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<mods:title id="B0F2CC41D4281C0C2305A3788A31F880">Taxonomic Revision of the Spider Genus Actinopus Perty, 1833 (Araneae, Mygalomorphae, Actinopodidae)</mods:title>
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<mods:namePart id="3E0B8DF70B3D94B1D2497FEB46B25365">Miglio, Laura Tavares</mods:namePart>
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<taxonomicName id="46DCD9FFFF80CE50FCD5F9A9DFC43ABC" ID-CoL="64PM6" authority="Perty, 1833" authorityName="Perty" authorityYear="1833" box="[810,1175,1656,1683]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B3A87E6EFF80CE50FCD5F9A9DFC43ABC" bold="true" box="[810,1175,1656,1683]" pageId="13" pageNumber="14">
<emphasis id="B3A87E6EFF80CE50FCD5F9A9DF503ABD" bold="true" box="[810,1027,1656,1683]" italics="true" pageId="13" pageNumber="14">Actinopus tarsalis</emphasis>
<bibRefCitation id="E54DDF8DFF80CE50FBF4F9A9DFC43ABC" author="Perty, M." box="[1035,1175,1656,1682]" pageId="13" pageNumber="14" pagination="191 - 209" refId="ref139886" refString="Perty, M. (1833) Arachnides Brasilienses. In: de Spix, J. B. &amp; Martius, F. P. (Eds.), Delectus animalium articulatorum quae in itinere per Braziliam ann. 1817 et 1820 colligerunt. Monachii, pp. 191 - 209." type="book chapter" year="1833">Perty, 1833</bibRefCitation>
</emphasis>
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<figureCitation id="19E7BEF9FF80CE50FCD5F94DD8383A98" box="[810,875,1692,1718]" captionStart="FIGURE 9" captionStartId="14.[136,235,952,976]" captionTargetBox="[170,1419,201,919]" captionTargetId="figure-21@14.[151,1436,189,929]" captionTargetPageId="14" captionText="FIGURE 9. Actinopus tarsalis Perty, 1833 (drawings from Miglio et al., 2012), male: AC MPEG 11717: A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral. PA-Paraembolic apophysis; PAc-ProlateralAccessory keel; PI-Prolateral Inferior keel; PS-Prolateral Superior keel; Arrow: Serrated Area. Scale line: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5597255" httpUri="https://zenodo.org/record/5597255/files/figure.png" pageId="13" pageNumber="14">Fig. 9</figureCitation>
AC,
<figureCitation id="19E7BEF9FF80CE50FC4CF94CD8AF3A98" box="[947,1020,1692,1719]" captionStart="MAP 1" captionStartId="14.[136,192,2041,2065]" captionTargetBox="[157,1428,1114,2008]" captionTargetId="figure-76@14.[151,1435,1108,2016]" captionTargetPageId="14" captionText="MAP 1. Distribution map of Actinopus tarsalis Perty, 1833, and Actinopus castelo sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5655402" httpUri="https://zenodo.org/record/5655402/files/figure.png" pageId="13" pageNumber="14">Map 1</figureCitation>
</paragraph>
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<treatmentCitation id="007D846DFF80CE50FCD5F90FDFC33ADC" author="Perty, M." box="[810,1168,1757,1779]" page="198" pageId="13" pageNumber="14" year="1833">
<taxonomicName id="46DCD9FFFF80CE50FCD5F90FDFC33ADC" ID-CoL="64PM6" authority="Perty, 1833: 198" authorityName="Perty" authorityPageNumber="198" authorityYear="1833" box="[810,1168,1757,1779]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B3A87E6EFF80CE50FCD5F90FD88F3ADC" box="[810,988,1757,1779]" italics="true" pageId="13" pageNumber="14">Actinopus tarsalis</emphasis>
<bibRefCitation id="E54DDF8DFF80CE50FC18F90FDFC33ADC" author="Perty, M." box="[999,1168,1757,1779]" pageId="13" pageNumber="14" pagination="191 - 209" refId="ref139886" refString="Perty, M. (1833) Arachnides Brasilienses. In: de Spix, J. B. &amp; Martius, F. P. (Eds.), Delectus animalium articulatorum quae in itinere per Braziliam ann. 1817 et 1820 colligerunt. Monachii, pp. 191 - 209." type="book chapter" year="1833">Perty, 1833: 198</bibRefCitation>
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, pl. 39, fig. 6 (
<typeStatus id="5E671CDEFF80CE50FACCF90CDEDA3ADD" box="[1331,1417,1757,1779]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
♂,
<collectingRegion id="43186C9EFF80CE50FCA0F92DD8C03B3C" box="[863,915,1788,1810]" country="Brazil" name="Piaui" pageId="13" pageNumber="14">Piauí</collectingRegion>
,
<collectingCountry id="F9CBE2ECFF80CE50FC64F92DD8843B3C" box="[923,983,1788,1810]" name="Brazil" pageId="13" pageNumber="14">Brasil</collectingCountry>
, MNHN, not located); C.L.
