treatments-xml/data/D0/B9/A2/D0B9A23F51045CDEA8EE3DE5A93BF24E.xml

<document ID-CLB-Dataset="93882" ID-DOI="http://dx.doi.org/10.3897/zookeys.1111.77541" ID-GBIF-Dataset="73ed8545-b7b1-462c-aa42-caa435b3a60b" ID-PMC="PMC9848980" ID-Pensoft-Pub="1313-2970-1111-215" ID-Pensoft-UUID="811D9FEFE793533D8D80C41B2333FAF6" ID-PubMed="36760847" ID-ZooBank="AA05385C92F64597BFCC7723A373EC02" ModsDocID="1313-2970-1111-215" checkinTime="1657577066635" checkinUser="pensoft" docAuthor="Cavalcante-Silva, Amanda, Pereira, Rafael &amp; Calor, Adolfo Ricardo" docDate="2022" docId="D0B9A23F51045CDEA8EE3DE5A93BF24E" docLanguage="en" docName="ZooKeys 1111: 215-244" docOrigin="ZooKeys 1111" docPubDate="2022-07-11" docSource="http://dx.doi.org/10.3897/zookeys.1111.77541" docTitle="Helicopsyche (Feropsyche) ralphi Cavalcante-Silva &amp; Pereira &amp; Calor 2022, sp. nov." docType="treatment" docUuid="15602B59-C063-4007-82C0-9141BD06D2BE" docUuidSource="ZooBank" docVersion="6" id="811D9FEFE793533D8D80C41B2333FAF6" lastPageNumber="215" masterDocId="811D9FEFE793533D8D80C41B2333FAF6" masterDocTitle="Caddisflies (Trichoptera) checklist and a new species of Helicopsyche von Siebold, 1856, from the Brejo de Altitude de Triunfo, a relict rainforest within the Caatinga domain, Northeast Brazil" masterLastPageNumber="244" masterPageNumber="215" pageNumber="215" updateTime="1720549258475" updateUser="ExternalLinkService">
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<mods:title id="B2A34D4393575197AECAE7CE26F1214B">Caddisflies (Trichoptera) checklist and a new species of Helicopsyche von Siebold, 1856, from the Brejo de Altitude de Triunfo, a relict rainforest within the Caatinga domain, Northeast Brazil</mods:title>
</mods:titleInfo>
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<mods:namePart id="160C837BDCE19E4536EF660E32AEE03A">Cavalcante-Silva, Amanda</mods:namePart>
<mods:affiliation id="ABEB69DFB6CE522EE09B0D066057B248">Laboratorio de Entomologia Aquatica, PPG Biodiversidade e Evolucao, Instituto de Biologia, Universidade Federal da Bahia, Rua Barao de Jeremoabo, 147, Campus Ondina, CEP 40170 - 115, Salvador, Bahia, Brazil</mods:affiliation>
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<mods:namePart id="E5E344B1A14C0C543427ADBA270ACFBD">Pereira, Rafael</mods:namePart>
<mods:nameIdentifier id="D2B00A8204B71E640575252B02419A45" type="ORCID">https://orcid.org/0000-0001-7578-6042</mods:nameIdentifier>
<mods:affiliation id="C146D868C28F4A4CC27BB0F78C560036">Laboratorio de Entomologia Aquatica, PPG Biodiversidade e Evolucao, Instituto de Biologia, Universidade Federal da Bahia, Rua Barao de Jeremoabo, 147, Campus Ondina, CEP 40170 - 115, Salvador, Bahia, Brazil</mods:affiliation>
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<mods:namePart id="34F93EF12A8ECA828A6AE961874DD8E6">Calor, Adolfo Ricardo</mods:namePart>
<mods:nameIdentifier id="582363AF0004C1E76BCE5B96BCE90FBA" type="ORCID">https://orcid.org/0000-0003-3975-3176</mods:nameIdentifier>
<mods:affiliation id="1293710E28C8619BDFEC8A8DA969442F">Laboratorio de Entomologia Aquatica, PPG Biodiversidade e Evolucao, Instituto de Biologia, Universidade Federal da Bahia, Rua Barao de Jeremoabo, 147, Campus Ondina, CEP 40170 - 115, Salvador, Bahia, Brazil</mods:affiliation>
<mods:nameIdentifier id="D305E816B1CBF59664DE077F95B59224" type="email">acalor@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="288F1271DA175A387FCAF0CEA4D2D90A">text</mods:typeOfResource>
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<mods:part id="FB19635FBEE057D97859030007288B0A">
<mods:date id="BFE3A00030ED1AF930D13CD5919BDB8C">2022</mods:date>
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<mods:number id="9860B8B0EB791A8E9185472DD105AA95">2022-07-11</mods:number>
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<mods:number id="158F382C6CD72EA38074E4C090C69A83">1111</mods:number>
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<mods:start id="BB31DF36319C00F741C012CEC1A37320">215</mods:start>
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<mods:url id="072399602446A2DC89C233083AE7426B">http://dx.doi.org/10.3897/zookeys.1111.77541</mods:url>
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<mods:identifier id="C55E4D189A24829113868DE9300DC928" type="DOI">http://dx.doi.org/10.3897/zookeys.1111.77541</mods:identifier>
<mods:identifier id="059F2E6AC854BC64364377784C95ACEA" type="Pensoft-Pub">1313-2970-1111-215</mods:identifier>
<mods:identifier id="03F9A9E64B3679C561AC10D4147EE675" type="ZooBank">AA05385C92F64597BFCC7723A373EC02</mods:identifier>
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<treatment id="D0B9A23F51045CDEA8EE3DE5A93BF24E" ID-GBIF-Taxon="197225302" LSID="urn:lsid:zoobank.org:act:15602B59-C063-4007-82C0-9141BD06D2BE" httpUri="http://treatment.plazi.org/id/D0B9A23F51045CDEA8EE3DE5A93BF24E" lastPageNumber="215" pageId="0" pageNumber="215">
<subSubSection id="F0463C6041112C99318652F626DF3281" pageId="0" pageNumber="215" type="nomenclature">
<paragraph id="0E02DF3F07564781E0E26FA3DB120179" pageId="0" pageNumber="215">
<taxonomicName id="F785A16251C04E833EB14A1EEEA519FC" LSID="http://zoobank.org/15602B59-C063-4007-82C0-9141BD06D2BE" authority="Cavalcante-Silva &amp; Pereira &amp; Calor, 2022" authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Helicopsyche (Feropsyche) ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="species" species="ralphi" status="sp. nov." subGenus="Feropsyche">Helicopsyche (Feropsyche) ralphi</taxonomicName>
<taxonomicNameLabel id="3ACD1E7607F63CE0ABE5EE83F54D9671" pageId="0" pageNumber="215">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="269748F0A34A6591B5C85FE5D9F107D5" pageId="0" pageNumber="215" type="description">
<paragraph id="CF31D2559EB198EF78A61ACA57449EB7" pageId="0" pageNumber="215">
<figureCitation id="2A646894C3FFB3BD6CD6D3CD8B9FA400" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">Figs 2</figureCitation>
<figureCitation id="DA50266028A007AF33F8CD4A3846C05D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">, 3</figureCitation>
<figureCitation id="57BE4AD5AA6BABD6C1B7DC00FE4B31D0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">, 4</figureCitation>
<figureCitation id="05BE982BC1A3A050640E237DB84DF0CF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">, 5</figureCitation>
