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<document id="668F7DA423D5DDD4A1DEEA69E74F6E79" ID-DOI="10.11646/zootaxa.3946.4.3" ID-GBIF-Dataset="46db289d-0964-4a71-8b5c-ab81e23e961f" ID-ISSN="1175-5326" ID-Zenodo-Dep="238608" ID-ZooBank="7A5F142A-EEDE-453A-9CB5-241917A83921" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461038442617" checkinUser="plazi" docAuthor="Rifkind, Jacques" docDate="2015" docId="0151C53EFFA2FFA49DDCFB3B45EE8DA6" docLanguage="en" docName="zt03946p552.pdf" docOrigin="Zootaxa 3946 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cymatodera monticola Rifkind, n. sp." docType="treatment" docVersion="8" lastPageNumber="537" masterDocId="FD68BD46FFB3FFB69D4BFF9B462E8F6A" masterDocTitle="New species of Cymatodera Gray (Coleoptera: Cleridae: Tillinae) from México and Central America, with notes on others" masterLastPageNumber="552" masterPageNumber="519" pageNumber="536" updateTime="1698574954642" updateUser="plazi">
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<mods:title id="DF3DFEE24CECF6DDE88806A9D5844899">New species of Cymatodera Gray (Coleoptera: Cleridae: Tillinae) from México and Central America, with notes on others</mods:title>
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<mods:namePart id="1EC804C4AACEC52B9E231D589BF169C9">Rifkind, Jacques</mods:namePart>
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<mods:date id="918D59C4ACBE8B7EB29D3E0C40DA0C32">2015</mods:date>
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<treatment id="0151C53EFFA2FFA49DDCFB3B45EE8DA6" ID-DOI="http://doi.org/10.5281/zenodo.6110757" ID-GBIF-Taxon="119609544" ID-Zenodo-Dep="6110757" LSID="urn:lsid:plazi:treatment:0151C53EFFA2FFA49DDCFB3B45EE8DA6" httpUri="http://treatment.plazi.org/id/0151C53EFFA2FFA49DDCFB3B45EE8DA6" lastPageId="18" lastPageNumber="537" pageId="17" pageNumber="536">
<subSubSection id="C1E227A3FFA2FFA79DDCFB3B47008BB6" pageId="17" pageNumber="536" type="nomenclature">
<paragraph id="89477428FFA2FFA79DDCFB3B444F8BD0" blockId="17.[151,609,1184,1244]" box="[151,609,1184,1210]" pageId="17" pageNumber="536">
<heading id="D20FC344FFA2FFA79DDCFB3B444F8BD0" bold="true" box="[151,609,1184,1210]" fontSize="11" level="1" pageId="17" pageNumber="536" reason="1">
<emphasis id="BB8CA83AFFA2FFA79DDCFB3B444F8BD0" bold="true" box="[151,609,1184,1210]" pageId="17" pageNumber="536">
<taxonomicName id="4EF80FABFFA2FFA79DDCFB3B443C8BD0" authority="Rifkind" authorityName="Rifkind" box="[151,530,1184,1210]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="536" phylum="Arthropoda" rank="species" species="monticola" status="sp. nov.">
<emphasis id="BB8CA83AFFA2FFA79DDCFB3B47848BD0" bold="true" box="[151,426,1184,1210]" italics="true" pageId="17" pageNumber="536">Cymatodera monticola</emphasis>
Rifkind
</taxonomicName>
,
<taxonomicNameLabel id="A0BF1541FFA2FFA79F54FB3B444F8BD0" box="[543,609,1184,1210]" pageId="17" pageNumber="536" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="89477428FFA2FFA79DDCFB5847008BB6" blockId="17.[151,609,1184,1244]" box="[151,302,1219,1244]" pageId="17" pageNumber="536">
(
<figureCitation id="11C368ADFFA2FFA79DD4FB5847088BB6" box="[159,294,1219,1244]" captionStart="FIGURES 34 36. 34" captionStartId="18.[151,264,1750,1773]" captionTargetBox="[162,1415,784,1717]" captionTargetId="figure@18.[150,1436,772,1730]" captionTargetPageId="18" captionText="FIGURES 34 36. 34. Habitus of Cymatodera monticola Rifkind, n. sp. (holotype male). 35. Pygidium (ventral view) of male Cymatodera monticola. 36. Pygidium (dorsal view) of male Cymatodera monticola." httpUri="https://zenodo.org/record/238617/files/figure.png" pageId="17" pageNumber="536">Figs. 3436</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFA2FFA79DDCFA90470A8AE5" pageId="17" pageNumber="536" type="materials_examined">
<paragraph id="89477428FFA2FFA79DDCFA90470A8AE5" blockId="17.[151,1437,1291,2036]" pageId="17" pageNumber="536">
<emphasis id="BB8CA83AFFA2FFA79DDCFA9047778A4E" bold="true" box="[151,345,1291,1316]" pageId="17" pageNumber="536">
<typeStatus id="5643CA8AFFA2FFA79DDCFA9046FC8A4E" box="[151,210,1291,1316]" pageId="17" pageNumber="536">Type</typeStatus>
specimens.