<bibRefCitation id="E54DDF8DFF80CE50FB1EF92DDED13B3F" author="Koch, C. L." box="[1249,1410,1788,1810]" pageId="13" pageNumber="14" refId="ref138989" refString="Koch, C. L. (1842) Die Arachniden. Band 9; Band 10. Zeh'schen Buchhandlung, Nurnberg, 108 pp; 36 pp." type="book" year="1842">Koch, 1842: 101</bibRefCitation>
, fig. 753;
<bibRefCitation id="E54DDF8DFF80CE50FC6DF8CADF643B1E" author="Simon, E." box="[914,1079,1819,1841]" pageId="13" pageNumber="14" refId="ref140707" refString="Simon, E. (1892) Histoire naturelle des araignees. Roret, Paris, 256 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1892">Simon, 1892: 80</bibRefCitation>
, figs 80, 82;
<bibRefCitation id="E54DDF8DFF80CE50FB43F8CADE3C3B1E" author="Lucas, S. &amp; Cirelli, A. &amp; Knysak, I. &amp; Zveibil, L." box="[1212,1391,1819,1840]" pageId="13" pageNumber="14" pagination="127 - 138" refId="ref139265" refString="Lucas, S., Cirelli, A., Knysak, I. &amp; Zveibil, L. (1978) Aracnideos coletados no Piaui durante a realizac &quot; o do Projeto Rondon XXII. Memorias do Instituto Butantan, 42, 127 - 138." type="journal article" year="1978">
Lucas
<emphasis id="B3A87E6EFF80CE50FB01F8CADE623B1E" box="[1278,1329,1819,1840]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
, 1978
</bibRefCitation>
/
<bibRefCitation id="E54DDF8DFF80CE50FA88F8CADEFF3B1E" author="Lucas, S. &amp; Cirelli, A. &amp; Knysak, I. &amp; Zveibil, L." box="[1399,1452,1819,1840]" pageId="13" pageNumber="14" pagination="127 - 138" refId="ref139312" refString="Lucas, S., Cirelli, A., Knysak, I. &amp; Zveibil, L. (1979) Aracnideos coletados no Piaui durante a realizac &quot; o do Projeto Rondon XXII. Memorias do Instituto Butantan, 43, 127 - 138." type="journal article" year="1979">1979:</bibRefCitation>
133, figs 36;
<bibRefCitation id="E54DDF8DFF80CE50FC17F8E8DF983B61" author="Miglio, L. T. &amp; Lucas, S. M. &amp; Bonaldo, A. B." box="[1000,1227,1849,1871]" pageId="13" pageNumber="14" pagination="275 - 379" refId="ref139626" refString="Miglio, L. T., Lucas, S. M. &amp; Bonaldo, A. B. (2012) On the identity of the type species Actinopus tarsalis (Araneae: Actinopodidae). Zoologia, 29 (4), 275 - 379. https: // doi. org / 10.1590 / S 1984 - 46702012000400009" type="journal article" year="2012">
Miglio
<emphasis id="B3A87E6EFF80CE50FBD1F8EBDF0D3B60" box="[1070,1118,1849,1871]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
, 2012: 376
</bibRefCitation>
, figs 112;
<bibRefCitation id="E54DDF8DFF80CE50FAD3F8E8DEFF3B61" author="World Spider Catalog" box="[1324,1452,1849,1871]" pageId="13" pageNumber="14" refId="ref140996" refString="World Spider Catalog (2020) World Spider Catalog. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 22 July 2020)" type="url" year="2020">World Spider</bibRefCitation>
Catalog, 2020.
