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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1086.77408" ID-GBIF-Dataset="dd81d919-f6d9-4fda-bbd1-7c45de307fc2" ID-PMC="PMC8863766" ID-Pensoft-Pub="1313-2970-1086-33" ID-Pensoft-UUID="43AFBD7D61105F598ADA74076D751C11" ID-PubMed="35210912" ID-ZooBank="6C32148CA0144AC488EA82A1509E44F8" ModsDocID="1313-2970-1086-33" checkinTime="1644966834654" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Meng, Kaibaryer &amp; Fekete, Judit" docDate="2022" docId="4CBFCF0DA92355C880ADF3DEEF019CB6" docLanguage="en" docName="ZooKeys 1086: 33-76" docOrigin="ZooKeys 1086" docPubDate="2022-02-15" docSource="http://dx.doi.org/10.3897/zookeys.1086.77408" docTitle="Laeocathaica amdoana Moellendorff 1899" docType="treatment" docVersion="5" id="43AFBD7D61105F598ADA74076D751C11" lastPageNumber="33" masterDocId="43AFBD7D61105F598ADA74076D751C11" masterDocTitle="A review of the genus Laeocathaica Moellendorff, 1899 (Gastropoda, Pulmonata, Camaenidae)" masterLastPageNumber="76" masterPageNumber="33" pageNumber="33" updateTime="1668151486723" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the genus Laeocathaica Moellendorff, 1899 (Gastropoda, Pulmonata, Camaenidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022, Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., Budapest 1148, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Meng, Kaibaryer</mods:namePart>
<mods:affiliation>National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, H- 8200 Veszprem, Egyetem u. 10, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-02-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1086</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>33</mods:start>
<mods:end>76</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1086.77408</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1086.77408</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1086-33</mods:identifier>
<mods:identifier type="ZooBank">6C32148CA0144AC488EA82A1509E44F8</mods:identifier>
<mods:identifier type="Pensoft-UUID">43AFBD7D61105F598ADA74076D751C11</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="193377849" LSID="urn:lsid:plazi:treatment:4CBFCF0DA92355C880ADF3DEEF019CB6" httpUri="http://treatment.plazi.org/id/4CBFCF0DA92355C880ADF3DEEF019CB6" lastPageNumber="33" pageId="0" pageNumber="33">
<subSubSection pageId="0" pageNumber="33" type="nomenclature">
<paragraph pageId="0" pageNumber="33">
<taxonomicName LSID="4CBFCF0D-A923-55C8-80AD-F3DEEF019CB6" authority="Moellendorff, 1899" authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
Laeocathaica amdoana
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="description">
<paragraph pageId="0" pageNumber="33">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Laeocathaica amdoana A, B lectotype (SMF 8952) C, D 2016 / 74, specimen 2 E 2016 / 74, specimen 1. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure3" httpUri="https://binary.pensoft.net/fig/646378" pageId="0" pageNumber="33">Figures 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Laeocathaica amdoana Moellendorff, 1899, 2016 / 75 (4 different shells of the same sample). Scale bar: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure4" httpUri="https://binary.pensoft.net/fig/646379" pageId="0" pageNumber="33">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Laeocathaica amdoana Moellendorff, 1899 A lectotype of L. distinguenda (SMF 8959) B 2016 / 72, specimen 1 C 2016 / 72, specimen 2 D 2016 / 76. Scale bar: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure5" httpUri="https://binary.pensoft.net/fig/646380" pageId="0" pageNumber="33">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Laeocathaica amdoana Moellendorff, 1899 A, B 2016 / 70 a C, D 2016 / 71 E 2016 / 69. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure6" httpUri="https://binary.pensoft.net/fig/646381" pageId="0" pageNumber="33">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Laeocathaica amdoana Moellendorff, 1899 A lectotype of Laeocathaica tropidorhaphe (SMF 9074) B 2016 / 88 C 2016 / 89 D paratype of Laeocathaica dangchangensis Chen &amp; Zhang, 2004 (labelled as holotype). Scale bar: 10 mm. Photographs: K. Meng (D) and B. Pall-Gergely (A-C)." figureDoi="10.3897/zookeys.1086.77408.figure7" httpUri="https://binary.pensoft.net/fig/646382" pageId="0" pageNumber="33">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="reference_group">
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">Laeocathaica amdoana</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899: 92-93, pl. 5, fig. 5.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="distinguenda">Laeocathaica distinguenda</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899: 93, pl. 5, fig. 6.
<emphasis bold="true" pageId="0" pageNumber="33">new synonym</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="tropidorhaphe">Laeocathaica tropidorhaphe</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899: 94, pl. 5, fig. 7.