</emphasis>
<typeStatus id="5643CA8AFFA2FFA79C2BFA9047E68A4E" box="[352,456,1291,1316]" pageId="17" pageNumber="536" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F1EF34B8FFA2FFA79F5FFA9044438A4E" box="[532,621,1291,1316]" name="Mexico" pageId="17" pageNumber="536">México</collectingCountry>
, Oaxaca,
<quantity id="4E00D9CDFFA2FFA79F92FA9045228A4E" box="[729,780,1291,1316]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="17" pageNumber="536" unit="mi" value="3.0">3 mi</quantity>
N. Suchixtepec, 9500, km 144, Rt. 175, S. Oaxaca,
<date id="FD4652E8FFA2FFA79815FA9046888A22" pageId="17" pageNumber="536">VI-4- 6</date>
-[19]71, H. Howden.
<typeStatus id="5643CA8AFFA2FFA79CD2FAB4442F8A22" box="[409,513,1327,1352]" pageId="17" pageNumber="536" type="holotype">Holotype</typeStatus>
deposited in
<collectionCode id="EFE9ECEDFFA2FFA79FD1FAB444DE8A2D" box="[666,752,1327,1351]" pageId="17" pageNumber="536">CMNC</collectionCode>
.
<typeStatus id="5643CA8AFFA2FFA79FB7FAAB45438A22" box="[764,877,1328,1352]" pageId="17" pageNumber="536" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F1EF34B8FFA2FFA79E32FAB445C48A2D" box="[889,1002,1327,1351]" name="Mexico" pageId="17" pageNumber="536">MEXICO</collectingCountry>
, OAXACA: 1, same data as
<typeStatus id="5643CA8AFFA2FFA79878FAB443B68A22" box="[1331,1432,1327,1352]" pageId="17" pageNumber="536" type="holotype">holotype</typeStatus>
; 1, Hwy.
<quantity id="4E00D9CDFFA2FFA79DB3FAC847478A06" box="[248,361,1363,1388]" metricMagnitude="5" metricUnit="m" metricValue="2.821180032" pageId="17" pageNumber="536" unit="mi" value="175.3">175, 3 mi</quantity>
. N. Suchixtepec, 9500,
<date id="FD4652E8FFA2FFA79FC9FAC844938A01" box="[642,701,1363,1387]" pageId="17" pageNumber="536">VI-4</date>
-[19]71, D. E. Bright, collector.
