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<document id="E25EFD9475E8E220795C769C6F6B33A5" ID-CLB-Dataset="39130" ID-DOI="10.11646/zootaxa.4109.1.6" ID-GBIF-Dataset="6875db92-ff8d-4e6e-9289-eab37f2e6061" ID-ISSN="1175-5326" ID-Zenodo-Dep="260655" ID-ZooBank="E2CABF95-7527-42BD-B895-CD5A2B5F7CD1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1462430178508" checkinUser="plazi" docAuthor="Wielstra, B. &amp; Arntzen, J. W." docDate="2016" docId="163C2C30006EFF8DFF35F80F3F6A0423" docLanguage="en" docName="zootaxa.4109.1.6.pdf.imf" docOrigin="Zootaxa 4109 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Triturus anatolicus Wielstra &amp; Arntzen, 2016, sp. nov." docType="treatment" docVersion="10" lastPageNumber="79" masterDocId="EA055448006FFF8BFFA2FF8F3D420563" masterDocTitle="Description of a new species of crested newt, previously subsumed in Triturus ivanbureschi (Amphibia: Caudata: Salamandridae)" masterLastPageNumber="80" masterPageNumber="73" pageNumber="74" updateTime="1698662290673" updateUser="ExternalLinkService">
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<mods:title id="DDE3160888945D154954A1CF52A92195">Description of a new species of crested newt, previously subsumed in Triturus ivanbureschi (Amphibia: Caudata: Salamandridae)</mods:title>
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<mods:namePart id="60194B74E967D4FB6208EAAFA2A1EE27">Wielstra, B.</mods:namePart>
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<mods:namePart id="EAC0286EBB5889CA18798CEE52F4560F">Arntzen, J. W.</mods:namePart>
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<mods:title id="3DE331E4EE13DC5E328B93841F2D3B98">Zootaxa</mods:title>
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<mods:date id="52C6D94460F9B13E26EC665A45513FF2">2016</mods:date>
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<treatment id="163C2C30006EFF8DFF35F80F3F6A0423" ID-DOI="http://doi.org/10.5281/zenodo.5674936" ID-GBIF-Taxon="120624239" ID-Zenodo-Dep="5674936" LSID="urn:lsid:plazi:treatment:163C2C30006EFF8DFF35F80F3F6A0423" httpUri="http://treatment.plazi.org/id/163C2C30006EFF8DFF35F80F3F6A0423" lastPageId="6" lastPageNumber="79" pageId="1" pageNumber="74">
<subSubSection id="D68FCEAD006EFF8AFF35F80F3CA602F9" box="[151,484,1920,1946]" pageId="1" pageNumber="74" type="nomenclature">
<paragraph id="9E2A9D26006EFF8AFF35F80F3CA602F9" blockId="1.[151,484,1920,1946]" box="[151,484,1920,1946]" pageId="1" pageNumber="74">
<heading id="C5622A4A006EFF8AFF35F80F3CA602F9" bold="true" box="[151,484,1920,1946]" fontSize="11" level="1" pageId="1" pageNumber="74" reason="1">
<emphasis id="ACE14134006EFF8AFF35F80F3CA602F9" bold="true" box="[151,484,1920,1946]" pageId="1" pageNumber="74">
<taxonomicName id="5995E6A5006EFF8AFF35F80F3C3D02F9" ID-CoL="7D889" box="[151,383,1920,1946]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="1" pageNumber="74" phylum="Chordata" rank="species" species="anatolicus" status="sp. nov.">
<emphasis id="ACE14134006EFF8AFF35F80F3C3D02F9" bold="true" box="[151,383,1920,1946]" italics="true" pageId="1" pageNumber="74">Triturus anatolicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7D2FC4F006EFF8AFE25F80F3CA602F9" box="[391,484,1920,1946]" pageId="1" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D68FCEAD006EFF89FF35F8483CF10423" lastPageId="2" lastPageNumber="75" pageId="1" pageNumber="74" type="materials_examined">
<paragraph id="9E2A9D26006EFF89FF35F8483CF10423" blockId="1.[151,1436,1991,2016]" lastBlockId="2.[151,1436,151,320]" lastPageId="2" lastPageNumber="75" pageId="1" pageNumber="74">
<emphasis id="ACE14134006EFF8AFF35F8483C8E0283" bold="true" box="[151,460,1991,2016]" pageId="1" pageNumber="74">
<typeStatus id="412E2384006EFF8AFF35F8483D900283" box="[151,210,1991,2016]" pageId="1" pageNumber="74">Type</typeStatus>
material.
<typeStatus id="412E2384006EFF8AFEFAF8483C850283" box="[344,455,1991,2016]" pageId="1" pageNumber="74" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="F88405E3006EFF8AFE7BF8483F7302BC" LSID="urn:lsid:biocol.org:col:34992" box="[473,561,1991,2015]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="74">RMNH</collectionCode>
.
<collectionCode id="F88405E3006EFF8AFD96F8483FC602BC" box="[564,644,1991,2015]" pageId="1" pageNumber="74">RENA</collectionCode>
.48232, an adult male (
<figureCitation id="06AE81A3006EFF8AFC38F8483EA102BC" box="[922,995,1991,2016]" captionStart="FIGURE 3" captionStartId="4.[151,250,1948,1970]" captionTargetBox="[153,1433,195,1926]" captionTargetId="figure@4.[153,1433,195,1927]" captionTargetPageId="4" captionText="FIGURE 3. The holotype and one of the paratypes of Triturus anatolicus sp. nov. in life. Lateral (a) and ventral (b) view of a male (below, holotype, RMNH. RENA. 48232) and a female (above, paratype, RMNH. RENA. 48239)." httpUri="https://zenodo.org/record/260658/files/figure.png" pageId="1" pageNumber="74">Fig. 3</figureCitation>
) from Gölköy, near Kalecik,
<collectingCountry id="E682DDB6006EFF8AFAEEF84838DE0283" box="[1356,1436,1991,2016]" name="Turkey" pageId="1" pageNumber="74">Turkey</collectingCountry>
(coordinates:
<geoCoordinate id="FBA1FBE1006DFF89FE96FF173CE105D3" box="[308,419,152,176]" direction="north" orientation="latitude" pageId="2" pageNumber="75" precision="55" value="40.077">40.077°N</geoCoordinate>
,
<geoCoordinate id="FBA1FBE1006DFF89FE11FF173F5C05D3" box="[435,542,152,176]" direction="east" orientation="longitude" pageId="2" pageNumber="75" precision="55" value="33.341">33.341°E</geoCoordinate>
, elevation:
<quantity id="596D30C3006DFF89FD05FF173E6805CC" box="[679,810,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="2" pageNumber="75" unit="m" value="1230.0">1230 meter</quantity>
a.s.l.;
<figureCitation id="06AE81A3006DFF89FCDEFF183E8105D3" box="[892,963,151,176]" captionStart="FIGURE 4" captionStartId="5.[151,250,1139,1161]" captionTargetBox="[176,1422,201,1102]" captionTargetId="figure@5.[176,1422,201,1103]" captionTargetPageId="5" captionText="FIGURE 4. The type locality of Triturus anatolicus sp. nov. at Gölköy, Turkey." httpUri="https://zenodo.org/record/260659/files/figure.png" pageId="2" pageNumber="75">Fig. 4</figureCitation>
), collected
<date id="EA2BBBE6006DFF89FBEFFF1839A605D3" box="[1101,1252,151,176]" pageId="2" pageNumber="75" value="2014-04-06">6 April 2014</date>
and donated to Naturalis Biodiversity Center by Kurtuluş Olgun from the Department of Biology at Adnan Menderes University, Aydın,
<collectingCountry id="E682DDB6006DFF89FF49FF503C7F059B" box="[235,317,223,248]" name="Turkey" pageId="2" pageNumber="75">Turkey</collectingCountry>
.
<emphasis id="ACE14134006DFF89FEE8FF503C8A059B" bold="true" box="[330,456,223,248]" pageId="2" pageNumber="75">
<typeStatus id="412E2384006DFF89FEE8FF503C86059B" box="[330,452,223,248]" pageId="2" pageNumber="75" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Six males (
<collectionCode id="F88405E3006DFF89FDF6FF6F3FEF059B" LSID="urn:lsid:biocol.org:col:34992" box="[596,685,224,248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="75">RMNH</collectionCode>
.
<collectionCode id="F88405E3006DFF89FD12FF6F3E43059B" box="[688,769,224,248]" pageId="2" pageNumber="75">RENA</collectionCode>
.48233-48238), six females (
<collectionCode id="F88405E3006DFF89FBE5FF5039E2059B" LSID="urn:lsid:biocol.org:col:34992" box="[1095,1184,223,248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="75">RMNH</collectionCode>
.
<collectionCode id="F88405E3006DFF89FB01FF5039B6059B" box="[1187,1268,223,248]" pageId="2" pageNumber="75">RENA</collectionCode>
.48239-48245), and a juvenile (
<collectionCode id="F88405E3006DFF89FEE0FE8B3CD9047E" LSID="urn:lsid:biocol.org:col:34992" box="[322,411,260,285]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="75">RMNH</collectionCode>
.
<collectionCode id="F88405E3006DFF89FE3DFE8B3CB2047E" box="[415,496,260,285]" pageId="2" pageNumber="75">RENA</collectionCode>
.48246), with collection details identical to the
<typeStatus id="412E2384006DFF89FC59FE8B3923047E" box="[1019,1121,260,285]" pageId="2" pageNumber="75" type="holotype">holotype</typeStatus>
. For imagery, see
<figureCitation id="06AE81A3006DFF89FA89FE8B382F047F" box="[1323,1389,260,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1948,1970]" captionTargetBox="[153,1433,195,1926]" captionTargetId="figure@4.[153,1433,195,1927]" captionTargetPageId="4" captionText="FIGURE 3. The holotype and one of the paratypes of Triturus anatolicus sp. nov. in life. Lateral (a) and ventral (b) view of a male (below, holotype, RMNH. RENA. 48232) and a female (above, paratype, RMNH. RENA. 48239)." httpUri="https://zenodo.org/record/260658/files/figure.png" pageId="2" pageNumber="75">Fig. 3</figureCitation>
and online Appendix 1 and 2.