<emphasis bold="true" pageId="0" pageNumber="33">new synonym</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">Laeocathaica amdoana</taxonomicName>
. -
<bibRefCitation author="Yen, T-C" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" pageId="0" pageNumber="33" refId="B19" refString="Yen, T-C, 1939. Die chinesischen Land- und Suesswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" title="Die chinesischen Land- und Suesswasser-Gastropoden des Natur-Museums Senckenberg." year="1939">Yen 1939</bibRefCitation>
: 148, pl. 15, fig. 31.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="distinguenda">Laeocathaica distinguenda</taxonomicName>
. -
<bibRefCitation author="Yen, T-C" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" pageId="0" pageNumber="33" refId="B19" refString="Yen, T-C, 1939. Die chinesischen Land- und Suesswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" title="Die chinesischen Land- und Suesswasser-Gastropoden des Natur-Museums Senckenberg." year="1939">Yen 1939</bibRefCitation>
: 149, pl. 15, fig. 32.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="tropidorhaphe">Laeocathaica tropidorhaphe</taxonomicName>
. -
<bibRefCitation author="Yen, T-C" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" pageId="0" pageNumber="33" refId="B19" refString="Yen, T-C, 1939. Die chinesischen Land- und Suesswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" title="Die chinesischen Land- und Suesswasser-Gastropoden des Natur-Museums Senckenberg." year="1939">Yen 1939</bibRefCitation>
: 149, pl. 15, fig. 33.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica (Laeocathaica) amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana" subGenus="Laeocathaica">Laeocathaica (Laeocathaica) amdoana</taxonomicName>
. -
<bibRefCitation author="Zilch, A" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" pageId="0" pageNumber="33" refId="B20" refString="Zilch, A, 1968. Die Typen und Typoide des Natur-Museums Senckenberg, 41: Mollusca, Bradybaenidae, Bradybaeninae. Archiv fuer Molluskenkunde 98(3/4): 155-207." title="Die Typen und Typoide des Natur-Museums Senckenberg, 41: Mollusca, Bradybaenidae, Bradybaeninae. Archiv fuer Molluskenkunde 98 (3 / 4): 155 - 207." year="1968">Zilch 1968</bibRefCitation>
: 173.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica (Laeocathaica) distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="distinguenda" subGenus="Laeocathaica">Laeocathaica (Laeocathaica) distinguenda</taxonomicName>
. -
<bibRefCitation author="Zilch, A" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" pageId="0" pageNumber="33" refId="B20" refString="Zilch, A, 1968. Die Typen und Typoide des Natur-Museums Senckenberg, 41: Mollusca, Bradybaenidae, Bradybaeninae. Archiv fuer Molluskenkunde 98(3/4): 155-207." title="Die Typen und Typoide des Natur-Museums Senckenberg, 41: Mollusca, Bradybaenidae, Bradybaeninae. Archiv fuer Molluskenkunde 98 (3 / 4): 155 - 207." year="1968">Zilch 1968</bibRefCitation>
: 173.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica (Laeocathaica) tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="tropidorhaphe" subGenus="Laeocathaica">Laeocathaica (Laeocathaica) tropidorhaphe</taxonomicName>
. -
<bibRefCitation author="Zilch, A" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main" pageId="0" pageNumber="33" refId="B20" refString="Zilch, A, 1968. Die Typen und Typoide des Natur-Museums Senckenberg, 41: Mollusca, Bradybaenidae, Bradybaeninae. Archiv fuer Molluskenkunde 98(3/4): 155-207." title="Die Typen und Typoide des Natur-Museums Senckenberg, 41: Mollusca, Bradybaenidae, Bradybaeninae. Archiv fuer Molluskenkunde 98 (3 / 4): 155 - 207." year="1968">Zilch 1968</bibRefCitation>
: 175.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">Laeocathaica amdoana</taxonomicName>
. - Chen &amp; Zhang 2004: 316, fig. 303.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="distinguenda">Laeocathaica distinguenda</taxonomicName>
. - Chen &amp; Zhang, 2004: 318, fig. 305.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="tropidorhaphe">Laeocathaica tropidorhaphe</taxonomicName>
. - Chen &amp; Zhang 2004: 319, fig. 307.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="D. Chen &amp; G. Zhang" authorityYear="2004" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica dangchangensis" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="dangchangensis">Laeocathaica dangchangensis</taxonomicName>
Chen &amp; Zhang, 2004: 339 [Chinese description], 443 [English description], fig. 332.
<emphasis bold="true" pageId="0" pageNumber="33">new synonym</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="type material">
<paragraph pageId="0" pageNumber="33">Type material.</paragraph>
<paragraph pageId="0" pageNumber="33">
China (Gansu), Ho-dshi-gou, coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Potanin 853, SMF 8952 (lectotype of
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
, Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Laeocathaica amdoana A, B lectotype (SMF 8952) C, D 2016 / 74, specimen 2 E 2016 / 74, specimen 1. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure3" httpUri="https://binary.pensoft.net/fig/646378" pageId="0" pageNumber="33">3A, B</figureCitation>
) • China (SO-Gansu), Wen-Hsien, coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Potanin 907, SMF 8953 (paralectotype of
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
) • China (Sy-tshuan): Thal des Pui-hob. Lum-du, coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Potanin 906, SMF 8959 (lectotype of
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica distinguenda</emphasis>
</taxonomicName>
, Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Laeocathaica amdoana Moellendorff, 1899 A lectotype of L. distinguenda (SMF 8959) B 2016 / 72, specimen 1 C 2016 / 72, specimen 2 D 2016 / 76. Scale bar: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure5" httpUri="https://binary.pensoft.net/fig/646380" pageId="0" pageNumber="33">5A</figureCitation>
