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<document ID-DOI="10.11646/zootaxa.5277.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7889695" ID-ZooBank="73ED69F3-3887-460D-86DE-C9F1302C8EC3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683099613832" checkinUser="plazi" docAuthor="Kluge, Nikita, Srinivasan, Pandiarajan, Sivaruban, T., Barathy, S. &amp; Isack, Rajasekaran" docDate="2023" docId="03F487D0A36C4D55A2DBFF43FC92E925" docLanguage="en" docName="zootaxa.5277.2.1.pdf" docOrigin="Zootaxa 5277 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Baetis (Tenuibaetis) fujitanii Kaltenbach &amp; Gattolliat 2019" docType="treatment" docVersion="2" lastPageNumber="228" masterDocId="FFCDFFA8A3764D4EA24CFFD4FFABE864" masterDocTitle="Contribution to the knowledge of the subgenus Tenuibaetis Kang &amp; Yang 1994 (Ephemeroptera, Baetidae, Baetis s. l.)" masterLastPageNumber="258" masterPageNumber="201" pageNumber="227" updateTime="1683100342607" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Contribution to the knowledge of the subgenus Tenuibaetis Kang &amp; Yang 1994 (Ephemeroptera, Baetidae, Baetis s. l.)</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kluge, Nikita</mods:namePart>
<mods:affiliation>Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia. Website: http: // insecta. bio. spbu. ru</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Srinivasan, Pandiarajan</mods:namePart>
<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sivaruban, T.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8997-9355</mods:nameIdentifier>
<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India &amp; sivaruban 270 @ gmail. com; https: // orcid. org / 0000 - 0001 - 8997 - 9355</mods:affiliation>
<mods:nameIdentifier type="email">sivaruban270@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Barathy, S.</mods:namePart>
<mods:affiliation>Department of Zoology, Fatima College, Madurai- 625018, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Isack, Rajasekaran</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-9952-4335</mods:nameIdentifier>
<mods:affiliation>PG and Research department of Zoology, the American College, Madurai- 625002, India &amp; iceisack 143 @ gmail. com; https: // orcid. org / 0000 - 0002 - 9952 - 4335</mods:affiliation>
<mods:nameIdentifier type="email">iceisack143@gmail.com</mods:nameIdentifier>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-02</mods:number>
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<mods:number>5277</mods:number>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5277.2.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7889695</mods:identifier>
<mods:identifier type="ZooBank">73ED69F3-3887-460D-86DE-C9F1302C8EC3</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7889709" ID-Zenodo-Dep="7889709" LSID="urn:lsid:plazi:treatment:03F487D0A36C4D55A2DBFF43FC92E925" httpUri="http://treatment.plazi.org/id/03F487D0A36C4D55A2DBFF43FC92E925" lastPageId="27" lastPageNumber="228" pageId="26" pageNumber="227">
<subSubSection box="[151,905,151,178]" pageId="26" pageNumber="227" type="nomenclature">
<paragraph blockId="26.[151,905,151,214]" box="[151,905,151,178]" pageId="26" pageNumber="227">
<heading bold="true" box="[151,905,151,178]" fontSize="11" level="1" pageId="26" pageNumber="227" reason="1">
<emphasis bold="true" box="[151,905,151,178]" pageId="26" pageNumber="227">
4.
<taxonomicName authority="Kaltenbach &amp; Gattolliat 2019" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2019" box="[181,905,151,178]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="fujitanii" subGenus="Tenuibaetis">
<emphasis bold="true" box="[181,256,151,177]" italics="true" pageId="26" pageNumber="227">Baetis</emphasis>
(
<emphasis bold="true" box="[273,412,151,178]" italics="true" pageId="26" pageNumber="227">Tenuibaetis</emphasis>
)
<emphasis bold="true" box="[429,527,151,178]" italics="true" pageId="26" pageNumber="227">fujitanii</emphasis>
<bibRefCitation author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[534,905,151,177]" pageId="26" pageNumber="227" pagination="13 - 23" refId="ref17903" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2019) A new species of Tenuibaetis Kang &amp; Yang, 1994 from Indonesia (Ephemeroptera, Baetidae). ZooKeys, 820, 13 - 23. https: // doi. org / 10.3897 / zookeys. 820.31487" type="journal article" year="2019">Kaltenbach &amp; Gattolliat 2019</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,323,187,214]" pageId="26" pageNumber="227" type="description">
<paragraph blockId="26.[151,905,151,214]" box="[151,323,187,214]" pageId="26" pageNumber="227">
(