<figureCitation id="ECA45DBA1870554A74ED3EDF58DE353A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Helicopsyche ralphi sp. nov., larvae and case A larval case, dorsal view B larval case, ventral view C larval lateral habitus." figureDoi="10.3897/zookeys.1111.77541.figure6" httpUri="https://binary.pensoft.net/fig/712454" pageId="0" pageNumber="215">, 6</figureCitation>
<figureCitation id="D5D0FFB84BAB6F65656C2290679DFCBF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Helicopsyche ralphi sp. nov., pupa and case A pupa front B abdominal segment IX and anal processes, dorsal view C pupa lateral habitus D-H abdominal segments I-V, dorsal, with details of dorsal hook plates I pupa case, ventral view J pupa case with sieve membrane highlighted, ventral view." figureDoi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="94D0457DDF06BA72B61368E43B3ECBB7" pageId="0" pageNumber="215" type="materials_examined">
<paragraph id="A7477FF0D859E01FEABD69DA9E7FBDB5" pageId="0" pageNumber="215">Material examined.</paragraph>
<paragraph id="516580CE6C1BD16B08EE9958A80FA84D" pageId="0" pageNumber="215">
<materialsCitation id="045222CF87234B823B303B941B207D0F" collectingDate="2017-09-21" collectingMethod="UV light pan trap" collectorName="Cavalcante-Silva, A." country="Brazil" elevation="890" latitude="-7.8675003" location="Pinga" longLatPrecision="21" longitude="-38.120277" municipality="Triunfo" specimenCount="1" specimenCount-male="1" stateProvince="Pernambuco" typeStatus="Holotype">
<emphasis id="A0C1EAF1A33C130877AEEA9FAFE127BD" bold="true" pageId="0" pageNumber="215">
<emphasis id="4BE29AC2E8ED45F4283162CE767DEA84" italics="true" pageId="0" pageNumber="215">
<typeStatus id="8B9DE11CC57816E4F9C9517C19204464">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="11A66682E60A6723416259E251F204D1" name="Brazil">Brazil</collectingCountry>
, 
<specimenCount id="018EBA92996DA2E640B98C38005F66D8" type="male">1 male</specimenCount>
; 
<collectingRegion id="21207754E35300F48806629DF1B142E2" country="Brazil" name="Pernambuco">Pernambuco</collectingRegion>
, 
<collectingMunicipality id="5A650AE2AF944076C208237A3B50BE1B">Triunfo</collectingMunicipality>
, 
<location id="A3CD7196BE49228CAEEFBC93D5D638F8" LSID="urn:lsid:plazi:treatment:D0B9A23F51045CDEA8EE3DE5A93BF24E:A3CD7196BE49228CAEEFBC93D5D638F8" country="Brazil" latitude="-7.8675003" longLatPrecision="21" longitude="-38.120277" municipality="Triunfo" name="Pinga" stateProvince="Pernambuco">Pinga</location>
stream; 
<geoCoordinate id="5283234339060B09E7D65243353D236B" degrees="7" direction="south" minutes="52" orientation="latitude" precision="15" seconds="3" value="-7.8675003">7°52'3&quot;S</geoCoordinate>
, 
<geoCoordinate id="ADEF241EF4C5EC867E107FCDCC6B65DA" degrees="38" direction="west" minutes="7" orientation="longitude" precision="15" seconds="13" value="-38.120277">38°7'13&quot;W</geoCoordinate>
, el. 
<quantity id="9167B31AD1B01F1A89732B03977B61A3" metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">
<elevation id="3836C927F6D5F1C74CC0E26D306BE0A3" metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">890 m</elevation>
</quantity>
; 
<collectingDate id="7ECF13AB2371CC1DB939DEB2BAAE6BF9" value="2017-09-21">21.ix.2017</collectingDate>
; 
<collectorName id="0D98F9731956D7A791B1A8509FF333C4">Cavalcante-Silva, A.</collectorName>
leg.; 
<collectingMethod id="CE0FEE057CB14E3B3F671FD825EAC98B">UV light pan trap</collectingMethod>
(MZSP)
</materialsCitation>
. 
<materialsCitation id="B51A99A2F6AC0FDA4A7E46D9305AC7DE" collectingDate="2017-09-21" collectingDateMax="2019-05-01" collectingDateMin="2017-09-21" collectorName="Cavalcante-Silva, A, Pereira, R." country="Brazil" elevation="740" latitude="-7.8681946" location="Pinga" longLatPrecision="1" longitude="-38.121025" municipality="Triunfo" specimenCount="44" specimenCount-female="24" specimenCount-male="20" stateProvince="Pernambuco" typeStatus="Paratypes">
<emphasis id="9199F4F3DE81B8A154FD93210869EBFB" bold="true" pageId="0" pageNumber="215">
<emphasis id="B1303506DDDAED9BF18776F2A72C0CC2" italics="true" pageId="0" pageNumber="215">
<typeStatus id="15B771A7C09F53154C8873B9E7C42FD9">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
Same data as holotype, except 
<specimenCount id="FE83FA4E0A6A57CBE9B36964F16F2B6C" type="male">5 males</specimenCount>
; Grito stream; 
<geoCoordinate id="A82ED256BEB02D49FEFD0F587FC617A7" degrees="7" direction="south" minutes="51" orientation="latitude" precision="15" seconds="41" value="-7.8613887">7°51'41&quot;S</geoCoordinate>
, 
<geoCoordinate id="64D3AC3B33464E2B26F8E37627215835" degrees="38" direction="west" minutes="5" orientation="longitude" precision="15" seconds="25" value="-38.090275">38°5'25&quot;W</geoCoordinate>
, el. 
<quantity id="DA88C8E74BD057FF51ACA861A11BED8D" metricMagnitude="2" metricUnit="m" metricValue="7.4" unit="m" value="740.0">
<elevation id="D57B99EDBEE2994E0D15719A84633175" metricMagnitude="2" metricUnit="m" metricValue="7.4" unit="m" value="740.0">740 m</elevation>
</quantity>
; 
<collectingDate id="809EB963425B739A00A1E15C5C6101FC" value="2018-08-09">09.viii.2018</collectingDate>
(UFBA); same except 
<specimenCount id="3E80401ED184DA8804353C220766548B" type="male">3 males</specimenCount>
; Laje stream; 
<geoCoordinate id="67F6A4035344E66197FBE76C8A879568" degrees="7" direction="south" minutes="52" orientation="latitude" precision="15" seconds="13" value="-7.870278">7°52'13&quot;S</geoCoordinate>
, 
<geoCoordinate id="38AFFD4E7DEDA6ADA011A17309711E5D" degrees="38" direction="west" minutes="5" orientation="longitude" precision="15" seconds="18" value="-38.088333">38°5'18&quot;W</geoCoordinate>
, el. 
<quantity id="8689D38581BAA266F55389AC1B3345CC" metricMagnitude="2" metricUnit="m" metricValue="5.8" unit="m" value="580.0">
<elevation id="EDCA6B80C17C970DD9E88CDAE5E804AA" metricMagnitude="2" metricUnit="m" metricValue="5.8" unit="m" value="580.0">580 m</elevation>
</quantity>
; 
<collectingDate id="CEE8F8967BFD66A2EDB82533D0A01C69" value="2018-08-07">07.viii.2018</collectingDate>
(DZRJ); same except 
<specimenCount id="5B75FB761E0249CB1015A4DE82446F35" type="male">6 males</specimenCount>
; Pinga stream; 
<geoCoordinate id="189950E8AF912D261BCD65D4C903F32B" degrees="7" direction="south" minutes="52" orientation="latitude" precision="15" seconds="3" value="-7.8675003">7°52'3&quot;S</geoCoordinate>
, 
<geoCoordinate id="BC6098BF3E80C3BAEDD2E37AC5C3335C" degrees="38" direction="west" minutes="7" orientation="longitude" precision="15" seconds="13" value="-38.120277">38°7'13&quot;W</geoCoordinate>
, el. 