</paragraph>
<paragraph id="8163A27CFF80CE50FCD5F8A6DFE63BA2" blockId="13.[810,1452,1756,2092]" box="[810,1205,1911,1932]" pageId="13" pageNumber="14">
<emphasis id="B3A87E6EFF80CE50FCD5F8A6D8FD3BA2" box="[810,942,1911,1932]" italics="true" pageId="13" pageNumber="14">Pachyloscelis</emphasis>
<taxonomicName id="46DCD9FFFF80CE50FC4BF8A6DFE23BA2" ID-CoL="64PM6" authority=": Lucas, 1837: 377" authorityName="Lucas" authorityPageNumber="377" authorityYear="1837" box="[948,1201,1911,1932]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B3A87E6EFF80CE50FC4BF8A6D8AF3BA2" box="[948,1020,1911,1932]" italics="true" pageId="13" pageNumber="14">tarsalis</emphasis>
:
<bibRefCitation id="E54DDF8DFF80CE50FBF7F8A6DFE23BA2" author="Lucas, H." box="[1032,1201,1911,1932]" pageId="13" pageNumber="14" pagination="369 - 392" refId="ref139111" refString="Lucas, H. (1837) Observations sur les araneides du genre Pachyloscelis, et synonymie de ce genre. Annales de la Societe Entomologique de France, 6, 369 - 392." type="journal article" year="1837">Lucas, 1837: 377</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8163A27CFF80CE50FCD5F847DF923B85" blockId="13.[810,1452,1756,2092]" box="[810,1217,1942,1964]" pageId="13" pageNumber="14">
<taxonomicName id="46DCD9FFFF80CE50FCD5F847DFEE3B85" authority=": Walckenaer, 1842: 437" authorityName="Walckenaer" authorityPageNumber="437" authorityYear="1842" box="[810,1213,1942,1964]" class="Arachnida" family="Atypidae" genus="Sphodros" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B3A87E6EFF80CE50FCD5F847D8823B85" box="[810,977,1942,1963]" italics="true" pageId="13" pageNumber="14">Sphodros tarsalis</emphasis>
:
<bibRefCitation id="E54DDF8DFF80CE50FC21F847DFEE3B85" author="Walckenaer, C. A." box="[990,1213,1942,1964]" pageId="13" pageNumber="14" refId="ref140850" refString="Walckenaer, C. A. (1842) Histoire naturelle des Insects. Apteres. Roret, Paris, v. 2, 549 pp." type="book" year="1842">Walckenaer, 1842: 437</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8163A27CFF80CE50FCD5F865DE5F3BE4" blockId="13.[810,1452,1756,2092]" box="[810,1292,1972,1994]" pageId="13" pageNumber="14">
<emphasis id="B3A87E6EFF80CE50FCD5F865D8FD3BE7" box="[810,942,1972,1993]" italics="true" pageId="13" pageNumber="14">Pachyloscelis</emphasis>
<taxonomicName id="46DCD9FFFF80CE50FC4BF864DFD43BE4" ID-CoL="64PKR" authority=": Mello-Leit" authorityName="Mello-Leit" box="[948,1159,1972,1994]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="B3A87E6EFF80CE50FC4BF864DF5E3BE4" box="[948,1037,1973,1994]" italics="true" pageId="13" pageNumber="14">crassipes</emphasis>
: Mello-Leit
</taxonomicName>
„o, 1943: 149.
</paragraph>
<paragraph id="8163A27CFF80CE50FCD5F802DFAE3BC6" blockId="13.[810,1452,1756,2092]" box="[810,1277,2003,2025]" pageId="13" pageNumber="14">
<emphasis id="B3A87E6EFF80CE50FCD5F802D8FD3BC6" box="[810,942,2003,2024]" italics="true" pageId="13" pageNumber="14">Pachyloscelis</emphasis>
<taxonomicName id="46DCD9FFFF80CE50FC4BF802DF2B3BC7" authority=": Mello-Leit" authorityName="Mello-Leit" box="[948,1144,2003,2025]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="B3A87E6EFF80CE50FC4BF802D8AC3BC6" box="[948,1023,2003,2024]" italics="true" pageId="13" pageNumber="14">luteipes</emphasis>
: Mello-Leit
</taxonomicName>
„o, 1943: 149.
