears to be variation in the ornamentation within this species itself and among the synonymies. For example, the test in pl. 2, fig. 10 closely resembles the sketched imaged of
<taxonomicName id="56DE4D2FFFDC93217D4798DBFCD7FC79" box="[537,787,983,1009]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="bifurcata">
<emphasis id="A3AAEABEFFDC93217D4798DBFCD7FC79" box="[537,787,983,1009]" italics="true" pageId="15" pageNumber="16">Marginulina bifurcata</emphasis>
</taxonomicName>
in
<bibRefCitation id="F54F4B5DFFDC93217C6898DBFC30FC79" author="Fornasini, C." box="[822,1012,983,1009]" pageId="15" pageNumber="16" pagination="1 - 68" refId="ref31860" refString="Fornasini, C. (1902) Sinossi metodica dei foraminiferi sin qui rinvenuti nella sabbia del Lido di Rimini. Memorie della Accademia della scienze dell'Istituto di Bologna, Series, 5, 1 - 68." type="journal article" year="1902">Fornasini (1902)</bibRefCitation>
, with the terminal chamber being just as strongly costate as the preceding chambers.
<taxonomicName id="56DE4D2FFFDC93217DE298F7FC7DFB9D" box="[700,953,1019,1045]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="bifurcata">
<emphasis id="A3AAEABEFFDC93217DE298F7FC7DFB9D" box="[700,953,1019,1045]" italics="true" pageId="15" pageNumber="16">Marginulina bifurcata</emphasis>
</taxonomicName>
has, however, been synonymised into
<taxonomicName id="56DE4D2FFFDC93217A2098F0FF2CFBB1" authorityName="Brady" authorityYear="1884" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="costata">
<emphasis id="A3AAEABEFFDC93217A2098F0FF2CFBB1" italics="true" pageId="15" pageNumber="16">M. costata</emphasis>
</taxonomicName>
. The specimens in figures 11 and 12 resemble
<taxonomicName id="56DE4D2FFFDC93217C599F13FB1AFBB2" authority="(d'Orbigny, 1826)" baseAuthorityName="d'Orbigny" baseAuthorityYear="1826" box="[775,1246,1055,1082]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="raphanus">
<emphasis id="A3AAEABEFFDC93217C599F13FBC2FBB1" box="[775,1030,1055,1081]" italics="true" pageId="15" pageNumber="16">Marginulina raphanus</emphasis>
(
<bibRefCitation id="F54F4B5DFFDC93217B489F13FB13FBB2" author="d'Orbigny, A. D." box="[1046,1239,1055,1082]" pageId="15" pageNumber="16" pagination="245 - 314" refId="ref30526" refString="d'Orbigny, A. D. (1826) Tableau methodique de la classe des Cephalopodes. Annales des Sciences Naturelles, 7, 245 - 314." type="journal article" year="1826">dOrbigny, 1826</bibRefCitation>
)
</taxonomicName>
, but that species has a pointed initial end, whereas specimens from this study area is rounded along the initial portion. There has also been confusion in the identification of this taxon.
<bibRefCitation id="F54F4B5DFFDC93217C5C9F6BFC65FB0A" author="Barker, R. W." box="[770,929,1127,1154]" pageId="15" pageNumber="16" pagination="1 - 238" refId="ref27766" refString="Barker, R. W. (1960) Taxonomic notes on the species figured by H. B. Brady in his report on the Foraminifera dredged by H. M. S. Challenger during the years 1873 - 1876. Society of Economic Palaeontologists and Mineralogists, Special Publication, 9, 1 - 238." type="journal article" year="1960">Barker (1960)</bibRefCitation>
re-identified this species as
<emphasis id="A3AAEABEFFDC93217BBA9F6BFF23FB2D" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="56DE4D2FFFDC93217BBA9F6BFF27FB2D" baseAuthorityName="Goes" baseAuthorityYear="1894" class="Nodosariata" family="Vaginulinidae" genus="Marginulinopsis" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="bradyi">Marginulinopsis bradyi</taxonomicName>
,
</emphasis>
whereas
<bibRefCitation id="F54F4B5DFFDC93217E089F80FE2DFB2D" author="Jones, R. W." box="[342,489,1163,1190]" pageId="15" pageNumber="16" refId="ref33343" refString="Jones, R. W. (1994) The Challenger Foraminifera. Oxford University Press, Oxford, 149 pp." type="book" year="1994">Jones (1994)</bibRefCitation>
split the figured identifications of
<taxonomicName id="56DE4D2FFFDC93217C209F80FC33FB2D" authorityName="Brady" authorityYear="1884" box="[894,1015,1164,1189]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="costata">
<emphasis id="A3AAEABEFFDC93217C209F80FC33FB2D" box="[894,1015,1164,1189]" italics="true" pageId="15" pageNumber="16">M. costata</emphasis>
</taxonomicName>
in
<bibRefCitation id="F54F4B5DFFDC93217B7E9F87FB7CFB2D" author="Brady, H. B." box="[1056,1208,1163,1189]" pageId="15" pageNumber="16" pagination="1 - 814" refId="ref28770" refString="Brady, H. B. (1884) Report of the foraminifera dredged by H. M. S. Challenger during the years 1873 - 1876. Zoology, 9, 1 - 814." type="journal article" year="1884">Brady (1884)</bibRefCitation>
into
<taxonomicName id="56DE4D2FFFDC93217BA99F87FF24FB41" authorityName="Jones" authorityYear="1994" class="Nodosariata" family="Vaginulinidae" genus="Vaginulinopsis" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="bradyi">