<quantity id="B305BE212B81C16A8DDC9EBC1996DB76" metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">
<elevation id="A1D54FAD1C7974942C014399B5204671" metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">890 m</elevation>
</quantity>
; 
<collectingDate id="20FFBB6E0BBEBFDD35AE5E23DF127A2D" value="2017-09-21">21.ix.2017</collectingDate>
(MZSP); same except 
<specimenCount id="A7B8FBFB76DC5C24FB021687E095F4EA" type="male">6 males</specimenCount>
(INPA); same except 
<specimenCount id="EEA9060E9CED9A8504358034EA8C4852" type="female">1 female</specimenCount>
; 
<geoCoordinate id="BD7A9D7397058CB9D4C26DF18F5B4A14" degrees="7" direction="south" minutes="52" orientation="latitude" precision="15" seconds="3" value="-7.8675003">7°52'3&quot;S</geoCoordinate>
, 
<geoCoordinate id="918B3DE94513F21281263BF3E75B1C30" degrees="38" direction="west" minutes="7" orientation="longitude" precision="15" seconds="13" value="-38.120277">38°7'13&quot;W</geoCoordinate>
, el. 
<quantity id="36B7CD038A6219125370B70CE2337304" metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">
<elevation id="D76E30903A43232B5B7FEC811BF51BFC" metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">890 m</elevation>
</quantity>
; 
<collectingDate id="09DA557BA2C82416471A62D6972A1F91" value="2019-02-03">03.ii.2019</collectingDate>
(MZSP); same except 
<specimenCount id="63E4362AA3C7AB5F6271ADFF7164B298" type="female">5 females</specimenCount>
; 
<geoCoordinate id="3023D0D1F0B8650BAD16CCF5F60909C9" degrees="7" direction="south" minutes="52" orientation="latitude" precision="1" seconds="5,5" value="-7.8681946">7°52'5,5&quot;S</geoCoordinate>
, 
<geoCoordinate id="33C34F02D08B9D9B9FC8CB574AAD89D5" degrees="38" direction="west" minutes="7" orientation="longitude" precision="1" seconds="15,6" value="-38.121">38°7'15,6&quot;W</geoCoordinate>
, el. 
<quantity id="9A2DF78134FCDE4EDAEC58EAF18C152A" metricMagnitude="2" metricUnit="m" metricValue="8.7" unit="m" value="870.0">
<elevation id="DD75FBAAC7A9BE254D570E2E7EB409A9" metricMagnitude="2" metricUnit="m" metricValue="8.7" unit="m" value="870.0">870 m</elevation>
</quantity>
; 
<collectingDate id="1C6A23D32A0ED52321D467EFAF5FF797" value="2019-05-01">01.v.2019</collectingDate>
; 
<collectorName id="E34B045689EAB63965149C3DFAE7E36A">Cavalcante-Silva, A</collectorName>
, 
<collectorName id="D18D039A764201F8EBC7208872796199">Pereira, R.</collectorName>
leg. (MZSP); same except 
<specimenCount id="D2910C98DCEB4BF722E0093118D0FEC1" type="female">6 females</specimenCount>
; 
<geoCoordinate id="2F5B0D157B9E91B7F7410C2266FE83EB" degrees="7" direction="south" minutes="52" orientation="latitude" precision="1" seconds="5,5" value="-7.8681946">7°52'5,5&quot;S</geoCoordinate>
, 
<geoCoordinate id="25BF89BBF2B94E39CAD2C0852788515C" degrees="38" direction="west" minutes="7" orientation="longitude" precision="1" seconds="15,7" value="-38.121025">38°7'15,7&quot;W</geoCoordinate>
, el. 
<quantity id="F2C57F495A15FD4A5C53FEDE09D93AB3" metricMagnitude="2" metricUnit="m" metricValue="8.65" unit="m" value="865.0">
<elevation id="1A02141DADB9971B72689A9CBF7897F9" metricMagnitude="2" metricUnit="m" metricValue="8.65" unit="m" value="865.0">865 m</elevation>
</quantity>
; 
<collectingDate id="7A16EBC81C4257DC4E2725B2B77FA811" value="2019-05-01">01.v.2019</collectingDate>
(UFBA); same except 
<specimenCount id="1A69E2542769EEB837A47B339AB7FD6E" type="female">6 females</specimenCount>
(DZRJ); same except 
<specimenCount id="4C72171BB42636B6F52979F1178B50CF" type="female">6 females</specimenCount>
(INPA)
</materialsCitation>
.
</paragraph>
<caption id="05B4DD89B034A39FFDF59131FDB1C82D" doi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215" start="Figure 2" startId="F2">
<paragraph id="8F5B1529707AA04E1404D0A991F793BE" pageId="0" pageNumber="215">
<emphasis id="917DE5F9537C7CE0BEA52037B3C53EE2" bold="true" pageId="0" pageNumber="215">Figure 2.</emphasis>
<taxonomicName id="2EDDA6C7FDD54F0E8DA89B97C43C270F" authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="species" species="ralphi">
<emphasis id="9F9E3C1B3228D3DBAA123EF9B589806A" italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., male 
<emphasis id="3FA30FEA41B4E7796625B2F177BCD10F" bold="true" pageId="0" pageNumber="215">A</emphasis>
genitalia, lateral view 
<emphasis id="409EF7A6ED0070AB69D65489F58F93E8" bold="true" pageId="0" pageNumber="215">B</emphasis>
segments IX and X and inferior appendages, dorsal view 
<emphasis id="314E9593AC8BD37D3DB0A263F9B9BD7B" bold="true" pageId="0" pageNumber="215">C</emphasis>
inferior appendage, ventral view 
<emphasis id="BE3B7D990A9FB609D6190BC970C45983" bold="true" pageId="0" pageNumber="215">D</emphasis>
phallus, lateral view 
<emphasis id="1F331FAA93B3D0C654EDD1BBCBF740EE" bold="true" pageId="0" pageNumber="215">E</emphasis>
phallus, ventral view 
<emphasis id="E64EB36CC2D224875A8193C333161A20" bold="true" pageId="0" pageNumber="215">F</emphasis>
sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a.seg. IX = apodeme of abdominal segment IX; sup.a = superior appendage; seg. X = abdominal segment X; bas.l = basomesal lobe; inf.a = inferior appendage; pha.b = phallobase; pha.scl. = phallotremal sclerite.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DF1D5CBEE0A34CD0CCE0D0E62FAAA390" pageId="0" pageNumber="215" type="diagnosis">
<paragraph id="7A255672B98205D9E28B74A23C823BB6" pageId="0" pageNumber="215">Diagnosis.</paragraph>
<paragraph id="D5238FB45D7ECEDBD64EDB1596473E90" pageId="0" pageNumber="215">
The new species is distinguished from all other congeners by the following characters of the male genitalia: inferior appendage subtriangular, acuminated in posterior region, basomesal lobe subtriangular ~ 1/2 the length of the inferior appendage, in lateral view, trapezoid, with spine-like setae in posterior margin, in ventral view; abdominal segment X slender, slightly cleft at the apex, in dorsal view. The characters of the genitalia of new species are morphologically similar to 
<taxonomicName id="A8F0645B04A8A0978633C5BD52ED90D2" authorityName="Johanson" authorityYear="1999" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche flinti" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="species" species="flinti">
<emphasis id="4CCC6FF6F32C4A8109FD32C9A44ADE25" italics="true" pageId="0" pageNumber="215">Helicopsyche flinti</emphasis>
</taxonomicName>
<bibRefCitation id="E9160C63543CE20DF50F5210B05BDFB8" DOI="https://doi.org/10.1076/aqin.21.2.127.4535" author="Johanson, KA" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="215" pagination="127 - 132" refId="B36" refString="Johanson, KA, 1999. Description of two new Neotropical Helicopsyche (Trichoptera: Helicopsychidae). Aquatic Insects 21 (2): 127 - 132, DOI: https://doi.org/10.1076/aqin.21.2.127.4535" title="Description of two new Neotropical Helicopsyche (Trichoptera: Helicopsychidae)." url="https://doi.org/10.1076/aqin.21.2.127.4535" volume="21" year="1999">Johanson (1999)</bibRefCitation>
. The new species presents abdominal segment X with a rounded apex, and a medial row of spine-like setae, in dorsal view (while 
<taxonomicName id="E1C3C3C3F6FD4CD56C913FF470D9CECB" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="H. flinti" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="species" species="flinti">
<emphasis id="C96160AB751D58EDA20CE94F42146EAB" italics="true" pageId="0" pageNumber="215">H. flinti</emphasis>
</taxonomicName>
presents abdominal segment X with apex nearly straight, side row of spine-like setae), and inferior appendage with strongly projecting mesal margin, forming a large, rounded lobe, in dorsal view (while 
<taxonomicName id="D15AB555EE60F28D7BB95C6C77DD2377" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="H. flinti" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="species" species="flinti">
<emphasis id="EEC51542B28A2AE9E9BBADCE2E4D8B45" italics="true" pageId="0" pageNumber="215">H. flinti</emphasis>
</taxonomicName>
presents an inferior appendage without a large mesal lobe).