</paragraph>
</subSubSection>
<caption id="CAEACDAE006DFF89FF35FC0939B5018D" httpUri="https://zenodo.org/record/260657/files/figure.png" pageId="2" pageNumber="75" targetBox="[151,1436,397,857]" targetPageId="2">
<paragraph id="9E2A9D26006DFF89FF35FC0939B5018D" blockId="2.[151,1436,902,1262]" pageId="2" pageNumber="75">
<emphasis id="ACE14134006DFF89FF35FC093C5406FF" bold="true" box="[151,278,902,924]" pageId="2" pageNumber="75">FIGURE 2.</emphasis>
Genetic data for
<emphasis id="ACE14134006DFF89FE77FC073E6506FF" box="[469,807,902,925]" italics="true" pageId="2" pageNumber="75">
<taxonomicName id="5995E6A5006DFF89FE77FC073FB706FE" box="[469,757,902,925]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="subSpecies" species="ivanbureschi" subSpecies="sensu">
Triturus ivanbureschi
<taxonomicNameLabel id="B7D2FC4F006DFF89FD1FFC073FB706FE" box="[701,757,904,925]" pageId="2" pageNumber="75" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato
</emphasis>
presented in Wielstra
<emphasis id="ACE14134006DFF89FBB8FC07391306FF" box="[1050,1105,902,925]" italics="true" pageId="2" pageNumber="75">et al.</emphasis>
(submitted) illustrate the sharp geographical overturn between the genotypes representing its two constituent species. Panel (a) shows the 67 sampled populations as dots with the colored background representing a rough distribution outline of the two species based on nuclear DNA. Populations are colored according to the mtDNA type present; western populations of
<taxonomicName id="5995E6A5006DFF89FBDAFC6B387C069A" box="[1144,1342,995,1017]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="species" species="anatolicus" status="sp. nov.">
<emphasis id="ACE14134006DFF89FBDAFC6B387C069A" box="[1144,1342,995,1017]" italics="true" pageId="2" pageNumber="75">Triturus anatolicus</emphasis>
</taxonomicName>
<emphasis id="ACE14134006DFF89FAE5FC6C38D9069B" bold="true" box="[1351,1435,995,1016]" pageId="2" pageNumber="75">
<taxonomicNameLabel id="B7D2FC4F006DFF89FAE5FC6C38D9069B" box="[1351,1435,995,1016]" pageId="2" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
possess mtDNA of
<emphasis id="ACE14134006DFF89FEC7FB8D3F8C0174" box="[357,718,1025,1047]" italics="true" pageId="2" pageNumber="75">
<taxonomicName id="5995E6A5006DFF89FEC7FB8D3FC60174" box="[357,644,1025,1047]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="subSpecies" species="ivanbureschi" subSpecies="sensu">
Triturus ivanbureschi
<taxonomicNameLabel id="B7D2FC4F006DFF89FDEEFB8D3FC60174" box="[588,644,1026,1047]" pageId="2" pageNumber="75" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
. Those populations harboring individuals showing introgression and genetically admixed individuals based on nuclear DNA (as defined below) are highlighted. Panel (b) shows a Structure plot representing
<emphasis id="ACE14134006DFF89FEBEFBB03C6C0137" box="[284,302,1087,1108]" italics="true" pageId="2" pageNumber="75">Q</emphasis>
scores with which the 428 sampled individuals were allocated to two different species (
<emphasis id="ACE14134006DFF89FB1EFBB1398A0137" box="[1212,1224,1086,1108]" italics="true" pageId="2" pageNumber="75">k</emphasis>
= 2 was the optimal number of gene pools inferred in the Structure analysis). The individuals are ordered based on the
<emphasis id="ACE14134006DFF89FBDCFBD139D20110" box="[1150,1168,1118,1139]" italics="true" pageId="2" pageNumber="75">Q</emphasis>
score with which they are allocated to
<emphasis id="ACE14134006DFF89FEB4FBF23F7901F1" box="[278,571,1148,1170]" italics="true" pageId="2" pageNumber="75">
<taxonomicName id="5995E6A5006DFF89FEB4FBF23CB001F1" box="[278,498,1148,1170]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="subSpecies" species="ivanbureschi" subSpecies="sensu">
T. ivanbureschi
<taxonomicNameLabel id="B7D2FC4F006DFF89FE18FBF23CB001F1" box="[442,498,1149,1170]" pageId="2" pageNumber="75" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
, from high to low values. Individuals allocated to their respective species with
<emphasis id="ACE14134006DFF89FAD6FBF238C401F1" box="[1396,1414,1149,1170]" italics="true" pageId="2" pageNumber="75">Q</emphasis>
≥ 0.987 are considered genetically pure. Individuals showing introgression are defined as being allocated to a species with 0.987&gt;
<emphasis id="ACE14134006DFF89FF0EFB353DFC01AC" box="[172,190,1210,1231]" italics="true" pageId="2" pageNumber="75">Q</emphasis>
&gt; 0.85 and genetically admixed individuals as being allocated to either species with
<emphasis id="ACE14134006DFF89FB8BFB35397901AC" box="[1065,1083,1210,1231]" italics="true" pageId="2" pageNumber="75">Q</emphasis>
≤ 0.85. Those individuals that are not genetically pure are highlighted in panel (b). Note that nuclear introgression is geographically asymmetric.
</paragraph>
</caption>
<subSubSection id="D68FCEAD006DFF89FF65FA9A3CF50261" pageId="2" pageNumber="75" type="diagnosis">
<paragraph id="9E2A9D26006DFF89FF65FA9A3CF50261" blockId="2.[151,1436,1301,2010]" pageId="2" pageNumber="75">
<emphasis id="ACE14134006DFF89FF65FA9A3C03004D" bold="true" box="[199,321,1301,1326]" pageId="2" pageNumber="75">Diagnosis.</emphasis>
The new species is assigned to the genus
<taxonomicName id="5995E6A5006DFF89FCB7FA993E2C004E" box="[789,878,1302,1325]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="ACE14134006DFF89FCB7FA993E2C004E" box="[789,878,1302,1325]" italics="true" pageId="2" pageNumber="75">Triturus</emphasis>
</taxonomicName>
because it possesses the characteristics of crested newts (the
<taxonomicName id="5995E6A5006DFF89FEB5FAB53CD40032" box="[279,406,1338,1361]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="species" species="cristatus">
<emphasis id="ACE14134006DFF89FEB5FAB53CD40032" box="[279,406,1338,1361]" italics="true" pageId="2" pageNumber="75">T. cristatus</emphasis>
</taxonomicName>
superspecies), in particular the combination of a denticulated crest (in males in breeding costume), a dark brown dorsal coloration and an orange ventral side covered with black spots. Based on mtDNA sequence data, crested newts from the
<typeStatus id="412E2384006DFF89FDFEFA0D3FCE00F9" box="[604,652,1410,1434]" pageId="2" pageNumber="75">type</typeStatus>
locality are firmly placed in the genus
<taxonomicName id="5995E6A5006DFF89FBFDFA0D39FA00FA" box="[1119,1208,1410,1433]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="ACE14134006DFF89FBFDFA0D39FA00FA" box="[1119,1208,1410,1433]" italics="true" pageId="2" pageNumber="75">Triturus</emphasis>
</taxonomicName>
, within the distinct central
<emphasis id="ACE14134006DFF89FF51FA293CA600DD" box="[243,484,1445,1470]" italics="true" pageId="2" pageNumber="75">
<taxonomicName id="5995E6A5006DFF89FF51FA293CF000DE" box="[243,434,1445,1470]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="subSpecies" species="karelinii" subSpecies="sensu">
T. karelinii
<taxonomicNameLabel id="B7D2FC4F006DFF89FED6FA293CF000DE" box="[372,434,1446,1469]" pageId="2" pageNumber="75" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato
</emphasis>
mtDNA lineage (Wielstra
<emphasis id="ACE14134006DFF89FCBEFA293E1600DD" box="[796,852,1445,1470]" italics="true" pageId="2" pageNumber="75">et al.</emphasis>
, 2010). Although identification based on mtDNA is possible across the majority of the range of the new species, mtDNA derived from
<taxonomicName id="5995E6A5006DFF89FB97FA4539A20081" box="[1077,1248,1481,1506]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="species" species="ivanbureschi">
<emphasis id="ACE14134006DFF89FB97FA4539A20081" box="[1077,1248,1481,1506]" italics="true" pageId="2" pageNumber="75">T. ivanbureschi</emphasis>
</taxonomicName>
has introgressed at the western part of the range (Wielstra
<emphasis id="ACE14134006DFF89FD20FA613FFC0365" box="[642,702,1517,1542]" italics="true" pageId="2" pageNumber="75">et al.</emphasis>
, submitted;
<figureCitation id="06AE81A3006DFF89FCF2FA623EDA0365" box="[848,920,1517,1542]" captionStart="FIGURE 2" captionStartId="2.[151,250,902,924]" captionTargetBox="[151,1436,397,857]" captionTargetId="figure@2.[151,1436,397,857]" captionTargetPageId="2" captionText="FIGURE 2. Genetic data for Triturus ivanbureschi sensu lato presented in Wielstra et al. (submitted) illustrate the sharp geographical overturn between the genotypes representing its two constituent species. Panel (a) shows the 67 sampled populations as dots with the colored background representing a rough distribution outline of the two species based on nuclear DNA. Populations are colored according to the mtDNA type present; western populations of Triturus anatolicus sp. nov. possess mtDNA of Triturus ivanbureschi sensu stricto. Those populations harboring individuals showing introgression and genetically admixed individuals based on nuclear DNA (as defined below) are highlighted. Panel (b) shows a Structure plot representing Q scores with which the 428 sampled individuals were allocated to two different species (k = 2 was the optimal number of gene pools inferred in the Structure analysis). The individuals are ordered based on the Q score with which they are allocated to T. ivanbureschi sensu stricto, from high to low values. Individuals allocated to their respective species with Q ≥ 0.987 are considered genetically pure. Individuals showing introgression are defined as being allocated to a species with 0.987&gt; Q&gt; 0.85 and genetically admixed individuals as being allocated to either species with Q ≤ 0.85. Those individuals that are not genetically pure are highlighted in panel (b). Note that nuclear introgression is geographically asymmetric." httpUri="https://zenodo.org/record/260657/files/figure.png" pageId="2" pageNumber="75">Fig. 2</figureCitation>
). The pattern of asymmetric introgression is mirrored by certain individual nuclear DNA markers and we consider this to represent a genomic footprint of hybrid zone movement (Wielstra
<emphasis id="ACE14134006DFF89FDB9F9B93F14032D" box="[539,598,1589,1614]" italics="true" pageId="2" pageNumber="75">et al.</emphasis>
, submitted;
<figureCitation id="06AE81A3006DFF89FD4AF9BA3E6C032D" box="[744,814,1589,1614]" captionStart="FIGURE 2" captionStartId="2.[151,250,902,924]" captionTargetBox="[151,1436,397,857]" captionTargetId="figure@2.[151,1436,397,857]" captionTargetPageId="2" captionText="FIGURE 2. Genetic data for Triturus ivanbureschi sensu lato presented in Wielstra et al. (submitted) illustrate the sharp geographical overturn between the genotypes representing its two constituent species. Panel (a) shows the 67 sampled populations as dots with the colored background representing a rough distribution outline of the two species based on nuclear DNA. Populations are colored according to the mtDNA type present; western populations of Triturus anatolicus sp. nov. possess mtDNA of Triturus ivanbureschi sensu stricto. Those populations harboring individuals showing introgression and genetically admixed individuals based on nuclear DNA (as defined below) are highlighted. Panel (b) shows a Structure plot representing Q scores with which the 428 sampled individuals were allocated to two different species (k = 2 was the optimal number of gene pools inferred in the Structure analysis). The individuals are ordered based on the Q score with which they are allocated to T. ivanbureschi sensu stricto, from high to low values. Individuals allocated to their respective species with Q ≥ 0.987 are considered genetically pure. Individuals showing introgression are defined as being allocated to a species with 0.987&gt; Q&gt; 0.85 and genetically admixed individuals as being allocated to either species with Q ≤ 0.85. Those individuals that are not genetically pure are highlighted in panel (b). Note that nuclear introgression is geographically asymmetric." httpUri="https://zenodo.org/record/260657/files/figure.png" pageId="2" pageNumber="75">Fig. 2</figureCitation>
).