<emphasis id="A3AAEABEFFDC93217BA99F87FF24FB41" italics="true" pageId="15" pageNumber="16">Vaginulinopsis bradyi</emphasis>
</taxonomicName>
and
<taxonomicName id="56DE4D2FFFDC93217E499FBCFE49FB41" authorityName="Brady" authorityYear="1884" box="[279,397,1200,1225]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="costata">
<emphasis id="A3AAEABEFFDC93217E499FBCFE49FB41" box="[279,397,1200,1225]" italics="true" pageId="15" pageNumber="16">M. costata</emphasis>
</taxonomicName>
. The original specimen of
<taxonomicName id="56DE4D2FFFDC93217DE59FA3FC53FB41" box="[699,919,1199,1225]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="bradyi">
<emphasis id="A3AAEABEFFDC93217DE59FA3FC53FB41" box="[699,919,1199,1225]" italics="true" pageId="15" pageNumber="16">Marginulina bradyi</emphasis>
</taxonomicName>
appears to be more initially coiled and flatter, broadening towards the terminal end, whereas
<taxonomicName id="56DE4D2FFFDC93217DF59FD8FCE7FB65" authorityName="Brady" authorityYear="1884" box="[683,803,1236,1261]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="costata">
<emphasis id="A3AAEABEFFDC93217DF59FD8FCE7FB65" box="[683,803,1236,1261]" italics="true" pageId="15" pageNumber="16">M. costata</emphasis>
</taxonomicName>
does not show the same initial portion and later stages. In this study, the figured specimens have been identified as
<emphasis id="A3AAEABEFFDC93217C6E9FFBFB83FA99" box="[816,1095,1271,1297]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="56DE4D2FFFDC93217C6E9FFBFB86FA99" baseAuthorityName="Batsch" baseAuthorityYear="1791" box="[816,1090,1271,1297]" class="Nodosariata" family="Vaginulinidae" genus="Marginulinopsis" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="costata">Marginulinopsis costata</taxonomicName>
.
</emphasis>
The figured specimens 11 and 12 also resemble that of
<taxonomicName id="56DE4D2FFFDC93217EEF9E17FD03FABD" authorityName="Asano" authorityYear="1937" box="[433,711,1307,1333]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="sendaiensis">
<emphasis id="A3AAEABEFFDC93217EEF9E17FD03FABD" box="[433,711,1307,1333]" italics="true" pageId="15" pageNumber="16">Marginulina sendaiensis</emphasis>
</taxonomicName>
in
<bibRefCitation id="F54F4B5DFFDC93217DB39E17FC45FABD" author="Asano, K." box="[749,897,1307,1333]" pageId="15" pageNumber="16" pagination="28 - 35" refId="ref27449" refString="Asano, K. (1937) Fossil Foraminifera from the Moniwa Shell-Beds in the Vicinity of Sendai. Rikuzen Province. The Journal of the Geological Society of Japan, 44, 28 - 35. https: // doi. org / 10.5575 / geosoc. 44.28" type="journal article" year="1937">Asano (1937</bibRefCitation>
,
<bibRefCitation id="F54F4B5DFFDC93217CD09E17FC0EFABD" author="Asano, K." box="[910,970,1307,1333]" pageId="15" pageNumber="16" pagination="423 - 430" refId="ref27498" refString="Asano, K. (1949) New Miocene foraminifera from Japan. Journal of Paleontology, 23, 423 - 430." type="journal article" year="1949">1949</bibRefCitation>
), possessing less or no ornamentation on their terminal chamber and are more abundant (up to 5% in some of the samples) than those having tests completely covered in costae.The difference between
<taxonomicName id="56DE4D2FFFDC93217D379E6FFCBFFAF5" authorityName="Asano" authorityYear="1937" box="[617,891,1379,1405]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="sendaiensis">
<emphasis id="A3AAEABEFFDC93217D379E6FFCBFFAF5" box="[617,891,1379,1405]" italics="true" pageId="15" pageNumber="16">Marginulina sendaiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="56DE4D2FFFDC93217CF39E68FBE5FAF5" baseAuthorityName="Batsch" baseAuthorityYear="1791" box="[941,1057,1380,1405]" class="Nodosariata" family="Vaginulinidae" genus="Marginulinopsis" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="costata">
<emphasis id="A3AAEABEFFDC93217CF39E68FBE5FAF5" box="[941,1057,1380,1405]" italics="true" pageId="15" pageNumber="16">M. costata</emphasis>
</taxonomicName>
is that the aperture of
<taxonomicName id="56DE4D2FFFDC93217A4E9E6FFEE7FA29" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="sendaisensis">
<emphasis id="A3AAEABEFFDC93217A4E9E6FFEE7FA29" italics="true" pageId="15" pageNumber="16">Marginulina sendaisensis</emphasis>
</taxonomicName>
is centrally located on the terminal chamber, but the aperture of
<taxonomicName id="56DE4D2FFFDC93217CA99E84FBAAFA29" baseAuthorityName="Batsch" baseAuthorityYear="1791" box="[1015,1134,1416,1441]" class="Nodosariata" family="Vaginulinidae" genus="Marginulinopsis" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="costata">
<emphasis id="A3AAEABEFFDC93217CA99E84FBAAFA29" box="[1015,1134,1416,1441]" italics="true" pageId="15" pageNumber="16">M. costata</emphasis>
</taxonomicName>
is located excentrically.