</paragraph>
<caption id="828AEC1B90949DBD105103ECC563349C" doi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215" start="Figure 3" startId="F3">
<paragraph id="E65A25598D0248FDA3D6DA1DBD017789" pageId="0" pageNumber="215">
<emphasis id="97F81C03A65D0F02C36788854D04BD64" bold="true" pageId="0" pageNumber="215">Figure 3.</emphasis>
<taxonomicName id="C878D10F38A4EA4A118391BFA2766A16" authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="species" species="ralphi">
<emphasis id="68905AEE2C6F3F7D8F96C0C10667765E" italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., female 
<emphasis id="9DBDBDAC1632A8064203CE511487618B" bold="true" pageId="0" pageNumber="215">A</emphasis>
genitalia, lateral view 
<emphasis id="25C71FBBCB1B86FCFCD299BE3573A73A" bold="true" pageId="0" pageNumber="215">B</emphasis>
genitalia, ventral view 
<emphasis id="9FCD39A339182053D42B6F41F355A5F4" bold="true" pageId="0" pageNumber="215">C</emphasis>
genitalia, dorsal view 
<emphasis id="33E325FEA3BCE5B2B4145AE760A236E4" bold="true" pageId="0" pageNumber="215">D</emphasis>
sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e.seg. IX = external part of abdominal segment IX; pre.a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="66BFB327DA1A17ACAAF9DA86EF97BC4A" pageId="0" pageNumber="215" type="description">
<paragraph id="B56DF730A22349A8D8075A957BDF6DF1" pageId="0" pageNumber="215">Description.</paragraph>
<paragraph id="2045DDA642D0F2C737885417439EF9D9" pageId="0" pageNumber="215">
<emphasis id="1CABF6A09EF4C77A89C801D65BD2143F" bold="true" pageId="0" pageNumber="215">Adults</emphasis>
(Fig. 
<figureCitation id="213875DD088FB084D1617E56061DFF8C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4</figureCitation>
): length of forewing 4.1-5.2 mm (
<emphasis id="0442A00A3306359EEDF31F1D3066E9DB" italics="true" pageId="0" pageNumber="215">n</emphasis>
= 20). Wings: forewing without discoidal cell, without medial cell, with thyridial cells; hind wing without discoidal cell, without thyridial cell. Head: brownish; antennae yellowish, shorter than forewing length, scape yellowish, shorter than head length, covered with long setae (Fig. 
<figureCitation id="B103DB7470B41AFFA8A90D4B50E63B13" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4E-G</figureCitation>
). Thorax: pronotum brownish, with warts, filiform, covered with small and ferruginous setae; mesoscutum brownish, with mesoscutal warts spherical and not covered with setae; mesoscutellum brownish, with mesoscutellar warts spherical and not covered with setae (Fig. 
<figureCitation id="4BB938A6B695C278200F6E9AC39593D6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4G</figureCitation>
); legs yellowish, tibial spur formula 2,2,4.
</paragraph>
<caption id="A50436FAC326824B234310734118AA34" doi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215" start="Figure 4" startId="F4">
<paragraph id="B9888ED3B71099EDE113ED47C8044AAA" pageId="0" pageNumber="215">
<emphasis id="849208280779A3588E95A1FF623E1065" bold="true" pageId="0" pageNumber="215">Figure 4.</emphasis>
<taxonomicName id="EFC29CA03B7C07AA8CE7E31E93FD821F" authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="species" species="ralphi">
<emphasis id="DC1B7655290791F6A95151FC87FB53E4" italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., adult 
<emphasis id="E0F3CC50066C49C0FBAD2F47654DFA19" bold="true" pageId="0" pageNumber="215">A</emphasis>
male head, dorsal view 
<emphasis id="FD9A6778B04ED5FEE1C9AE89FDE25343" bold="true" pageId="0" pageNumber="215">B</emphasis>
male head, frontal view 
<emphasis id="ECDDE8EAB906B714FEF387D3770D9426" bold="true" pageId="0" pageNumber="215">C</emphasis>
male habitus, dorsal view 
<emphasis id="81755316821DD3A0948AAD50CA816472" bold="true" pageId="0" pageNumber="215">D</emphasis>
female head, dorsal view 
<emphasis id="08201B6ADF9DA5FB6CCEA0B7412F6718" bold="true" pageId="0" pageNumber="215">E</emphasis>
male head, frontal view 
<emphasis id="1D5BC234C1E53AC70ACCD8C3EE1B9D04" bold="true" pageId="0" pageNumber="215">F</emphasis>
female habitus, dorsal view 
<emphasis id="96A1533FF4782EB329DAE8AE8B20431F" bold="true" pageId="0" pageNumber="215">G</emphasis>
female habitus, lateral view.
</paragraph>
</caption>
<paragraph id="075AB884C2453ECD25FB5D93CE505A78" pageId="0" pageNumber="215">
<emphasis id="7DF6E7681E2C5001FEB618E0C0C32EEB" bold="true" pageId="0" pageNumber="215">Male</emphasis>
(Figs 
<figureCitation id="71D7B85C44C00B938638EA4619F9BDA1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2</figureCitation>
, 
<figureCitation id="7F10A7E5AA904FBFF6D54ECC2543D23C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4A-C</figureCitation>
): body length ~ 3.3-4.6 mm (
<emphasis id="7918D757706BBCCF992C85F340BC0E9E" italics="true" pageId="0" pageNumber="215">n</emphasis>
= 20).
</paragraph>
<paragraph id="91ED62D1856A983B5D97A11897C6D680" pageId="0" pageNumber="215">
<emphasis id="CF59893414C418BF4BA2CC70551E4561" bold="true" italics="true" pageId="0" pageNumber="215">Head</emphasis>
: interantennal warts present, brownish, spherical, covered with small setae; posteroantennal warts present, brownish, club shaped, covered with long setae; cephalic warts present, brownish, subtriangular, covered with long setae; postocular warts present, filiform, brownish, covered with long setae (Fig. 