<typeStatus id="5643CA8AFFA2FFA7996DFACF42BD8A06" box="[1062,1171,1364,1388]" pageId="17" pageNumber="536" type="paratype">Paratypes</typeStatus>
are deposited in
<collectionCode id="EFE9ECEDFFA2FFA79813FAC843B28A01" box="[1368,1436,1363,1387]" pageId="17" pageNumber="536">JNRC</collectionCode>
and
<collectionCode id="EFE9ECEDFFA2FFA79D8DFAE3470E8AE5" box="[198,288,1400,1423]" httpUri="http://grbio.org/cool/brtj-zfcn" name="W.F. Barr Entomological Collection" pageId="17" pageNumber="536">WFBM</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFA2FFA79D8CFA00477C8902" pageId="17" pageNumber="536" type="diagnosis">
<paragraph id="89477428FFA2FFA79D8CFA00477C8902" blockId="17.[151,1437,1291,2036]" pageId="17" pageNumber="536">
<emphasis id="BB8CA83AFFA2FFA79D8CFA00476F8ADE" bold="true" box="[199,321,1435,1460]" pageId="17" pageNumber="536">Diagnosis.</emphasis>
Separable from other Mexican brachypterous
<taxonomicName id="4EF80FABFFA2FFA79EC8FA0042208AD9" box="[899,1038,1435,1459]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="BB8CA83AFFA2FFA79EC8FA0042208AD9" box="[899,1038,1435,1459]" italics="true" pageId="17" pageNumber="536">Cymatodera</emphasis>
</taxonomicName>
species by virtue of a singular combination of relatively large size (&gt;
<quantity id="4E00D9CDFFA2FFA79F07FA24448F8ABD" box="[588,673,1471,1495]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="536" unit="mm" value="10.0">10 mm</quantity>
), elongate, subflattened, partially translucent elytra with a strongly bisinuate posterior margin (
<figureCitation id="11C368ADFFA2FFA79C99FA78440B8A91" box="[466,549,1507,1532]" captionStart="FIGURES 34 36. 34" captionStartId="18.[151,264,1750,1773]" captionTargetBox="[162,1415,784,1717]" captionTargetId="figure@18.[150,1436,772,1730]" captionTargetPageId="18" captionText="FIGURES 34 36. 34. Habitus of Cymatodera monticola Rifkind, n. sp. (holotype male). 35. Pygidium (ventral view) of male Cymatodera monticola. 36. Pygidium (dorsal view) of male Cymatodera monticola." httpUri="https://zenodo.org/record/238617/files/figure.png" pageId="17" pageNumber="536">Fig. 34</figureCitation>
), and unique pygidial characteristics.
<taxonomicName id="4EF80FABFFA2FFA79EAFFA7842C98A91" box="[996,1255,1507,1531]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="536" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="BB8CA83AFFA2FFA79EAFFA7842C98A91" box="[996,1255,1507,1531]" italics="true" pageId="17" pageNumber="536">Cymatodera monticola</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4EF80FABFFA2FFA798C9FA78471E894A" authority="Rifkind" authorityName="Rifkind" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="536" phylum="Arthropoda" rank="species" species="barri">
<emphasis id="BB8CA83AFFA2FFA798C9FA7846FF8975" italics="true" pageId="17" pageNumber="536">C. barri</emphasis>
Rifkind
</taxonomicName>
, also a Oaxacan endemic, but in that species the elytral apices are rounded, with the male abdominal tergite 5 deeply incised at the posterior margin, and tergite 6 with the posterior margin usually transverse rather than emarginate.
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFA2FFA49D8CF9E845EA8D0A" lastPageId="18" lastPageNumber="537" pageId="17" pageNumber="536" type="description">
<paragraph id="89477428FFA2FFA49D8CF9E842C48EBA" blockId="17.[151,1437,1291,2036]" lastBlockId="18.[151,1437,151,717]" lastPageId="18" lastPageNumber="537" pageId="17" pageNumber="536">
<emphasis id="BB8CA83AFFA2FFA79D8CF9E8477789E6" bold="true" box="[199,345,1651,1676]" pageId="17" pageNumber="536">Description.</emphasis>
(
<typeStatus id="5643CA8AFFA2FFA79C2CF9E847F889E6" box="[359,470,1651,1676]" pageId="17" pageNumber="536" type="holotype">Holotype</typeStatus>
). Length: 12.0 mm. Form: elongate, dorsoventrally compressed, brachypterous. Color: reddish brown; head and pronotum a little darker; elytra testaceous and partially translucent, with indistinct brown markings (
<figureCitation id="11C368ADFFA2FFA79C5BF920474A89B9" box="[272,356,1723,1748]" captionStart="FIGURES 34 36. 34" captionStartId="18.[151,264,1750,1773]" captionTargetBox="[162,1415,784,1717]" captionTargetId="figure@18.[150,1436,772,1730]" captionTargetPageId="18" captionText="FIGURES 34 36. 34. Habitus of Cymatodera monticola Rifkind, n. sp. (holotype male). 35. Pygidium (ventral view) of male Cymatodera monticola. 36. Pygidium (dorsal view) of male Cymatodera monticola." httpUri="https://zenodo.org/record/238617/files/figure.png" pageId="17" pageNumber="536">Fig. 34</figureCitation>
