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<document id="F0A3FFA2BEB38C70CA15420329785AD5" ID-CLB-Dataset="3948" ID-DOI="10.5852/ejt.2021.775.1551" ID-GBIF-Dataset="cc10c087-bec3-4947-81d7-b3e5580a5c94" ID-ISSN="2118-9773" ID-Zenodo-Dep="5589689" ID-ZooBank="976D7020-5904-4951-97CE-B4FE58DA12A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634822039012" checkinUser="felipe" docAuthor="González-Alvarado, Arturo &amp; Vaz-De, Fernando Z." docDate="2021" docId="03F48795DC50FFBCA614A6C2FB8BFDF7" docLanguage="en" docName="ejt-775_86-106.pdf" docOrigin="European Journal of Taxonomy 775" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Deltochilum nonstriatum González-Alvarado &amp; Vaz-De 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="101" masterDocId="FFCDFFEDDC5DFFB3A45AA753FFCDFFC7" masterDocTitle="Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum)" masterLastPageNumber="106" masterPageNumber="86" pageNumber="99" updateTime="1699464201688" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="4B946C240F154845F26B0BCB74003C3E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BC1BF336C7BB851DDC9721579D94371E">
<mods:title id="715F8CCF82F29A3BB5C17F26CE0D05D9">Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum)</mods:title>
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<mods:name id="29184BF58DE82B00CB34703127F4CC76" type="personal">
<mods:role id="DC08DA0DC175120BDC2E66BDAEABC9A0">
<mods:roleTerm id="3A4CC276A838FFAE9FD755711C65D1D3">Author</mods:roleTerm>
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<mods:namePart id="0CB6E7973E235B5766C3194E63333518">González-Alvarado, Arturo</mods:namePart>
<mods:nameIdentifier id="36BCC3AD45BD82D189A1229C088CAEC2" type="LSID">2D58A702-73A9-4C66-8CED-6B8B9A2AB10B</mods:nameIdentifier>
<mods:affiliation id="42BE4904A7BF359928F48FFF4704A60A">Programa de Pós-graduação em Ecologia e Conservação da Biodiversidade, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="18A14E6291C3D5A29BFF5C29E629558A" type="email">fagakorn@gmail.com</mods:nameIdentifier>
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<mods:name id="BCFE75CD11362A36CF4013E404AD66DD" type="personal">
<mods:role id="BD756500B7C17CE2BD21C776A55F2F02">
<mods:roleTerm id="3CD4F8276ED13B8F1DBD8F90AB95FF2B">Author</mods:roleTerm>
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<mods:namePart id="681DD0BD000AC63D64DB0B441801ED23">Vaz-De, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="7BB943AF00A9B1FF2786B1C0C3270D85" type="LSID">2FF2B7D6-1A6B-43C1-9966-A1A949FB2B05</mods:nameIdentifier>
<mods:affiliation id="6638330ABE4696290CE0AFC886DD5945">Instituto de Biociências, Departamento de Biologia e Zoologia, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="848AD0EA83606BD5800F4E1CC4885F6F" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:title id="CF4B59F454C5ED0A321A9B73519C1A13">European Journal of Taxonomy</mods:title>
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<mods:part id="09E3EF5E704C334A6455F38DE9C19493">
<mods:date id="E81F1E7ACD96FBB26EA1BC0A5EA02B37">2021</mods:date>
<mods:detail id="126BE8DC5E817409EF133974FC388FB0" type="pubDate">
<mods:number id="F4A71EF368D6E0E987D7FA0929148426">2021-10-20</mods:number>
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<mods:number id="56EC5FFA1DEB1D6F66F3577A3E22C3E7">775</mods:number>
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<mods:classification id="1674A40759FD8A919E4AB5259F8EC0D3">journal article</mods:classification>
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<mods:identifier id="E14846E50E8E24B0B51199AD2183B545" type="DOI">10.5852/ejt.2021.775.1551</mods:identifier>
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<treatment id="03F48795DC50FFBCA614A6C2FB8BFDF7" ID-DOI="http://doi.org/10.5281/zenodo.5589709" ID-GBIF-Taxon="189151491" ID-Zenodo-Dep="5589709" LSID="urn:lsid:plazi:treatment:03F48795DC50FFBCA614A6C2FB8BFDF7" httpUri="http://treatment.plazi.org/id/03F48795DC50FFBCA614A6C2FB8BFDF7" lastPageId="15" lastPageNumber="101" pageId="13" pageNumber="99">
<subSubSection id="C3476508DC50FFBEA614A6C2FC2BFE6B" box="[590,998,401,428]" pageId="13" pageNumber="99" type="nomenclature">
<paragraph id="8BE23683DC50FFBEA614A6C2FC2BFE6B" blockId="13.[363,1224,401,502]" box="[590,998,401,428]" pageId="13" pageNumber="99">
<heading id="D0AA81EFDC50FFBEA614A6C2FC2BFE6B" box="[590,998,401,428]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="99" reason="2">
<taxonomicName id="4C5D4D00DC50FFBEA614A6C2FC4EFE6B" ID-CoL="8SSP2" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[590,899,401,428]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="99" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