<collectingCountry id="E682DDB6006DFF89FCE5F9B93E24032D" box="[839,870,1590,1614]" name="American Samoa" pageId="2" pageNumber="75">As</collectingCountry>
yet only multilocus nuclear DNA sequence data distinguish
<taxonomicName id="5995E6A5006DFF89FEB8F9D53CE90311" box="[282,427,1625,1650]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="species" species="anatolicus" status="sp. nov.">
<emphasis id="ACE14134006DFF89FEB8F9D53CE90311" box="[282,427,1625,1650]" italics="true" pageId="2" pageNumber="75">T. anatolicus</emphasis>
</taxonomicName>
<emphasis id="ACE14134006DFF89FE10F9D63F4F0311" bold="true" box="[434,525,1625,1650]" pageId="2" pageNumber="75">
<taxonomicNameLabel id="B7D2FC4F006DFF89FE10F9D63F4F0311" box="[434,525,1625,1650]" pageId="2" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
unambiguously from
<taxonomicName id="5995E6A5006DFF89FCA5F9D53EF10311" box="[775,947,1625,1650]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="species" species="ivanbureschi">
<emphasis id="ACE14134006DFF89FCA5F9D53EF10311" box="[775,947,1625,1650]" italics="true" pageId="2" pageNumber="75">T. ivanbureschi</emphasis>
</taxonomicName>
and from
<taxonomicName id="5995E6A5006DFF89FB9DF9D539FB0311" box="[1087,1209,1625,1650]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="species" species="karelinii">
<emphasis id="ACE14134006DFF89FB9DF9D539FB0311" box="[1087,1209,1625,1650]" italics="true" pageId="2" pageNumber="75">T. karelinii</emphasis>
</taxonomicName>
, with which both
<taxonomicName id="5995E6A5006DFF89FA25F9D53C4E03F5" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="species" species="anatolicus" status="sp. nov.">
<emphasis id="ACE14134006DFF89FA25F9D53C4E03F5" italics="true" pageId="2" pageNumber="75">T. anatolicus</emphasis>
</taxonomicName>
<emphasis id="ACE14134006DFF89FEB1F9F23C2C03F5" bold="true" box="[275,366,1661,1686]" pageId="2" pageNumber="75">
<taxonomicNameLabel id="B7D2FC4F006DFF89FEB1F9F23C2C03F5" box="[275,366,1661,1686]" pageId="2" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="5995E6A5006DFF89FE07F9F13F1003F5" box="[421,594,1661,1686]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="species" species="ivanbureschi">
<emphasis id="ACE14134006DFF89FE07F9F13F1003F5" box="[421,594,1661,1686]" italics="true" pageId="2" pageNumber="75">T. ivanbureschi</emphasis>
</taxonomicName>
were until recently considered conspecific (Wielstra
<emphasis id="ACE14134006DFF89FB0EF9F139A703F5" box="[1196,1253,1661,1686]" italics="true" pageId="2" pageNumber="75">et al.</emphasis>
, submitted). No morphological features are currently known to distinguish the three species comprising
<emphasis id="ACE14134006DFF89FBCAF92D381A03D9" box="[1128,1368,1697,1722]" italics="true" pageId="2" pageNumber="75">
<taxonomicName id="5995E6A5006DFF89FBCAF92D386403DA" box="[1128,1318,1697,1722]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="subSpecies" species="karelinii" subSpecies="sensu">
T. karelinii
<taxonomicNameLabel id="B7D2FC4F006DFF89FB4AF92D386403DA" box="[1256,1318,1698,1721]" pageId="2" pageNumber="75" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato
</emphasis>
. How to separate
<taxonomicName id="5995E6A5006DFF89FEBAF9493CEB03BD" box="[280,425,1733,1758]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="species" species="anatolicus" status="sp. nov.">
<emphasis id="ACE14134006DFF89FEBAF9493CEB03BD" box="[280,425,1733,1758]" italics="true" pageId="2" pageNumber="75">T. anatolicus</emphasis>
</taxonomicName>
<emphasis id="ACE14134006DFF89FE12F94A3F4903BD" bold="true" box="[432,523,1733,1758]" pageId="2" pageNumber="75">
<taxonomicNameLabel id="B7D2FC4F006DFF89FE12F94A3F4903BD" box="[432,523,1733,1758]" pageId="2" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="5995E6A5006DFF89FDBBF9493F8703BD" box="[537,709,1733,1758]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="species" species="ivanbureschi">
<emphasis id="ACE14134006DFF89FDBBF9493F8703BD" box="[537,709,1733,1758]" italics="true" pageId="2" pageNumber="75">T. ivanbureschi</emphasis>
</taxonomicName>
and
<taxonomicName id="5995E6A5006DFF89FD5FF9493E3703BD" box="[765,885,1733,1758]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="2" pageNumber="75" phylum="Chordata" rank="species" species="karelinii">
<emphasis id="ACE14134006DFF89FD5FF9493E3703BD" box="[765,885,1733,1758]" italics="true" pageId="2" pageNumber="75">T. karelinii</emphasis>
</taxonomicName>
from the other crested newt species is discussed in
<bibRefCitation id="FA04E0D7006DFF89FF16F9663CF10261" box="[180,435,1769,1794]" pageId="2" pageNumber="75" refString="Wielstra, B., Litvinchuk, S. N., Naumov, B., Tzankov, N. &amp; Arntzen, J. W. (2013 b) A revised taxonomy of crested newts in the Triturus karelinii group (Amphibia: Caudata: Salamandridae), with the description of a new species. Zootaxa, 3682, 441 - 453. http: // dx. doi. org / 10.11646 / zootaxa. 3682.3.5" type="journal article">
Wielstra
<emphasis id="ACE14134006DFF89FEBAF9653C130261" box="[280,337,1769,1794]" italics="true" pageId="2" pageNumber="75">et al.</emphasis>
(2013b)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D68FCEAD006DFF8EFF65F88238DA0355" lastPageId="5" lastPageNumber="78" pageId="2" pageNumber="75" type="description">
<paragraph id="9E2A9D26006DFF88FF65F8823EC3047F" blockId="2.[151,1436,1301,2010]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="76" pageId="2" pageNumber="75">
<emphasis id="ACE14134006DFF89FF65F8823CAF0245" bold="true" box="[199,493,1805,1830]" pageId="2" pageNumber="75">
Description of
<typeStatus id="412E2384006DFF89FEDDF8823CAA0245" box="[383,488,1805,1830]" pageId="2" pageNumber="75" type="holotype">holotype</typeStatus>
.
</emphasis>
A relatively robust crested newt, with a broad head (wider than neck) and well developed limbs. Four fingers and five toes. Toes fringed but interdigital webbing absent. Relative length of fingers left 1 &lt;4 &lt;2 &lt;3, right 4 &lt;1 &lt;3 &lt;2 (3 and 4 possibly regenerated). Relative length of toes on both sides 1 &lt;5 &lt;2 &lt;4 &lt;3. Skin granulated on dorsal and lateral sides, tail base and throat; tail and venter smooth. Gular fold inconspicuous. Swollen cloaca with papillae bordering cloacal slit. Denticulated crest, indented over the position of the legs and cloaca. Laterally compressed tail with evident dorsal fin and less conspicuous ventral fin. In life a brownish-blackish base color on the dorsolateral side, with scattered black flecks, most pronounced and marbled on the head. Bluish-white streak along the lateral side of the tail along the caudal vertebrae. A deep-orange ventral side and throat, dotted with small angular black spots; throat spots smaller and denser than belly spots. In the preserved state colors have slightly faded (see online Appendix 1).
</paragraph>
<caption id="CAEACDAE006CFF88FF35FEC73C99041C" ID-Table-UUID="CAEACDAE006CFF88FF35FEC73C99041C" httpUri="http://table.plazi.org/id/CAEACDAE006CFF88FF35FEC73C99041C" pageId="3" pageNumber="76" targetBox="[160,1383,410,1053]" targetIsTable="true" targetPageId="3">
<paragraph id="9E2A9D26006CFF88FF35FEC73C99041C" blockId="3.[151,1435,328,383]" pageId="3" pageNumber="76">
<emphasis id="ACE14134006CFF88FF35FEC73C4B043C" bold="true" box="[151,265,328,351]" pageId="3" pageNumber="76">TABLE 1.</emphasis>
Number of rib-bearing pre-sacral vertebrae and measurements (in millimeters) for the holotype and paratypes of
<taxonomicName id="5995E6A5006CFF88FF16FEE73C3D041C" box="[180,383,360,383]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="76" phylum="Chordata" rank="species" species="anatolicus" status="sp. nov.">
<emphasis id="ACE14134006CFF88FF16FEE73C3D041C" box="[180,383,360,383]" italics="true" pageId="3" pageNumber="76">Triturus anatolicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7D2FC4F006CFF88FE24FEE73C99041C" box="[390,475,360,383]" pageId="3" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="9E2A9D26006CFF88FF02FE143814017E" pageId="3" pageNumber="76">
<table id="EC956F86006C0074FF02FE153825017E" box="[160,1383,410,1053]" gridcols="8" gridrows="16" pageId="3" pageNumber="76">
<tr id="20A59F64006C0074FF02FE15382504D3" box="[160,1383,410,432]" gridrow="0" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FE153C3C04D3" box="[160,382,410,432]" gridcol="0" gridrow="0" pageId="3" pageNumber="76">Museum number</th>
<th id="6374F618006C0074FE50FE153F0604D3" box="[498,580,410,432]" gridcol="1" gridrow="0" pageId="3" pageNumber="76">Sex</th>
<th id="6374F618006C0074FD2AFE153FA304D3" box="[648,737,410,432]" gridcol="2" gridrow="0" pageId="3" pageNumber="76">Material</th>
<th id="6374F618006C0074FC92FE153E3404D3" box="[816,886,410,432]" gridcol="3" gridrow="0" pageId="3" pageNumber="76">NRBV</th>
<th id="6374F618006C0074FC1BFE153EB404D3" box="[953,1014,410,432]" gridcol="4" gridrow="0" pageId="3" pageNumber="76">SVL1</th>
<th id="6374F618006C0074FB97FE15393004D3" box="[1077,1138,410,432]" gridcol="5" gridrow="0" pageId="3" pageNumber="76">SVL2</th>
<th id="6374F618006C0074FB10FE1539B404D3" box="[1202,1270,410,432]" gridcol="6" gridrow="0" pageId="3" pageNumber="76">TL</th>
<th id="6374F618006C0074FA9FFE15382504D3" box="[1341,1383,410,432]" gridcol="7" gridrow="0" pageId="3" pageNumber="76">ILD</th>
</tr>
<tr id="20A59F64006C0074FF02FE4B381404B9" box="[160,1366,452,474]" gridrow="1" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FE4B3C3C04B9" box="[160,382,452,474]" gridcol="0" gridrow="1" pageId="3" pageNumber="76">RMNH.RENA.48232</th>
<td id="6374F618006C0074FE50FE4B3F0604B9" box="[498,580,452,474]" gridcol="1" gridrow="1" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FD2AFE4B3FA304B9" box="[648,737,452,474]" gridcol="2" gridrow="1" pageId="3" pageNumber="76">holotype</td>
<td id="6374F618006C0074FC92FE4B3E3404B9" box="[816,886,452,474]" gridcol="3" gridrow="1" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFE4B3EB404B9" box="[953,1014,452,474]" gridcol="4" gridrow="1" pageId="3" pageNumber="76">53</td>
<td id="6374F618006C0074FB97FE4B393004B9" box="[1077,1138,452,474]" gridcol="5" gridrow="1" pageId="3" pageNumber="76">61</td>
<td id="6374F618006C0074FB10FE4B39B404B9" box="[1202,1270,452,474]" gridcol="6" gridrow="1" pageId="3" pageNumber="76">108</td>
<td id="6374F618006C0074FA9FFE4B382504B9" box="[1341,1383,452,474]" gridcol="7" gridrow="1" pageId="3" pageNumber="76">27</td>
</tr>
<tr id="20A59F64006C0074FF02FE6138140767" box="[160,1366,494,516]" gridrow="2" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FE613C3C0767" box="[160,382,494,516]" gridcol="0" gridrow="2" pageId="3" pageNumber="76">RMNH.RENA.48233</th>
<td id="6374F618006C0074FE50FE613F060767" box="[498,580,494,516]" gridcol="1" gridrow="2" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FD2AFE613FA30767" box="[648,737,494,516]" gridcol="2" gridrow="2" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FE613E340767" box="[816,886,494,516]" gridcol="3" gridrow="2" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFE613EB40767" box="[953,1014,494,516]" gridcol="4" gridrow="2" pageId="3" pageNumber="76">63</td>