</paragraph>
<paragraph id="8163A27CFF80CE50FCD5F823DFBB3429" blockId="13.[810,1452,1756,2092]" box="[810,1256,2034,2056]" pageId="13" pageNumber="14">
<taxonomicName id="46DCD9FFFF80CE50FCD5F823DF303426" ID-CoL="64PKR" authority=": Mello-Leit" authorityName="Mello-Leit" box="[810,1123,2034,2056]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="B3A87E6EFF80CE50FCD5F823D8BA3426" box="[810,1001,2034,2056]" italics="true" pageId="13" pageNumber="14">Actinopus crassipes</emphasis>
: Mello-Leit
</taxonomicName>
„o, 1943: 149.
</paragraph>
<paragraph id="8163A27CFF80CE50FCD5F7C7DF8A3405" blockId="13.[810,1452,1756,2092]" box="[810,1241,2070,2092]" pageId="13" pageNumber="14">
<taxonomicName id="46DCD9FFFF80CE50FCD5F7C7DF073402" authority=": Mello-Leit" authorityName="Mello-Leit" box="[810,1108,2070,2092]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="B3A87E6EFF80CE50FCD5F7C7D8883405" box="[810,987,2070,2091]" italics="true" pageId="13" pageNumber="14">Actinopus luteipes</emphasis>
: Mello-Leit
</taxonomicName>
„o, 1943: 149.
</paragraph>
</subSubSection>
<caption id="D5A3F2F4FF83CE53FF77FC69DF0A3836" ID-DOI="http://doi.org/10.5281/zenodo.5597255" ID-Zenodo-Dep="5597255" httpUri="https://zenodo.org/record/5597255/files/figure.png" pageId="14" pageNumber="15" startId="14.[136,235,952,976]" targetBox="[170,1419,201,919]" targetPageId="14">
<paragraph id="8163A27CFF83CE53FF77FC69DF0A3836" blockId="14.[136,1452,952,1048]" pageId="14" pageNumber="15">
<emphasis id="B3A87E6EFF83CE53FF77FC69DA563FFE" bold="true" box="[136,261,952,976]" pageId="14" pageNumber="15">FIGURE 9.</emphasis>
<taxonomicName id="46DCD9FFFF83CE53FEF1FC69D9173FFE" authority="Perty, 1833" authorityName="Perty" authorityYear="1833" box="[270,580,952,977]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B3A87E6EFF83CE53FEF1FC69DA943FFE" box="[270,455,952,976]" italics="true" pageId="14" pageNumber="15">Actinopus tarsalis</emphasis>
<bibRefCitation id="E54DDF8DFF83CE53FE30FC68D9173FFE" author="Perty, M." box="[463,580,952,977]" pageId="14" pageNumber="15" pagination="191 - 209" refId="ref139886" refString="Perty, M. (1833) Arachnides Brasilienses. In: de Spix, J. B. &amp; Martius, F. P. (Eds.), Delectus animalium articulatorum quae in itinere per Braziliam ann. 1817 et 1820 colligerunt. Monachii, pp. 191 - 209." type="book chapter" year="1833">Perty, 1833</bibRefCitation>
</taxonomicName>
(drawings from
<bibRefCitation id="E54DDF8DFF83CE53FD0CFC69D8EB3FFE" author="Miglio, L. T. &amp; Lucas, S. M. &amp; Bonaldo, A. B." box="[755,952,952,976]" pageId="14" pageNumber="15" pagination="275 - 379" refId="ref139626" refString="Miglio, L. T., Lucas, S. M. &amp; Bonaldo, A. B. (2012) On the identity of the type species Actinopus tarsalis (Araneae: Actinopodidae). Zoologia, 29 (4), 275 - 379. https: // doi. org / 10.1590 / S 1984 - 46702012000400009" type="journal article" year="2012">
Miglio
<emphasis id="B3A87E6EFF83CE53FCBFFC68D8263FFE" box="[832,885,952,976]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
, 2012
</bibRefCitation>
), male: AC MPEG 11717: A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral.
<emphasis id="B3A87E6EFF83CE53FCACFC0DD8283FDA" bold="true" box="[851,891,988,1012]" pageId="14" pageNumber="15">PA-</emphasis>
Paraembolic apophysis;
<emphasis id="B3A87E6EFF83CE53FB8EFC0DDFCF3FDA" bold="true" box="[1137,1180,988,1012]" pageId="14" pageNumber="15">PAc</emphasis>
-Prolateral Accessory keel;
<emphasis id="B3A87E6EFF83CE53FF77FBD1DBF23836" bold="true" box="[136,161,1024,1048]" pageId="14" pageNumber="15">PI</emphasis>
-Prolateral Inferior keel;
<emphasis id="B3A87E6EFF83CE53FE61FBD1DAEF3836" bold="true" box="[414,444,1024,1048]" pageId="14" pageNumber="15">PS</emphasis>
-Prolateral Superior keel; Arrow: Serrated Area. Scale line: 1 mm.