<emphasis id="B929EA91DC50FFBEA614A6C2FC4EFE6B" bold="true" box="[590,899,401,428]" italics="true" pageId="13" pageNumber="99">Deltochilum nonstriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC50FFBEA7D1A6C1FC2BFE6B" box="[907,998,402,428]" pageId="13" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3476508DC50FFBEA531A6E5FBE1FE32" pageId="13" pageNumber="99" type="description">
<paragraph id="8BE23683DC50FFBEA531A6E5FB05FE16" blockId="13.[363,1224,401,502]" box="[363,1224,438,465]" pageId="13" pageNumber="99">
<uri id="FFCC3A81DC50FFBEA531A6E5FB05FE16" box="[363,1224,438,465]" pageId="13" pageNumber="99">
urn:lsid:zoobank.org:act:
<uuid id="FFFB0C56DC50FFBEA6D3A6E5FB05FE16" box="[649,1224,438,465]" pageId="13" pageNumber="99">563D19C2-0450-4A9E-A4DC-B4AD395F25F0</uuid>
</uri>
</paragraph>
<paragraph id="8BE23683DC50FFBEA65DA688FBE1FE32" blockId="13.[363,1224,401,502]" box="[519,1068,475,502]" pageId="13" pageNumber="99">
<figureCitation id="13662A06DC50FFBEA65DA688FDA9FE32" box="[519,612,475,501]" captionStart="Fig" captionStartId="3.[189,232,1892,1918]" captionTargetBox="[222,1362,786,1773]" captionTargetId="figure-203@3.[219,1368,777,1781]" captionTargetPageId="3" captionText="Fig. 1. Habitus of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="13" pageNumber="99">Figs 1C</figureCitation>
,
<figureCitation id="13662A06DC50FFBEA62BA688FD5BFE32" box="[625,662,475,501]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[237,1351,525,1826]" captionTargetId="figure-106@4.[237,1351,525,1826]" captionTargetPageId="4" captionText="Fig. 2. Head of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589695" httpUri="https://zenodo.org/record/5589695/files/figure.png" pageId="13" pageNumber="99">2C</figureCitation>
,
<figureCitation id="13662A06DC50FFBEA6F8A688FD26FE32" box="[674,747,475,501]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="13" pageNumber="99">3C, G</figureCitation>
,
<figureCitation id="13662A06DC50FFBEA6ADA688FCD1FE32" box="[759,796,475,501]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="13" pageNumber="99">4C</figureCitation>
,
<figureCitation id="13662A06DC50FFBEA772A688FC80FE31" box="[808,845,475,502]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="13" pageNumber="99">5C</figureCitation>
,
<figureCitation id="13662A06DC50FFBEA703A688FCB0FE32" box="[857,893,475,501]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="13" pageNumber="99">6C</figureCitation>
,
<figureCitation id="13662A06DC50FFBEA7D0A688FC1EFE32" box="[906,979,475,501]" captionStart="Fig" captionStartId="9.[189,232,1822,1848]" captionTargetBox="[206,1383,744,1716]" captionTargetId="figure-193@9.[203,1383,744,1716]" captionTargetPageId="9" captionText="Fig. 7.Aedeagus of members of thegilli species group.A–E. Lateral view.F–G. Dorsal view of paramera. Arrows show the apical-dorsal notches of the paramera. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov. CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Holotype of D. jocelynae sp. nov. G. Holotype of D. nonstriatum sp. nov., CMNEN WDS00041746." figureDoi="http://doi.org/10.5281/zenodo.5589711" httpUri="https://zenodo.org/record/5589711/files/figure.png" pageId="13" pageNumber="99">7C, G</figureCitation>
,
<figureCitation id="13662A06DC50FFBEA785A688FBCEFE32" box="[991,1027,475,501]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[233,1362,650,1777]" captionTargetId="figure-181@10.[218,1370,644,1787]" captionTargetPageId="10" captionText="Fig. 8. Endophallus of members of thegilli species group.Arrows show the scales on the sub-medial area of the endophallus. A. Elongate scales. C. Regular scales. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589713" httpUri="https://zenodo.org/record/5589713/files/figure.png" pageId="13" pageNumber="99">8C</figureCitation>
,
<figureCitation id="13662A06DC50FFBEA055A688FBE1FE32" box="[1039,1068,475,501]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado &amp; Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="13" pageNumber="99">10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3476508DC50FFBEA4E7A54EFB40FD12" pageId="13" pageNumber="99" type="diagnosis">
<paragraph id="8BE23683DC50FFBEA4E7A54EFEFBFDF0" blockId="13.[189,310,541,567]" box="[189,310,541,567]" pageId="13" pageNumber="99">
<heading id="D0AA81EFDC50FFBEA4E7A54EFEFBFDF0" bold="true" box="[189,310,541,567]" fontSize="11" level="3" pageId="13" pageNumber="99" reason="3">
<emphasis id="B929EA91DC50FFBEA4E7A54EFEFBFDF0" bold="true" box="[189,310,541,567]" pageId="13" pageNumber="99">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE23683DC50FFBEA4E7A51EFB40FD12" blockId="13.[189,1399,589,726]" pageId="13" pageNumber="99">
Close to
<taxonomicName id="4C5D4D00DC50FFBEA573A51DFE72FDA0" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[297,447,589,616]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="99" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
<emphasis id="B929EA91DC50FFBEA573A51DFEF0FDA0" box="[297,317,590,615]" italics="true" pageId="13" pageNumber="99">D</emphasis>
.
