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<mods:title id="F56D45F32B1D9CDD9C1A930366AAE5B7">Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="A45A1248E2D3CBAB5C8A94E446DAD46C">Liu, Tian-tian</mods:namePart>
<mods:affiliation id="802AD844924944CAFC2D4055F3B12D53">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395, Japan &amp; The Kyushu University Museum, Hakozaki</mods:affiliation>
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<mods:namePart id="66F78A1B425056C1E630A04D1A335291">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="24D49E541738BACC941EE52C83D54B17">The Kyushu University Museum, Hakozaki</mods:affiliation>
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<emphasis id="B9166F3FAC07FF8F4CC6FA3213F7FA2C" bold="true" box="[910,1192,1481,1505]" italics="true" pageId="13" pageNumber="176">Bryothinusa sakishimana</emphasis>
<bibRefCitation id="EFF3CEDCAC07FF8F4BE7FA311215FA2C" author="SAWADA K." box="[1199,1354,1482,1505]" pageId="13" pageNumber="176" pagination="141 - 152" refId="ref26218" refString="SAWADA K. 1991: On new genera and species of intertidal Aleocharinae (Coleoptera: Staphylinidae) and Goniacerinae (Pselaphidae) from Singapore and Japan. Raffles Bulletin of Zoology 39 (1): 141 - 152." type="journal article" year="1991">Sawada, 1991</bibRefCitation>
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(
<figureCitation id="1359AFA8AC07FF8F4CBEFA131361FA36" box="[1014,1086,1512,1531]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="13" pageNumber="176">Figs 1G</figureCitation>
; 
<figureCitation id="1359AFA8AC07FF8F4B0FFA13133CFA36" box="[1095,1123,1512,1531]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="13" pageNumber="176">2G</figureCitation>
; 
<figureCitation id="1359AFA8AC07FF8F4B24FA131326FA36" box="[1132,1145,1512,1531]" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="13" pageNumber="176">9</figureCitation>
; 
<figureCitation id="1359AFA8AC07FF8F4BCAFA13139DFA36" box="[1154,1218,1512,1531]" captionStart="Fig" captionStartId="34.[151,182,1955,1974]" captionTargetBox="[167,1408,193,1935]" captionTargetId="figure-13@34.[161,1426,189,1938]" captionTargetPageId="34" captionText="Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn &amp; Jeon, 2004 and B. japonica Liu, Ono &amp; Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn &amp; Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka." figureDoi="http://doi.org/10.5281/zenodo.5037374" httpUri="https://zenodo.org/record/5037374/files/figure.png" pageId="13" pageNumber="176">21B, D</figureCitation>
; 
<figureCitation id="1359AFA8AC07FF8F4B83FA1313BEFA36" box="[1227,1249,1512,1531]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn &amp; Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono &amp; Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn &amp; Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="13" pageNumber="176">22</figureCitation>
)
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<taxonomicName id="4C62C8AEAC07FF8F4C74F9DF13E4F9F5" ID-CoL="NJWY" authority="Sawada, 1991: 144" authorityName="Sawada" authorityPageNumber="144" authorityYear="1991" box="[828,1211,1572,1592]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="sakishimana">
<emphasis id="B9166F3FAC07FF8F4C74F9DF1346F9FA" box="[828,1049,1572,1591]" italics="true" pageId="13" pageNumber="176">Bryothinusa sakishimana</emphasis>
<bibRefCitation id="EFF3CEDCAC07FF8F4B50F9DE13E4F9F5" author="SAWADA K." box="[1048,1211,1573,1592]" pageId="13" pageNumber="176" pagination="141 - 152" refId="ref26218" refString="SAWADA K. 1991: On new genera and species of intertidal Aleocharinae (Coleoptera: Staphylinidae) and Goniacerinae (Pselaphidae) from Singapore and Japan. Raffles Bulletin of Zoology 39 (1): 141 - 152." type="journal article" year="1991">Sawada, 1991: 144</bibRefCitation>
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(original description; 
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locality: 
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, Nansei-shotô, Iriomote-jima), 145 (compared with 
<taxonomicName id="4C62C8AEAC07FF8F4AC2F9BB14B8F9A0" ID-CoL="NJWQ" authority="Sawada" authorityName="Sawada" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="nakanei">
<emphasis id="B9166F3FAC07FF8F4AC2F9BB14C1F9A0" italics="true" pageId="13" pageNumber="176">B. nakanei</emphasis>
Sawada
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);Aඌ*üൾ (2004): 594 (in annotated catalogue of world species); Sආൾඍൺඇൺ (2004): 464 (in catalogue of Palearctic species); F*©ൺඇκ &amp;A*üඇ (2011): 27 (in checklist); Sർ*ü*“අκൾ &amp; Sආൾඍൺඇൺ (2015): 676 (in catalogue of Palearctic species); Lංඎ et al. (2020): 589 (in key of Japanese species), 592 (in checklist).