) • SO-Gansu (NW China), coll. C.R. Boettger ex coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, SMF 95024/1 paralectotype of
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica distinguenda</emphasis>
</taxonomicName>
• SO-Gansu: Nanping, coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Potanin 8, 64, 846, SMF 8958/5 paratypes of
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica distinguenda</emphasis>
</taxonomicName>
• SO-Gansu,
<normalizedToken originalValue="Yü-Lin-Guam">Yue-Lin-Guam</normalizedToken>
, u. Wen-hsien, coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Potanin 11, 521, 565, SMF 8955/3 paralectotypes of
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica distinguenda</emphasis>
</taxonomicName>
• China: SO-Gansu, coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Potanin 725a, SMF 8957/3 paralectotypes of
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica distinguenda</emphasis>
</taxonomicName>
• Shy-Pu am Pui-hu, coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Potanin 69, 653, SMF 8956/2 paralectotypes of
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica distinguenda</emphasis>
</taxonomicName>
• SO-Gansu, zw. Li-dshia-pu u. Hsi-gu-tsheng, coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Potanin 923, SMF 9074 (lectotype of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">L. tropidorhaphe</emphasis>
</taxonomicName>
, Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Laeocathaica amdoana Moellendorff, 1899 A lectotype of Laeocathaica tropidorhaphe (SMF 9074) B 2016 / 88 C 2016 / 89 D paratype of Laeocathaica dangchangensis Chen &amp; Zhang, 2004 (labelled as holotype). Scale bar: 10 mm. Photographs: K. Meng (D) and B. Pall-Gergely (A-C)." figureDoi="10.3897/zookeys.1086.77408.figure7" httpUri="https://binary.pensoft.net/fig/646382" pageId="0" pageNumber="33">7A</figureCitation>
) • Same data, SMF 9077/2 paralectotypes • NW-China, SO-Gansu, coll. C.R. Boettger 1904 ex coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
SMF 95126/1 (paralectotype of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">L. tropidorhaphe</emphasis>
</taxonomicName>
) • SO-Gansu, Tan-tshang, coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Potanin 545, 623, 808, SMF 9075/4 (paralectotypes of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">L. tropidorhaphe</emphasis>
</taxonomicName>
) • Gansu: Dshie-dshou, coll.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Potanin 119, SMF 9076/1 (paralectotype of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">L. tropidorhaphe</emphasis>
</taxonomicName>
) • IZCAS TM 095895 (labelled as holotype of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. dangchangensis" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="dangchangensis">
<emphasis italics="true" pageId="0" pageNumber="33">L. dangchangensis</emphasis>
</taxonomicName>
, but its measurements do not match with the ones given in the original description, see also remarks, Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Laeocathaica amdoana Moellendorff, 1899 A lectotype of Laeocathaica tropidorhaphe (SMF 9074) B 2016 / 88 C 2016 / 89 D paratype of Laeocathaica dangchangensis Chen &amp; Zhang, 2004 (labelled as holotype). Scale bar: 10 mm. Photographs: K. Meng (D) and B. Pall-Gergely (A-C)." figureDoi="10.3897/zookeys.1086.77408.figure7" httpUri="https://binary.pensoft.net/fig/646382" pageId="0" pageNumber="33">7D</figureCitation>
) • IZCAS TM 095843-095894 + IZCAS TM 095896-095950 (103 paratypes of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. dangchangensis" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="dangchangensis">
<emphasis italics="true" pageId="0" pageNumber="33">L. dangchangensis</emphasis>
</taxonomicName>
in ethanol).
</paragraph>
<caption doi="10.3897/zookeys.1086.77408.figure3" httpUri="https://binary.pensoft.net/fig/646378" pageId="0" pageNumber="33" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Figure 3.</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="33">A, B</emphasis>
lectotype (SMF 8952)
<emphasis bold="true" pageId="0" pageNumber="33">C, D</emphasis>
2016/74, specimen2
<emphasis bold="true" pageId="0" pageNumber="33">E</emphasis>
2016/74, specimen1. Scale bars: 10 mm. All photographs: B.
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="materials_examined">
<paragraph pageId="0" pageNumber="33">Museum material.</paragraph>
<paragraph pageId="0" pageNumber="33">
<materialsCitation accessionNumber="SMF 42564, SMF 24269" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Szechwan, Moellendorff" specimenCount="1">
<collectorName>Szechwan</collectorName>
, coll.
<normalizedToken originalValue="Möllendorff">
<collectorName>Moellendorff</collectorName>
</normalizedToken>
,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF42564">SMF 42564</accessionNumber>
/1 • Tan-Tschan, coll.
<normalizedToken originalValue="Möllendorff">
<collectorName>Moellendorff</collectorName>
</normalizedToken>
,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF24269">SMF 24269</accessionNumber>
/1 (&quot;
<emphasis italics="true" pageId="0" pageNumber="33">tropidorhape</emphasis>
&quot;)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1086.77408.figure4" httpUri="https://binary.pensoft.net/fig/646379" pageId="0" pageNumber="33" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Figure 4.</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, 2016/75 (4 different shells of the same sample). Scale bar:
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
. All photographs: B.
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="materials_examined">
<paragraph pageId="0" pageNumber="33">
New material (typical
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="tropidorhaphe">Laeocathaica tropidorhaphe</taxonomicName>
).