<emphasis id="B929EA91DC50FFBEA517A51EFE72FDA0" box="[333,447,589,615]" italics="true" pageId="13" pageNumber="99">jocelynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC50FFBEA59DA51DFDEEFDAF" box="[455,547,590,616]" pageId="13" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
by having the striae I–VII inconspicuous (
<figureCitation id="13662A06DC50FFBEA07AA51EFB69FDAF" box="[1056,1188,589,616]" captionStart="Fig" captionStartId="3.[189,232,1892,1918]" captionTargetBox="[222,1362,786,1773]" captionTargetId="figure-203@3.[219,1368,777,1781]" captionTargetPageId="3" captionText="Fig. 1. Habitus of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="13" pageNumber="99">Figs 1B–C</figureCitation>
,
<figureCitation id="13662A06DC50FFBEA0EBA51DFB3AFDAF" box="[1201,1271,590,616]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="13" pageNumber="99">4B–C</figureCitation>
) including apically (
<figureCitation id="13662A06DC50FFBEA576A521FE61FD4A" box="[300,428,626,653]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="13" pageNumber="99">Fig. 5B–C</figureCitation>
). However, it can be distinguished by presenting the largest and densest interstrial punctures (
<figureCitation id="13662A06DC50FFBEA51BA5C4FE53FD76" box="[321,414,663,689]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="13" pageNumber="99">Fig. 4C</figureCitation>
), as well as densest punctures on the head frons (
<figureCitation id="13662A06DC50FFBEA7B6A5C4FB8AFD76" box="[1004,1095,663,689]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[237,1351,525,1826]" captionTargetId="figure-106@4.[237,1351,525,1826]" captionTargetPageId="4" captionText="Fig. 2. Head of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589695" httpUri="https://zenodo.org/record/5589695/files/figure.png" pageId="13" pageNumber="99">Fig. 2C</figureCitation>
) and on the pronotal disc (
<figureCitation id="13662A06DC50FFBEA49DA5E8FE85FD12" box="[199,328,699,725]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="13" pageNumber="99">Fig. 3C, G</figureCitation>
); finally, by the irregular shiny points on the pronotal disc (
<figureCitation id="13662A06DC50FFBEA7A5A5E8FB4DFD12" box="[1023,1152,699,725]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="13" pageNumber="99">Fig. 3C, G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3476508DC50FFBEA4E7A5ADFBF5FC8F" box="[189,1080,766,840]" pageId="13" pageNumber="99" type="etymology">
<paragraph id="8BE23683DC50FFBEA4E7A5ADFE89FCDF" blockId="13.[189,324,766,792]" box="[189,324,766,792]" pageId="13" pageNumber="99">
<heading id="D0AA81EFDC50FFBEA4E7A5ADFE89FCDF" bold="true" box="[189,324,766,792]" fontSize="11" level="3" pageId="13" pageNumber="99" reason="3">
<emphasis id="B929EA91DC50FFBEA4E7A5ADFE89FCDF" bold="true" box="[189,324,766,792]" pageId="13" pageNumber="99">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BE23683DC50FFBEA4E7A47DFBF5FC8F" blockId="13.[189,1080,814,840]" box="[189,1080,814,840]" pageId="13" pageNumber="99">
From Latin
<emphasis id="B929EA91DC50FFBEA513A47CFEB8FC8F" box="[329,373,815,840]" italics="true" pageId="13" pageNumber="99">non</emphasis>
- 'not' +
<emphasis id="B929EA91DC50FFBEA58CA47CFDC1FC8F" box="[470,524,815,840]" italics="true" pageId="13" pageNumber="99">stria</emphasis>
in reference to the inconspicuous elytral striae.