<td id="6374F618006C0074FB97FE6139300767" box="[1077,1138,494,516]" gridcol="5" gridrow="2" pageId="3" pageNumber="76">71</td>
<td id="6374F618006C0074FB10FE6139B40767" box="[1202,1270,494,516]" gridcol="6" gridrow="2" pageId="3" pageNumber="76">126</td>
<td id="6374F618006C0074FA9FFE6138250767" box="[1341,1383,494,516]" gridcol="7" gridrow="2" pageId="3" pageNumber="76">34</td>
</tr>
<tr id="20A59F64006C0074FF02FD983814074E" box="[160,1366,535,557]" gridrow="3" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FD983C3C074E" box="[160,382,535,557]" gridcol="0" gridrow="3" pageId="3" pageNumber="76">RMNH.RENA.48234</th>
<td id="6374F618006C0074FE50FD983F06074E" box="[498,580,535,557]" gridcol="1" gridrow="3" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FD2AFD983FA3074E" box="[648,737,535,557]" gridcol="2" gridrow="3" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FD983E34074E" box="[816,886,535,557]" gridcol="3" gridrow="3" pageId="3" pageNumber="76">12</td>
<td id="6374F618006C0074FC1BFD983EB4074E" box="[953,1014,535,557]" gridcol="4" gridrow="3" pageId="3" pageNumber="76">53</td>
<td id="6374F618006C0074FB97FD983930074E" box="[1077,1138,535,557]" gridcol="5" gridrow="3" pageId="3" pageNumber="76">60</td>
<td id="6374F618006C0074FB10FD9839B4074E" box="[1202,1270,535,557]" gridcol="6" gridrow="3" pageId="3" pageNumber="76">110</td>
<td id="6374F618006C0074FA9FFD983825074E" box="[1341,1383,535,557]" gridcol="7" gridrow="3" pageId="3" pageNumber="76">29</td>
</tr>
<tr id="20A59F64006C0074FF02FDCF38140735" box="[160,1366,576,598]" gridrow="4" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FDCF3C3C0735" box="[160,382,576,598]" gridcol="0" gridrow="4" pageId="3" pageNumber="76">RMNH.RENA.48235</th>
<td id="6374F618006C0074FE50FDCF3F060735" box="[498,580,576,598]" gridcol="1" gridrow="4" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FD2AFDCF3FA30735" box="[648,737,576,598]" gridcol="2" gridrow="4" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FDCF3E340735" box="[816,886,576,598]" gridcol="3" gridrow="4" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFDCF3EB40735" box="[953,1014,576,598]" gridcol="4" gridrow="4" pageId="3" pageNumber="76">53</td>
<td id="6374F618006C0074FB97FDCF39300735" box="[1077,1138,576,598]" gridcol="5" gridrow="4" pageId="3" pageNumber="76">61</td>
<td id="6374F618006C0074FB10FDCF39B40735" box="[1202,1270,576,598]" gridcol="6" gridrow="4" pageId="3" pageNumber="76">109</td>
<td id="6374F618006C0074FA9FFDCF38250735" box="[1341,1383,576,598]" gridcol="7" gridrow="4" pageId="3" pageNumber="76">26</td>
</tr>
<tr id="20A59F64006C0074FF02FDE5381407E3" box="[160,1366,618,640]" gridrow="5" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FDE53C3C07E3" box="[160,382,618,640]" gridcol="0" gridrow="5" pageId="3" pageNumber="76">RMNH.RENA.48236</th>
<td id="6374F618006C0074FE50FDE53F0607E3" box="[498,580,618,640]" gridcol="1" gridrow="5" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FD2AFDE53FA307E3" box="[648,737,618,640]" gridcol="2" gridrow="5" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FDE53E3407E3" box="[816,886,618,640]" gridcol="3" gridrow="5" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFDE53EB407E3" box="[953,1014,618,640]" gridcol="4" gridrow="5" pageId="3" pageNumber="76">50</td>
<td id="6374F618006C0074FB97FDE5393007E3" box="[1077,1138,618,640]" gridcol="5" gridrow="5" pageId="3" pageNumber="76">58</td>
<td id="6374F618006C0074FB10FDE539B407E3" box="[1202,1270,618,640]" gridcol="6" gridrow="5" pageId="3" pageNumber="76">100</td>
<td id="6374F618006C0074FA9FFDE5382507E3" box="[1341,1383,618,640]" gridcol="7" gridrow="5" pageId="3" pageNumber="76">27</td>
</tr>
<tr id="20A59F64006C0074FF02FD1C381407CA" box="[160,1366,659,681]" gridrow="6" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FD1C3C3C07CA" box="[160,382,659,681]" gridcol="0" gridrow="6" pageId="3" pageNumber="76">RMNH.RENA.48237</th>
<td id="6374F618006C0074FE50FD1C3F0607CA" box="[498,580,659,681]" gridcol="1" gridrow="6" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FD2AFD1C3FA307CA" box="[648,737,659,681]" gridcol="2" gridrow="6" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FD1C3E3407CA" box="[816,886,659,681]" gridcol="3" gridrow="6" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFD1C3EB407CA" box="[953,1014,659,681]" gridcol="4" gridrow="6" pageId="3" pageNumber="76">56</td>
<td id="6374F618006C0074FB97FD1C393007CA" box="[1077,1138,659,681]" gridcol="5" gridrow="6" pageId="3" pageNumber="76">64</td>
<td id="6374F618006C0074FB10FD1C39B407CA" box="[1202,1270,659,681]" gridcol="6" gridrow="6" pageId="3" pageNumber="76">115</td>
<td id="6374F618006C0074FA9FFD1C382507CA" box="[1341,1383,659,681]" gridcol="7" gridrow="6" pageId="3" pageNumber="76">30</td>
</tr>
<tr id="20A59F64006C0074FF02FD33381407B1" box="[160,1366,700,722]" gridrow="7" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FD333C3C07B1" box="[160,382,700,722]" gridcol="0" gridrow="7" pageId="3" pageNumber="76">RMNH.RENA.48238</th>
<td id="6374F618006C0074FE50FD333F0607B1" box="[498,580,700,722]" gridcol="1" gridrow="7" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FD2AFD333FA307B1" box="[648,737,700,722]" gridcol="2" gridrow="7" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FD333E3407B1" box="[816,886,700,722]" gridcol="3" gridrow="7" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFD333EB407B1" box="[953,1014,700,722]" gridcol="4" gridrow="7" pageId="3" pageNumber="76">53</td>
<td id="6374F618006C0074FB97FD33393007B1" box="[1077,1138,700,722]" gridcol="5" gridrow="7" pageId="3" pageNumber="76">60</td>
<td id="6374F618006C0074FB10FD3339B407B1" box="[1202,1270,700,722]" gridcol="6" gridrow="7" pageId="3" pageNumber="76">110</td>
<td id="6374F618006C0074FA9FFD33382507B1" box="[1341,1383,700,722]" gridcol="7" gridrow="7" pageId="3" pageNumber="76">29</td>
</tr>
<tr id="20A59F64006C0074FF02FD693814079F" box="[160,1366,742,764]" gridrow="8" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FD693C3C079F" box="[160,382,742,764]" gridcol="0" gridrow="8" pageId="3" pageNumber="76">RMNH.RENA.48239</th>
<td id="6374F618006C0074FE50FD693F06079F" box="[498,580,742,764]" gridcol="1" gridrow="8" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FD2AFD693FA3079F" box="[648,737,742,764]" gridcol="2" gridrow="8" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FD693E34079F" box="[816,886,742,764]" gridcol="3" gridrow="8" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFD693EB4079F" box="[953,1014,742,764]" gridcol="4" gridrow="8" pageId="3" pageNumber="76">57</td>
<td id="6374F618006C0074FB97FD693930079F" box="[1077,1138,742,764]" gridcol="5" gridrow="8" pageId="3" pageNumber="76">62</td>
<td id="6374F618006C0074FB10FD6939B4079F" box="[1202,1270,742,764]" gridcol="6" gridrow="8" pageId="3" pageNumber="76">118</td>
<td id="6374F618006C0074FA9FFD693825079F" box="[1341,1383,742,764]" gridcol="7" gridrow="8" pageId="3" pageNumber="76">31</td>
</tr>
<tr id="20A59F64006C0074FF02FC8038140646" box="[160,1366,783,805]" gridrow="9" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FC803C3C0646" box="[160,382,783,805]" gridcol="0" gridrow="9" pageId="3" pageNumber="76">RMNH.RENA.48240</th>
<td id="6374F618006C0074FE50FC803F060646" box="[498,580,783,805]" gridcol="1" gridrow="9" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FD2AFC803FA30646" box="[648,737,783,805]" gridcol="2" gridrow="9" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FC803E340646" box="[816,886,783,805]" gridcol="3" gridrow="9" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFC803EB40646" box="[953,1014,783,805]" gridcol="4" gridrow="9" pageId="3" pageNumber="76">62</td>
<td id="6374F618006C0074FB97FC8039300646" box="[1077,1138,783,805]" gridcol="5" gridrow="9" pageId="3" pageNumber="76">68</td>
<td id="6374F618006C0074FB10FC8039B40646" box="[1202,1270,783,805]" gridcol="6" gridrow="9" pageId="3" pageNumber="76">133</td>
<td id="6374F618006C0074FA9FFC8038250646" box="[1341,1383,783,805]" gridcol="7" gridrow="9" pageId="3" pageNumber="76">32</td>
</tr>
<tr id="20A59F64006C0074FF02FCB73814062D" box="[160,1366,824,846]" gridrow="10" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FCB73C3C062D" box="[160,382,824,846]" gridcol="0" gridrow="10" pageId="3" pageNumber="76">RMNH.RENA.48241</th>
<td id="6374F618006C0074FE50FCB73F06062D" box="[498,580,824,846]" gridcol="1" gridrow="10" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FD2AFCB73FA3062D" box="[648,737,824,846]" gridcol="2" gridrow="10" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FCB73E34062D" box="[816,886,824,846]" gridcol="3" gridrow="10" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFCB73EB4062D" box="[953,1014,824,846]" gridcol="4" gridrow="10" pageId="3" pageNumber="76">58</td>
<td id="6374F618006C0074FB97FCB73930062D" box="[1077,1138,824,846]" gridcol="5" gridrow="10" pageId="3" pageNumber="76">63</td>
<td id="6374F618006C0074FB10FCB739B4062D" box="[1202,1270,824,846]" gridcol="6" gridrow="10" pageId="3" pageNumber="76">122</td>
<td id="6374F618006C0074FA9FFCB73825062D" box="[1341,1383,824,846]" gridcol="7" gridrow="10" pageId="3" pageNumber="76">33</td>
</tr>
<tr id="20A59F64006C0074FF02FCED3814061B" box="[160,1366,866,888]" gridrow="11" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FCED3C3C061B" box="[160,382,866,888]" gridcol="0" gridrow="11" pageId="3" pageNumber="76">RMNH.RENA.48242</th>
<td id="6374F618006C0074FE50FCED3F06061B" box="[498,580,866,888]" gridcol="1" gridrow="11" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FD2AFCED3FA3061B" box="[648,737,866,888]" gridcol="2" gridrow="11" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FCED3E34061B" box="[816,886,866,888]" gridcol="3" gridrow="11" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFCED3EB4061B" box="[953,1014,866,888]" gridcol="4" gridrow="11" pageId="3" pageNumber="76">58</td>
<td id="6374F618006C0074FB97FCED3930061B" box="[1077,1138,866,888]" gridcol="5" gridrow="11" pageId="3" pageNumber="76">62</td>
<td id="6374F618006C0074FB10FCED39B4061B" box="[1202,1270,866,888]" gridcol="6" gridrow="11" pageId="3" pageNumber="76">123</td>
<td id="6374F618006C0074FA9FFCED3825061B" box="[1341,1383,866,888]" gridcol="7" gridrow="11" pageId="3" pageNumber="76">30</td>
</tr>
<tr id="20A59F64006C0074FF02FC04381406C2" box="[160,1366,907,929]" gridrow="12" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FC043C3C06C2" box="[160,382,907,929]" gridcol="0" gridrow="12" pageId="3" pageNumber="76">RMNH.RENA.48243</th>
<td id="6374F618006C0074FE50FC043F0606C2" box="[498,580,907,929]" gridcol="1" gridrow="12" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FD2AFC043FA306C2" box="[648,737,907,929]" gridcol="2" gridrow="12" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FC043E3406C2" box="[816,886,907,929]" gridcol="3" gridrow="12" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFC043EB406C2" box="[953,1014,907,929]" gridcol="4" gridrow="12" pageId="3" pageNumber="76">62</td>