</paragraph>
<paragraph pageId="0" pageNumber="33">
<materialsCitation collectingDate="2016-06-02" collectingDateMax="2016-06-04" collectingDateMin="2016-06-02" collectorName="A. Hunyadi" country="China" county="Longnan Shi" elevation="1835" latitude="33.532066" location="Chengguan Zhen" longLatPrecision="1" longitude="104.65477" municipality="Dangchang Xian" specimenCount="1" stateProvince="Gansu">
<collectingCountry name="China">China</collectingCountry>
• 2 shells;
<collectingRegion country="China" name="Gansu">Gansu</collectingRegion>
,
<collectingCounty>Longnan Shi</collectingCounty>
,
<collectingMunicipality>Dangchang Xian</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:4CBFCF0DA92355C880ADF3DEEF019CB6:7ED28200563FF75538426A9C90C9FFF9" country="China" county="Longnan Shi" latitude="33.532066" longLatPrecision="1" longitude="104.65477" municipality="Dangchang Xian" name="Chengguan Zhen" stateProvince="Gansu">Chengguan Zhen</location>
,
<location LSID="urn:lsid:plazi:treatment:4CBFCF0DA92355C880ADF3DEEF019CB6:6CD88EE6F88226A9F66F78D614BAB4FE" country="China" county="Longnan Shi" latitude="33.532066" longLatPrecision="1" longitude="104.65477" municipality="Dangchang Xian" name="Wenchangmiao" stateProvince="Gansu">Wenchangmiao</location>
, temple hill (locality code: 2016/87);
<geoCoordinate degrees="34" direction="north" minutes="02.394" orientation="latitude" precision="1" value="34.0399">34°02.394'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="23.499" orientation="longitude" precision="1" value="104.39165">104°23.499'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.835" unit="m" value="1835.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.835" unit="m" value="1835.0">1835 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-06-02">02 June 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 2 shells;
<collectingRegion country="China" name="Gansu">Gansu</collectingRegion>
, Longnan Shi, Dangchang Xian, Lianghekou Xiang, Lianghekou Cun, rock above the intersection (locality code: 2016/89);
<geoCoordinate degrees="33" direction="north" minutes="41.808" orientation="latitude" precision="1" value="33.6968">33°41.808'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="29.182" orientation="longitude" precision="1" value="104.48637">104°29.182'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" unit="m" value="1245.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" unit="m" value="1245.0">1245 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-06-02">02 June 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution of Laeocathaica amdoana Moellendorff, 1899 in China (detail from Fig. 2 A)." figureDoi="10.3897/zookeys.1086.77408.figure8" httpUri="https://binary.pensoft.net/fig/646383" pageId="0" pageNumber="33">8C</figureCitation>
) • 2 shells;
<collectingRegion country="China" name="Gansu">Gansu</collectingRegion>
, Longnan Shi, Dangchang Xian, Guanting Zhen,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="km" value="1.5">1.5 km</quantity>
north of
<location LSID="urn:lsid:plazi:treatment:4CBFCF0DA92355C880ADF3DEEF019CB6:49D0B26259BBDE1ECDE7D950FBA39BAA" country="China" county="Longnan Shi" latitude="33.532066" longLatPrecision="1" longitude="104.65477" municipality="Dangchang Xian" name="Guanting" stateProvince="Gansu">Guanting</location>
towards Dangchang (locality code: 2016/88);
<geoCoordinate degrees="33" direction="north" minutes="50.803" orientation="latitude" precision="1" value="33.846718">33°50.803'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="32.470" orientation="longitude" precision="1" value="104.54117">104°32.470'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.815" unit="m" value="1815.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.815" unit="m" value="1815.0">1815 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-06-02">02 June 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Laeocathaica amdoana Moellendorff, 1899 A lectotype of Laeocathaica tropidorhaphe (SMF 9074) B 2016 / 88 C 2016 / 89 D paratype of Laeocathaica dangchangensis Chen &amp; Zhang, 2004 (labelled as holotype). Scale bar: 10 mm. Photographs: K. Meng (D) and B. Pall-Gergely (A-C)." figureDoi="10.3897/zookeys.1086.77408.figure7" httpUri="https://binary.pensoft.net/fig/646382" pageId="0" pageNumber="33">7B</figureCitation>
) • 2 shells;
<collectingRegion country="China" name="Gansu">Gansu</collectingRegion>
, Longnan Shi, Zhugqu Xian,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="km" value="2.5">2.5 km</quantity>
west of
<location LSID="urn:lsid:plazi:treatment:4CBFCF0DA92355C880ADF3DEEF019CB6:474D2225939E38098727C9EFEFEEF8B3" country="China" county="Longnan Shi" latitude="33.532066" longLatPrecision="1" longitude="104.65477" municipality="Dangchang Xian" name="Suoertou Cun" stateProvince="Gansu">Suoertou Cun</location>
, northern bank of Bailong He (locality code: 2016/91);
<geoCoordinate degrees="33" direction="north" minutes="46.906" orientation="latitude" precision="1" value="33.781765">33°46.906'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="20.106" orientation="longitude" precision="1" value="104.3351">104°20.106'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.2349999999999999" unit="m" value="1235.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2349999999999999" unit="m" value="1235.0">1235 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-06-03">03 June 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 2 shells;
<collectingRegion country="China" name="Gansu">Gansu</collectingRegion>
, Longnan Shi, Wudu Xian, Jiaogong Zhen,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="km" value="1.5">1.5 km</quantity>
west of
<location LSID="urn:lsid:plazi:treatment:4CBFCF0DA92355C880ADF3DEEF019CB6:A3FEE8EFDDF415E9C96B688B4170549E" country="China" county="Longnan Shi" latitude="33.532066" longLatPrecision="1" longitude="104.65477" municipality="Dangchang Xian" name="Chenjiaba Cun" stateProvince="Gansu">Chenjiaba Cun</location>
, Zhaoyangdong, below the cave (locality code: 2016/95);
<geoCoordinate degrees="33" direction="north" minutes="31.924" orientation="latitude" precision="1" value="33.532066">33°31.924'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="39.286" orientation="longitude" precision="1" value="104.65477">104°39.286'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.175" unit="m" value="1175.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.175" unit="m" value="1175.0">1175 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-06-04">04 June 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1086.77408.figure5" httpUri="https://binary.pensoft.net/fig/646380" pageId="0" pageNumber="33" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Figure 5.</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
<typeStatus>lectotype</typeStatus>
of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">L. distinguenda</emphasis>
</taxonomicName>
(SMF 8959)
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
2016/72, specimen1
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
2016/72, specimen2
<emphasis bold="true" pageId="0" pageNumber="33">D</emphasis>
2016/76. Scale bar:
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
. All photographs: B.
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1086.77408.figure6" httpUri="https://binary.pensoft.net/fig/646381" pageId="0" pageNumber="33" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Figure 6.</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899
<emphasis bold="true" pageId="0" pageNumber="33">A, B</emphasis>
2016/70a
<emphasis bold="true" pageId="0" pageNumber="33">C, D</emphasis>
2016/71
<emphasis bold="true" pageId="0" pageNumber="33">E</emphasis>
2016/69. Scale bars:
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
. All photographs: B.