</paragraph>
</subSubSection>
<subSubSection id="C3476508DC50FFBEA4E7A422FEB6FBEF" pageId="13" pageNumber="99" type="materials_examined">
<paragraph id="8BE23683DC50FFBEA4E7A422FEA1FC4D" blockId="13.[189,364,880,907]" box="[189,364,880,907]" pageId="13" pageNumber="99">
<heading id="D0AA81EFDC50FFBEA4E7A422FEA1FC4D" bold="true" box="[189,364,880,907]" fontSize="11" level="3" pageId="13" pageNumber="99" reason="3">
<emphasis id="B929EA91DC50FFBEA4E7A422FEA1FC4D" bold="true" box="[189,364,880,907]" pageId="13" pageNumber="99">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BE23683DC50FFBEA4B0A4F2FEB6FBEF" blockId="13.[189,1399,929,1064]" pageId="13" pageNumber="99">
<materialsCitation id="3B353CDEDC50FFBEA4B0A4F2FEB6FBEF" ID-GBIF-Occurrence="3391047307" collectingDate="1986-08-04" collectingDateMax="1986-08-06" collectingDateMin="1986-08-04" collectionCode="CMNEN" collectorName="B. D. Gill" country="Venezuela" elevation="1000" latitude="7.438889" location="Bolivar" longLatPrecision="21" longitude="-61.558334" municipality="Elena" pageId="13" pageNumber="99" specimenCode="WSD00041746" specimenCount="1" specimenCount-male="1" stateProvince="Bolivar" typeStatus="holotype">
<emphasis id="B929EA91DC50FFBEA4B0A4F2FE91FC7C" bold="true" box="[234,348,929,955]" pageId="13" pageNumber="99">
<typeStatus id="54E68821DC50FFBEA4B0A4F2FE91FC7C" box="[234,348,929,955]" pageId="13" pageNumber="99" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F34A7613DC50FFBEA4E7A496FEA6FC18" box="[189,363,965,991]" name="Venezuela" pageId="13" pageNumber="99">VENEZUELA</collectingCountry>
•
<specimenCount id="9D5BFD0ADC50FFBEA5D7A496FE70FC25" box="[397,445,965,994]" pageId="13" pageNumber="99" type="male">
1
<emphasis id="B929EA91DC50FFBEA5FDA495FE70FC25" bold="true" box="[423,445,966,994]" pageId="13" pageNumber="99">♂</emphasis>
</specimenCount>
;
<collectingRegion id="4999F861DC50FFBEA58AA496FDE7FC18" box="[464,554,965,991]" country="Venezuela" name="Bolivar" pageId="13" pageNumber="99">Bolívar</collectingRegion>
,
<locationDeviation id="9A502FA1DC50FFBEA660A496FCBFFC18" box="[570,882,964,991]" pageId="13" pageNumber="99">km 40 Sta. [Santa] Elena</locationDeviation>
,
<locationDeviation id="9A502FA1DC50FFBEA7D8A496FBE5FC18" box="[898,1064,965,991]" pageId="13" pageNumber="99">Icabaru Road</locationDeviation>
;
<geoCoordinate id="EE695044DC50FFBEA06DA490FB0FFC27" box="[1079,1218,963,992]" degrees="7" direction="north" minutes="26" orientation="latitude" pageId="13" pageNumber="99" precision="15" seconds="20" value="7.438889">7°26′20″ N</geoCoordinate>
,
<geoCoordinate id="EE695044DC50FFBEA088A490FABFFC27" box="[1234,1394,963,992]" degrees="61" direction="west" minutes="33" orientation="longitude" pageId="13" pageNumber="99" precision="15" seconds="30" value="-61.558334">61°33′30″ W</geoCoordinate>
; alt.