) as follows: on each elytron a small, oblique macula posterior to base and a small, irregularly sinuate, transverse fascia posterior to middle. A random scattering of elytral punctations are similarly infuscate. Head: measured across eyes, wider than pronotum; surface rather finely punctate and roughened; vestiture inconspicuous, pale, and mostly reclinate; antennae of moderate length. Pronotum: widest at anterior margin; longer than broad (ratio of length to width 3:2); surface shining, finely, transversely rugulose, coarsely but shallowly punctate anteriorly; vested as on head. Elytra: typical brachypterous form; elongate (ratio of length to width 7:3); widest at posterior 4/7; disk subflattened above; anterior margin narrow, arcuately emarginate; humeri obsolete; posterior margin bisinuate, apices subacuminate; surface shining, with rather deep, moderately to significantly coarse punctures, subserially arranged but not regularly assorted by size; vestiture inconspicuous, rather sparsely arrayed, composed of fine, pale, suberect and erect setae of moderate length. Metaventrite: short, sulcate posteriorly, subconical on either side; surface shining, finely, sparsely, punctulate. Abdomen: ventrite 5 (
<figureCitation id="11C368ADFFA1FFA49DD4FF4446DF8F92" box="[159,241,223,248]" captionStart="FIGURES 34 36. 34" captionStartId="18.[151,264,1750,1773]" captionTargetBox="[162,1415,784,1717]" captionTargetId="figure@18.[150,1436,772,1730]" captionTargetPageId="18" captionText="FIGURES 34 36. 34. Habitus of Cymatodera monticola Rifkind, n. sp. (holotype male). 35. Pygidium (ventral view) of male Cymatodera monticola. 36. Pygidium (dorsal view) of male Cymatodera monticola." httpUri="https://zenodo.org/record/238617/files/figure.png" pageId="18" pageNumber="537">Fig. 35</figureCitation>
) with sides oblique; posterior margin deeply, arcuately emarginate; ventrite 6 (
<figureCitation id="11C368ADFFA1FFA49926FF4442918F92" box="[1133,1215,223,248]" captionStart="FIGURES 34 36. 34" captionStartId="18.[151,264,1750,1773]" captionTargetBox="[162,1415,784,1717]" captionTargetId="figure@18.[150,1436,772,1730]" captionTargetPageId="18" captionText="FIGURES 34 36. 34. Habitus of Cymatodera monticola Rifkind, n. sp. (holotype male). 35. Pygidium (ventral view) of male Cymatodera monticola. 36. Pygidium (dorsal view) of male Cymatodera monticola." httpUri="https://zenodo.org/record/238617/files/figure.png" pageId="18" pageNumber="537">Fig. 35</figureCitation>
) with sides slightly inflected at middle, bowed outwardly posteriorly, each side with a sinuate, posteriorly divergent carina forming a flange distally that narrows to become part of a slightly downcurved, blunt tipped, lateral projection; surface rugose, without a longitudinal carina at middle; posterior margin rather broadly, arcuately emarginate at middle, extended laterally to form interior border of the lateral projections; tergite 5 (
<figureCitation id="11C368ADFFA1FFA49949FEF4427B8EE2" box="[1026,1109,367,392]" captionStart="FIGURES 34 36. 34" captionStartId="18.[151,264,1750,1773]" captionTargetBox="[162,1415,784,1717]" captionTargetId="figure@18.[150,1436,772,1730]" captionTargetPageId="18" captionText="FIGURES 34 36. 34. Habitus of Cymatodera monticola Rifkind, n. sp. (holotype male). 35. Pygidium (ventral view) of male Cymatodera monticola. 36. Pygidium (dorsal view) of male Cymatodera monticola." httpUri="https://zenodo.org/record/238617/files/figure.png" pageId="18" pageNumber="537">Fig. 36</figureCitation>
) with hind margin bisinuate, slightly, arcuately inflected at middle; tergite 6 (
<figureCitation id="11C368ADFFA1FFA49F8EFE0F45378EC7" box="[709,793,404,429]" captionStart="FIGURES 34 36. 34" captionStartId="18.[151,264,1750,1773]" captionTargetBox="[162,1415,784,1717]" captionTargetId="figure@18.[150,1436,772,1730]" captionTargetPageId="18" captionText="FIGURES 34 36. 34. Habitus of Cymatodera monticola Rifkind, n. sp. (holotype male). 35. Pygidium (ventral view) of male Cymatodera monticola. 36. Pygidium (dorsal view) of male Cymatodera monticola." httpUri="https://zenodo.org/record/238617/files/figure.png" pageId="18" pageNumber="537">Fig. 36</figureCitation>
) with sides broadly arcuate, hind angles subacute, hind margin with a broadly Vshaped emargination. Aedeagus: apex of phallus rather sharply upturned.