<figureCitation box="[159,315,187,214]" captionStart-0="FIGURES 125128" captionStart-1="FIGURES 129134" captionStart-2="FIGURES 135143" captionStart-3="FIGURES 144145" captionStartId-0="27.[151,264,1955,1979]" captionStartId-1="28.[151,264,1712,1736]" captionStartId-2="29.[151,264,1766,1790]" captionStartId-3="30.[151,264,1131,1155]" captionTargetBox-0="[151,1436,345,1928]" captionTargetBox-1="[151,1436,185,1688]" captionTargetBox-2="[151,1435,181,1742]" captionTargetBox-3="[151,1436,181,1107]" captionTargetId-0="figure-113@27.[151,1436,345,1932]" captionTargetId-1="figure-36@28.[151,1436,181,1688]" captionTargetId-2="figure-19@29.[151,1435,181,1742]" captionTargetId-3="figure-36@30.[151,1436,181,1107]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionTargetPageId-2="29" captionTargetPageId-3="30" captionText-0="FIGURES 125128. Baetis (Tenuibaetis) fujitanii. 125127, larval exuviae: 125, head; 126, half of pronotum and mesonotum; 127, abdomen; 128, young larva in natural pose." captionText-1="FIGURES 129134. Baetis (Tenuibaetis) fujitanii, exuviae of larval legs. 129131, fore, middle and hind legs; 132, anterior surface of hind femur; 133134, middle tibia." captionText-2="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." captionText-3="FIGURES 144145. Baetis (Tenuibaetis) fujitanii, genitalia of male imagines." figureDoi-0="http://doi.org/10.5281/zenodo.7889761" figureDoi-1="http://doi.org/10.5281/zenodo.7889763" figureDoi-2="http://doi.org/10.5281/zenodo.7889765" figureDoi-3="http://doi.org/10.5281/zenodo.7889767" httpUri-0="https://zenodo.org/record/7889761/files/figure.png" httpUri-1="https://zenodo.org/record/7889763/files/figure.png" httpUri-2="https://zenodo.org/record/7889765/files/figure.png" httpUri-3="https://zenodo.org/record/7889767/files/figure.png" pageId="26" pageNumber="227">Figs 125145</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,841,256,280]" pageId="26" pageNumber="227" type="reference_group">
<paragraph blockId="26.[151,841,256,280]" box="[151,841,256,280]" pageId="26" pageNumber="227">
<taxonomicName authority="Kaltenbach &amp; Gattolliat, 2019" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2019" box="[151,758,256,280]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="fujitanii" subGenus="Tenuibaetis">
<emphasis box="[151,214,256,279]" italics="true" pageId="26" pageNumber="227">Baetis</emphasis>
(
<emphasis box="[229,345,256,280]" italics="true" pageId="26" pageNumber="227">Tenuibaetis</emphasis>
)
<emphasis box="[360,440,256,280]" italics="true" pageId="26" pageNumber="227">fujitanii</emphasis>
<bibRefCitation author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[446,758,256,280]" pageId="26" pageNumber="227" pagination="13 - 23" refId="ref17903" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2019) A new species of Tenuibaetis Kang &amp; Yang, 1994 from Indonesia (Ephemeroptera, Baetidae). ZooKeys, 820, 13 - 23. https: // doi. org / 10.3897 / zookeys. 820.31487" type="journal article" year="2019">Kaltenbach &amp; Gattolliat, 2019</bibRefCitation>
</taxonomicName>
: (larva).
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="227" type="materials_examined">
<paragraph blockId="26.[151,1437,325,2008]" pageId="26" pageNumber="227">
<emphasis bold="true" box="[151,380,325,351]" pageId="26" pageNumber="227">Material examined</emphasis>
(
<collectionCode box="[399,451,326,352]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="26" pageNumber="227" type="Museum">ZIN</collectionCode>
).
<materialsCitation collectingDate="2012-08-31" collectingDateMax="2012-09-02" collectingDateMin="2012-08-31" collectorName="N. Kluge &amp; L. Sheyko" country="Indonesia" location="river Liwung" municipality="Bogor" pageId="26" pageNumber="227" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Timur">
<collectingCountry box="[473,625,326,352]" name="Indonesia" pageId="26" pageNumber="227">INDONESIA</collectingCountry>
,
<collectingRegion box="[642,706,326,352]" country="Indonesia" name="Jawa Timur" pageId="26" pageNumber="227">JAVA</collectingRegion>
:
<collectingMunicipality box="[725,797,326,352]" pageId="26" pageNumber="227">Bogor</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03F487D0A36C4D55A2DBFF43FC92E925:8E82601DA36C4D54A167FE92FC68E904" box="[811,963,326,352]" country="Indonesia" municipality="Bogor" name="river Liwung" pageId="26" pageNumber="227" stateProvince="Jawa Timur">river Liwung</location>
,
<date box="[978,1191,326,352]" pageId="26" pageNumber="227" value="2012-08-31" valueMax="2012-09-02" valueMin="2012-08-31">
<collectingDate box="[978,1191,326,352]" pageId="26" pageNumber="227" value="2012-08-31" valueMax="2012-09-02" valueMin="2012-08-31">31.VIII2.IX.2012</collectingDate>
</date>
, coll.