<figureCitation id="D06015E289AD8B2196EAD4F2B9E303C9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4A-C</figureCitation>
); maxillary palps yellowish, with two segments, covered with long ferruginous setae; labial palps yellowish, with three segments, covered with long ferruginous setae. Abdomen: abdominal sternum VI process present, almost same length as segment, tubular, apically rounded, covered with small microtrichiae (Fig. 
<figureCitation id="9C5327B25C6424FA2EA014E210B85624" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2F</figureCitation>
).
</paragraph>
<paragraph id="E597EFC3BD7F1D8E62F7482D3BDF8E52" pageId="0" pageNumber="215">
<emphasis id="60742A75C849941FB0E5DA519A184B5C" bold="true" pageId="0" pageNumber="215">
<emphasis id="63A95F2242536A1676486AD47926905C" italics="true" pageId="0" pageNumber="215">Genitalia</emphasis>
.
</emphasis>
Abdominal segment IX with slightly concave anteroventral margin in ventral half; apodeme well developed laterally, located midlaterally on segment; posterior margin nearly straight, in lateral view (Fig. 
<figureCitation id="A5011D6FC0EA16532FD35A584DFD60FB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2A</figureCitation>
), anterior margin strongly concave, in dorsal view (Fig. 
<figureCitation id="4AA437DE6B63C5456C9321A3B6A2B900" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2B</figureCitation>
); preanal appendages setose, rounded in lateral view (Fig. 
<figureCitation id="9105A1D7088743D184302F465619E4C4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2A</figureCitation>
), clavate in dorsal view (Fig. 
<figureCitation id="91A08E47CB911C6BED3133CB850A7D79" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2B</figureCitation>
). Abdominal segment X tubular, dorsal margin slightly curved, in lateral view (Fig. 
<figureCitation id="4B080B88A38C1EF5A8D31796C3FCF17C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2A</figureCitation>
); slender, mesodorsal borders inverted Y-shaped, bearing two rows of short setae, near the center, in dorsal view (Fig. 
<figureCitation id="E6E4A032A61245449A5250DDD6428E2D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2B</figureCitation>
). Inferior appendage subtriangular, acuminated in posterior region, in lateral view (Fig. 
<figureCitation id="5EDB5C0D8CAA8762C7A0F36F6F087613" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2A</figureCitation>
); anterior margin slightly convex, posterior margin undulated and tapered apex, in dorsal view (Fig. 
<figureCitation id="0D9A673E2244BBE871F59F226419CB35" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2B</figureCitation>
); basomesal lobe of inferior appendage, in lateral view well developed, with ventral margin covered with long setae and dorsal margin with spine-like setae, in ventral view (Fig. 
<figureCitation id="6D675963EC9F1208ACBA651F6C419AE8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2C</figureCitation>
). Phallus tubular, phallobase rounded, ventral view (Fig. 
<figureCitation id="41FB894E53990C1F99A1D7708972C590" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2E</figureCitation>
), acuminate at anteroventral border, in lateral view (Fig. 
<figureCitation id="AF80156E069922F5B906B97E61053210" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2D</figureCitation>
), slightly down curved; phallotremal sclerite conspicuous, moon shaped in lateral view (Fig. 
<figureCitation id="4D079C3966B762F6CDE23367EEC719D5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Helicopsyche ralphi sp. nov., male A genitalia, lateral view B segments IX and X and inferior appendages, dorsal view C inferior appendage, ventral view D phallus, lateral view E phallus, ventral view F sternum VI, lateral view. Abbreviations: seg. IX = abdominal segment IX; a. seg. IX = apodeme of abdominal segment IX; sup. a = superior appendage; seg. X = abdominal segment X; bas. l = basomesal lobe; inf. a = inferior appendage; pha. b = phallobase; pha. scl. = phallotremal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure2" httpUri="https://binary.pensoft.net/fig/712450" pageId="0" pageNumber="215">2D</figureCitation>
).
</paragraph>
<paragraph id="14BD7B1B4A406603ACE6FD4F31A41FC7" pageId="0" pageNumber="215">
<emphasis id="30BFF1AC84B205BAEE8FB32BAB1D318B" bold="true" pageId="0" pageNumber="215">Female</emphasis>
(Figs 
<figureCitation id="0754B6473C64034DE4446490CCE6E7E1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3</figureCitation>
, 
<figureCitation id="95E2603F3D67AB3A60D07F91BA503807" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4D-G</figureCitation>
): body length ~ 3.9-5.4 mm (
<emphasis id="1DF88B4427522C88DCD268BE1566CD60" italics="true" pageId="0" pageNumber="215">n</emphasis>
= 20).
</paragraph>
<paragraph id="642DBEBE8A0AF4633FE56DBCE9CB4FE7" pageId="0" pageNumber="215">
<emphasis id="44BFF326C3C5E0231F9F431329DE310E" bold="true" italics="true" pageId="0" pageNumber="215">Head</emphasis>
: interantennal warts present, brownish, spherical, covered with small setae; postero-antennal warts present, brownish, covered with long setae; cephalic warts present, brownish, subtriangular, covered with long setae (Fig. 
<figureCitation id="260991B74B84FF3FE6E1A7AE8D531C36" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4D-F</figureCitation>
); postocular warts present, filiform, brownish, covered with long setae (Fig. 
<figureCitation id="595B52F4D2805EE8142EC95E0801BF8C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4F</figureCitation>
); maxillary palps yellowish, with 5-segments, covered with long and yellowish setae; labial palps yellowish, with 3-segments, covered with long yellowish setae (Fig. 
<figureCitation id="DC69DA24A69E1381190EE6B7F01F2471" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Helicopsyche ralphi sp. nov., adult A male head, dorsal view B male head, frontal view C male habitus, dorsal view D female head, dorsal view E male head, frontal view F female habitus, dorsal view G female habitus, lateral view." figureDoi="10.3897/zookeys.1111.77541.figure4" httpUri="https://binary.pensoft.net/fig/712452" pageId="0" pageNumber="215">4G</figureCitation>
). Abdomen: abdominal sternum VI process present, ~ 1/3 segment length, tubular and apically rounded, covered with small microtrichiae (Fig. 
<figureCitation id="2C3B2FC94BE9CCC00840D796C92A5E44" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3D</figureCitation>
).
</paragraph>
<paragraph id="3697F75DA45061AD7234D8739AE9157D" pageId="0" pageNumber="215">
<emphasis id="2A63F0C9126A93B87FD20A80F5A01B3F" bold="true" pageId="0" pageNumber="215">
<emphasis id="1663CD2C7084261EF491394FBC790E37" italics="true" pageId="0" pageNumber="215">Genitalia</emphasis>
.
</emphasis>
Abdominal segment IX is well separated from abdominal segment VIII and indistinctly separated from abdominal segment X, anterior margin convex, in lateral view (Fig. 
<figureCitation id="6871024646736EE3DB53BBF906DAF1C4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3A</figureCitation>
); external part of abdominal segment IX apically incised, in ventral view (Fig. 
<figureCitation id="B3AEA6FAB700180FF97B5ADD1B75FD99" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3B</figureCitation>
). Preanal appendage long and filiform, in lateral view (Fig. 
<figureCitation id="C05F6ACB33D240F744235E70424EB956" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3A</figureCitation>
). Abdominal segment X with two branches; dorsal branch narrow, base with apex broad, rounded and covered with long setae, in lateral view (Fig. 