<td id="6374F618006C0074FB97FC04393006C2" box="[1077,1138,907,929]" gridcol="5" gridrow="12" pageId="3" pageNumber="76">67</td>
<td id="6374F618006C0074FB10FC0439B406C2" box="[1202,1270,907,929]" gridcol="6" gridrow="12" pageId="3" pageNumber="76">117 **</td>
<td id="6374F618006C0074FA9FFC04382506C2" box="[1341,1383,907,929]" gridcol="7" gridrow="12" pageId="3" pageNumber="76">33</td>
</tr>
<tr id="20A59F64006C0074FF02FC3B381406A9" box="[160,1366,948,970]" gridrow="13" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FC3B3C3C06A9" box="[160,382,948,970]" gridcol="0" gridrow="13" pageId="3" pageNumber="76">RMNH.RENA.48244</th>
<td id="6374F618006C0074FE50FC3B3F0606A9" box="[498,580,948,970]" gridcol="1" gridrow="13" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FD2AFC3B3FA306A9" box="[648,737,948,970]" gridcol="2" gridrow="13" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FC3B3E3406A9" box="[816,886,948,970]" gridcol="3" gridrow="13" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFC3B3EB406A9" box="[953,1014,948,970]" gridcol="4" gridrow="13" pageId="3" pageNumber="76">65</td>
<td id="6374F618006C0074FB97FC3B393006A9" box="[1077,1138,948,970]" gridcol="5" gridrow="13" pageId="3" pageNumber="76">70</td>
<td id="6374F618006C0074FB10FC3B39B406A9" box="[1202,1270,948,970]" gridcol="6" gridrow="13" pageId="3" pageNumber="76">131</td>
<td id="6374F618006C0074FA9FFC3B382506A9" box="[1341,1383,948,970]" gridcol="7" gridrow="13" pageId="3" pageNumber="76">36</td>
</tr>
<tr id="20A59F64006C0074FF02FC5138140697" box="[160,1366,990,1012]" gridrow="14" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FC513C3C0697" box="[160,382,990,1012]" gridcol="0" gridrow="14" pageId="3" pageNumber="76">RMNH.RENA.48245</th>
<td id="6374F618006C0074FE50FC513F060697" box="[498,580,990,1012]" gridcol="1" gridrow="14" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FD2AFC513FA30697" box="[648,737,990,1012]" gridcol="2" gridrow="14" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FC513E340697" box="[816,886,990,1012]" gridcol="3" gridrow="14" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFC513EB40697" box="[953,1014,990,1012]" gridcol="4" gridrow="14" pageId="3" pageNumber="76">56</td>
<td id="6374F618006C0074FB97FC5139300697" box="[1077,1138,990,1012]" gridcol="5" gridrow="14" pageId="3" pageNumber="76">60</td>
<td id="6374F618006C0074FB10FC5139B40697" box="[1202,1270,990,1012]" gridcol="6" gridrow="14" pageId="3" pageNumber="76">111</td>
<td id="6374F618006C0074FA9FFC5138250697" box="[1341,1383,990,1012]" gridcol="7" gridrow="14" pageId="3" pageNumber="76">31</td>
</tr>
<tr id="20A59F64006C0074FF02FB883814017E" box="[160,1366,1031,1053]" gridrow="15" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF02FB883C3C017E" box="[160,382,1031,1053]" gridcol="0" gridrow="15" pageId="3" pageNumber="76">RMNH.RENA.48246</th>
<td id="6374F618006C0074FE50FB883F06017E" box="[498,580,1031,1053]" gridcol="1" gridrow="15" pageId="3" pageNumber="76">juvenile</td>
<td id="6374F618006C0074FD2AFB883FA3017E" box="[648,737,1031,1053]" gridcol="2" gridrow="15" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FC92FB883E34017E" box="[816,886,1031,1053]" gridcol="3" gridrow="15" pageId="3" pageNumber="76">13</td>
<td id="6374F618006C0074FC1BFB883EB4017E" box="[953,1014,1031,1053]" gridcol="4" gridrow="15" pageId="3" pageNumber="76">51</td>
<td id="6374F618006C0074FB97FB883930017E" box="[1077,1138,1031,1053]" gridcol="5" gridrow="15" pageId="3" pageNumber="76">55</td>
<td id="6374F618006C0074FB10FB8839B4017E" box="[1202,1270,1031,1053]" gridcol="6" gridrow="15" pageId="3" pageNumber="76">102</td>
<td id="6374F618006C0074FA9FFB883825017E" box="[1341,1383,1031,1053]" gridcol="7" gridrow="15" pageId="3" pageNumber="76">29</td>
</tr>
</table>
</paragraph>
<paragraph id="9E2A9D26006CFF88FF35FBCD3C4A013A" blockId="3.[151,264,1090,1113]" box="[151,264,1090,1113]" pageId="3" pageNumber="76">
<tableNote id="63739CA8006CFF88FF35FBCD3C4A013A" box="[151,264,1090,1113]" pageId="3" pageNumber="76" targetBox="[160,1383,410,1053]" targetPageId="3">continued.</tableNote>
</paragraph>
<paragraph id="9E2A9D26006CFF88FF03FBFA38170394" pageId="3" pageNumber="76">
<table id="EC956F86006C0074FF03FBFB382A0394" box="[161,1384,1140,1783]" gridcols="9" gridrows="16" pageId="3" pageNumber="76">
<tr id="20A59F64006C0074FF03FBFB382A01E9" box="[161,1384,1140,1162]" gridrow="0" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03FBFB3C3C01E9" box="[161,382,1140,1162]" gridcol="0" gridrow="0" pageId="3" pageNumber="76">Museum number</th>
<th id="6374F618006C0074FE47FBFB3F7401E9" box="[485,566,1140,1162]" gridcol="1" gridrow="0" pageId="3" pageNumber="76">Sex</th>
<th id="6374F618006C0074FDD4FBFB3F8D01E9" box="[630,719,1140,1162]" gridcol="2" gridrow="0" pageId="3" pageNumber="76">Material</th>
<th id="6374F618006C0074FCB5FBFB3E0101E9" box="[791,835,1140,1162]" gridcol="3" gridrow="0" pageId="3" pageNumber="76">HL</th>
<th id="6374F618006C0074FCD5FBFB3EE101E9" box="[887,931,1140,1162]" gridcol="4" gridrow="0" pageId="3" pageNumber="76">HW</th>
<th id="6374F618006C0074FC7AFBFB394401E9" box="[984,1030,1140,1162]" gridcol="5" gridrow="0" pageId="3" pageNumber="76">Arm</th>
<th id="6374F618006C0074FBE3FBFB39D201E9" box="[1089,1168,1140,1162]" gridcol="6" gridrow="0" pageId="3" pageNumber="76">Finger3</th>
<th id="6374F618006C0074FB77FBFB384301E9" box="[1237,1281,1140,1162]" gridcol="7" gridrow="0" pageId="3" pageNumber="76">Leg</th>
<th id="6374F618006C0074FA97FBFB382A01E9" box="[1333,1384,1140,1162]" gridcol="8" gridrow="0" pageId="3" pageNumber="76">Toe4</th>
</tr>
<tr id="20A59F64006C0074FF03FB11382A01D7" box="[161,1384,1182,1204]" gridrow="1" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03FB113C3C01D7" box="[161,382,1182,1204]" gridcol="0" gridrow="1" pageId="3" pageNumber="76">RMNH.RENA.48232</th>
<td id="6374F618006C0074FE47FB113F7401D7" box="[485,566,1182,1204]" gridcol="1" gridrow="1" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FDD4FB113F8D01D7" box="[630,719,1182,1204]" gridcol="2" gridrow="1" pageId="3" pageNumber="76">holotype</td>
<td id="6374F618006C0074FCB5FB113E0101D7" box="[791,835,1182,1204]" gridcol="3" gridrow="1" pageId="3" pageNumber="76">16.2</td>
<td id="6374F618006C0074FCD5FB113EE101D7" box="[887,931,1182,1204]" gridcol="4" gridrow="1" pageId="3" pageNumber="76">10.6</td>
<td id="6374F618006C0074FC7AFB11394401D7" box="[984,1030,1182,1204]" gridcol="5" gridrow="1" pageId="3" pageNumber="76">18.7</td>
<td id="6374F618006C0074FBE3FB1139D201D7" box="[1089,1168,1182,1204]" gridcol="6" gridrow="1" pageId="3" pageNumber="76">5.6 *</td>
<td id="6374F618006C0074FB77FB11384301D7" box="[1237,1281,1182,1204]" gridcol="7" gridrow="1" pageId="3" pageNumber="76">22.1</td>
<td id="6374F618006C0074FA97FB11382A01D7" box="[1333,1384,1182,1204]" gridcol="8" gridrow="1" pageId="3" pageNumber="76">9.5</td>
</tr>
<tr id="20A59F64006C0074FF03FB47382A01BD" box="[161,1384,1224,1246]" gridrow="2" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03FB473C3C01BD" box="[161,382,1224,1246]" gridcol="0" gridrow="2" pageId="3" pageNumber="76">RMNH.RENA.48233</th>
<td id="6374F618006C0074FE47FB473F7401BD" box="[485,566,1224,1246]" gridcol="1" gridrow="2" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FDD4FB473F8D01BD" box="[630,719,1224,1246]" gridcol="2" gridrow="2" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5FB473E0101BD" box="[791,835,1224,1246]" gridcol="3" gridrow="2" pageId="3" pageNumber="76">18.4</td>
<td id="6374F618006C0074FCD5FB473EE101BD" box="[887,931,1224,1246]" gridcol="4" gridrow="2" pageId="3" pageNumber="76">12.0</td>
<td id="6374F618006C0074FC7AFB47394401BD" box="[984,1030,1224,1246]" gridcol="5" gridrow="2" pageId="3" pageNumber="76">26.9</td>
<td id="6374F618006C0074FBE3FB4739D201BD" box="[1089,1168,1224,1246]" gridcol="6" gridrow="2" pageId="3" pageNumber="76">10.4</td>
<td id="6374F618006C0074FB77FB47384301BD" box="[1237,1281,1224,1246]" gridcol="7" gridrow="2" pageId="3" pageNumber="76">26.6</td>
<td id="6374F618006C0074FA97FB47382A01BD" box="[1333,1384,1224,1246]" gridcol="8" gridrow="2" pageId="3" pageNumber="76">11.6</td>
</tr>
<tr id="20A59F64006C0074FF03FB7E382A0064" box="[161,1384,1265,1287]" gridrow="3" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03FB7E3C3C0064" box="[161,382,1265,1287]" gridcol="0" gridrow="3" pageId="3" pageNumber="76">RMNH.RENA.48234</th>
<td id="6374F618006C0074FE47FB7E3F740064" box="[485,566,1265,1287]" gridcol="1" gridrow="3" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FDD4FB7E3F8D0064" box="[630,719,1265,1287]" gridcol="2" gridrow="3" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5FB7E3E010064" box="[791,835,1265,1287]" gridcol="3" gridrow="3" pageId="3" pageNumber="76">16.5</td>
<td id="6374F618006C0074FCD5FB7E3EE10064" box="[887,931,1265,1287]" gridcol="4" gridrow="3" pageId="3" pageNumber="76">11.5</td>
<td id="6374F618006C0074FC7AFB7E39440064" box="[984,1030,1265,1287]" gridcol="5" gridrow="3" pageId="3" pageNumber="76">22.7</td>
<td id="6374F618006C0074FBE3FB7E39D20064" box="[1089,1168,1265,1287]" gridcol="6" gridrow="3" pageId="3" pageNumber="76">8.6</td>
<td id="6374F618006C0074FB77FB7E38430064" box="[1237,1281,1265,1287]" gridcol="7" gridrow="3" pageId="3" pageNumber="76">25.9</td>
<td id="6374F618006C0074FA97FB7E382A0064" box="[1333,1384,1265,1287]" gridcol="8" gridrow="3" pageId="3" pageNumber="76">10.4</td>
</tr>
<tr id="20A59F64006C0074FF03FA94382A0052" box="[161,1384,1307,1329]" gridrow="4" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03FA943C3C0052" box="[161,382,1307,1329]" gridcol="0" gridrow="4" pageId="3" pageNumber="76">RMNH.RENA.48235</th>
<td id="6374F618006C0074FE47FA943F740052" box="[485,566,1307,1329]" gridcol="1" gridrow="4" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FDD4FA943F8D0052" box="[630,719,1307,1329]" gridcol="2" gridrow="4" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5FA943E010052" box="[791,835,1307,1329]" gridcol="3" gridrow="4" pageId="3" pageNumber="76">16.3</td>
<td id="6374F618006C0074FCD5FA943EE10052" box="[887,931,1307,1329]" gridcol="4" gridrow="4" pageId="3" pageNumber="76">11.2</td>
<td id="6374F618006C0074FC7AFA9439440052" box="[984,1030,1307,1329]" gridcol="5" gridrow="4" pageId="3" pageNumber="76">22.0</td>
<td id="6374F618006C0074FBE3FA9439D20052" box="[1089,1168,1307,1329]" gridcol="6" gridrow="4" pageId="3" pageNumber="76">7.9</td>
<td id="6374F618006C0074FB77FA9438430052" box="[1237,1281,1307,1329]" gridcol="7" gridrow="4" pageId="3" pageNumber="76">22.9</td>
<td id="6374F618006C0074FA97FA94382A0052" box="[1333,1384,1307,1329]" gridcol="8" gridrow="4" pageId="3" pageNumber="76">8.8</td>
</tr>
<tr id="20A59F64006C0074FF03FACB382A0039" box="[161,1384,1348,1370]" gridrow="5" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03FACB3C3C0039" box="[161,382,1348,1370]" gridcol="0" gridrow="5" pageId="3" pageNumber="76">RMNH.RENA.48236</th>