</paragraph>
</caption>
<caption id="D5A3F2F4FF83CE53FF77F828DF4C343E" ID-DOI="http://doi.org/10.5281/zenodo.5655402" ID-Zenodo-Dep="5655402" box="[136,1055,2041,2065]" httpUri="https://zenodo.org/record/5655402/files/figure.png" pageId="14" pageNumber="15" startId="14.[136,192,2041,2065]" targetBox="[157,1428,1114,2008]" targetPageId="14">
<paragraph id="8163A27CFF83CE53FF77F828DF4C343E" blockId="14.[136,1055,2041,2065]" box="[136,1055,2041,2065]" pageId="14" pageNumber="15">
<emphasis id="B3A87E6EFF83CE53FF77F828DB8A343F" bold="true" box="[136,217,2041,2065]" pageId="14" pageNumber="15">MAP 1.</emphasis>
Distribution map of
<taxonomicName id="46DCD9FFFF83CE53FE51F828D9B1343F" authority="Perty, 1833" authorityName="Perty" authorityYear="1833" box="[430,738,2041,2065]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B3A87E6EFF83CE53FE51F828D935343F" box="[430,614,2041,2065]" italics="true" pageId="14" pageNumber="15">Actinopus tarsalis</emphasis>
<bibRefCitation id="E54DDF8DFF83CE53FD93F828D9B1343F" author="Perty, M." box="[620,738,2041,2065]" pageId="14" pageNumber="15" pagination="191 - 209" refId="ref139886" refString="Perty, M. (1833) Arachnides Brasilienses. In: de Spix, J. B. &amp; Martius, F. P. (Eds.), Delectus animalium articulatorum quae in itinere per Braziliam ann. 1817 et 1820 colligerunt. Monachii, pp. 191 - 209." type="book chapter" year="1833">Perty, 1833</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="46DCD9FFFF83CE53FCE8F828D898343F" box="[791,971,2041,2065]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="castelo" status="sp. nov.">
<emphasis id="B3A87E6EFF83CE53FCE8F828D898343F" box="[791,971,2041,2065]" italics="true" pageId="14" pageNumber="15">Actinopus castelo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A89BC315FF83CE53FC2EF828DF4C343E" box="[977,1055,2041,2064]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="C9C6F1F7FF82CE52FF77FF4FDAB23DC2" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8163A27CFF82CE52FF77FF4FDAB23DC2" blockId="15.[136,778,158,531]" pageId="15" pageNumber="16">
<emphasis id="B3A87E6EFF82CE52FF77FF4FDA513C96" bold="true" box="[136,258,158,184]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
Males of
<taxonomicName id="46DCD9FFFF82CE52FE82FF4EDAA03C96" authorityName="Perty" authorityYear="1833" box="[381,499,158,185]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="B3A87E6EFF82CE52FE82FF4EDAA03C96" box="[381,499,158,185]" italics="true" pageId="15" pageNumber="16">A. tarsalis</emphasis>
</taxonomicName>
differ from those of all species, except
<taxonomicName id="46DCD9FFFF82CE52FECCFF14DACC3CF1" box="[307,415,197,223]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="castelo">
<emphasis id="B3A87E6EFF82CE52FECCFF14DACC3CF1" box="[307,415,197,223]" italics="true" pageId="15" pageNumber="16">A. castelo</emphasis>
</taxonomicName>
(
<figureCitation id="19E7BEF9FF82CE52FE52FF14DAAF3CF1" box="[429,508,197,223]" captionStart="FIGURE 12" captionStartId="17.[136,235,1115,1139]" captionTargetBox="[158,1417,199,1057]" captionTargetId="figure-364@17.[151,1436,189,1092]" captionTargetPageId="17" captionText="FIGURE 12. Actinopus castelo sp. nov., male: AC MPEG 22277: A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral. PA-Paraembolic apophysis; PAc-Prolateral Accessory keel; PI-Prolateral Inferior keel; PS-Prolateral Superior keel; Arrows: Serrated Area. Scale line: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5597263" httpUri="https://zenodo.org/record/5597263/files/figure.png" pageId="15" pageNumber="16">Fig. 12</figureCitation>
AC), by the copulatory bulb without BTA and by the robust embolar base, inserted basally at a right angle (90°); the embolar apex is flattened and expanded, arrow-shaped in dorsal view. They differ from those of
<taxonomicName id="46DCD9FFFF82CE52FE5CFE8ED95C3D56" box="[419,527,350,377]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="castelo">
<emphasis id="B3A87E6EFF82CE52FE5CFE8EDAE73D57" box="[419,436,351,377]" italics="true" pageId="15" pageNumber="16">A</emphasis>
.