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material.
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Not examined.
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Material examined. 
<collectingCountry id="F375F3BDAC07FF8F4CBAF8F21365F8D1" box="[1010,1082,1801,1820]" name="Japan" pageId="13" pageNumber="176">JAPAN</collectingCountry>
: Nൺඇඌൾං-ඌΗඈඍත:
</emphasis>
3 JJ 
<specimenCount id="9D6478A4AC07FF8F4A4CF8F2126DF8D1" box="[1284,1330,1795,1820]" pageId="13" pageNumber="176" type="female">2 ♀♀</specimenCount>
, 11 unsexed spec., Ôgomori, Takara-jima, Tokara-rettô, Kagoshima-ken, 
<date id="FFDC95EDAC07FF8F4A08F8DF12C6F8FA" box="[1344,1433,1828,1847]" pageId="13" pageNumber="176" value="2019-07-03">3.vii.2019</date>
, 
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. Liu, N. Tsuji &amp; H. Ono; 1 unsexed spec., Mouth of Honera-gawa, Takana, Taketomi-chô, Iriomote-jima, Okinawa-ken, 
<date id="FFDC95EDAC07FF8F4BA3F8A21263F8A1" box="[1259,1340,1881,1900]" pageId="13" pageNumber="176" value="2019-04-02">2.iv.2019</date>
, 
<collectionCode id="ED732BE8AC07FF8F4A0CF8A2120DF8A1" box="[1348,1362,1881,1900]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="176">T</collectionCode>
. Nozaki. Deposited at 
<collectionCode id="ED732BE8AC07FF8F4CE4F88F1481F84A" box="[940,990,1908,1927]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="13" pageNumber="176" type="Herbarium">KUM</collectionCode>
but some will be distributed to 
<collectionCode id="ED732BE8AC07FF8F4BA5F88F1276F84A" box="[1261,1321,1908,1927]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="13" pageNumber="176">NMPC</collectionCode>
and 
<collectionCode id="ED732BE8AC07FF8F4A1BF88F12C9F84A" box="[1363,1430,1908,1927]" pageId="13" pageNumber="176">BMNH</collectionCode>
.
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<emphasis id="B9166F3FAC07FF8F4C74F85B14BEF87A" bold="true" box="[828,993,1952,1975]" pageId="13" pageNumber="176">Redescription.</emphasis>
Body (
<figureCitation id="1359AFA8AC07FF8F4B78F85B13DCF87A" box="[1072,1155,1952,1975]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="13" pageNumber="176">Fig. 1G</figureCitation>
) small, about 
<quantity id="4C9A1EC8AC07FF8F4A54F85B12C3F87A" box="[1308,1436,1952,1975]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="13" pageNumber="176" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6–1.9 mm</quantity>
in length, covered with minute setae. Abdomen almost black and darker than head and pronotum; elytra lighter than pronotum; antenna and legs yellowish brown.
</paragraph>
<caption id="DF1DE3A5AC04FF8C4FDFF82B1579F7D4" ID-DOI="http://doi.org/10.5281/zenodo.5037348" ID-Zenodo-Dep="5037348" httpUri="https://zenodo.org/record/5037348/files/figure.png" pageId="14" pageNumber="177" startId="14.[151,182,2000,2019]" targetBox="[159,1400,189,1927]" targetPageId="14">
<paragraph id="8BDDB32DAC04FF8C4FDFF82B1579F7D4" blockId="14.[151,1437,2000,2073]" pageId="14" pageNumber="177">
Fig. 8. 