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="materials_examined">
<paragraph pageId="0" pageNumber="33">New material (other morphs).</paragraph>
<paragraph pageId="0" pageNumber="33">
<materialsCitation collectingDate="2016-05-28" collectingDateMax="2016-05-31" collectingDateMin="2016-05-28" collectorName="A. Hunyadi, Sichuan, Aba, Jiuzhaigou Xian, Baihe Xiang, Taiping Cun, Baishui He, Anle Xiang, Jiuzhaigou Shi" county="Lvjiaba" elevation="1445" latitude="33.061867" location="Zhongtianshan Cun" longLatPrecision="1" longitude="104.836815" municipality="Jianshan" specimenCount="1">
6 shells; Sichuan, Aba, Jiuzhaigou Xian, Baihe Xiang, southern edge of Taiping Cun, rock wall facing north (locality code: 2016/72);
<geoCoordinate degrees="33" direction="north" minutes="18.026" orientation="latitude" precision="1" value="33.300434">33°18.026'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="09.500" orientation="longitude" precision="1" value="104.15833">104°09.500'E</geoCoordinate>
;
<collectingDate value="2016-05-30">30 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Laeocathaica amdoana Moellendorff, 1899 A lectotype of L. distinguenda (SMF 8959) B 2016 / 72, specimen 1 C 2016 / 72, specimen 2 D 2016 / 76. Scale bar: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure5" httpUri="https://binary.pensoft.net/fig/646380" pageId="0" pageNumber="33">5B, C</figureCitation>
) • 1 shell;
<collectorName>Sichuan</collectorName>
,
<collectorName>Aba</collectorName>
,
<collectorName>Jiuzhaigou Xian</collectorName>
,
<collectorName>Baihe Xiang</collectorName>
,
<collectorName>Taiping Cun</collectorName>
, eastern bank of
<collectorName>Baishui He</collectorName>
(locality code: 2016/73);
<geoCoordinate degrees="33" direction="north" minutes="18.366" orientation="latitude" precision="1" value="33.3061">33°18.366'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="09.413" orientation="longitude" precision="1" value="104.15688">104°09.413'E</geoCoordinate>
;
<collectingDate value="2016-05-30">30 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 4 shells;
<collectorName>Sichuan</collectorName>
,
<collectorName>Aba</collectorName>
,
<collectorName>Jiuzhaigou Xian</collectorName>
,
<collectorName>Anle Xiang</collectorName>
, ca.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="km" value="1.5">1.5 km</quantity>
east of
<location LSID="urn:lsid:plazi:treatment:4CBFCF0DA92355C880ADF3DEEF019CB6:1165F005A3593F658C95103F6335CDEC" county="Lvjiaba" latitude="33.061867" longLatPrecision="1" longitude="104.836815" municipality="Jianshan" name="Zhongtianshan Cun">Zhongtianshan Cun</location>
towards
<collectorName>Jiuzhaigou Shi</collectorName>
(locality code: 2016/74);
<geoCoordinate degrees="33" direction="north" minutes="17.279" orientation="latitude" precision="1" value="33.287983">33°17.279'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="12.702" orientation="longitude" precision="1" value="104.2117">104°12.702'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.4449999999999998" unit="m" value="1445.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.4449999999999998" unit="m" value="1445.0">1445 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-30">30 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Laeocathaica amdoana A, B lectotype (SMF 8952) C, D 2016 / 74, specimen 2 E 2016 / 74, specimen 1. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure3" httpUri="https://binary.pensoft.net/fig/646378" pageId="0" pageNumber="33">3C-E</figureCitation>
) • 4 shells; Sichuan, Aba, Jiuzhaigou Xian, Guoyuan Xiang,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
from Guoyuanyi Cun towards Lengshuishan Cun (locality code: 2016/75);
<geoCoordinate degrees="33" direction="north" minutes="07.681" orientation="latitude" precision="1" value="33.128017">33°07.681'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="18.489" orientation="longitude" precision="1" value="104.30815">104°18.489'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.22" unit="m" value="1220.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.22" unit="m" value="1220.0">1220 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-30">30 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Laeocathaica amdoana Moellendorff, 1899, 2016 / 75 (4 different shells of the same sample). Scale bar: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure4" httpUri="https://binary.pensoft.net/fig/646379" pageId="0" pageNumber="33">4</figureCitation>
) • 1 shell; Sichuan, Aba, Jiuzhaigou Xian, Guoyuan Xiang, Guoyuaner Cun, environment of the bridge (locality code: 2016/76);
<geoCoordinate degrees="33" direction="north" minutes="06.922" orientation="latitude" precision="1" value="33.115368">33°06.922'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="19.617" orientation="longitude" precision="1" value="104.32695">104°19.617'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-30">30 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Laeocathaica amdoana Moellendorff, 1899 A lectotype of L. distinguenda (SMF 8959) B 2016 / 72, specimen 1 C 2016 / 72, specimen 2 D 2016 / 76. Scale bar: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure5" httpUri="https://binary.pensoft.net/fig/646380" pageId="0" pageNumber="33">5D</figureCitation>
) • 3 shells; Gansu, Longnan Shi, Wenxian, Buziba Xiang, southern edge of Buziba Cun, western bank of the river (locality code: 2016/77);
<geoCoordinate degrees="33" direction="north" minutes="03.592" orientation="latitude" precision="1" value="33.059868">33°03.592'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="37.094" orientation="longitude" precision="1" value="104.61823">104°37.094'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.215" unit="m" value="1215.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.215" unit="m" value="1215.0">1215 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-31">31 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 1 shell; Gansu, Longnan Shi, Wenxian, Buziba Xiang, northern edge of Taojiaba Cun,
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
</quantity>
towards Buziba (locality code: 2016/78);
<geoCoordinate degrees="33" direction="north" minutes="02.706" orientation="latitude" precision="1" value="33.0451">33°02.706'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="37.157" orientation="longitude" precision="1" value="104.619286">104°37.157'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-31">31 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 4 shells; Gansu, Longnan Shi, Wenxian, Buziba Xiang,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
south of
<location LSID="urn:lsid:plazi:treatment:4CBFCF0DA92355C880ADF3DEEF019CB6:F3BB8D24A13203D3786501BED8DE5DB3" county="Lvjiaba" latitude="33.061867" longLatPrecision="1" longitude="104.836815" municipality="Jianshan" name="Taojiaba Cun">Taojiaba Cun</location>