<collectorName box="[1264,1370,326,352]" pageId="26" pageNumber="227">N. Kluge</collectorName>
&amp;
<collectorName pageId="26" pageNumber="227">L. Sheyko</collectorName>
: 2 L-S-I♁, 1 L-S-I
<specimenCount box="[457,471,362,388]" pageId="26" pageNumber="227" type="female"></specimenCount>
</materialsCitation>
, 1 L-S
<specimenCount box="[554,569,362,388]" pageId="26" pageNumber="227" type="female"></specimenCount>
,
<specimenCount box="[581,702,362,388]" pageId="26" pageNumber="227" type="larva">247 larvae</specimenCount>
;
<materialsCitation box="[713,1345,362,388]" collectingDate="2012-09-02" collectorName="N. Kluge" country="Indonesia" location="river Sadane" municipality="Bogor" pageId="26" pageNumber="227" specimenCount="4" specimenCount-larva="4" stateProvince="Jawa Timur">
<collectingMunicipality box="[713,785,362,388]" pageId="26" pageNumber="227">Bogor</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03F487D0A36C4D55A2DBFF43FC92E925:8E82601DA36C4D54A150FEBEFC07E9E0" box="[796,940,362,388]" country="Indonesia" municipality="Bogor" name="river Sadane" pageId="26" pageNumber="227" stateProvince="Jawa Timur">river Sadane</location>
,
<date box="[951,1067,362,388]" pageId="26" pageNumber="227" value="2012-09-02">
<collectingDate box="[951,1067,362,388]" pageId="26" pageNumber="227" value="2012-09-02">2.IX.2012</collectingDate>
</date>
, coll.
<collectorName box="[1135,1242,362,388]" pageId="26" pageNumber="227">N. Kluge</collectorName>
:
<specimenCount box="[1252,1345,362,388]" pageId="26" pageNumber="227" type="larva">4 larvae</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" box="[199,479,397,423]" pageId="26" pageNumber="227">
<emphasis bold="true" box="[199,479,397,423]" pageId="26" pageNumber="227">Additional descriptions</emphasis>
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" pageId="26" pageNumber="227">
<emphasis bold="true" box="[199,275,434,460]" pageId="26" pageNumber="227">
<emphasis bold="true" box="[199,267,434,460]" italics="true" pageId="26" pageNumber="227">Larva</emphasis>
.
</emphasis>
CUTICULAR COLORATION: Head brown (
<figureCitation box="[817,916,434,461]" captionStart="FIGURES 125128" captionStartId="27.[151,264,1955,1979]" captionTargetBox="[151,1436,345,1928]" captionTargetId="figure-113@27.[151,1436,345,1932]" captionTargetPageId="27" captionText="FIGURES 125128. Baetis (Tenuibaetis) fujitanii. 125127, larval exuviae: 125, head; 126, half of pronotum and mesonotum; 127, abdomen; 128, young larva in natural pose." figureDoi="http://doi.org/10.5281/zenodo.7889761" httpUri="https://zenodo.org/record/7889761/files/figure.png" pageId="26" pageNumber="227">Fig. 125</figureCitation>
). Pronotum brown to brown with blanks, with sigilla lighter than background; mesonotum and fore protoptera brown, with more or less contrasting blanks (
<figureCitation captionStart="FIGURES 125128" captionStartId="27.[151,264,1955,1979]" captionTargetBox="[151,1436,345,1928]" captionTargetId="figure-113@27.[151,1436,345,1932]" captionTargetPageId="27" captionText="FIGURES 125128. Baetis (Tenuibaetis) fujitanii. 125127, larval exuviae: 125, head; 126, half of pronotum and mesonotum; 127, abdomen; 128, young larva in natural pose." figureDoi="http://doi.org/10.5281/zenodo.7889761" httpUri="https://zenodo.org/record/7889761/files/figure.png" pageId="26" pageNumber="227">Fig. 126</figureCitation>
). Thoracic pleura brownish, sterna colorless. Legs with brown and colorless areas (
<figureCitation box="[1176,1339,506,532]" captionStart="FIGURES 129134" captionStartId="28.[151,264,1712,1736]" captionTargetBox="[151,1436,185,1688]" captionTargetId="figure-36@28.[151,1436,181,1688]" captionTargetPageId="28" captionText="FIGURES 129134. Baetis (Tenuibaetis) fujitanii, exuviae of larval legs. 129131, fore, middle and hind legs; 132, anterior surface of hind femur; 133134, middle tibia." figureDoi="http://doi.org/10.5281/zenodo.7889763" httpUri="https://zenodo.org/record/7889763/files/figure.png" pageId="26" pageNumber="227">Figs 129131</figureCitation>
); if fore femur with expressed proximal blank, shape of proximal blank subtriangular or comma-like, with inner-distal angle stretched distally (