<figureCitation id="CC61694B253590B67B7E4EC3ECFEACD9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3A</figureCitation>
), bilobed with U-shaped with apical incision, in dorsal view (Fig. 
<figureCitation id="0F66F1D059311432C0B276AA7EE78B35" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3C</figureCitation>
); ventral branch with sinuous margin, in lateral view (Fig. 
<figureCitation id="0A745AF33BAEF33672C7F0A4D136C394" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3A</figureCitation>
), and apex obtuse in ventral and dorsal view (Fig. 
<figureCitation id="E655A0A78FBC2198DF72DF0768D2D078" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3B, C</figureCitation>
). Vagina with thick anterior margin, in ventral view (Fig. 
<figureCitation id="28E8DAF5A02632655CD473F636A37313" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3B</figureCitation>
); vaginal sclerite slender along its length, in lateral view (Fig. 
<figureCitation id="55829298A3EA5C8331819933A4C045B8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3A</figureCitation>
), finger-shaped projection on the anterior margin, internal sclerite long, with sclerotized lateral margins, in ventral view (Fig. 
<figureCitation id="A0D9981D61459160485D5AD1A57FB59C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Helicopsyche ralphi sp. nov., female A genitalia, lateral view B genitalia, ventral view C genitalia, dorsal view D sternum VI, lateral view. Abbreviations: seg. VIII = abdominal segment VIII; seg. IX = abdominal segment IX; e. seg. IX = external part of abdominal segment IX; pre. a = preanal appendage; Xd = dorsal branch abdominal segment X; Xv = ventral branch abdominal segment X; vag. = vagina; vag. scl. = vaginal sclerite." figureDoi="10.3897/zookeys.1111.77541.figure3" httpUri="https://binary.pensoft.net/fig/712451" pageId="0" pageNumber="215">3B</figureCitation>
).
</paragraph>
<paragraph id="BD2A4933F0673F7E0E1EB34F737A8152" pageId="0" pageNumber="215">
<emphasis id="E7811F4EDCFCFBFFFE90FAB0B3E44EE0" bold="true" pageId="0" pageNumber="215">Larva (5th instar)</emphasis>
(Figs 
<figureCitation id="CB64F9D8CC24D78657DE058AB6E9C937" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A-L</figureCitation>
, 
<figureCitation id="864BD5813A87940F2285F427A3DDFA78" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Helicopsyche ralphi sp. nov., larvae and case A larval case, dorsal view B larval case, ventral view C larval lateral habitus." figureDoi="10.3897/zookeys.1111.77541.figure6" httpUri="https://binary.pensoft.net/fig/712454" pageId="0" pageNumber="215">6C</figureCitation>
): Body total length 2.9-3.6 mm (
<emphasis id="10938973DC8F4A8D3CA8CD3179784556" italics="true" pageId="0" pageNumber="215">n</emphasis>
= 10).
</paragraph>
<caption id="D3572B659CE3565D29736A3558DDDB99" doi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215" start="Figure 5" startId="F5">
<paragraph id="95E9B27AA913506D5FFC428AEC637574" pageId="0" pageNumber="215">
<emphasis id="6566512ED409CAE9025FC88C2E335007" bold="true" pageId="0" pageNumber="215">Figure 5.</emphasis>
<taxonomicName id="56462AE23DFC9A068D037ED4558EB438" authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="species" species="ralphi">
<emphasis id="0D0AD8ABABAE84CA6AC6C41C75494271" italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., larvae 
<emphasis id="D6BEB164E341C372FF704CB1F41703C4" bold="true" pageId="0" pageNumber="215">A</emphasis>
head, dorsal view 
<emphasis id="A25A417F450694E82281B984CC3DC6BE" bold="true" pageId="0" pageNumber="215">B</emphasis>
head, ventral view 
<emphasis id="E29091E09CFC27E59504C9771D4044FA" bold="true" pageId="0" pageNumber="215">C</emphasis>
head, left lateral view 
<emphasis id="341B96547E67AE6B6BC9BF5AE5DB7201" bold="true" pageId="0" pageNumber="215">D</emphasis>
mandibles, dorsal view 
<emphasis id="81858F745D0605160DFACD6C1E161765" bold="true" pageId="0" pageNumber="215">E</emphasis>
labrum, dorsal view 
<emphasis id="8FAEC725C3B6CE516396931E875431C1" bold="true" pageId="0" pageNumber="215">F</emphasis>
notos, dorsal view 
<emphasis id="B5C1074139A3E87DB12893704672DED0" bold="true" pageId="0" pageNumber="215">G</emphasis>
proleg, lateral view 
<emphasis id="8E19B0078DB3C8A8132F180479E125E1" bold="true" pageId="0" pageNumber="215">H</emphasis>
mesoleg, lateral view 
<emphasis id="E9CEBB10B6401CB38C1D904D97EA1111" bold="true" pageId="0" pageNumber="215">I</emphasis>
metaleg, lateral view 
<emphasis id="14CE06CD7152D8FE5FF93F9C036B231D" bold="true" pageId="0" pageNumber="215">J</emphasis>
lateral hump 
<emphasis id="84B5FDE957ED7EED08F8F7D6EFB932A8" bold="true" pageId="0" pageNumber="215">L</emphasis>
anal legs.
</paragraph>
</caption>
<paragraph id="60EB0D0C2523FAEE6896B2135E8B194F" pageId="0" pageNumber="215">
<emphasis id="EE9460CD87F9F1CFF9111185BD0A71EB" bold="true" italics="true" pageId="0" pageNumber="215">Head</emphasis>
: oval, with anterior margin 1.3 
<normalizedToken id="00A394FD3A9305F6211DD83CB00EC2BE" originalValue="×">x</normalizedToken>
broader than posterior margin, in dorsal view (Fig. 
<figureCitation id="2EBA54B395B26A96CD5D218024A9ADCC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A</figureCitation>
), mostly light brown, with pale region on anterolateral margin of the head capsule until antenna region, around stemmata, medial region of frontoclypeus margin, and posteromedial region of head capsule, in dorsal view (Fig. 
<figureCitation id="6B85ED4A979CFD4BCE9E855D7E7695BE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A</figureCitation>
), with lateral region light brown, in ventral view (Fig. 
<figureCitation id="8C4E76AFE846D58FF599EB1DB31DF532" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5B</figureCitation>
), cardo and anterior ventral apotome dark brown, six and eight brown muscle scars in left and right, respectively, in dorsal view (Fig. 
<figureCitation id="CBB28B758F21F592255C5B05E22500CE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A</figureCitation>
), head capsule with muscle scar light brown in posterolateral region, in dorsal view (Fig. 
<figureCitation id="DA707B50B1D04EE99F3A11D3FE19CBE2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A</figureCitation>
), with frontal area flattened, muscle scars in basal region, cardo and ventral apotome sclerotized, ventral view (Fig. 
<figureCitation id="0735856B92E4D00746AD085800649B91" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5B</figureCitation>
); with muscle scar light brown in posterior region, in lateral view (Fig. 
<figureCitation id="58E4EE33B4836CD44D1BDDA2D662C200" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5C</figureCitation>
); frontoclypeus and adjacent areas nearly flat and margined with semicircular carina, frontoclypeal suture with strongly delimited margin, frontoclypeal with one muscle scar brown in medial region, and three in posterior region, in dorsal view (Fig. 
<figureCitation id="CC4133B6A623058353984D758114BC48" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A</figureCitation>
); labrum translucent, with short setae covering the anterior margin, in ventral view (Fig. 