<taxonomicName id="4C62C8AEAC04FF8C4F9CF82B15D1F82E" authority="Liu, Ono &amp; Maruyama, 2020" authorityName="Liu, Ono &amp; Maruyama" authorityYear="2020" box="[212,654,2000,2019]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="177" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B9166F3FAC04FF8C4F9CF82B16D3F82F" box="[212,396,2000,2019]" italics="true" pageId="14" pageNumber="177">Bryothinusa japonica</emphasis>
<bibRefCitation id="EFF3CEDCAC04FF8C4ED9F82B15D1F82E" author="LIU T. T. &amp; ONO H. &amp; MARUYAMA M." box="[401,654,2000,2019]" pageId="14" pageNumber="177" pagination="585 - 593" refId="ref25551" refString="LIU T. T., ONO H. &amp; MARUYAMA M. 2020: A new species and new record of the intertidal rove beetle genus Bryothinusa Casey (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 4743 (4): 585 - 593." type="journal article" year="2020">Liu, Ono &amp; Maruyama, 2020</bibRefCitation>
</taxonomicName>
. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm. Modified from Lංඎ et al. (2020).
</paragraph>
</caption>
<caption id="DF1DE3A5AC05FF8D4FDEF81612C1F7D7" ID-DOI="http://doi.org/10.5281/zenodo.5037350" ID-Zenodo-Dep="5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="15" pageNumber="178" startId="15.[150,181,2029,2048]" targetBox="[158,1411,189,1970]" targetPageId="15">
<paragraph id="8BDDB32DAC05FF8D4FDEF81612C1F7D7" blockId="15.[150,1438,2028,2074]" pageId="15" pageNumber="178">
Fig. 9. 
<taxonomicName id="4C62C8AEAC05FF8D4F9CF8171571F7CD" authority="Sawada, 1991" authorityName="Sawada" authorityYear="1991" box="[212,558,2028,2048]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="178" phylum="Arthropoda" rank="species" species="sakishimana">
<emphasis id="B9166F3FAC05FF8D4F9CF81716F1F832" box="[212,430,2028,2047]" italics="true" pageId="15" pageNumber="178">Bryothinusa sakishimana</emphasis>
<bibRefCitation id="EFF3CEDCAC05FF8D4EFCF8161571F7CD" author="SAWADA K." box="[436,558,2029,2048]" pageId="15" pageNumber="178" pagination="141 - 152" refId="ref26218" refString="SAWADA K. 1991: On new genera and species of intertidal Aleocharinae (Coleoptera: Staphylinidae) and Goniacerinae (Pselaphidae) from Singapore and Japan. Raffles Bulletin of Zoology 39 (1): 141 - 152." type="journal article" year="1991">Sawada, 1991</bibRefCitation>
</taxonomicName>
. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BDDB32DAC1AFF924FFDFF45161DFE78" blockId="16.[151,761,190,1653]" pageId="16" pageNumber="179">
Head evenly rounded, more or less produced anteriorly, wider than long, shorter than pronotum and elytra; eyes (
<figureCitation id="1359AFA8AC1AFF924FD7FF0517AEFED8" box="[159,241,254,277]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn &amp; Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono &amp; Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn &amp; Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="16" pageNumber="179">Fig. 2G</figureCitation>
) large, setae present between about 21–23 facets, slightly longer than tempora length; antenna yellowish brown, longer than total length of head and pronotum; segment I 1.4 times longer than II, almost 3.8 times longer than III, and longer than XI; antennal segments V–X almost as long as wide.