<figureCitation box="[429,514,578,604]" captionStart="FIGURES 2230" captionStartId="8.[151,264,1852,1876]" captionTargetBox="[151,1436,181,1827]" captionTargetId="figure-36@8.[151,1436,181,1828]" captionTargetPageId="8" captionText="FIGURES 2230. Baetis (Tenuibaetis) ursinus, larval exuviae (2429, one and the same magnification). 22, labrum; 23, maxilla and labium; 24, abdomen, 2527, middle and hind legs; 28, half of pronotum and mesonotum; 29, head; 30, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7889707" httpUri="https://zenodo.org/record/7889707/files/figure.png" pageId="26" pageNumber="227">Fig. 29</figureCitation>
). Abdominal terga nearly uniformly brown; tergum
<collectionCode box="[1105,1135,578,604]" pageId="26" pageNumber="227">IV</collectionCode>
lighter laterally (
<figureCitation box="[1333,1431,578,604]" captionStart="FIGURES 125128" captionStartId="27.[151,264,1955,1979]" captionTargetBox="[151,1436,345,1928]" captionTargetId="figure-113@27.[151,1436,345,1932]" captionTargetPageId="27" captionText="FIGURES 125128. Baetis (Tenuibaetis) fujitanii. 125127, larval exuviae: 125, head; 126, half of pronotum and mesonotum; 127, abdomen; 128, young larva in natural pose." figureDoi="http://doi.org/10.5281/zenodo.7889761" httpUri="https://zenodo.org/record/7889761/files/figure.png" pageId="26" pageNumber="227">Fig. 127</figureCitation>
;
<bibRefCitation author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[151,487,614,640]" pageId="26" pageNumber="227" pagination="13 - 23" refId="ref17903" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2019) A new species of Tenuibaetis Kang &amp; Yang, 1994 from Indonesia (Ephemeroptera, Baetidae). ZooKeys, 820, 13 - 23. https: // doi. org / 10.3897 / zookeys. 820.31487" type="journal article" year="2019">Kaltenbach &amp; Gattolliat 2019</bibRefCitation>
: fig. 1a). Caudalii brownish.
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" pageId="26" pageNumber="227">
HYPODERMAL COLORATION: Either not expressed, or abdominal terga with dark brown line on posterior margin and other markings as in imago (as in
<figureCitation box="[665,821,686,713]" captionStart="FIGURES 135143" captionStartId="29.[151,264,1766,1790]" captionTargetBox="[151,1435,181,1742]" captionTargetId="figure-19@29.[151,1435,181,1742]" captionTargetPageId="29" captionText="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." figureDoi="http://doi.org/10.5281/zenodo.7889765" httpUri="https://zenodo.org/record/7889765/files/figure.png" pageId="26" pageNumber="227">Figs 135, 141</figureCitation>
).
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" box="[200,1437,722,749]" pageId="26" pageNumber="227">
SHAPEAND
<collectionCode box="[358,496,722,748]" pageId="26" pageNumber="227">SETATION</collectionCode>
:Mouthpartsasinoriginaldescription(
<figureCitation box="[906,997,722,749]" captionStart="FIGURES 125128" captionStartId="27.[151,264,1955,1979]" captionTargetBox="[151,1436,345,1928]" captionTargetId="figure-113@27.[151,1436,345,1932]" captionTargetPageId="27" captionText="FIGURES 125128. Baetis (Tenuibaetis) fujitanii. 125127, larval exuviae: 125, head; 126, half of pronotum and mesonotum; 127, abdomen; 128, young larva in natural pose." figureDoi="http://doi.org/10.5281/zenodo.7889761" httpUri="https://zenodo.org/record/7889761/files/figure.png" pageId="26" pageNumber="227">Fig.125</figureCitation>
;
<bibRefCitation author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[1003,1321,722,748]" pageId="26" pageNumber="227" pagination="13 - 23" refId="ref17903" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2019) A new species of Tenuibaetis Kang &amp; Yang, 1994 from Indonesia (Ephemeroptera, Baetidae). ZooKeys, 820, 13 - 23. https: // doi. org / 10.3897 / zookeys. 820.31487" type="journal article" year="2019">Kaltenbach&amp;Gattolliat2019</bibRefCitation>
:fig.2ah).
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" box="[199,918,758,784]" pageId="26" pageNumber="227">
Hind protoptera present (
<bibRefCitation author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[481,818,758,784]" pageId="26" pageNumber="227" pagination="13 - 23" refId="ref17903" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2019) A new species of Tenuibaetis Kang &amp; Yang, 1994 from Indonesia (Ephemeroptera, Baetidae). ZooKeys, 820, 13 - 23. https: // doi. org / 10.3897 / zookeys. 820.31487" type="journal article" year="2019">Kaltenbach &amp; Gattolliat 2019</bibRefCitation>
: fig. 3d).