<quantity id="4CA59B66DC50FFBEA4A9A4B9FE97FBC3" box="[243,346,1002,1028]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="99" unit="m" value="1000.0">
<elevation id="0070D1B0DC50FFBEA4A9A4B9FE97FBC3" box="[243,346,1002,1028]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="99" unit="m" value="1000.0">1000 m</elevation>
</quantity>
;
<date id="FFE31043DC50FFBEA52BA4BAFDFBFBC4" box="[369,566,1001,1028]" pageId="13" pageNumber="99" value="1986-08-04" valueMax="1986-08-06" valueMin="1986-08-04">
<collectingDate id="EFA7E9ABDC50FFBEA52BA4BAFDFBFBC4" box="[369,566,1001,1028]" pageId="13" pageNumber="99" value="1986-08-04" valueMax="1986-08-06" valueMin="1986-08-04">4–6 Aug. 1986</collectingDate>
</date>
;
<collectorName id="26A85355DC50FFBEA617A4B9FD09FBC3" box="[589,708,1002,1028]" pageId="13" pageNumber="99">B.D. Gill</collectorName>
leg.; [aedeagus and endophallus extracted];
<collectionCode id="ED4CAE46DC50FFBEA157A4B9FABAFBC3" box="[1293,1399,1002,1028]" pageId="13" pageNumber="99">CMNEN</collectionCode>
<specimenCode id="DBFB9EF8DC50FFBEA4E7A35DFEB5FBEF" box="[189,376,1038,1064]" collectionCode="CMNEN" pageId="13" pageNumber="99">WSD00041746</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3476508DC50FFBDA4E7A302FC22FE08" lastPageId="14" lastPageNumber="100" pageId="13" pageNumber="99" type="description">
<paragraph id="8BE23683DC50FFBEA4E7A302FE83FBAC" blockId="13.[189,334,1105,1131]" box="[189,334,1105,1131]" pageId="13" pageNumber="99">
<heading id="D0AA81EFDC50FFBEA4E7A302FE83FBAC" bold="true" box="[189,334,1105,1131]" fontSize="11" level="3" pageId="13" pageNumber="99" reason="3">
<emphasis id="B929EA91DC50FFBEA4E7A302FE83FBAC" bold="true" box="[189,334,1105,1131]" pageId="13" pageNumber="99">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BE23683DC50FFBEA4E7A3D2FCAEFB22" blockId="13.[189,1398,1153,1253]" pageId="13" pageNumber="99">
MEASUREMENTS AND COLOR.
<typeStatus id="54E68821DC50FFBEA5A5A3D2FDA1FB5C" box="[511,620,1153,1179]" pageId="13" pageNumber="99" type="holotype">Holotype</typeStatus>
male, length
<quantity id="4CA59B66DC50FFBEA751A3D2FCA5FB5C" box="[779,872,1153,1179]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="13" pageNumber="99" unit="mm" value="8.4">8.4 mm</quantity>
, humeral width
<quantity id="4CA59B66DC50FFBEA07EA3D2FB4CFB5C" box="[1060,1153,1153,1180]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="13" pageNumber="99" unit="mm" value="5.2">5.2 mm</quantity>
. Dark green with red reflections dorsally (
<figureCitation id="13662A06DC50FFBEA5F6A3F6FDC5FB78" box="[428,520,1189,1215]" captionStart="Fig" captionStartId="3.[189,232,1892,1918]" captionTargetBox="[222,1362,786,1773]" captionTargetId="figure-203@3.[219,1368,777,1781]" captionTargetPageId="3" captionText="Fig. 1. Habitus of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="13" pageNumber="99">Fig. 1C</figureCitation>
). Black ventrally, with shiny red reflections on anterior area of metaventral process, meso- and metafemora and ventrite VI (
<figureCitation id="13662A06DC50FFBEA6A3A399FC98FB22" box="[761,853,1226,1253]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="13" pageNumber="99">Fig. 6C</figureCitation>
).
</paragraph>
<paragraph id="8BE23683DC50FFBEA4E7A25EFCFCFAB7" blockId="13.[189,1398,1293,1392]" pageId="13" pageNumber="99">
HEAD (
<figureCitation id="13662A06DC50FFBEA554A25EFEA7FAE0" box="[270,362,1293,1319]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[237,1351,525,1826]" captionTargetId="figure-106@4.[237,1351,525,1826]" captionTargetPageId="4" captionText="Fig. 2. Head of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589695" httpUri="https://zenodo.org/record/5589695/files/figure.png" pageId="13" pageNumber="99">Fig. 2C</figureCitation>
). Dorsal inter-ocular distance approximately seven times width of one eye. Punctures on frons separated by less than one diameter of each puncture, almost contiguous. Punctures on head disc separated by less than one diameter of each puncture.
</paragraph>
<paragraph id="8BE23683DC50FFBEA4E7A2CAFB07FA1F" blockId="13.[189,1398,1432,1496]" pageId="13" pageNumber="99">
PRONOTUM (
<figureCitation id="13662A06DC50FFBEA517A2CBFE1AFA74" box="[333,471,1432,1459]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="13" pageNumber="99">Fig. 3C, G</figureCitation>
). Medial angle projected. Punctures on the disc separated by less than one diameter, almost contiguous. Shiny points on disc irregular and contiguous to punctures.