<td id="6374F618006C0074FE47FACB3F740039" box="[485,566,1348,1370]" gridcol="1" gridrow="5" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FDD4FACB3F8D0039" box="[630,719,1348,1370]" gridcol="2" gridrow="5" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5FACB3E010039" box="[791,835,1348,1370]" gridcol="3" gridrow="5" pageId="3" pageNumber="76">15.5</td>
<td id="6374F618006C0074FCD5FACB3EE10039" box="[887,931,1348,1370]" gridcol="4" gridrow="5" pageId="3" pageNumber="76">10.7</td>
<td id="6374F618006C0074FC7AFACB39440039" box="[984,1030,1348,1370]" gridcol="5" gridrow="5" pageId="3" pageNumber="76">20.2</td>
<td id="6374F618006C0074FBE3FACB39D20039" box="[1089,1168,1348,1370]" gridcol="6" gridrow="5" pageId="3" pageNumber="76">7.7</td>
<td id="6374F618006C0074FB77FACB38430039" box="[1237,1281,1348,1370]" gridcol="7" gridrow="5" pageId="3" pageNumber="76">21.3</td>
<td id="6374F618006C0074FA97FACB382A0039" box="[1333,1384,1348,1370]" gridcol="8" gridrow="5" pageId="3" pageNumber="76">8.6</td>
</tr>
<tr id="20A59F64006C0074FF03FAE2382A00E0" box="[161,1384,1389,1411]" gridrow="6" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03FAE23C3C00E0" box="[161,382,1389,1411]" gridcol="0" gridrow="6" pageId="3" pageNumber="76">RMNH.RENA.48237</th>
<td id="6374F618006C0074FE47FAE23F7400E0" box="[485,566,1389,1411]" gridcol="1" gridrow="6" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FDD4FAE23F8D00E0" box="[630,719,1389,1411]" gridcol="2" gridrow="6" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5FAE23E0100E0" box="[791,835,1389,1411]" gridcol="3" gridrow="6" pageId="3" pageNumber="76">16.7</td>
<td id="6374F618006C0074FCD5FAE23EE100E0" box="[887,931,1389,1411]" gridcol="4" gridrow="6" pageId="3" pageNumber="76">11.9</td>
<td id="6374F618006C0074FC7AFAE2394400E0" box="[984,1030,1389,1411]" gridcol="5" gridrow="6" pageId="3" pageNumber="76">19.1</td>
<td id="6374F618006C0074FBE3FAE239D200E0" box="[1089,1168,1389,1411]" gridcol="6" gridrow="6" pageId="3" pageNumber="76">5.5</td>
<td id="6374F618006C0074FB77FAE2384300E0" box="[1237,1281,1389,1411]" gridcol="7" gridrow="6" pageId="3" pageNumber="76">23.7</td>
<td id="6374F618006C0074FA97FAE2382A00E0" box="[1333,1384,1389,1411]" gridcol="8" gridrow="6" pageId="3" pageNumber="76">9.3</td>
</tr>
<tr id="20A59F64006C0074FF03FA19382A00CF" box="[161,1384,1430,1452]" gridrow="7" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03FA193C3C00CF" box="[161,382,1430,1452]" gridcol="0" gridrow="7" pageId="3" pageNumber="76">RMNH.RENA.48238</th>
<td id="6374F618006C0074FE47FA193F7400CF" box="[485,566,1430,1452]" gridcol="1" gridrow="7" pageId="3" pageNumber="76">male</td>
<td id="6374F618006C0074FDD4FA193F8D00CF" box="[630,719,1430,1452]" gridcol="2" gridrow="7" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5FA193E0100CF" box="[791,835,1430,1452]" gridcol="3" gridrow="7" pageId="3" pageNumber="76">16.6</td>
<td id="6374F618006C0074FCD5FA193EE100CF" box="[887,931,1430,1452]" gridcol="4" gridrow="7" pageId="3" pageNumber="76">11.5</td>
<td id="6374F618006C0074FC7AFA19394400CF" box="[984,1030,1430,1452]" gridcol="5" gridrow="7" pageId="3" pageNumber="76">20.8</td>
<td id="6374F618006C0074FBE3FA1939D200CF" box="[1089,1168,1430,1452]" gridcol="6" gridrow="7" pageId="3" pageNumber="76">7.8</td>
<td id="6374F618006C0074FB77FA19384300CF" box="[1237,1281,1430,1452]" gridcol="7" gridrow="7" pageId="3" pageNumber="76">22.8</td>
<td id="6374F618006C0074FA97FA19382A00CF" box="[1333,1384,1430,1452]" gridcol="8" gridrow="7" pageId="3" pageNumber="76">9.4</td>
</tr>
<tr id="20A59F64006C0074FF03FA4F382A00B5" box="[161,1384,1472,1494]" gridrow="8" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03FA4F3C3C00B5" box="[161,382,1472,1494]" gridcol="0" gridrow="8" pageId="3" pageNumber="76">RMNH.RENA.48239</th>
<td id="6374F618006C0074FE47FA4F3F7400B5" box="[485,566,1472,1494]" gridcol="1" gridrow="8" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FDD4FA4F3F8D00B5" box="[630,719,1472,1494]" gridcol="2" gridrow="8" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5FA4F3E0100B5" box="[791,835,1472,1494]" gridcol="3" gridrow="8" pageId="3" pageNumber="76">16.8</td>
<td id="6374F618006C0074FCD5FA4F3EE100B5" box="[887,931,1472,1494]" gridcol="4" gridrow="8" pageId="3" pageNumber="76">11.2</td>
<td id="6374F618006C0074FC7AFA4F394400B5" box="[984,1030,1472,1494]" gridcol="5" gridrow="8" pageId="3" pageNumber="76">21.5</td>
<td id="6374F618006C0074FBE3FA4F39D200B5" box="[1089,1168,1472,1494]" gridcol="6" gridrow="8" pageId="3" pageNumber="76">6.8</td>
<td id="6374F618006C0074FB77FA4F384300B5" box="[1237,1281,1472,1494]" gridcol="7" gridrow="8" pageId="3" pageNumber="76">21.9</td>
<td id="6374F618006C0074FA97FA4F382A00B5" box="[1333,1384,1472,1494]" gridcol="8" gridrow="8" pageId="3" pageNumber="76">8.1</td>
</tr>
<tr id="20A59F64006C0074FF03FA66382A009C" box="[161,1384,1513,1535]" gridrow="9" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03FA663C3C009C" box="[161,382,1513,1535]" gridcol="0" gridrow="9" pageId="3" pageNumber="76">RMNH.RENA.48240</th>
<td id="6374F618006C0074FE47FA663F74009C" box="[485,566,1513,1535]" gridcol="1" gridrow="9" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FDD4FA663F8D009C" box="[630,719,1513,1535]" gridcol="2" gridrow="9" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5FA663E01009C" box="[791,835,1513,1535]" gridcol="3" gridrow="9" pageId="3" pageNumber="76">18.5</td>
<td id="6374F618006C0074FCD5FA663EE1009C" box="[887,931,1513,1535]" gridcol="4" gridrow="9" pageId="3" pageNumber="76">13.0</td>
<td id="6374F618006C0074FC7AFA663944009C" box="[984,1030,1513,1535]" gridcol="5" gridrow="9" pageId="3" pageNumber="76">23.6</td>
<td id="6374F618006C0074FBE3FA6639D2009C" box="[1089,1168,1513,1535]" gridcol="6" gridrow="9" pageId="3" pageNumber="76">8.9</td>
<td id="6374F618006C0074FB77FA663843009C" box="[1237,1281,1513,1535]" gridcol="7" gridrow="9" pageId="3" pageNumber="76">24.1</td>
<td id="6374F618006C0074FA97FA66382A009C" box="[1333,1384,1513,1535]" gridcol="8" gridrow="9" pageId="3" pageNumber="76">9.5</td>
</tr>
<tr id="20A59F64006C0074FF03F99C382A034A" box="[161,1384,1555,1577]" gridrow="10" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03F99C3C3C034A" box="[161,382,1555,1577]" gridcol="0" gridrow="10" pageId="3" pageNumber="76">RMNH.RENA.48241</th>
<td id="6374F618006C0074FE47F99C3F74034A" box="[485,566,1555,1577]" gridcol="1" gridrow="10" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FDD4F99C3F8D034A" box="[630,719,1555,1577]" gridcol="2" gridrow="10" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5F99C3E01034A" box="[791,835,1555,1577]" gridcol="3" gridrow="10" pageId="3" pageNumber="76">17.2</td>
<td id="6374F618006C0074FCD5F99C3EE1034A" box="[887,931,1555,1577]" gridcol="4" gridrow="10" pageId="3" pageNumber="76">11.5</td>
<td id="6374F618006C0074FC7AF99C3944034A" box="[984,1030,1555,1577]" gridcol="5" gridrow="10" pageId="3" pageNumber="76">20.8</td>
<td id="6374F618006C0074FBE3F99C39D2034A" box="[1089,1168,1555,1577]" gridcol="6" gridrow="10" pageId="3" pageNumber="76">7.9</td>
<td id="6374F618006C0074FB77F99C3843034A" box="[1237,1281,1555,1577]" gridcol="7" gridrow="10" pageId="3" pageNumber="76">22.2</td>
<td id="6374F618006C0074FA97F99C382A034A" box="[1333,1384,1555,1577]" gridcol="8" gridrow="10" pageId="3" pageNumber="76">7.8</td>
</tr>
<tr id="20A59F64006C0074FF03F9B3382A0331" box="[161,1384,1596,1618]" gridrow="11" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03F9B33C3C0331" box="[161,382,1596,1618]" gridcol="0" gridrow="11" pageId="3" pageNumber="76">RMNH.RENA.48242</th>
<td id="6374F618006C0074FE47F9B33F740331" box="[485,566,1596,1618]" gridcol="1" gridrow="11" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FDD4F9B33F8D0331" box="[630,719,1596,1618]" gridcol="2" gridrow="11" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5F9B33E010331" box="[791,835,1596,1618]" gridcol="3" gridrow="11" pageId="3" pageNumber="76">17.1</td>
<td id="6374F618006C0074FCD5F9B33EE10331" box="[887,931,1596,1618]" gridcol="4" gridrow="11" pageId="3" pageNumber="76">11.4</td>
<td id="6374F618006C0074FC7AF9B339440331" box="[984,1030,1596,1618]" gridcol="5" gridrow="11" pageId="3" pageNumber="76">20.9</td>
<td id="6374F618006C0074FBE3F9B339D20331" box="[1089,1168,1596,1618]" gridcol="6" gridrow="11" pageId="3" pageNumber="76">7.8</td>
<td id="6374F618006C0074FB77F9B338430331" box="[1237,1281,1596,1618]" gridcol="7" gridrow="11" pageId="3" pageNumber="76">21.1</td>
<td id="6374F618006C0074FA97F9B3382A0331" box="[1333,1384,1596,1618]" gridcol="8" gridrow="11" pageId="3" pageNumber="76">8.1</td>
</tr>
<tr id="20A59F64006C0074FF03F9EA382A0318" box="[161,1384,1637,1659]" gridrow="12" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03F9EA3C3C0318" box="[161,382,1637,1659]" gridcol="0" gridrow="12" pageId="3" pageNumber="76">RMNH.RENA.48243</th>
<td id="6374F618006C0074FE47F9EA3F740318" box="[485,566,1637,1659]" gridcol="1" gridrow="12" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FDD4F9EA3F8D0318" box="[630,719,1637,1659]" gridcol="2" gridrow="12" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5F9EA3E010318" box="[791,835,1637,1659]" gridcol="3" gridrow="12" pageId="3" pageNumber="76">18.6</td>
<td id="6374F618006C0074FCD5F9EA3EE10318" box="[887,931,1637,1659]" gridcol="4" gridrow="12" pageId="3" pageNumber="76">13.3</td>
<td id="6374F618006C0074FC7AF9EA39440318" box="[984,1030,1637,1659]" gridcol="5" gridrow="12" pageId="3" pageNumber="76">23.4</td>
<td id="6374F618006C0074FBE3F9EA39D20318" box="[1089,1168,1637,1659]" gridcol="6" gridrow="12" pageId="3" pageNumber="76">9.0</td>
<td id="6374F618006C0074FB77F9EA38430318" box="[1237,1281,1637,1659]" gridcol="7" gridrow="12" pageId="3" pageNumber="76">22.9</td>
<td id="6374F618006C0074FA97F9EA382A0318" box="[1333,1384,1637,1659]" gridcol="8" gridrow="12" pageId="3" pageNumber="76">7.1</td>
</tr>
<tr id="20A59F64006C0074FF03F901382A03C7" box="[161,1384,1678,1700]" gridrow="13" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03F9013C3C03C7" box="[161,382,1678,1700]" gridcol="0" gridrow="13" pageId="3" pageNumber="76">RMNH.RENA.48244</th>
<td id="6374F618006C0074FE47F9013F7403C7" box="[485,566,1678,1700]" gridcol="1" gridrow="13" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FDD4F9013F8D03C7" box="[630,719,1678,1700]" gridcol="2" gridrow="13" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5F9013E0103C7" box="[791,835,1678,1700]" gridcol="3" gridrow="13" pageId="3" pageNumber="76">18.7</td>
<td id="6374F618006C0074FCD5F9013EE103C7" box="[887,931,1678,1700]" gridcol="4" gridrow="13" pageId="3" pageNumber="76">13.5</td>
<td id="6374F618006C0074FC7AF901394403C7" box="[984,1030,1678,1700]" gridcol="5" gridrow="13" pageId="3" pageNumber="76">24.4</td>
<td id="6374F618006C0074FBE3F90139D203C7" box="[1089,1168,1678,1700]" gridcol="6" gridrow="13" pageId="3" pageNumber="76">9.5</td>
<td id="6374F618006C0074FB77F901384303C7" box="[1237,1281,1678,1700]" gridcol="7" gridrow="13" pageId="3" pageNumber="76">23.0</td>
<td id="6374F618006C0074FA97F901382A03C7" box="[1333,1384,1678,1700]" gridcol="8" gridrow="13" pageId="3" pageNumber="76">8.6</td>