</paragraph>
<paragraph id="916136ACFFDC93217F999EA7FD75FA61" blockId="15.[151,1437,551,1622]" pageId="15" pageNumber="16">
<emphasis id="A3AAEABEFFDC93217F999EA7FEA3FA4D" bold="true" box="[199,359,1451,1477]" pageId="15" pageNumber="16">Life strategy:</emphasis>
Species of this genus are generally shallow-infaunal in environments with variable conditions of low to high oxygen (
<bibRefCitation id="F54F4B5DFFDC93217E219EC3FD61FA61" author="Milker, Y. &amp; Schmiedl, G." box="[383,677,1487,1513]" pageId="15" pageNumber="16" pagination="1 - 134" refId="ref35283" refString="Milker, Y. &amp; Schmiedl, G. (2012) A taxonomic guide to modern benthic shelf foraminifera of the western Mediterranean Sea. Palaeontologica Electronica, 15, 1 - 134. https: // doi. org / 10.26879 / 271" type="journal article" year="2012">Milker &amp; Schmiedl, 2012</bibRefCitation>
).
</paragraph>
<paragraph id="916136ACFFDC93217F999EFFFB0EF985" blockId="15.[151,1437,551,1622]" box="[199,1226,1523,1550]" pageId="15" pageNumber="16">
<emphasis id="A3AAEABEFFDC93217F999EFFFDCBF985" bold="true" box="[199,527,1523,1549]" pageId="15" pageNumber="16">Global stratigraphic range:</emphasis>
This species occurs from the Jurassic to Recent (
<bibRefCitation id="F54F4B5DFFDC93217B699EFFFB79F985" author="Bagg, R. M." box="[1079,1213,1523,1549]" pageId="15" pageNumber="16" pagination="1 - 153" refId="ref27521" refString="Bagg, R. M. (1912) Pliocene and Pleistocene foraminifera from Southern California. United States Geological Survey Bulletin, 513, 1 - 153. https: // doi. org / 10.3133 / b 513" type="journal article" year="1912">Bagg, 1912</bibRefCitation>
).
</paragraph>
<paragraph id="916136ACFFDC93217F999D1BFCA6F9DE" blockId="15.[151,1437,551,1622]" pageId="15" pageNumber="16">
<emphasis id="A3AAEABEFFDC93217F999D1BFE01F9B9" bold="true" box="[199,453,1559,1585]" pageId="15" pageNumber="16">Regional occurrence:</emphasis>
<taxonomicName id="56DE4D2FFFDC93217E959D14FD85F9B9" baseAuthorityName="Batsch" baseAuthorityYear="1791" box="[459,577,1560,1585]" class="Nodosariata" family="Vaginulinidae" genus="Marginulinopsis" kingdom="Chromista" order="Vaginulinida" pageId="15" pageNumber="16" phylum="Foraminifera" rank="species" species="costata">
<emphasis id="A3AAEABEFFDC93217E959D14FD85F9B9" box="[459,577,1560,1585]" italics="true" pageId="15" pageNumber="16">M. costata</emphasis>
</taxonomicName>
is documented to occur in middle Miocene sediments on the Namibian outer continental shelf, south of the
<collectingRegion id="531AF84EFFDC93217EB49D30FD42F9DD" box="[490,646,1595,1622]" isEnumeration="true" pageId="15" pageNumber="16">
<collectingRegion id="531AF84EFFDC93217EB49D30FD85F9DE" box="[490,577,1596,1622]" country="Namibia" name="Kunene" pageId="15" pageNumber="16">Kunene</collectingRegion>
<collectingRegion id="531AF84EFFDC93217D169D37FD42F9DD" box="[584,646,1595,1621]" country="Nigeria" name="Rivers" pageId="15" pageNumber="16">River</collectingRegion>
</collectingRegion>
mouth (this study).
</paragraph>
</subSubSection>
</treatment>
</document>