</paragraph>
<paragraph id="8BE23683DC50FFBEA4E7A153FDE0F96A" blockId="13.[189,1401,1536,1709]" pageId="13" pageNumber="99">
ELYTRA (
<figureCitation id="13662A06DC50FFBEA57FA153FE48F9DD" box="[293,389,1536,1562]" captionStart="Fig" captionStartId="3.[189,232,1892,1918]" captionTargetBox="[222,1362,786,1773]" captionTargetId="figure-203@3.[219,1368,777,1781]" captionTargetPageId="3" captionText="Fig. 1. Habitus of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="13" pageNumber="99">Figs 1C</figureCitation>
,
<figureCitation id="13662A06DC50FFBEA5CFA153FE75F9DD" box="[405,440,1536,1562]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="13" pageNumber="99">4C</figureCitation>
). Carina of the ninth interstria surpassing middle of elytral length. Striae I–VIII inconspicuous including apically (
<figureCitation id="13662A06DC50FFBEA60CA177FD7EF9F8" box="[598,691,1572,1599]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="13" pageNumber="99">Fig. 5C</figureCitation>
), only in some parts striae slightly visible and very narrow. Punctures of second and third interstriae on disc separated by less than one diameter. Punctures of third interstria on disc occupying about
<emphasis id="B929EA91DC50FFBEA6CDA13EFD6CF940" bold="true" box="[663,673,1645,1671]" pageId="13" pageNumber="99">1</emphasis>
⁄
<emphasis id="B929EA91DC50FFBEA6FCA13EFD76F940" bold="true" box="[678,699,1645,1671]" pageId="13" pageNumber="99">10</emphasis>
<superScript id="7C289BCBDC50FFBEA6E1A13EFD05F9BC" attach="left" box="[699,712,1645,1659]" fontSize="6" pageId="13" pageNumber="99">th</superScript>
the distance between striae II and III. Apical tubercles on interstriae III, V–VII (
<figureCitation id="13662A06DC50FFBEA599A1C1FDEDF96A" box="[451,544,1682,1709]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="13" pageNumber="99">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="8BE23683DC50FFBEA4E7A186FC84F89B" blockId="13.[189,1400,1749,1885]" pageId="13" pageNumber="99">
ABDOMEN (
<figureCitation id="13662A06DC50FFBEA565A186FE53F937" box="[319,414,1749,1776]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="13" pageNumber="99">Fig. 6C</figureCitation>
). Width of expansion of ventrite I, on ventrite III, narrower than distance between clypeal teeth; expansion reaching distal margin of ventrite IV. Margins of expansion between ventrites II–IV forming an acute angle. Apex of expansion rounded. Basal area of expansion with punctures separated by less than one diameter, almost contiguous.
</paragraph>
<paragraph id="8BE23683DC50FFBEA4E7A0D5FCB3F82F" blockId="13.[189,1398,1925,2024]" pageId="13" pageNumber="99">LEGS. Apex of mesotibia on ventral-internal margin with a small spatulate expansion. Expansion of metafemur twice wider than the width of metafemur basal to expansion. Internal margin of metatibia with large tubercles, occupying almost all metatibial length.</paragraph>
<paragraph id="8BE23683DC53FFBDA4E7A65AFDD0FEAE" blockId="14.[188,1399,265,362]" pageId="14" pageNumber="100">
PYGIDIUM (
<figureCitation id="13662A06DC53FFBDA560A65AFE5BFEE3" box="[314,406,265,292]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="14" pageNumber="100">Fig. 5C</figureCitation>
). Most of the punctures separated by less than one diameter; punctures basally denser that punctures on disc; basal punctures contiguous. Discal punctures occupying approximately
<emphasis id="B929EA91DC53FFBDA17AA67FFA8CFE81" bold="true" box="[1312,1345,300,326]" pageId="14" pageNumber="100">1/28</emphasis>
<superScript id="7C289BCBDC53FFBDA11BA678FA83FEFE" attach="left" box="[1345,1358,299,313]" fontSize="6" pageId="14" pageNumber="100">th</superScript>
the width on middle of pygidium.