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" pageId="26" pageNumber="227">
Stout two-channel setae forming row along outer side of femur (including two subapical setae) elongate, blunt, distally widened and flattened (
<figureCitation box="[503,662,830,856]" captionStart="FIGURES 129134" captionStartId="28.[151,264,1712,1736]" captionTargetBox="[151,1436,185,1688]" captionTargetId="figure-36@28.[151,1436,181,1688]" captionTargetPageId="28" captionText="FIGURES 129134. Baetis (Tenuibaetis) fujitanii, exuviae of larval legs. 129131, fore, middle and hind legs; 132, anterior surface of hind femur; 133134, middle tibia." figureDoi="http://doi.org/10.5281/zenodo.7889763" httpUri="https://zenodo.org/record/7889763/files/figure.png" pageId="26" pageNumber="227">Figs 129131</figureCitation>
). Anterior-apical margin of femur with small, pseudo-bifurcate setae, as on tibiae. Anterior side of femur with colorless oval scales in semicircular sockets (
<figureCitation box="[1128,1224,866,892]" captionStart="FIGURES 129134" captionStartId="28.[151,264,1712,1736]" captionTargetBox="[151,1436,185,1688]" captionTargetId="figure-36@28.[151,1436,181,1688]" captionTargetPageId="28" captionText="FIGURES 129134. Baetis (Tenuibaetis) fujitanii, exuviae of larval legs. 129131, fore, middle and hind legs; 132, anterior surface of hind femur; 133134, middle tibia." figureDoi="http://doi.org/10.5281/zenodo.7889763" httpUri="https://zenodo.org/record/7889763/files/figure.png" pageId="26" pageNumber="227">Fig. 132</figureCitation>
) (as on abdomen). Stout setae on anterior side of femur two-channel, blunt or clavate, mostly short; on each leg, one or few most distal setae long and clavate (
<figureCitation box="[412,571,938,964]" captionStart="FIGURES 129134" captionStartId="28.[151,264,1712,1736]" captionTargetBox="[151,1436,185,1688]" captionTargetId="figure-36@28.[151,1436,181,1688]" captionTargetPageId="28" captionText="FIGURES 129134. Baetis (Tenuibaetis) fujitanii, exuviae of larval legs. 129131, fore, middle and hind legs; 132, anterior surface of hind femur; 133134, middle tibia." figureDoi="http://doi.org/10.5281/zenodo.7889763" httpUri="https://zenodo.org/record/7889763/files/figure.png" pageId="26" pageNumber="227">Figs 129132</figureCitation>
;
<bibRefCitation author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[582,918,938,964]" pageId="26" pageNumber="227" pagination="13 - 23" refId="ref17903" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2019) A new species of Tenuibaetis Kang &amp; Yang, 1994 from Indonesia (Ephemeroptera, Baetidae). ZooKeys, 820, 13 - 23. https: // doi. org / 10.3897 / zookeys. 820.31487" type="journal article" year="2019">Kaltenbach &amp; Gattolliat 2019</bibRefCitation>
: fig. 3a).
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" pageId="26" pageNumber="227">
Tibia of each leg with small, pseudo-bifurcate setae on outer side; stout setae along patella-tibial suture small, two-channel (
<figureCitation box="[304,464,1010,1036]" captionStart="FIGURES 129134" captionStartId="28.[151,264,1712,1736]" captionTargetBox="[151,1436,185,1688]" captionTargetId="figure-36@28.[151,1436,181,1688]" captionTargetPageId="28" captionText="FIGURES 129134. Baetis (Tenuibaetis) fujitanii, exuviae of larval legs. 129131, fore, middle and hind legs; 132, anterior surface of hind femur; 133134, middle tibia." figureDoi="http://doi.org/10.5281/zenodo.7889763" httpUri="https://zenodo.org/record/7889763/files/figure.png" pageId="26" pageNumber="227">Figs 133134</figureCitation>
).
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" pageId="26" pageNumber="227">
Denticles on posterior margins of all terga IX blunt (
<bibRefCitation author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[795,1128,1046,1072]" pageId="26" pageNumber="227" pagination="13 - 23" refId="ref17903" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2019) A new species of Tenuibaetis Kang &amp; Yang, 1994 from Indonesia (Ephemeroptera, Baetidae). ZooKeys, 820, 13 - 23. https: // doi. org / 10.3897 / zookeys. 820.31487" type="journal article" year="2019">Kaltenbach &amp; Gattolliat 2019</bibRefCitation>
: fig. 3c), shorter on anterior terga and longer on posterior terga. Denticles on posterior margin of sternum triangular, present only on sternum
<collectionCode box="[1400,1432,1082,1108]" pageId="26" pageNumber="227">IX</collectionCode>
, sometimes also on sternum
<collectionCode box="[464,516,1118,1144]" pageId="26" pageNumber="227">VIII</collectionCode>
; in male, row of denticles not interrupted by protogonostyli. Paraproct with small denticles (
<bibRefCitation author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[270,613,1154,1180]" pageId="26" pageNumber="227" pagination="13 - 23" refId="ref17903" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2019) A new species of Tenuibaetis Kang &amp; Yang, 1994 from Indonesia (Ephemeroptera, Baetidae). ZooKeys, 820, 13 - 23. https: // doi. org / 10.3897 / zookeys. 820.31487" type="journal article" year="2019">Kaltenbach &amp; Gattolliat 2019</bibRefCitation>
: figs
<quantity box="[678,706,1154,1180]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="26" pageNumber="227" unit="g" value="3.0">3g</quantity>
h). Tergalii with oval scales on semicircular sockets. Tergalius I much smaller than others (
<bibRefCitation author="Kaltenbach, T. &amp; Gattolliat, J. - L." box="[475,817,1190,1216]" pageId="26" pageNumber="227" pagination="13 - 23" refId="ref17903" refString="Kaltenbach, T. &amp; Gattolliat, J. - L. (2019) A new species of Tenuibaetis Kang &amp; Yang, 1994 from Indonesia (Ephemeroptera, Baetidae). ZooKeys, 820, 13 - 23. https: // doi. org / 10.3897 / zookeys. 820.31487" type="journal article" year="2019">Kaltenbach &amp; Gattolliat 2019</bibRefCitation>
: figs 3ef). Paracercus significantly smaller than cerci (ibid., fig. 1).