</paragraph>
<paragraph id="8BDDB32DAC1AFF924FFDFE4515A7FCD8" blockId="16.[151,761,190,1653]" pageId="16" pageNumber="179">
Mouthparts (
<figureCitation id="1359AFA8AC1AFF924E0EFE45169EFE18" box="[326,449,446,469]" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="16" pageNumber="179">Figs 9A–E</figureCitation>
) with distribution of labral setae as in 
<figureCitation id="1359AFA8AC1AFF924E5BFE251635FE38" box="[275,362,478,501]" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="16" pageNumber="179">Fig. 9A</figureCitation>
, with large and thick spiniform seta antero-laterally; mandibles (
<figureCitation id="1359AFA8AC1AFF924E98FE051579FDD8" box="[464,550,510,533]" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="16" pageNumber="179">Fig. 9B</figureCitation>
) elongate triangle, asymmetrical, right mandible with median tooth; median tooth shorter than 1/4 of mandibles; left without tooth, blunt at apex; maxilla with apical margin of lacinia (
<figureCitation id="1359AFA8AC1AFF924D85FDA517E8FD58" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="16" pageNumber="179">Fig. 9C</figureCitation>
) with second spine very small, not obvious; 3 setae on inner margin of lacinia; labial palpus (
<figureCitation id="1359AFA8AC1AFF924D65FD651520FD78" box="[557,639,670,693]" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="16" pageNumber="179">Fig. 9D</figureCitation>
) with most setae located basally; mentum (
<figureCitation id="1359AFA8AC1AFF924E94FD451574FD18" box="[476,555,702,725]" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="16" pageNumber="179">Fig. 9E</figureCitation>
) with antero-lateral angles prolonged into spinose processes; anterior margin deeply truncate; small “v seta” far away from apical setae.
</paragraph>
<paragraph id="8BDDB32DAC1AFF924FFDFCE515F9FCF8" blockId="16.[151,761,190,1653]" box="[181,678,798,821]" pageId="16" pageNumber="179">Pronotum shorter than elytra, wider than head.</paragraph>
<paragraph id="8BDDB32DAC1AFF924FFDFCC517A0FC58" blockId="16.[151,761,190,1653]" pageId="16" pageNumber="179">Elytra subquadrate, narrower than pronotum; almost unicolour and hind wings present but small, apparently flightless.</paragraph>
<paragraph id="8BDDB32DAC1AFF924FFDFC651500FC38" blockId="16.[151,761,190,1653]" pageId="16" pageNumber="179">
Male abdominal sternite VIII (
<figureCitation id="1359AFA8AC1AFF924D49FC65150EFC78" box="[513,593,926,949]" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="16" pageNumber="179">Fig. 9F</figureCitation>
) with posterior margin slightly serrate. Median lobe of aedeagus as in 
<figureCitation id="1359AFA8AC1AFF924FDFFC251678FC38" box="[151,295,990,1013]" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="16" pageNumber="179">Figures 9H, I</figureCitation>
; paramere (
<figureCitation id="1359AFA8AC1AFF924EECFC2516B0FC38" box="[420,495,990,1013]" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="16" pageNumber="179">Fig. 9J</figureCitation>
) elongate.
</paragraph>
<paragraph id="8BDDB32DAC1AFF924FFDFC0515A6FB98" blockId="16.[151,761,190,1653]" pageId="16" pageNumber="179">
Female abdominal sternite VIII (
<figureCitation id="1359AFA8AC1AFF924D7DFC0515CFFBD8" box="[565,656,1022,1045]" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="16" pageNumber="179">Fig. 9G</figureCitation>
) slightly serrate alike male on posterior margin. Spermatheca (
<figureCitation id="1359AFA8AC1AFF924D85FBE517E6FB98" captionStart="Fig" captionStartId="15.[150,181,2029,2048]" captionTargetBox="[158,1411,189,1970]" captionTargetId="figure-14@15.[156,1430,189,1999]" captionTargetPageId="15" captionText="Fig. 9. Bryothinusa sakishimana Sawada, 1991. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037350" httpUri="https://zenodo.org/record/5037350/files/figure.png" pageId="16" pageNumber="179">Fig. 9K</figureCitation>
) about 1.5 times coiled near base, apical part elongate.
</paragraph>
<paragraph id="8BDDB32DAC1AFF924FDFFBA5159FFB18" blockId="16.[151,761,190,1653]" pageId="16" pageNumber="179">
<emphasis id="B9166F3FAC1AFF924FDFFBA5161AFBB8" bold="true" box="[151,325,1118,1141]" pageId="16" pageNumber="179">Measurements.</emphasis>
BL 1.60–1.90, FBL 0.70–1.00, HL 0.28– 0.33, HW 0.35–0.40, PL 0.33–0.38, PW 0.40–0.45, EL 0.35–0.40, EW 0.38–0.40, HTL 0.33–0.35, AL 0.70–0.83, AS-I 0.12–0.15, AS-II 0.09–0.10, AS-III 0.03–0.04.