</paragraph>
<paragraph id="8BE23683DC53FFBDA4E7A6C2FC22FE08" blockId="14.[189,1398,401,463]" pageId="14" pageNumber="100">
GENITALIA (
<figureCitation id="13662A06DC53FFBDA51EA6C2FE6FFE6C" box="[324,418,401,427]" captionStart="Fig" captionStartId="9.[189,232,1822,1848]" captionTargetBox="[206,1383,744,1716]" captionTargetId="figure-193@9.[203,1383,744,1716]" captionTargetPageId="9" captionText="Fig. 7.Aedeagus of members of thegilli species group.A–E. Lateral view.F–G. Dorsal view of paramera. Arrows show the apical-dorsal notches of the paramera. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov. CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Holotype of D. jocelynae sp. nov. G. Holotype of D. nonstriatum sp. nov., CMNEN WDS00041746." figureDoi="http://doi.org/10.5281/zenodo.5589711" httpUri="https://zenodo.org/record/5589711/files/figure.png" pageId="14" pageNumber="100">FigS 7C</figureCitation>
,
<figureCitation id="13662A06DC53FFBDA5EBA6C2FE19FE6C" box="[433,468,401,427]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[233,1362,650,1777]" captionTargetId="figure-181@10.[218,1370,644,1787]" captionTargetPageId="10" captionText="Fig. 8. Endophallus of members of thegilli species group.Arrows show the scales on the sub-medial area of the endophallus. A. Elongate scales. C. Regular scales. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589713" httpUri="https://zenodo.org/record/5589713/files/figure.png" pageId="14" pageNumber="100">8C</figureCitation>
). Aedeagus as described in the
<taxonomicName id="4C5D4D00DC53FFBDA716A6C2FCB1FE6C" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" box="[844,892,401,427]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="100" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC53FFBDA716A6C2FCB1FE6C" box="[844,892,401,427]" italics="true" pageId="14" pageNumber="100">gilli</emphasis>
</taxonomicName>
species group. Medial endophallite almost straight. Sub-medial area of endophallus with scales (
<figureCitation id="13662A06DC53FFBDA768A6E7FC40FE09" box="[818,909,436,462]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[233,1362,650,1777]" captionTargetId="figure-181@10.[218,1370,644,1787]" captionTargetPageId="10" captionText="Fig. 8. Endophallus of members of thegilli species group.Arrows show the scales on the sub-medial area of the endophallus. A. Elongate scales. C. Regular scales. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589713" httpUri="https://zenodo.org/record/5589713/files/figure.png" pageId="14" pageNumber="100">Fig. 8C</figureCitation>
, arrow).
</paragraph>
</subSubSection>
<caption id="DF22660BDC53FFBDA4E7A0DBFB60F82E" ID-DOI="http://doi.org/10.5281/zenodo.5589717" ID-Zenodo-Dep="5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="14" pageNumber="100" startId="14.[189,232,1928,1954]" targetBox="[336,1251,578,1870]" targetPageId="14">
<paragraph id="8BE23683DC53FFBDA4E7A0DBFB60F82E" blockId="14.[187,1400,1928,2025]" pageId="14" pageNumber="100">
<emphasis id="B929EA91DC53FFBDA4E7A0DBFED5F865" bold="true" box="[189,280,1928,1954]" pageId="14" pageNumber="100">Fig. 10.</emphasis>
Distribution map of the
<taxonomicName id="4C5D4D00DC53FFBDA677A0DBFD91F865" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" box="[557,604,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="100" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC53FFBDA677A0DBFD91F865" box="[557,604,1928,1954]" italics="true" pageId="14" pageNumber="100">gilli</emphasis>
</taxonomicName>
species group. Blue circle =
<taxonomicName id="4C5D4D00DC53FFBDA7FFA0DBFE67F802" authority="Gonzalez-Alvarado &amp; Vaz-de-Mello, 2021" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="100" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC53FFBDA7FFA0DBFBA2F865" box="[933,1135,1928,1954]" italics="true" pageId="14" pageNumber="100">Deltochilum gilli</emphasis>
<bibRefCitation id="EFCC4B72DC53FFBDA02FA0DBFE67F802" author="Gonzalez-Alvarado A. &amp; Vaz-de-Mello F. Z." pageId="14" pageNumber="100" pagination="0244657" refId="ref7969" refString="Gonzalez-Alvarado A. &amp; Vaz-de-Mello F. Z. 2021. Towards a comprehensive taxonomic revision of the Neotropical dung beetle subgenus Deltochilum (Deltohyboma) Lane, 1946 (Coleoptera: Scarabaeidae: Scarabaeinae): Division into species groups. PLOS ONE 16: e 0244657. https: // doi. org / 10.1371 / journal. pone. 0244657" type="journal article" year="2021">González-Alvarado &amp; Vaz-de-Mello, 2021</bibRefCitation>
</taxonomicName>
; yellow square =
<taxonomicName id="4C5D4D00DC53FFBDA6D8A0FFFCD7F802" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[642,794,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="100" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
<emphasis id="B929EA91DC53FFBDA6D8A0FFFCD7F802" box="[642,794,1963,1989]" italics="true" pageId="14" pageNumber="100">D. jocelynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC53FFBDA778A0FFFC4FF801" box="[802,898,1964,1990]" pageId="14" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
; red star =
<taxonomicName id="4C5D4D00DC53FFBDA056A0FFFB72F802" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[1036,1215,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="100" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
<emphasis id="B929EA91DC53FFBDA056A0FFFBEDF802" box="[1036,1056,1964,1989]" italics="true" pageId="14" pageNumber="100">D</emphasis>
.