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" pageId="26" pageNumber="227">
<emphasis bold="true" box="[199,323,1261,1288]" pageId="26" pageNumber="227">
<emphasis bold="true" box="[199,316,1261,1288]" italics="true" pageId="26" pageNumber="227">Subimago</emphasis>
.
</emphasis>
CUTICULAR COLORATION: Head colorless, antennae brown. Pronotum colorless. Mesonotum in median part mostly colorless and ochre, laterally with lighter and darker brown areas (
<figureCitation box="[1143,1241,1298,1324]" captionStart="FIGURES 135143" captionStartId="29.[151,264,1766,1790]" captionTargetBox="[151,1435,181,1742]" captionTargetId="figure-19@29.[151,1435,181,1742]" captionTargetPageId="29" captionText="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." figureDoi="http://doi.org/10.5281/zenodo.7889765" httpUri="https://zenodo.org/record/7889765/files/figure.png" pageId="26" pageNumber="227">Fig. 137</figureCitation>
). Thoracic pleura and sterna with light brownish and dark brown areas (as in
<figureCitation box="[843,943,1334,1361]" captionStart="FIGURES 215224" captionStartId="43.[151,264,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-19@43.[151,1435,181,1746]" captionTargetPageId="43" captionText="FIGURES 215224. Baetis (Tenuibaetis) bialatus sp. n. 215, subimaginal exuviae of mesopleuron; 216, subimaginal exuviae of half of mesonotum; 217218, exuviae of fore and hind legs of male subimago; 219220, fore and middle legs of male imago; 221, male imago; 222, female imago; 223, subimaginal exuviae of abdominal segments VIIIIX; 224, egg (215220, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889793" httpUri="https://zenodo.org/record/7889793/files/figure.png" pageId="26" pageNumber="227">Fig. 215</figureCitation>
). Legs with femora lighter, tibiae and tarsi darker (
<figureCitation box="[241,343,1370,1396]" captionStart="FIGURES 135143" captionStartId="29.[151,264,1766,1790]" captionTargetBox="[151,1435,181,1742]" captionTargetId="figure-19@29.[151,1435,181,1742]" captionTargetPageId="29" captionText="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." figureDoi="http://doi.org/10.5281/zenodo.7889765" httpUri="https://zenodo.org/record/7889765/files/figure.png" pageId="26" pageNumber="227">Fig. 140</figureCitation>
). Abdominal terga I
<collectionCode box="[589,637,1370,1396]" pageId="26" pageNumber="227">VIII</collectionCode>
medially nearly colorless, laterally diffusively darkened; terga
<collectionCode box="[1372,1401,1370,1396]" pageId="26" pageNumber="227">IX</collectionCode>
X entirely light (as in
<figureCitation box="[371,468,1406,1432]" captionStart="FIGURES 169180" captionStartId="34.[151,264,1880,1904]" captionTargetBox="[151,1436,181,1856]" captionTargetId="figure-36@34.[151,1436,181,1856]" captionTargetPageId="34" captionText="FIGURES 169180. Baetis (Tenuibaetis) kaltenbachi sp. n. 169170, male imago; 171, female imago; 172, subimaginal exuviae of abdomen; 173174, fore and middle legs of male imago; 175176, their subimaginal exuviae; 177, subimaginal exuviae of half of mesonotum; 178, hind wing of male imago; 179, the same, female; 180, blunt microlepides on fore tarsus of male subimago (169170, 172178, 180, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7889777" httpUri="https://zenodo.org/record/7889777/files/figure.png" pageId="26" pageNumber="227">Fig. 172</figureCitation>
).