</tr>
<tr id="20A59F64006C0074FF03F937382A03AD" box="[161,1384,1720,1742]" gridrow="14" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03F9373C3C03AD" box="[161,382,1720,1742]" gridcol="0" gridrow="14" pageId="3" pageNumber="76">RMNH.RENA.48245</th>
<td id="6374F618006C0074FE47F9373F7403AD" box="[485,566,1720,1742]" gridcol="1" gridrow="14" pageId="3" pageNumber="76">female</td>
<td id="6374F618006C0074FDD4F9373F8D03AD" box="[630,719,1720,1742]" gridcol="2" gridrow="14" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5F9373E0103AD" box="[791,835,1720,1742]" gridcol="3" gridrow="14" pageId="3" pageNumber="76">16.1</td>
<td id="6374F618006C0074FCD5F9373EE103AD" box="[887,931,1720,1742]" gridcol="4" gridrow="14" pageId="3" pageNumber="76">11.0</td>
<td id="6374F618006C0074FC7AF937394403AD" box="[984,1030,1720,1742]" gridcol="5" gridrow="14" pageId="3" pageNumber="76">20.7</td>
<td id="6374F618006C0074FBE3F93739D203AD" box="[1089,1168,1720,1742]" gridcol="6" gridrow="14" pageId="3" pageNumber="76">7.2</td>
<td id="6374F618006C0074FB77F937384303AD" box="[1237,1281,1720,1742]" gridcol="7" gridrow="14" pageId="3" pageNumber="76">22.6</td>
<td id="6374F618006C0074FA97F937382A03AD" box="[1333,1384,1720,1742]" gridcol="8" gridrow="14" pageId="3" pageNumber="76">8.0</td>
</tr>
<tr id="20A59F64006C0074FF03F96E382A0394" box="[161,1384,1761,1783]" gridrow="15" pageId="3" pageNumber="76">
<th id="6374F618006C0074FF03F96E3C3C0394" box="[161,382,1761,1783]" gridcol="0" gridrow="15" pageId="3" pageNumber="76">RMNH.RENA.48246</th>
<td id="6374F618006C0074FE47F96E3F740394" box="[485,566,1761,1783]" gridcol="1" gridrow="15" pageId="3" pageNumber="76">juvenile</td>
<td id="6374F618006C0074FDD4F96E3F8D0394" box="[630,719,1761,1783]" gridcol="2" gridrow="15" pageId="3" pageNumber="76">paratype</td>
<td id="6374F618006C0074FCB5F96E3E010394" box="[791,835,1761,1783]" gridcol="3" gridrow="15" pageId="3" pageNumber="76">15.2</td>
<td id="6374F618006C0074FCD5F96E3EE10394" box="[887,931,1761,1783]" gridcol="4" gridrow="15" pageId="3" pageNumber="76">10.3</td>
<td id="6374F618006C0074FC7AF96E39440394" box="[984,1030,1761,1783]" gridcol="5" gridrow="15" pageId="3" pageNumber="76">17.6</td>
<td id="6374F618006C0074FBE3F96E39D20394" box="[1089,1168,1761,1783]" gridcol="6" gridrow="15" pageId="3" pageNumber="76">6.6</td>
<td id="6374F618006C0074FB77F96E38430394" box="[1237,1281,1761,1783]" gridcol="7" gridrow="15" pageId="3" pageNumber="76">18.6</td>
<td id="6374F618006C0074FA97F96E382A0394" box="[1333,1384,1761,1783]" gridcol="8" gridrow="15" pageId="3" pageNumber="76">6.7</td>
</tr>
</table>
</paragraph>
<paragraph id="9E2A9D26006CFF88FF35F8973E2E02CC" blockId="3.[151,1435,1816,1967]" pageId="3" pageNumber="76">
<tableNote id="63739CA8006CFF88FF35F8973E2E02CC" pageId="3" pageNumber="76" targetBox="[161,1384,1140,1783]" targetPageId="3">
Abbreviations used are:
<collectionCode id="F88405E3006CFF88FE04F8973CB2024C" box="[422,496,1816,1839]" pageId="3" pageNumber="76">NRBV</collectionCode>
= Number of rib-bearing pre-sacral vertebrae; SVL1 = Snout-vent length up to and including the insertion of the hind-legs; SVL2 = Snout-vent length up to and including the cloaca; TL = Total length; ILD = Inter-limb distance; HL = Head length; HW = Head width; Arm = Length of the right arm including the third finger; Finger3 = Length of the right third finger; Leg = Length of the right leg including the fourth toe; Toe4 = Length of the right fourth toe. * Finger c3/4 regenerated; ** Deformed tail tip.
</tableNote>
</paragraph>
<caption id="CAEACDAE006BFF8FFF35F81339D802B2" httpUri="https://zenodo.org/record/260658/files/figure.png" pageId="4" pageNumber="77" targetBox="[153,1433,195,1926]" targetPageId="4">
<paragraph id="9E2A9D26006BFF8FFF35F81339D802B2" blockId="4.[151,1435,1948,2001]" pageId="4" pageNumber="77">
<emphasis id="ACE14134006BFF8FFF35F8133C5102D2" bold="true" box="[151,275,1948,1970]" pageId="4" pageNumber="77">FIGURE 3.</emphasis>
The holotype and one of the paratypes of
<taxonomicName id="5995E6A5006BFF8FFD66F8123EC402D1" box="[708,902,1948,1970]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="77" phylum="Chordata" rank="species" species="anatolicus" status="sp. nov.">
<emphasis id="ACE14134006BFF8FFD66F8123EC402D1" box="[708,902,1948,1970]" italics="true" pageId="4" pageNumber="77">Triturus anatolicus</emphasis>
</taxonomicName>
<emphasis id="ACE14134006BFF8FFC2EF8123E9C02D1" bold="true" box="[908,990,1949,1970]" pageId="4" pageNumber="77">
<taxonomicNameLabel id="B7D2FC4F006BFF8FFC2EF8123E9C02D1" box="[908,990,1949,1970]" pageId="4" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life. Lateral (a) and ventral (b) view of a male (below, holotype, RMNH.RENA.48232) and a female (above, paratype, RMNH.RENA.48239).
</paragraph>
</caption>
<caption id="CAEACDAE006AFF8EFF35FBFC3E8101EA" box="[151,963,1139,1161]" httpUri="https://zenodo.org/record/260659/files/figure.png" pageId="5" pageNumber="78" targetBox="[176,1422,201,1102]" targetPageId="5">
<paragraph id="9E2A9D26006AFF8EFF35FBFC3E8101EA" blockId="5.[151,963,1139,1161]" box="[151,963,1139,1161]" pageId="5" pageNumber="78">
<emphasis id="ACE14134006AFF8EFF35FBFC3C5001EB" bold="true" box="[151,274,1139,1161]" pageId="5" pageNumber="78">FIGURE 4.</emphasis>
The type locality of
<taxonomicName id="5995E6A5006AFF8EFE44FBFB3FE501EA" box="[486,679,1139,1161]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="78" phylum="Chordata" rank="species" species="anatolicus" status="sp. nov.">
<emphasis id="ACE14134006AFF8EFE44FBFB3FE501EA" box="[486,679,1139,1161]" italics="true" pageId="5" pageNumber="78">Triturus anatolicus</emphasis>
</taxonomicName>
<emphasis id="ACE14134006AFF8EFD0CFBFC3FBC01EB" bold="true" box="[686,766,1139,1160]" pageId="5" pageNumber="78">
<taxonomicNameLabel id="B7D2FC4F006AFF8EFD0CFBFC3FBC01EB" box="[686,766,1139,1160]" pageId="5" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at Gölköy, Turkey.
</paragraph>
</caption>
<paragraph id="9E2A9D26006AFF8EFF65FB3A3F4D00E1" blockId="5.[151,1437,1205,2022]" pageId="5" pageNumber="78">The number of rib-bearing pre-sacral vertebrae (NRBV) is 13, as determined from X-ray photographs (see online Appendix 2). The following measurements are in millimeters and have been determined with plastic Vernier calipers. Snout-vent length up to and including the hind-legs 53; Snout-vent length up to and including the cloaca 61; Total length 108; Inter-limb distance 27; Head length 16.2; Head width 10.6; Length of the right arm including the third finger 18.7; Length of the right third finger 5.6; Length of the right leg including the fourth toe 22.1; Length of the right fourth toe 9.5.</paragraph>
<paragraph id="9E2A9D26006AFF8EFF65FA0238DA0355" blockId="5.[151,1437,1205,2022]" pageId="5" pageNumber="78">
<emphasis id="ACE14134006AFF8EFF65FA023CB700C5" bold="true" box="[199,501,1421,1446]" pageId="5" pageNumber="78">
Description of
<typeStatus id="412E2384006AFF8EFED5FA023CB200C5" box="[375,496,1421,1446]" pageId="5" pageNumber="78" type="paratype">paratypes</typeStatus>
.
</emphasis>
The morphology of the
<typeStatus id="412E2384006AFF8EFCA7FA013E3200C5" box="[773,880,1422,1446]" pageId="5" pageNumber="78" type="paratype">paratypes</typeStatus>
resembles that of the
<typeStatus id="412E2384006AFF8EFBCAFA02398800C5" box="[1128,1226,1421,1446]" pageId="5" pageNumber="78" type="holotype">holotype</typeStatus>
in general, but the pattern of black spots on the ventral side is highly variable among individuals (online Appendix 1). Compared to males (such as the
<typeStatus id="412E2384006AFF8EFECBFA5A3C92008D" box="[361,464,1493,1518]" pageId="5" pageNumber="78" type="holotype">holotype</typeStatus>
), adult females have non-swollen cloacae, lack the denticulated crest and possess a less pronounced tail fin. All
<typeStatus id="412E2384006AFF8EFE05FA753F500371" box="[423,530,1530,1554]" pageId="5" pageNumber="78" type="paratype">paratypes</typeStatus>
have an NRBV count of 13 except one, which has an NRBV count of 12 (
<tableCitation id="D317A89D006AFF8EFAFEFA763DE40355" captionStart="TABLE 1" captionStartId="3.[151,239,328,351]" captionTargetBox="[160,1383,410,1053]" captionTargetPageId="3" captionText="TABLE 1. Number of rib-bearing pre-sacral vertebrae and measurements (in millimeters) for the holotype and paratypes of Triturus anatolicus sp. nov." httpUri="http://table.plazi.org/id/CAEACDAE006CFF88FF35FEC73C99041C" pageId="5" pageNumber="78" tableUuid="CAEACDAE006CFF88FF35FEC73C99041C">Table 1</tableCitation>
; online Appendix 2). In
<tableCitation id="D317A89D006AFF8EFE18F9923F4E0355" box="[442,524,1565,1590]" captionStart="TABLE 1" captionStartId="3.[151,239,328,351]" captionTargetBox="[160,1383,410,1053]" captionTargetPageId="3" captionText="TABLE 1. Number of rib-bearing pre-sacral vertebrae and measurements (in millimeters) for the holotype and paratypes of Triturus anatolicus sp. nov." httpUri="http://table.plazi.org/id/CAEACDAE006CFF88FF35FEC73C99041C" pageId="5" pageNumber="78" tableUuid="CAEACDAE006CFF88FF35FEC73C99041C">Table 1</tableCitation>
the same measurements as taken for the
<typeStatus id="412E2384006AFF8EFC75F992397B0355" box="[983,1081,1565,1590]" pageId="5" pageNumber="78" type="holotype">holotype</typeStatus>
are provided for the
<typeStatus id="412E2384006AFF8EFA87F99138D60355" box="[1317,1428,1566,1590]" pageId="5" pageNumber="78" type="paratype">paratypes</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="D68FCEAD006AFF8EFF65F9CE394F0235" pageId="5" pageNumber="78" type="distribution">
<paragraph id="9E2A9D26006AFF8EFF65F9CE394F0235" blockId="5.[151,1437,1205,2022]" pageId="5" pageNumber="78">
<emphasis id="ACE14134006AFF8EFF65F9CE3C220339" bold="true" box="[199,352,1601,1626]" pageId="5" pageNumber="78">Distribution.</emphasis>
The distribution encompasses Asiatic
<collectingCountry id="E682DDB6006AFF8EFC84F9CE3E340339" box="[806,886,1601,1626]" name="Turkey" pageId="5" pageNumber="78">Turkey</collectingCountry>
south of the Black Sea, reaching up to c. 200 kilometers inland (usually less), but not into inner
<collectingCountry id="E682DDB6006AFF8EFD73F9EA3E77031D" box="[721,821,1637,1662]" name="Turkey" pageId="5" pageNumber="78">Anatolia</collectingCountry>
. To the west the new species reaches the Bosphorus at the northern side of the Marmara Sea. On the southern side of the Marmara Sea it meets
<emphasis id="ACE14134006AFF8EFB04F9053D9D03A6" italics="true" pageId="5" pageNumber="78">
<taxonomicName id="5995E6A5006AFF8EFB04F90538DE03C2" box="[1190,1436,1673,1698]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="78" phylum="Chordata" rank="subSpecies" species="ivanbureschi" subSpecies="sensu">
T. ivanbureschi
<taxonomicNameLabel id="B7D2FC4F006AFF8EFAFCF90538DE03C2" box="[1374,1436,1674,1697]" pageId="5" pageNumber="78" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
, east of Lake Ulubat and west of the city of Bursa. The two species form a hybrid zone here (Wielstra