<emphasis id="B3A87E6EFF82CE52FE3FFE8FD95C3D56" box="[448,527,350,376]" italics="true" pageId="15" pageNumber="16">castelo</emphasis>
</taxonomicName>
by PAc inconspicuous at prolateral surface, embolus shorter, tegulum wider than in
<taxonomicName id="46DCD9FFFF82CE52FF58FE7DDA453DEB" box="[167,278,427,454]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="castelo">
<emphasis id="B3A87E6EFF82CE52FF58FE7DDA453DEB" box="[167,278,427,454]" italics="true" pageId="15" pageNumber="16">A. castelo</emphasis>
</taxonomicName>
and PA more conspicuous (
<figureCitation id="19E7BEF9FF82CE52FDA9FE7AD9C93DEB" box="[598,666,427,453]" captionStart="FIGURE 9" captionStartId="14.[136,235,952,976]" captionTargetBox="[170,1419,201,919]" captionTargetId="figure-21@14.[151,1436,189,929]" captionTargetPageId="14" captionText="FIGURE 9. Actinopus tarsalis Perty, 1833 (drawings from Miglio et al., 2012), male: AC MPEG 11717: A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral. PA-Paraembolic apophysis; PAc-ProlateralAccessory keel; PI-Prolateral Inferior keel; PS-Prolateral Superior keel; Arrow: Serrated Area. Scale line: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5597255" httpUri="https://zenodo.org/record/5597255/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
AC, see
<bibRefCitation id="E54DDF8DFF82CE52FF77FE03DA0F3DC2" author="Miglio, L. T. &amp; Lucas, S. M. &amp; Bonaldo, A. B." box="[136,348,466,492]" pageId="15" pageNumber="16" pagination="275 - 379" refId="ref139626" refString="Miglio, L. T., Lucas, S. M. &amp; Bonaldo, A. B. (2012) On the identity of the type species Actinopus tarsalis (Araneae: Actinopodidae). Zoologia, 29 (4), 275 - 379. https: // doi. org / 10.1590 / S 1984 - 46702012000400009" type="journal article" year="2012">
Miglio
<emphasis id="B3A87E6EFF82CE52FF24FE03DA473DC2" box="[219,276,466,492]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
, 2012
</bibRefCitation>
, figs 112).
</paragraph>
</subSubSection>
<subSubSection id="C9C6F1F7FF82CE52FF42FE29D9F13E3D" box="[189,674,504,531]" pageId="15" pageNumber="16" type="description">
<paragraph id="8163A27CFF82CE52FF42FE29D9F13E3D" blockId="15.[136,778,158,531]" box="[189,674,504,531]" pageId="15" pageNumber="16">
<emphasis id="B3A87E6EFF82CE52FF42FE29DA1C3E3C" bold="true" box="[189,335,504,530]" pageId="15" pageNumber="16">Description.</emphasis>
See
<bibRefCitation id="E54DDF8DFF82CE52FE7AFE29D9CD3E3D" author="Miglio, L. T. &amp; Lucas, S. M. &amp; Bonaldo, A. B." box="[389,670,504,531]" pageId="15" pageNumber="16" pagination="275 - 379" refId="ref139626" refString="Miglio, L. T., Lucas, S. M. &amp; Bonaldo, A. B. (2012) On the identity of the type species Actinopus tarsalis (Araneae: Actinopodidae). Zoologia, 29 (4), 275 - 379. https: // doi. org / 10.1590 / S 1984 - 46702012000400009" type="journal article" year="2012">
Miglio
<emphasis id="B3A87E6EFF82CE52FE27FE28D9423E3C" box="[472,529,504,530]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(2012: 376)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>