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" box="[200,756,1442,1468]" pageId="26" pageNumber="227">HYPODERMAL COLORATION: As in imago.</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" pageId="26" pageNumber="227">
<emphasis bold="true" box="[199,345,1477,1504]" italics="true" pageId="26" pageNumber="227">Imago, male</emphasis>
(
<figureCitation box="[361,461,1478,1505]" captionStart="FIGURES 135143" captionStartId="29.[151,264,1766,1790]" captionTargetBox="[151,1435,181,1742]" captionTargetId="figure-19@29.[151,1435,181,1742]" captionTargetPageId="29" captionText="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." figureDoi="http://doi.org/10.5281/zenodo.7889765" httpUri="https://zenodo.org/record/7889765/files/figure.png" pageId="26" pageNumber="227">Fig. 135</figureCitation>
). Head ochre. Antennae ochre. Turbinate eyes orange. Thorax ochre with light brown. Forewing with membrane entirely colorless; veins from ochre (nearer to wing base) to brown; pterostigma with few oblique cross veins and disconnected vein fragments. Hind wing full-sized, colorless, with short and non-hooked costal projection, with 2 longitudinal veins (
<figureCitation box="[645,743,1586,1612]" captionStart="FIGURES 135143" captionStartId="29.[151,264,1766,1790]" captionTargetBox="[151,1435,181,1742]" captionTargetId="figure-19@29.[151,1435,181,1742]" captionTargetPageId="29" captionText="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." figureDoi="http://doi.org/10.5281/zenodo.7889765" httpUri="https://zenodo.org/record/7889765/files/figure.png" pageId="26" pageNumber="227">Fig. 138</figureCitation>
). Legs uniformly ochre (
<figureCitation box="[1019,1178,1586,1612]" captionStart="FIGURES 135143" captionStartId="29.[151,264,1766,1790]" captionTargetBox="[151,1435,181,1742]" captionTargetId="figure-19@29.[151,1435,181,1742]" captionTargetPageId="29" captionText="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." figureDoi="http://doi.org/10.5281/zenodo.7889765" httpUri="https://zenodo.org/record/7889765/files/figure.png" pageId="26" pageNumber="227">Figs 142143</figureCitation>
). Middle and hind tarsi with 2 apical spines (on 1st+2nd and 3rd segments). Abdominal terga and sterna ochre with brownish or reddish, with lateral areas darker and median areas lighter; terga
<collectionCode box="[782,811,1658,1684]" pageId="26" pageNumber="227">IX</collectionCode>
X lighter than others; terga I
<collectionCode box="[1159,1189,1658,1684]" pageId="26" pageNumber="227">IX</collectionCode>
or some of them with contrastingly dark brown band bordering posterior margin and not reaching lateral margins (
<figureCitation box="[1195,1354,1694,1721]" captionStart="FIGURES 135143" captionStartId="29.[151,264,1766,1790]" captionTargetBox="[151,1435,181,1742]" captionTargetId="figure-19@29.[151,1435,181,1742]" captionTargetPageId="29" captionText="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." figureDoi="http://doi.org/10.5281/zenodo.7889765" httpUri="https://zenodo.org/record/7889765/files/figure.png" pageId="26" pageNumber="227">Figs 135, 141</figureCitation>
). Cerci ochre. Gonostyli ochre; 1st segment relatively narrow, only slightly narrowing toward apex (nearly parallel-sided); 3rd (terminal) segment short and petiolate (
<figureCitation box="[627,785,1766,1793]" captionStart="FIGURES 144145" captionStartId="30.[151,264,1131,1155]" captionTargetBox="[151,1436,181,1107]" captionTargetId="figure-36@30.[151,1436,181,1107]" captionTargetPageId="30" captionText="FIGURES 144145. Baetis (Tenuibaetis) fujitanii, genitalia of male imagines." figureDoi="http://doi.org/10.5281/zenodo.7889767" httpUri="https://zenodo.org/record/7889767/files/figure.png" pageId="26" pageNumber="227">Figs 144145</figureCitation>
). Gonovectes gently bent, light brownish. Sterno-styligeral muscle completely absent.
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" pageId="26" pageNumber="227">
<emphasis bold="true" box="[199,368,1837,1864]" italics="true" pageId="26" pageNumber="227">Imago, female</emphasis>
(
<figureCitation box="[386,487,1838,1864]" captionStart="FIGURES 135143" captionStartId="29.[151,264,1766,1790]" captionTargetBox="[151,1435,181,1742]" captionTargetId="figure-19@29.[151,1435,181,1742]" captionTargetPageId="29" captionText="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." figureDoi="http://doi.org/10.5281/zenodo.7889765" httpUri="https://zenodo.org/record/7889765/files/figure.png" pageId="26" pageNumber="227">Fig. 136</figureCitation>
). Coloration similar to that of male. Hind wing slightly smaller than in male (
<figureCitation captionStart="FIGURES 135143" captionStartId="29.[151,264,1766,1790]" captionTargetBox="[151,1435,181,1742]" captionTargetId="figure-19@29.[151,1435,181,1742]" captionTargetPageId="29" captionText="FIGURES 135143. Baetis (Tenuibaetis) fujitanii. 135, male imago; 136, female imago; 137, subimaginal exuviae of half of mesonotum; 138, male hind wing; 139, female hind wing; 140, exuviae of male fore leg; 141, abdomen of male imago; 142143, middle and fore legs of male imago." figureDoi="http://doi.org/10.5281/zenodo.7889765" httpUri="https://zenodo.org/record/7889765/files/figure.png" pageId="26" pageNumber="227">Fig. 139</figureCitation>
). All tarsi with 2 apical spines (fore tarsus on 2nd and 3rd segments, middle and hind tarsi on 1st+2nd and 3rd segments).