<emphasis id="ACE14134006AFF8EFAFFF92138D703A5" box="[1373,1429,1709,1734]" italics="true" pageId="5" pageNumber="78">et al.</emphasis>
, submitted). To the east the new species reaches the town of Yomra, just east of the city of Trabzon. The nearest known
<taxonomicName id="5995E6A5006AFF8EFF48F9793C06026E" box="[234,324,1782,1805]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="ACE14134006AFF8EFF48F9793C06026E" box="[234,324,1782,1805]" italics="true" pageId="5" pageNumber="78">Triturus</emphasis>
</taxonomicName>
localities further east are from the extreme NE of
<collectingCountry id="E682DDB6006AFF8EFCD6F97A3E87026D" box="[884,965,1781,1806]" name="Turkey" pageId="5" pageNumber="78">Turkey</collectingCountry>
, over
<quantity id="596D30C3006AFF8EFBAAF97A391F026D" box="[1032,1117,1781,1806]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="78" unit="km" value="150.0">150 km</quantity>
away, and probably concern
<emphasis id="ACE14134006AFF8EFF35F8953CEB0252" box="[151,425,1817,1842]" italics="true" pageId="5" pageNumber="78">
<taxonomicName id="5995E6A5006AFF8EFF35F8953C1B0252" box="[151,345,1817,1842]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="78" phylum="Chordata" rank="subSpecies" species="karelinii" subSpecies="sensu">
T. karelinii
<taxonomicNameLabel id="B7D2FC4F006AFF8EFEB9F8953C1B0252" box="[283,345,1818,1841]" pageId="5" pageNumber="78" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
(
<bibRefCitation id="FA04E0D7006AFF8EFE18F8963FF40251" author="Wielstra" box="[442,694,1817,1842]" pageId="5" pageNumber="78" refString="Wielstra, B., Baird, A. B. &amp; Arntzen, J. W. (2013 a) A multimarker phylogeography of crested newts (Triturus cristatus superspecies) reveals cryptic species. Molecular Phylogenetics and Evolution, 67, 167 - 175. http: // dx. doi. org / 10.1016 / j. ympev. 2013.01.009" type="journal article" year="2013" yearSuffix="a">
Wielstra
<emphasis id="ACE14134006AFF8EFD81F8953F1F0251" box="[547,605,1817,1842]" italics="true" pageId="5" pageNumber="78">et al.</emphasis>
, 2013a
</bibRefCitation>
). An outline of the distribution of the new species is provided in
<figureCitation id="06AE81A3006AFF8EFF35F8B23D9B0235" box="[151,217,1853,1878]" captionStart="FIGURE 1" captionStartId="1.[151,250,1500,1522]" captionTargetBox="[151,1436,1021,1478]" captionTargetId="figure@1.[151,1436,1021,1479]" captionTargetPageId="1" captionText="FIGURE 1. The two cryptic species comprising Triturus ivanbureschi sensu lato are distributed in Asiatic Turkey and the south-eastern Balkan Peninsula. A rough distribution outline for T. ivanbureschi sensu stricto is shown in green and for T. anatolicus sp. nov. in red. The two taxa meet at a hybrid zone in northwestern Asiatic Turkey. A rough distribution outline for T. karelinii sensu stricto with which Triturus ivanbureschi sensu lato until recently was considered to be conspecific is in blue. Records of Triturus newts are absent from most of NE Turkey, suggesting that the ranges of T. anatolicus sp. nov. and T. karelinii sensu stricto do not meet. White dots represent type localities for, from left to right, T. ivanbureschi sensu stricto, T. anatolicus sp. nov. and T. karelinii sensu stricto." httpUri="https://zenodo.org/record/260656/files/figure.png" pageId="5" pageNumber="78">Fig. 1</figureCitation>
. A database of distribution records is provided in
<bibRefCitation id="FA04E0D7006AFF8EFCA8F8B2394B0235" box="[778,1033,1853,1878]" pageId="5" pageNumber="78" refString="Wielstra, B., Sillero, N., Voros, J. &amp; Arntzen, J. W. (2014 b) The distribution of the crested and marbled newt species (Amphibia: Salamandridae: Triturus) - an addition to the New Atlas of Amphibians and Reptiles of Europe. Amphibia- Reptilia, 35, 376 - 381. http: // dx. doi. org / 10.1163 / 15685381 - 00002960" type="journal article">
Wielstra
<emphasis id="ACE14134006AFF8EFCCCF8B13EE40235" box="[878,934,1853,1878]" italics="true" pageId="5" pageNumber="78">et al.</emphasis>
(2014b)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D68FCEAD006AFF8DFF65F8EE3F6A0423" lastPageId="6" lastPageNumber="79" pageId="5" pageNumber="78" type="etymology">
<paragraph id="9E2A9D26006AFF8EFF65F8EE38500285" blockId="5.[151,1437,1205,2022]" pageId="5" pageNumber="78">
<emphasis id="ACE14134006AFF8EFF65F8EE3C0C0219" bold="true" box="[199,334,1889,1914]" pageId="5" pageNumber="78">Etymology.</emphasis>
The specific epithet reflects the distribution of the new
<taxonomicName id="5995E6A5006AFF8EFC77F8ED396D021A" box="[981,1071,1890,1913]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="ACE14134006AFF8EFC77F8ED396D021A" box="[981,1071,1890,1913]" italics="true" pageId="5" pageNumber="78">Triturus</emphasis>
</taxonomicName>
species.
<taxonomicName id="5995E6A5006AFF8EFB3BF8ED38300219" box="[1177,1394,1889,1914]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="78" phylum="Chordata" rank="species" species="anatolicus" status="sp. nov.">
<emphasis id="ACE14134006AFF8EFB3BF8ED38300219" box="[1177,1394,1889,1914]" italics="true" pageId="5" pageNumber="78">Triturus anatolicus</emphasis>
</taxonomicName>
<emphasis id="ACE14134006AFF8EFAD8F8EE3D8A02FD" bold="true" pageId="5" pageNumber="78">
<taxonomicNameLabel id="B7D2FC4F006AFF8EFAD8F8EE3D8A02FD" pageId="5" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is endemic to
<collectingCountry id="E682DDB6006AFF8EFEC8F80A3C8C02FD" box="[362,462,1925,1950]" name="Turkey" pageId="5" pageNumber="78">Anatolia</collectingCountry>
. It is the only
<taxonomicName id="5995E6A5006AFF8EFDC9F8093F8602FE" box="[619,708,1926,1949]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="ACE14134006AFF8EFDC9F8093F8602FE" box="[619,708,1926,1949]" italics="true" pageId="5" pageNumber="78">Triturus</emphasis>
</taxonomicName>
species to which this applies. It should be noted that the range of
<emphasis id="ACE14134006AFF8EFF35F8253C9F02A2" box="[151,477,1961,1986]" italics="true" pageId="5" pageNumber="78">
<taxonomicName id="5995E6A5006AFF8EFF35F8253CCE02A2" box="[151,396,1961,1986]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="78" phylum="Chordata" rank="subSpecies" species="ivanbureschi" subSpecies="sensu">
T. ivanbureschi
<taxonomicNameLabel id="B7D2FC4F006AFF8EFEECF8253CCE02A2" box="[334,396,1962,1985]" pageId="5" pageNumber="78" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
covers western
<collectingCountry id="E682DDB6006AFF8EFD38F8263FB802A1" box="[666,762,1961,1986]" name="Turkey" pageId="5" pageNumber="78">Anatolia</collectingCountry>
(
<bibRefCitation id="FA04E0D7006AFF8EFCA9F826394502A1" author="Wielstra" box="[779,1031,1961,1986]" pageId="5" pageNumber="78" refString="Wielstra, B., Baird, A. B. &amp; Arntzen, J. W. (2013 a) A multimarker phylogeography of crested newts (Triturus cristatus superspecies) reveals cryptic species. Molecular Phylogenetics and Evolution, 67, 167 - 175. http: // dx. doi. org / 10.1016 / j. ympev. 2013.01.009" type="journal article" year="2013" yearSuffix="a">
Wielstra
<emphasis id="ACE14134006AFF8EFCD6F8253EED02A1" box="[884,943,1961,1986]" italics="true" pageId="5" pageNumber="78">et al.</emphasis>
, 2013a
</bibRefCitation>
; Wielstra
<emphasis id="ACE14134006AFF8EFBDEF82539F402A1" box="[1148,1206,1961,1986]" italics="true" pageId="5" pageNumber="78">et al.</emphasis>
, submitted) and the range of
<emphasis id="ACE14134006AFF8EFF58F8413F4A0286" box="[250,520,1997,2022]" italics="true" pageId="5" pageNumber="78">
<taxonomicName id="5995E6A5006AFF8EFF58F8413CFB0286" box="[250,441,1997,2022]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="78" phylum="Chordata" rank="subSpecies" species="karelinii" subSpecies="sensu">
T. karelinii
<taxonomicNameLabel id="B7D2FC4F006AFF8EFED9F8413CFB0286" box="[379,441,1998,2021]" pageId="5" pageNumber="78" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
probably protrudes into northeastern
<collectingCountry id="E682DDB6006AFF8EFC0FF842394C0285" box="[941,1038,1997,2022]" name="Turkey" pageId="5" pageNumber="78">Anatolia</collectingCountry>
(Wielstra
<emphasis id="ACE14134006AFF8EFB21F84139F90285" box="[1155,1211,1997,2022]" italics="true" pageId="5" pageNumber="78">et al.</emphasis>
, 2010).
</paragraph>
<paragraph id="9E2A9D260069FF8DFF65FF183F6A0423" blockId="6.[151,1436,151,320]" pageId="6" pageNumber="79">
<emphasis id="ACE141340069FF8DFF65FF183F4C05D3" bold="true" box="[199,526,151,176]" pageId="6" pageNumber="79">Proposed vernacular name.</emphasis>
We propose to use the vernacular name Anatolian Crested Newt for
<taxonomicName id="5995E6A50069FF8DFAAEFF1638DE05D3" box="[1292,1436,151,176]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="79" phylum="Chordata" rank="species" species="anatolicus" status="sp. nov.">
<emphasis id="ACE141340069FF8DFAAEFF1638DE05D3" box="[1292,1436,151,176]" italics="true" pageId="6" pageNumber="79">T. anatolicus</emphasis>
</taxonomicName>
<emphasis id="ACE141340069FF8DFF35FF333DB005B6" bold="true" box="[151,242,188,213]" pageId="6" pageNumber="79">
<taxonomicNameLabel id="B7D2FC4F0069FF8DFF35FF333DB005B6" box="[151,242,188,213]" pageId="6" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
This name highlights its status as an Anatolian endemic. We suggest to use the vernacular name Balkan Crested Newt for
<emphasis id="ACE141340069FF8DFEFFFF6E3FD9059B" box="[349,667,223,248]" italics="true" pageId="6" pageNumber="79">
<taxonomicName id="5995E6A50069FF8DFEFFFF6E3F0E059B" box="[349,588,223,248]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="79" phylum="Chordata" rank="subSpecies" species="ivanbureschi" subSpecies="sensu">
T. ivanbureschi
<taxonomicNameLabel id="B7D2FC4F0069FF8DFDACFF6E3F0E059B" box="[526,588,225,248]" pageId="6" pageNumber="79" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
(rather than Balkan-Anatolian Crested Newt previously applied to
<emphasis id="ACE141340069FF8DFA25FF6E3CDD047E" italics="true" pageId="6" pageNumber="79">
<taxonomicName id="5995E6A50069FF8DFA25FF6E3C2E047F" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="79" phylum="Chordata" rank="subSpecies" species="ivanbureschi" subSpecies="sensu">
T. ivanbureschi
<taxonomicNameLabel id="B7D2FC4F0069FF8DFE8CFE8A3C2E047F" box="[302,364,261,284]" pageId="6" pageNumber="79" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato
</emphasis>
). Although
<emphasis id="ACE141340069FF8DFD8BFE8A3E28047F" box="[553,874,260,285]" italics="true" pageId="6" pageNumber="79">
<taxonomicName id="5995E6A50069FF8DFD8BFE8A3E58047F" box="[553,794,260,285]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="79" phylum="Chordata" rank="subSpecies" species="ivanbureschi" subSpecies="sensu">
T. ivanbureschi
<taxonomicNameLabel id="B7D2FC4F0069FF8DFD7EFE8A3E58047F" box="[732,794,261,284]" pageId="6" pageNumber="79" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
also partially occurs in
<collectingCountry id="E682DDB60069FF8DFBDBFE8B399E047E" box="[1145,1244,260,285]" name="Turkey" pageId="6" pageNumber="79">Anatolia</collectingCountry>
, the main part of its range is in the Balkan Peninsula.
</paragraph>
</subSubSection>
</treatment>
</document>