</paragraph>
<paragraph id="89477428FFA1FFA49D8CFE4745EA8D0A" blockId="18.[151,1437,151,717]" pageId="18" pageNumber="537">
<emphasis id="BB8CA83AFFA1FFA49D8CFE4747158E9F" bold="true" box="[199,315,476,501]" pageId="18" pageNumber="537">Variation.</emphasis>
The female has the abdominal sternites partially infuscate. Ventrite 5 has the posterior margin broadly, shallowly, arcuately emarginate; ventrite 6 has its sides slightly oblique, its hind margin shallowly, triangularly inflected, with a distinct depression at middle; tergite 6 is also shallowly, triangularly inflected at posterior margin, which appears to be folded downward along its length.
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFA1FFA49D8CFDF745C68DEF" box="[199,1000,620,645]" pageId="18" pageNumber="537" type="etymology">
<paragraph id="89477428FFA1FFA49D8CFDF745C68DEF" blockId="18.[151,1437,151,717]" box="[199,1000,620,645]" pageId="18" pageNumber="537">
<emphasis id="BB8CA83AFFA1FFA49D8CFDF747608DEF" bold="true" box="[199,334,620,645]" pageId="18" pageNumber="537">Etymology.</emphasis>
The specific epithet refers to this species montane habitat.
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFA1FFA49D8CFD1443B88DC2" box="[199,1430,655,680]" pageId="18" pageNumber="537" type="distribution">
<paragraph id="89477428FFA1FFA49D8CFD1443B88DC2" blockId="18.[151,1437,151,717]" box="[199,1430,655,680]" pageId="18" pageNumber="537">
<emphasis id="BB8CA83AFFA1FFA49D8CFD14474E8DC2" bold="true" box="[199,352,655,680]" pageId="18" pageNumber="537">Distribution.</emphasis>
Known only from the Sierra Sur of Oaxaca,
<collectingCountry id="F1EF34B8FFA1FFA49E13FD14459F8DC2" box="[856,945,655,680]" name="Mexico" pageId="18" pageNumber="537">México</collectingCountry>
, in the vicinity of San Miguel Suchixtepec.
</paragraph>
</subSubSection>
<subSubSection id="C1E227A3FFA1FFA49D8CFD2F45EE8DA6" box="[199,960,692,717]" pageId="18" pageNumber="537" type="biology_ecology">
<paragraph id="89477428FFA1FFA49D8CFD2F45EE8DA6" blockId="18.[151,1437,151,717]" box="[199,960,692,717]" pageId="18" pageNumber="537">
<emphasis id="BB8CA83AFFA1FFA49D8CFD2F47098DA7" bold="true" box="[199,295,692,717]" pageId="18" pageNumber="537">Biology.</emphasis>
Specimens were collected at 9500 (
<quantity id="4E00D9CDFFA1FFA49F88FD2F45348DA6" box="[707,794,692,717]" metricMagnitude="3" metricUnit="m" metricValue="2.895" pageId="18" pageNumber="537" unit="m" value="2895.0">2895 m</quantity>
) in early June.
</paragraph>
</subSubSection>
</treatment>
</document>