</paragraph>
<paragraph blockId="26.[151,1437,325,2008]" box="[199,604,1945,1972]" pageId="26" pageNumber="227">
<emphasis bold="true" box="[199,334,1945,1971]" pageId="26" pageNumber="227">Dimension.</emphasis>
Forewing length
<quantity box="[532,600,1946,1972]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="227" unit="mm" value="4.0">4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="27" lastPageNumber="228" pageId="26" pageNumber="227" type="distribution">
<paragraph blockId="26.[151,1437,325,2008]" box="[199,417,1981,2008]" pageId="26" pageNumber="227">
<emphasis bold="true" box="[199,353,1981,2007]" pageId="26" pageNumber="227">Distribution.</emphasis>
<collectingRegion box="[359,412,1982,2008]" country="Indonesia" name="Jawa Timur" pageId="26" pageNumber="227">Java</collectingRegion>
.
</paragraph>
<paragraph blockId="27.[151,1437,150,321]" pageId="27" pageNumber="228">
<emphasis bold="true" box="[199,354,150,176]" pageId="27" pageNumber="228">Comparison.</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2019" box="[363,534,151,177]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="fujitanii" subGenus="Tenuibaetis">
<emphasis box="[363,388,151,176]" italics="true" pageId="27" pageNumber="228">B.</emphasis>
(
<emphasis box="[406,427,151,176]" italics="true" pageId="27" pageNumber="228">T.</emphasis>
)
<emphasis box="[445,534,151,177]" italics="true" pageId="27" pageNumber="228">fujitanii</emphasis>
</taxonomicName>
is similar to
<taxonomicName authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[689,887,151,177]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="frequentus" subGenus="Tenuibaetis">
<emphasis box="[689,713,151,176]" italics="true" pageId="27" pageNumber="228">B.</emphasis>
(
<emphasis box="[731,752,151,176]" italics="true" pageId="27" pageNumber="228">T.</emphasis>
)
<emphasis box="[770,887,151,177]" italics="true" pageId="27" pageNumber="228">frequentus</emphasis>
</taxonomicName>
, and there are no characters allowing to clearly separate these species. Larvae of
<taxonomicName authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2019" box="[515,678,187,213]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="fujitanii" subGenus="Tenuibaetis">
<emphasis box="[515,540,187,212]" italics="true" pageId="27" pageNumber="228">B.</emphasis>
(
<emphasis box="[554,575,187,212]" italics="true" pageId="27" pageNumber="228">T.</emphasis>
)
<emphasis box="[589,678,187,213]" italics="true" pageId="27" pageNumber="228">fujitanii</emphasis>
</taxonomicName>
from Bogor (the
<typeStatus box="[866,914,187,213]" pageId="27" pageNumber="228">type</typeStatus>
locality) have rather uniform dorsal coloration, similar in all individuals, while larvae of Ceylonese populations of
<taxonomicName authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[940,1141,223,249]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="frequentus" subGenus="Tenuibaetis">
<emphasis box="[940,964,223,248]" italics="true" pageId="27" pageNumber="228">B.</emphasis>
(
<emphasis box="[984,1005,223,248]" italics="true" pageId="27" pageNumber="228">T.</emphasis>
)
<emphasis box="[1024,1141,223,249]" italics="true" pageId="27" pageNumber="228">frequentus</emphasis>
</taxonomicName>
have variable coloration. Probably, species-specific character of
<taxonomicName authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2019" box="[594,762,259,285]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="fujitanii" subGenus="Tenuibaetis">
<emphasis box="[594,618,259,284]" italics="true" pageId="27" pageNumber="228">B.</emphasis>
(
<emphasis box="[635,657,259,284]" italics="true" pageId="27" pageNumber="228">T.</emphasis>
)
<emphasis box="[673,762,259,285]" italics="true" pageId="27" pageNumber="228">fujitanii</emphasis>
</taxonomicName>
is the fact that the most distal two-channel seta on anterior surface of each femur is much longer than others (
<figureCitation box="[714,812,295,321]" captionStart="FIGURES 129134" captionStartId="28.[151,264,1712,1736]" captionTargetBox="[151,1436,185,1688]" captionTargetId="figure-36@28.[151,1436,181,1688]" captionTargetPageId="28" captionText="FIGURES 129134. Baetis (Tenuibaetis) fujitanii, exuviae of larval legs. 129131, fore, middle and hind legs; 132, anterior surface of hind femur; 133134, middle tibia." figureDoi="http://doi.org/10.5281/zenodo.7889763" httpUri="https://zenodo.org/record/7889763/files/figure.png" pageId="27" pageNumber="228">Fig. 132</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>