towards Dongyukou Cun (locality code: 2016/79);
<geoCoordinate degrees="33" direction="north" minutes="01.865" orientation="latitude" precision="1" value="33.031082">33°01.865'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="37.329" orientation="longitude" precision="1" value="104.62215">104°37.329'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">1150 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-31">31 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 4 shells; Gansu, Longnan Shi, Wenxian, Chengguan Zhen, next to a museum (locality code: 2016/64);
<geoCoordinate degrees="32" direction="north" minutes="56.471" orientation="latitude" precision="1" value="32.941185">32°56.471'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="40.379" orientation="longitude" precision="1" value="104.67298">104°40.379'E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.65" metricValueMax="9.7" metricValueMin="9.6" unit="m" value="965.0" valueMax="970.0" valueMin="960.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.65" metricValueMax="9.7" metricValueMin="9.6" unit="m" value="965.0" valueMax="970.0" valueMin="960.0">960-970 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-28">28 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 3 shells; Gansu, Longnan Shi, Wenxian, Chengguan Zhen, cemetery hill above the city (locality code: 2016/65);
<geoCoordinate degrees="32" direction="north" minutes="57.026" orientation="latitude" precision="1" value="32.95043">32°57.026'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="40.527" orientation="longitude" precision="1" value="104.67545">104°40.527'E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.09" unit="m" value="1090.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.09" unit="m" value="1090.0">1090 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-28">28 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 4 shells; Gansu, Longnan Shi, Wenxian, Jianshan Xiang,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
</quantity>
south of
<location LSID="urn:lsid:plazi:treatment:4CBFCF0DA92355C880ADF3DEEF019CB6:33E633A75CE8113B86ADA7CED395ACDA" county="Lvjiaba" latitude="33.061867" longLatPrecision="1" longitude="104.836815" municipality="Jianshan" name="Hekou Cun">Hekou Cun</location>
, eastern bank of Bailong He (locality code: 2016/68);
<geoCoordinate degrees="33" direction="north" minutes="01.703" orientation="latitude" precision="1" value="33.028385">33°01.703'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="53.602" orientation="longitude" precision="1" value="104.893364">104°53.602'E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.1" unit="m" value="810.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.1" unit="m" value="810.0">810 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-29">29 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 2 shells; Gansu, Longnan Shi, eastern edge of Wenxian, northern bank of the river (locality code: 2016/66);
<geoCoordinate degrees="32" direction="north" minutes="56.459" orientation="latitude" precision="1" value="32.940983">32°56.459'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="41.372" orientation="longitude" precision="1" value="104.68954">104°41.372'E</geoCoordinate>
;
<collectingDate value="2016-05-28">28 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 4 shells; Gansu, Longnan Shi, Wenxian, Jianshan Xiang, southern edge of Hekou Cun, western bank of Bailong He (locality code: 2016/67);
<geoCoordinate degrees="33" direction="north" minutes="02.014" orientation="latitude" precision="1" value="33.033566">33°02.014'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="53.478" orientation="longitude" precision="1" value="104.8913">104°53.478'E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-29">29 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 3 shells; Gansu, Longnan Shi, Wenxian, Shifang Xiang,
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
</quantity>
from the northwestern edge of Baiyiba Cun towards Dongyukou Cun, left bank of the river (locality code: 2016/83);
<geoCoordinate degrees="32" direction="north" minutes="58.985" orientation="latitude" precision="1" value="32.98308">32°58.985'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="37.503" orientation="longitude" precision="1" value="104.62505">104°37.503'E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.7" unit="m" value="970.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.7" unit="m" value="970.0">970 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-31">31 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 4 shells; Gansu, Longnan Shi, Wenxian, Shifang Xiang,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</elevation>
</quantity>
northwest from Baiyiba Cun towards Dongyukou Cun (locality code: 2016/82);
<geoCoordinate degrees="32" direction="north" minutes="59.346" orientation="latitude" precision="1" value="32.9891">32°59.346'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="37.233" orientation="longitude" precision="1" value="104.62055">104°37.233'E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.8" unit="m" value="980.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.8" unit="m" value="980.0">980 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-31">31 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 9 shells; Gansu, Longnan Shi, Wenxian, Jianshan Xiang, western edge of Hekou Cun towards Caojiaba, along road no. 212 (locality code: 2016/69);
<geoCoordinate degrees="33" direction="north" minutes="2.343" orientation="latitude" precision="1" value="33.03905">33°2.343'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="53.045" orientation="longitude" precision="1" value="104.88409">104°53.045'E</geoCoordinate>
;
<collectingDate value="2016-05-29">29 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Laeocathaica amdoana Moellendorff, 1899 A, B 2016 / 70 a C, D 2016 / 71 E 2016 / 69. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure6" httpUri="https://binary.pensoft.net/fig/646381" pageId="0" pageNumber="33">6E</figureCitation>
) • 8 shells; Gansu, Longnan Shi, Wenxian, Jianshan Xiang,
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</elevation>
</quantity>
west of
<collectingMunicipality>Jianshan</collectingMunicipality>
towards Diaohuya (locality code: 2016/70a);