<figureCitation id="52403ACBEAF597FD00F927EEEB072B24" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5B</figureCitation>
); mandibles asymmetrical, each internal margin with pale, long, thin setae; left mandible with three teeth, the apical tooth trilobed, mesal and basal tooth acute; right mandible with three teeth, the apical tooth trilobed, mesal and basal tooth obtuse, in dorsal view (Fig. 
<figureCitation id="E0578E9BD4A99983C2BBB2850A5B8397" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5D-E</figureCitation>
); chaetotaxy of head as in Figure 
<figureCitation id="E09DB357249BAF54EFAA75CEFF0D0200" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5A-E</figureCitation>
. Thorax: pronotum brown with dark muscle scars, anterior region with row of long setae at margin, covered long setae to near medial region, posterior margin sinuous and lighter with few and scattered setae, in dorsal view (Fig. 
<figureCitation id="93921C0FC3B23E7FCB40BEAAFFDC7F59" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5F</figureCitation>
), trochantin almost as long as foreleg coxae, finger shaped with one spinelike setae in apex (Fig. 
<figureCitation id="81CE8B16E75B742EF91E84D11276459C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5G</figureCitation>
); mesonotum lighter than pronotum, with pale regions in medial region, muscle scars in dark brown shades, four pairs in anterior region and one pair in posterior region, posterior margins angulate, in dorsal view (Fig. 
<figureCitation id="F005835D887A7911C828FBD9BDCEE585" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5F</figureCitation>
); metanotum with three pairs of sclerites, two pairs of anteromesal (sa1) sclerites small, one pairs anterior subtriangular, and one pairs irregular bearing one setae and one pair of posterior subtriangular sclerites (fused sa2 and sa3 sclerites), each bearing single seta posteromesally (sa2) and several setae anterolateral (sa3), in dorsal view (Fig. 
<figureCitation id="6416F8A4274A7BB55D20CF321FAD5D29" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5F</figureCitation>
); lateral hump oval, apical region mostly dark, one short setae in anteromedial region (Fig. 
<figureCitation id="AE119B8475651EF12CC6ECAC10FF9427" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5J</figureCitation>
); thoracic legs with chaetotaxy as in Figure 
<figureCitation id="946AF595A58CA2B165DBCC874E584F5F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5G-I</figureCitation>
; The foreleg has a length equivalent to 2/3 of the midleg and 1/2 of the hindleg, foreleg segments robust and short, mid and hind leg segments filiform and long (Fig. 
<figureCitation id="36CA094ACED4CECAE38E5928ECB64370" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5I</figureCitation>
). Abdomen: anal prolegs each with lateral sclerites curved, anal claw elongates, with accessory parallel teeth pectinate, arranged like comb (Fig. 
<figureCitation id="7730503C075940A4012AF3E0BC8CDA7A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5L</figureCitation>
).
</paragraph>
<paragraph id="3D55C1FB989B3DDB301E97769F7A96D5" pageId="0" pageNumber="215">
<emphasis id="348304A8F623973481A1C4553F67B9C3" bold="true" italics="true" pageId="0" pageNumber="215">Larval case</emphasis>
(Figs 
<figureCitation id="010111DDD9E53AF200605915BC0EA5B0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Helicopsyche ralphi sp. nov., larvae and case A larval case, dorsal view B larval case, ventral view C larval lateral habitus." figureDoi="10.3897/zookeys.1111.77541.figure6" httpUri="https://binary.pensoft.net/fig/712454" pageId="0" pageNumber="215">6A-B</figureCitation>
, 
<figureCitation id="444DB5828EFE72324211A8DF8969787C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Helicopsyche ralphi sp. nov., pupa and case A pupa front B abdominal segment IX and anal processes, dorsal view C pupa lateral habitus D-H abdominal segments I-V, dorsal, with details of dorsal hook plates I pupa case, ventral view J pupa case with sieve membrane highlighted, ventral view." figureDoi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215">7I-J</figureCitation>
): length 2.9-3.6 mm (
<emphasis id="6C164032FE91727218AE4DDCFA0DDB9B" italics="true" pageId="0" pageNumber="215">n</emphasis>
= 10). Made with cemented sand grains, forming a snail-like, helical case, case with two 1/2 whorls at the end of the phase, with umbilicus open and deep.
</paragraph>
<caption id="1CCD8DF235A3C8AFF64FF8E4101B3A05" doi="10.3897/zookeys.1111.77541.figure6" httpUri="https://binary.pensoft.net/fig/712454" pageId="0" pageNumber="215" start="Figure 6" startId="F6">
<paragraph id="3823EC5DA69E834220B2E8DCC558F74C" pageId="0" pageNumber="215">
<emphasis id="F47C7C9FDC07AF34328D198A53A68726" bold="true" pageId="0" pageNumber="215">Figure 6.</emphasis>
<taxonomicName id="37429A5D3EA0D2D60022AAFD01EFB290" authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="species" species="ralphi">
<emphasis id="A625372644DB4FD2EFA63F4ACC443E74" italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., larvae and case 
<emphasis id="5125160FC45F67FFDB20987F9C7810CA" bold="true" pageId="0" pageNumber="215">A</emphasis>
larval case, dorsal view 
<emphasis id="B2F489D2A34E3CF50F91846458FCEE6B" bold="true" pageId="0" pageNumber="215">B</emphasis>
larval case, ventral view 
<emphasis id="E8D19FAEB5EE427892E54DC29E2FFBAA" bold="true" pageId="0" pageNumber="215">C</emphasis>
larval lateral habitus.
</paragraph>
</caption>
<caption id="A2704A9C8B958881083A953C61693D46" doi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215" start="Figure 7" startId="F7">
<paragraph id="D567BAA9C0F8BAD26B864D87E2FF873B" pageId="0" pageNumber="215">
<emphasis id="246491B841CC9A775C053B74F72DBBF6" bold="true" pageId="0" pageNumber="215">Figure 7.</emphasis>
<taxonomicName id="16AAB37F5066A290527FAE3BE240733A" authorityName="Cavalcante-Silva &amp; Pereira &amp; Calor" authorityYear="2022" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" lsidName="Helicopsyche ralphi" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="species" species="ralphi">
<emphasis id="0ADDB2D4FF1BA2BDA02B0C1AC7F0B35A" italics="true" pageId="0" pageNumber="215">Helicopsyche ralphi</emphasis>
</taxonomicName>
sp. nov., pupa and case 
<emphasis id="8B6D7C81883CB814849095F367706159" bold="true" pageId="0" pageNumber="215">A</emphasis>
pupa front 
<emphasis id="C392134E3F8870F528A01DC76C5EB454" bold="true" pageId="0" pageNumber="215">B</emphasis>
abdominal segment IX and anal processes, dorsal view 
<emphasis id="6B84E147FC00219CA0D30B9B74740A81" bold="true" pageId="0" pageNumber="215">C</emphasis>
pupa lateral habitus 
<emphasis id="FDF49DBEF287BA577A948D500D88D82C" bold="true" pageId="0" pageNumber="215">D-H</emphasis>
abdominal segments I-V, dorsal, with details of dorsal hook plates 
<emphasis id="7BBE2D0727C944434490757F3FCF5E29" bold="true" pageId="0" pageNumber="215">I</emphasis>
pupa case, ventral view 
<emphasis id="9F51043850764EFF37639F68D9708FEF" bold="true" pageId="0" pageNumber="215">J</emphasis>
pupa case with sieve membrane highlighted, ventral view.
</paragraph>
</caption>
<paragraph id="DD83DA4A2B77F8075C50E18E96A40474" pageId="0" pageNumber="215">
<emphasis id="4F45C2310BD73B4A52B9994F3649C11E" bold="true" italics="true" pageId="0" pageNumber="215">Pupa</emphasis>
(Fig. 