<emphasis id="B929EA91DC53FFBDA06AA0FFFB72F802" box="[1072,1215,1964,1989]" italics="true" pageId="14" pageNumber="100">nonstriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC53FFBDA090A0FFFAE7F801" box="[1226,1322,1964,1990]" pageId="14" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
; green triangle =
<taxonomicName id="4C5D4D00DC53FFBDA56CA09CFE30F82F" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[310,509,1999,2025]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="100" phylum="Arthropoda" rank="species" species="quasistriatum" status="sp. nov.">
<emphasis id="B929EA91DC53FFBDA56CA09CFE86F82F" box="[310,331,1999,2024]" italics="true" pageId="14" pageNumber="100">D</emphasis>
.
<emphasis id="B929EA91DC53FFBDA500A09CFE30F82F" box="[346,509,1999,2024]" italics="true" pageId="14" pageNumber="100">quasistriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC53FFBDA65FA09CFDAEF82E" box="[517,611,1999,2025]" pageId="14" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
; white eight-point star =
<taxonomicName id="4C5D4D00DC53FFBDA7D2A09CFB87F82F" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[904,1098,1999,2025]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="100" phylum="Arthropoda" rank="species" species="tenuistriatum" status="sp. nov.">
<emphasis id="B929EA91DC53FFBDA7D2A09CFC50F82F" box="[904,925,1999,2024]" italics="true" pageId="14" pageNumber="100">D</emphasis>
.
<emphasis id="B929EA91DC53FFBDA7F6A09CFB87F82F" box="[940,1098,1999,2024]" italics="true" pageId="14" pageNumber="100">tenuistriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC53FFBDA008A09CFB60F82E" box="[1106,1197,1999,2025]" pageId="14" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="C3476508DC52FFBCA4E7A65AFDECFE78" pageId="15" pageNumber="101" type="discussion">
<paragraph id="8BE23683DC52FFBCA4E7A65AFEE2FEE4" blockId="15.[189,303,265,291]" box="[189,303,265,291]" pageId="15" pageNumber="101">
<heading id="D0AA81EFDC52FFBCA4E7A65AFEE2FEE4" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="15" pageNumber="101" reason="3">
<emphasis id="B929EA91DC52FFBCA4E7A65AFEE2FEE4" bold="true" box="[189,303,265,291]" pageId="15" pageNumber="101">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BE23683DC52FFBCA4E7A66BFDECFE78" blockId="15.[189,1400,312,447]" pageId="15" pageNumber="101">
This species is only known from the
<typeStatus id="54E68821DC52FFBCA623A66BFD29FE95" box="[633,740,312,338]" pageId="15" pageNumber="101" type="holotype">holotype</typeStatus>
. The
<typeStatus id="54E68821DC52FFBCA77CA66BFC40FE95" box="[806,909,312,338]" pageId="15" pageNumber="101" type="holotype">holotype</typeStatus>
has an identification label from François Génier (CMNEN), in which he recognised this specimen as probably being a new species since 1998. The transcription of the label as follows: [handwritten]
<taxonomicName id="4C5D4D00DC52FFBCA715A6D3FBEDFE5D" authorityName="Eschscholtz" authorityYear="1822" box="[847,1056,384,410]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="101" phylum="Arthropoda" rank="genus">DELTOCHILUM</taxonomicName>
/? SP. NOV. / [printed] Dét. F. Génier, 199 [handwritten]8.
</paragraph>
</subSubSection>
<subSubSection id="C3476508DC52FFBCA4E7A6B4FB8BFDF7" pageId="15" pageNumber="101" type="distribution">
<paragraph id="8BE23683DC52FFBCA4E7A6B4FE79FDC7" blockId="15.[189,436,486,513]" box="[189,436,486,513]" pageId="15" pageNumber="101">
<heading id="D0AA81EFDC52FFBCA4E7A6B4FE79FDC7" bold="true" box="[189,436,486,513]" fontSize="11" level="3" pageId="15" pageNumber="101" reason="3">
<emphasis id="B929EA91DC52FFBCA4E7A6B4FE79FDC7" bold="true" box="[189,436,486,513]" pageId="15" pageNumber="101">Known distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BE23683DC52FFBCA4E7A545FB8BFDF7" blockId="15.[189,1094,534,560]" box="[189,1094,534,560]" pageId="15" pageNumber="101">
<collectingCountry id="F34A7613DC52FFBCA4E7A545FEA2FDF7" box="[189,367,534,560]" name="Venezuela" pageId="15" pageNumber="101">VENEZUELA</collectingCountry>
.
<collectingRegion id="4999F861DC52FFBCA520A545FE1AFDF7" box="[378,471,534,560]" country="Venezuela" name="Bolivar" pageId="15" pageNumber="101">Bolívar</collectingRegion>
: km 40 Santa Elena, Icabarú Road (
<figureCitation id="13662A06DC52FFBCA720A545FC1CFDF7" box="[890,977,534,560]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado &amp; Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="15" pageNumber="101">Fig. 10</figureCitation>
, red star).
</paragraph>
</subSubSection>
</treatment>
</document>
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