<geoCoordinate degrees="33" direction="north" minutes="02.559" orientation="latitude" precision="1" value="33.04265">33°02.559'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="51.254" orientation="longitude" precision="1" value="104.85423">104°51.254'E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">850 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2016-05-29">29 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Laeocathaica amdoana Moellendorff, 1899 A, B 2016 / 70 a C, D 2016 / 71 E 2016 / 69. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure6" httpUri="https://binary.pensoft.net/fig/646381" pageId="0" pageNumber="33">6A, B</figureCitation>
) • 4 shells; Gansu, Longnan Shi, Wenxian, Jianshan Xiang,
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
</quantity>
north of
<collectingCounty>Lvjiaba</collectingCounty>
, along road no. 212 (locality code: 2016/71);
<geoCoordinate degrees="33" direction="north" minutes="03.712" orientation="latitude" precision="1" value="33.061867">33°03.712'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="50.209" orientation="longitude" precision="1" value="104.836815">104°50.209'E</geoCoordinate>
;
<collectingDate value="2016-05-29">29 May 2016</collectingDate>
;
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Laeocathaica amdoana Moellendorff, 1899 A, B 2016 / 70 a C, D 2016 / 71 E 2016 / 69. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure6" httpUri="https://binary.pensoft.net/fig/646381" pageId="0" pageNumber="33">6C, D</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1086.77408.figure7" httpUri="https://binary.pensoft.net/fig/646382" pageId="0" pageNumber="33" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Figure 7.</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
<typeStatus>lectotype</typeStatus>
of
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica tropidorhaphe</emphasis>
</taxonomicName>
(SMF 9074)
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
2016/88
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
2016/89
<emphasis bold="true" pageId="0" pageNumber="33">D</emphasis>
<typeStatus>paratype</typeStatus>
of
<taxonomicName authorityName="D. Chen &amp; G. Zhang" authorityYear="2004" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica dangchangensis" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="dangchangensis">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica dangchangensis</emphasis>
</taxonomicName>
Chen &amp; Zhang, 2004 (labelled as
<typeStatus>holotype</typeStatus>
). Scale bar:
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
. Photographs: K. Meng (
<emphasis bold="true" pageId="0" pageNumber="33">D</emphasis>
) and B.
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="33">A-C</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="distribution">
<paragraph pageId="0" pageNumber="33">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="33">
This species is known from several sites in southwestern Gansu Province and the neighbouring areas in Sichuan (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution of Laeocathaica amdoana Moellendorff, 1899 in China (detail from Fig. 2 A)." figureDoi="10.3897/zookeys.1086.77408.figure8" httpUri="https://binary.pensoft.net/fig/646383" pageId="0" pageNumber="33">8</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1086.77408.figure8" httpUri="https://binary.pensoft.net/fig/646383" pageId="0" pageNumber="33" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Figure 8.</emphasis>
Distribution of
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899 in China (detail from Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of the genus Laeocathaica in China A see Figs 8, 11, 17, 24 for details B see Fig. 14 for detail." figureDoi="10.3897/zookeys.1086.77408.figure2" httpUri="https://binary.pensoft.net/fig/646377" pageId="0" pageNumber="33">2A</figureCitation>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="remarks">
<paragraph pageId="0" pageNumber="33">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
was described from Pass Ho-Dshi-Gou bei Mu-gua-tshi (the exact locality could not be located on the map) near Wenxian at the border with Sichuan province, and characterised by a domed, brown, dorsal side with a relatively broad, sharp, distinctly bordering, white band of the keel. We did not find shells identical to typical
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
, but have found similar ones that can be identified as conspecific (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Laeocathaica amdoana A, B lectotype (SMF 8952) C, D 2016 / 74, specimen 2 E 2016 / 74, specimen 1. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure3" httpUri="https://binary.pensoft.net/fig/646378" pageId="0" pageNumber="33">3C, D</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Laeocathaica amdoana Moellendorff, 1899, 2016 / 75 (4 different shells of the same sample). Scale bar: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure4" httpUri="https://binary.pensoft.net/fig/646379" pageId="0" pageNumber="33">4</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica distinguenda</emphasis>
</taxonomicName>
is represented in the Senckenberg Museum by several samples. Strangely, the lectotype is the most
<normalizedToken originalValue="“untypical”">&quot;untypical&quot;</normalizedToken>
among the lots labelled as
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">L. distinguenda</emphasis>
</taxonomicName>
due to its pale caramel colour, the blurry border of light and dark stripes, and the rounded body whorl. Our samples from the vicinity of Wenxian (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Laeocathaica amdoana Moellendorff, 1899 A lectotype of L. distinguenda (SMF 8959) B 2016 / 72, specimen 1 C 2016 / 72, specimen 2 D 2016 / 76. Scale bar: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure5" httpUri="https://binary.pensoft.net/fig/646380" pageId="0" pageNumber="33">5B-D</figureCitation>
) are more similar to the paralectotypes of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">L. distinguenda</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica tropidorhaphe</emphasis>
</taxonomicName>
was described from the north (Tanchang and its vicinity), and is characterised by a large, keeled shell with a flat dorsal side and a thick brown spiral band. The northernmost populations we collected (samples 2016/87, 2016/88, 2016/89, 2016/91, 2016/95) agree with the types of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">L. tropidorhaphe</emphasis>
</taxonomicName>
in size, shape, and colouration, but their spire height is variable. However, some shells from much further south are also similar (i.e., samples 2016/74, 2016/75, see Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Laeocathaica amdoana Moellendorff, 1899, 2016 / 75 (4 different shells of the same sample). Scale bar: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure4" httpUri="https://binary.pensoft.net/fig/646379" pageId="0" pageNumber="33">4</figureCitation>