<figureCitation id="7FD43F8BB5238517A654A39ECB18413B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Helicopsyche ralphi sp. nov., pupa and case A pupa front B abdominal segment IX and anal processes, dorsal view C pupa lateral habitus D-H abdominal segments I-V, dorsal, with details of dorsal hook plates I pupa case, ventral view J pupa case with sieve membrane highlighted, ventral view." figureDoi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215">7A-J</figureCitation>
): body length 3.3-4.1 mm (
<emphasis id="5823F8D2A9A9D49FE18D42FCF484BC66" italics="true" pageId="0" pageNumber="215">n</emphasis>
= 10). Generally dark brown, almost black, with yellowish abdomen; Head: mandibles curved with wide bases, each with length 2.5 
<normalizedToken id="73C4496A4762F88A99FAE8A273DACD92" originalValue="×">x</normalizedToken>
basal width, apex pointed and internal margin smooth (Fig. 
<figureCitation id="2F1B904FB420017FBC9AAB353A9E5C42" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Helicopsyche ralphi sp. nov., pupa and case A pupa front B abdominal segment IX and anal processes, dorsal view C pupa lateral habitus D-H abdominal segments I-V, dorsal, with details of dorsal hook plates I pupa case, ventral view J pupa case with sieve membrane highlighted, ventral view." figureDoi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215">7A</figureCitation>
). Abdomen: paired anterior dorsal hook plates on segments II-V, pair of posterior dorsal hook plates on segment IV asymmetrical; general morphology of dorsal hook plates as in (Fig. 
<figureCitation id="A2BABD6DACF4AE963064C23D54009032" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Helicopsyche ralphi sp. nov., larvae A head, dorsal view B head, ventral view C head, left lateral view D mandibles, dorsal view E labrum, dorsal view F notos, dorsal view G proleg, lateral view H mesoleg, lateral view I metaleg, lateral view J lateral hump L anal legs." figureDoi="10.3897/zookeys.1111.77541.figure5" httpUri="https://binary.pensoft.net/fig/712453" pageId="0" pageNumber="215">5C-H</figureCitation>
); terminal abdominal segment rounded, with two divergent digitate processes, each process bearing one subapical and three apical setae (Fig. 
<figureCitation id="A1F3D3158BDE4A64A065981AAF262431" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Helicopsyche ralphi sp. nov., pupa and case A pupa front B abdominal segment IX and anal processes, dorsal view C pupa lateral habitus D-H abdominal segments I-V, dorsal, with details of dorsal hook plates I pupa case, ventral view J pupa case with sieve membrane highlighted, ventral view." figureDoi="10.3897/zookeys.1111.77541.figure7" httpUri="https://binary.pensoft.net/fig/712455" pageId="0" pageNumber="215">7B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E9EB4C51DDED0E490BD8DE7147B6B53E" pageId="0" pageNumber="215" type="etymology">
<paragraph id="60A6432B5A5D70D18B273F4F383BF6A3" pageId="0" pageNumber="215">Etymology.</paragraph>
<paragraph id="A6D1EE965430BDEB1C75729264908E87" pageId="0" pageNumber="215">This species is named in honor of Dr. Ralph W. Holzenthal, for his outstanding contribution to the study of Neotropical caddisflies, and also as an acknowledgment for collaboration and his contributing to the training of young researchers.</paragraph>
</subSubSection>
<subSubSection id="FEA25364C4D0492C8A3CB49F3B3BE589" pageId="0" pageNumber="215" type="materials_examined">
<paragraph id="B1F60B99323E5766C76E2C53367C426A" pageId="0" pageNumber="215">Material additional.</paragraph>
<paragraph id="762E66A3C1896E437491E15677D5EFE7" pageId="0" pageNumber="215">
A1 (
<specimenCount id="906C397864CE3FDBD277195D85852821" type="female">1 female</specimenCount>
), C6 (
<specimenCount id="4AB5ADC8303DD8B10703E73F8714F01E" type="male">643 males</specimenCount>
); C7 (
<specimenCount id="1B40C61A59354668811E3871DB2DBEE0" type="larva">10 larvae</specimenCount>
, 
<specimenCount id="4F9F9334C581DB14BEB28D69137A9E07" type="pupa">10 pupae</specimenCount>
); D2 (
<specimenCount id="3D17E0E74A16D09C8A4A25583BD51426" type="male">34 males</specimenCount>
); D7 (
<specimenCount id="DF2CC032F89B37B54F563214F2DB9D3A" type="male">1 male</specimenCount>
); D17 (
<specimenCount id="8E7EE66D331CDE08892E7132B2FA5835" type="male">14 males</specimenCount>
); D18 (
<specimenCount id="EAD5C69D073680B4E4E115F35F71365F" type="male">379 males</specimenCount>
); D18 (
<specimenCount id="BFA221177831696ADE1CFAF272D9167C" type="female">49 females</specimenCount>
); D20 (
<specimenCount id="9E5545200DCE45D297C445BC7F16BB03" type="female">32 females</specimenCount>
); D22 (
<specimenCount id="264E673933CBA209711001BB08E477CB" type="larva">1 larvae</specimenCount>
); E1 (
<specimenCount id="76453E34B466F0556036BB222B47C8F9" type="male">1 male</specimenCount>
); F2 (
<specimenCount id="0F70904EDE5071656BFDCDB8D99BC221" type="male">30 males</specimenCount>
); F3 (
<specimenCount id="86C778E0B4E5A1C2B5C6EEC1091269D2" type="male">17 males</specimenCount>
) (UFBA) (Table 
<tableCitation id="E86A2C3E5C9931AD61BF1F35933EF858" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Collection data from Brejo de Altitude de Triunfo, with the sample of each collection site, geographic coordinates, elevation, date, traps (LPT = UV Light Pan Trap, MAL = Malaise and WSA = White Sheet Attraction), and acronyms of collectors." httpUri="http://table.plazi.org/id/6FB051B14B382AEF1FDDD28788D65E4B" pageId="0" pageNumber="215" tableUuid="6FB051B14B382AEF1FDDD28788D65E4B">1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A6187673C3113283F43E4ACE28E095A3" pageId="0" pageNumber="215" type="distribution">
<paragraph id="8A64936670038867B1148C0F7A9D2219" pageId="0" pageNumber="215">Distribution.</paragraph>
<paragraph id="97C25D2873A123DC521A04DCCE42A1CD" pageId="0" pageNumber="215">Brazil (Pernambuco state).</paragraph>
</subSubSection>
<subSubSection id="E5E3745683104BA201A6695A834A0803" pageId="0" pageNumber="215" type="caddisflies from brejo de altitude de triunfo">
<paragraph id="FE4B3722776AE8E09D50AFD43DD08CE3" pageId="0" pageNumber="215">Caddisflies from Brejo de Altitude de Triunfo</paragraph>
</subSubSection>
<subSubSection id="D3CA71E6A5C9BF6FE6A33E3D2141BC75" pageId="0" pageNumber="215" type="hydropsychidae">
<paragraph id="8AEE9288DDA0B97FB0E65AB0369F8CA6" pageId="0" pageNumber="215">
<taxonomicName id="581F79F3CB25A7F28874EE3CA35A7DC5" authorityName="J Curtis" authorityYear="1835" class="Insecta" family="Hydropsychidae" kingdom="Animalia" lsidName="" order="Trichoptera" pageId="0" pageNumber="215" phylum="Arthropoda" rank="family">HYDROPSYCHIDAE</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>