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC1AFF924FDFFB25168CFA18" pageId="16" pageNumber="179" type="diagnosis">
<paragraph id="8BDDB32DAC1AFF924FDFFB25168CFA18" blockId="16.[151,761,190,1653]" pageId="16" pageNumber="179">
<emphasis id="B9166F3FAC1AFF924FDFFB2516CBFB38" bold="true" box="[151,404,1246,1269]" pageId="16" pageNumber="179">Differential diagnosis.</emphasis>
In Nansei-shotô, this species is especially similar to 
<taxonomicName id="4C62C8AEAC1AFF924E30FB0516B4FAD8" box="[376,491,1277,1301]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="179" phylum="Arthropoda" rank="species" species="nakanei">
<emphasis id="B9166F3FAC1AFF924E30FB0516B4FAD8" box="[376,491,1277,1301]" italics="true" pageId="16" pageNumber="179">B. nakanei</emphasis>
</taxonomicName>
, but easily distinguished from it by the paler elytral colour, the number of facets, the spermathecal shape and the posterior margin of the tergite X rounded medially. This species is also similar to 
<taxonomicName id="4C62C8AEAC1AFF924DA9FAA5165EFA58" authority="Liu &amp; Ono &amp; Maruyama, 2021" authorityName="Liu &amp; Ono &amp; Maruyama" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="179" phylum="Arthropoda" rank="species" species="constricta" status="sp. nov.">
<emphasis id="B9166F3FAC1AFF924DA9FAA5165EFA58" italics="true" pageId="16" pageNumber="179">B. constricta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225D244AC1AFF924E4EFA851608FA58" box="[262,343,1406,1429]" pageId="16" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
in general appearance but differentiated from it by the elytral shape not being almost square and the shape of the spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC1AFF924FDFFA251537F9D8" pageId="16" pageNumber="179" type="distribution">
<paragraph id="8BDDB32DAC1AFF924FDFFA251537F9D8" blockId="16.[151,761,190,1653]" pageId="16" pageNumber="179">
<emphasis id="B9166F3FAC1AFF924FDFFA25166EFA38" bold="true" box="[151,305,1502,1525]" pageId="16" pageNumber="179">Distribution.</emphasis>
<collectingCountry id="F375F3BDAC1AFF924E74FA251624FA38" box="[316,379,1502,1525]" name="Japan" pageId="16" pageNumber="179">Japan</collectingCountry>
(
<emphasis id="B9166F3FAC1AFF924ED8FA25156BFA38" bold="true" box="[400,564,1502,1525]" pageId="16" pageNumber="179">Nansei-shotô:</emphasis>
Kagoshima-ken: Takara-jima; Okinawa-ken: Iriomote-jima).
</paragraph>
</subSubSection>
<subSubSection id="C378E0A6AC1AFF924FDFF9E51502F9B8" pageId="16" pageNumber="179" type="discussion">
<paragraph id="8BDDB32DAC1AFF924FDFF9E51502F9B8" blockId="16.[151,761,190,1653]" pageId="16" pageNumber="179">
<emphasis id="B9166F3FAC1AFF924FDFF9E51659F9F8" bold="true" box="[151,262,1566,1589]" pageId="16" pageNumber="179">Remarks.</emphasis>
This species is distributed in Nansei-shotô of 
<collectingCountry id="F375F3BDAC1AFF924FDFF9C51788F998" box="[151,215,1598,1621]" name="Japan" pageId="16" pageNumber="179">Japan</collectingCountry>
. A distribution map of examined specimens of this species in 
<collectingCountry id="F375F3BDAC1AFF924E4EF9A5161CF9B8" box="[262,323,1630,1653]" name="Japan" pageId="16" pageNumber="179">Japan</collectingCountry>
is illustrated in 
<figureCitation id="1359AFA8AC1AFF924EA7F9A51506F9B8" box="[495,601,1630,1653]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn &amp; Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono &amp; Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn &amp; Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="16" pageNumber="179">Figure 22</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>