). The characteristic colouration of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">L. tropidorhaphe</emphasis>
</taxonomicName>
(brown dorsal side with slender white band on the keel) is also not unique to the northern
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">L. tropidorhaphe</emphasis>
</taxonomicName>
populations, but can be found in more southern populations as well (compare Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Laeocathaica amdoana Moellendorff, 1899, 2016 / 75 (4 different shells of the same sample). Scale bar: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure4" httpUri="https://binary.pensoft.net/fig/646379" pageId="0" pageNumber="33">4a</figureCitation>
with Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Laeocathaica amdoana Moellendorff, 1899 A lectotype of Laeocathaica tropidorhaphe (SMF 9074) B 2016 / 88 C 2016 / 89 D paratype of Laeocathaica dangchangensis Chen &amp; Zhang, 2004 (labelled as holotype). Scale bar: 10 mm. Photographs: K. Meng (D) and B. Pall-Gergely (A-C)." figureDoi="10.3897/zookeys.1086.77408.figure7" httpUri="https://binary.pensoft.net/fig/646382" pageId="0" pageNumber="33">7</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="33">
Overall, there is a continuous variation across most of the historical and newly collected samples in terms of shape of dorsal side, shape of body whorl, size, colour, and sculpture (see Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Laeocathaica amdoana A, B lectotype (SMF 8952) C, D 2016 / 74, specimen 2 E 2016 / 74, specimen 1. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure3" httpUri="https://binary.pensoft.net/fig/646378" pageId="0" pageNumber="33">3B, C</figureCitation>
;
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Laeocathaica amdoana Moellendorff, 1899 A, B 2016 / 70 a C, D 2016 / 71 E 2016 / 69. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure6" httpUri="https://binary.pensoft.net/fig/646381" pageId="0" pageNumber="33">6B, C</figureCitation>
).
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
and the lectotype of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">L. distinguenda</emphasis>
</taxonomicName>
(but not the paralectotypes!) seem to be slightly out of the morphological continuum, but not to a degree that a species-level distinction should be applied.
</paragraph>
<paragraph pageId="0" pageNumber="33">
Colouration can be extremely variable even within a single population (see Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Laeocathaica amdoana Moellendorff, 1899, 2016 / 75 (4 different shells of the same sample). Scale bar: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure4" httpUri="https://binary.pensoft.net/fig/646379" pageId="0" pageNumber="33">4</figureCitation>
). Therefore, colour is of minor importance in species distinction within this group of
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica</emphasis>
</taxonomicName>
. It is a general trend that towards the southeastern part of the distribution of these
<normalizedToken originalValue="“species”">&quot;species&quot;</normalizedToken>
, the peripheral keel disappears and the body whorl becomes rounded.
</paragraph>
<paragraph pageId="0" pageNumber="33">
Because of the aforementioned reasons, and until anatomical and DNA sequence data become available, we do not find the names
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. distinguenda" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="33">L. distinguenda</emphasis>
</taxonomicName>
and
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">L. tropidorhaphe</emphasis>
</taxonomicName>
meaningful, and so we provisionally synonymise them with
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. amdoana" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">L. amdoana</emphasis>
</taxonomicName>
. Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Shell morphological traits of Laeocathaica amdoana Moellendorff, 1899 populations." httpUri="http://table.plazi.org/id/A68A5616E42A25407B14AD4933F57581" pageId="0" pageNumber="33" tableUuid="A68A5616E42A25407B14AD4933F57581">1</tableCitation>
summarises the key traits that are variable across and within newly collected populations.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="D. Chen &amp; G. Zhang" authorityYear="2004" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica dangchangensis" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="dangchangensis">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica dangchangensis</emphasis>
</taxonomicName>
Chen &amp; Zhang, 2004 is also a junior synonym of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. amdoana" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">L. amdoana</emphasis>
</taxonomicName>
, because it shows the same characteristic conchological features (large shell size, acute, white keel, almost flat dorsal side) as
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">L. tropidorhaphe</emphasis>
</taxonomicName>
. Moreover, its type locality (Shawan town, Dangshang county (
<geoCoordinate degrees="34" direction="north" minutes="0" orientation="latitude" precision="925" value="34.0">34°0'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="3" orientation="longitude" precision="925" value="104.05">104°3'E</geoCoordinate>
), Gansu Province, China) is situated close to the known sites of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="33">L. tropidorhaphe</emphasis>
</taxonomicName>
, whose two closest populations are situated at ca. 31 km and 35 km from the type locality of
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. dangchangensis" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="dangchangensis">
<emphasis italics="true" pageId="0" pageNumber="33">L. dangchangensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="33">
According to the original description of
<taxonomicName authorityName="D. Chen &amp; G. Zhang" authorityYear="2004" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica dangchangensis" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="dangchangensis">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica dangchangensis</emphasis>
</taxonomicName>
, the holotype has a shell width of 27.22 mm. However, the shell labelled as the holotype is 23 mm wide. Moreover, the number 6 is written on that
<normalizedToken originalValue="specimen’s">specimen's</normalizedToken>
dorsal side, whereas &quot;Holotype: Sp8&quot; is written on the label. We have not found any shells bearing the number 8. Consequently, the shell labelled as the holotype is a paratype, and the real holotype is probably one of the specimens labelled as paratypes, or lost.
</paragraph>
</subSubSection>
</treatment>
</document>
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