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<document id="A2D3113C6443F8412349F0E355A84538" ID-CLB-Dataset="20208" ID-DOI="10.11646/zootaxa.5115.4.1" ID-GBIF-Dataset="b99d2dc5-e0a6-4f73-b2c6-b21b6350c69f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6361552" ID-ZooBank="320C2E00-8035-431A-8AD7-D483D81923F6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647419732391" checkinUser="plazi" docAuthor="Ullah, Mishkat, Naeem, Muhammad, Mahmood, Khalid &amp; Garner, Beulah" docDate="2022" docId="291CD3215304DC69FF6DFAA38791FE65" docLanguage="en" docName="zootaxa.5115.4.1.pdf" docOrigin="Zootaxa 5115 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlaenius (Lithochlaenius) agilis Chaudoir 1856" docType="treatment" docVersion="7" lastPageNumber="471" masterDocId="D525AB595317DC7DFFFAFFFA836DFF91" masterDocTitle="Addition to the knowledge of Tribe Chlaeniini Brullé, 1834 (Coleoptera: Carabidae) from Pakistan" masterLastPageNumber="486" masterPageNumber="451" pageNumber="470" updateTime="1699318106451" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="41DA9BCCFB6A60433C6B756DCC9C1B27">Addition to the knowledge of Tribe Chlaeniini Brullé, 1834 (Coleoptera: Carabidae) from Pakistan</mods:title>
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<mods:namePart id="D929AC53A3B9ECD6549B9A6C5033F447">Ullah, Mishkat</mods:namePart>
<mods:affiliation id="EB09F28EA4A66C2AE98E8F273BC2FE6D">Insects Division, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom.</mods:affiliation>
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<mods:namePart id="756AB6BC4B0CE73E2695DDD530A2F8D8">Naeem, Muhammad</mods:namePart>
<mods:nameIdentifier id="A7B7A8BF1D746A6F5691D6A2562F6D68" type="ORCID">0000-0002-2025-2024</mods:nameIdentifier>
<mods:affiliation id="221528BEC02E247FE154DF693604A58D">muhammad. naeem @ uaar. edu. pk; https: // orcid. org / 0000 - 0002 - 2025 - 2024</mods:affiliation>
<mods:nameIdentifier id="8D400AF42737EDFB514171D97B8D1E48" type="email">muhammad.naeem@uaar.edu.pk</mods:nameIdentifier>
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<mods:namePart id="AF1D6473BBCDF72EA5520BAF3D3C2820">Mahmood, Khalid</mods:namePart>
<mods:nameIdentifier id="7C4BF2521CBC8F5DF1859005F645B0CD" type="ORCID">0000-0001-7162-3430</mods:nameIdentifier>
<mods:affiliation id="078DA248A126EE75B5E403299C4B4D0C">dkmahmood @ msn. com; https: // orcid. org / 0000 - 0001 - 7162 - 3430</mods:affiliation>
<mods:nameIdentifier id="C9ABE907C0BC4A01506B13925E715DDF" type="email">dkmahmood@msn.com</mods:nameIdentifier>
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<mods:namePart id="2F1FAB9D05E136624A5072158A375298">Garner, Beulah</mods:namePart>
<mods:nameIdentifier id="413E6AAC05AA707F9E431DF0910555BA" type="ORCID">0000-0002-5229-2450</mods:nameIdentifier>
<mods:affiliation id="0E0B54C92B5EF4BE20844995CA921550">Insects Division, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom. &amp; b. garner @ nhm. ac. uk; https: // orcid. org / 0000 - 0002 - 5229 - 2450</mods:affiliation>
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<mods:title id="C817EEFF3D22139F3C60F19F6FE0CCDF">Zootaxa</mods:title>
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<mods:date id="E3E47F4E5854B033201D0AFA6186E370">2022</mods:date>
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<treatment id="291CD3215304DC69FF6DFAA38791FE65" ID-DOI="http://doi.org/10.5281/zenodo.6365171" ID-GBIF-Taxon="194051279" ID-Zenodo-Dep="6365171" LSID="urn:lsid:plazi:treatment:291CD3215304DC69FF6DFAA38791FE65" httpUri="http://treatment.plazi.org/id/291CD3215304DC69FF6DFAA38791FE65" lastPageId="20" lastPageNumber="471" pageId="19" pageNumber="470">
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<taxonomicName id="66B519B45304DC6EFF6DFAA38199FAE5" ID-CoL="5Y7LJ" authority="Chaudoir, 1856" authorityName="Chaudoir" authorityYear="1856" box="[151,756,1369,1396]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="agilis" subGenus="Lithochlaenius">
<emphasis id="93C1BE255304DC6EFF6DFAA38199FAE5" bold="true" box="[151,756,1369,1396]" pageId="19" pageNumber="470">
<emphasis id="93C1BE255304DC6EFF6DFAA3827EFAE5" bold="true" box="[151,275,1369,1396]" italics="true" pageId="19" pageNumber="470">Chlaenius</emphasis>
(
<emphasis id="93C1BE255304DC6EFEDEFAA382B6FAE5" bold="true" box="[292,475,1369,1396]" italics="true" pageId="19" pageNumber="470">Lithochlaenius</emphasis>
)
<emphasis id="93C1BE255304DC6EFE16FAA38140FAE5" bold="true" box="[492,557,1369,1396]" italics="true" pageId="19" pageNumber="470">agilis</emphasis>
<bibRefCitation id="C5241FC65304DC6EFDCEFAA38199FAE5" author="Chaudoir, M." box="[564,756,1369,1396]" pageId="19" pageNumber="470" pagination="187 - 291" refId="ref26044" refString="Chaudoir, M. (1856) Memoire sur la famille des carabiques. 6 - e partie. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 29 (3), 187 - 291." type="journal article" year="1856">Chaudoir, 1856</bibRefCitation>
</emphasis>
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<paragraph id="A10A62375304DC6EFF6DFA848394FA08" blockId="19.[151,756,1369,1433]" box="[151,249,1406,1433]" pageId="19" pageNumber="470">
(
<figureCitation id="398E7EB25304DC6EFF5AFA84839CFA08" box="[160,241,1406,1433]" captionStart="FIGURE 6" captionStartId="18.[151,250,1943,1967]" captionTargetBox="[211,1376,181,1919]" captionTargetId="figure-25@18.[211,1376,181,1919]" captionTargetPageId="18" captionText="FIGURE 6. Dorsal habitus (a) Chlaenius (Epomis) nigricans Wiedemann, 1821; (b) Chlaenius (Lissauchenius) rufifemoratus rayotus Bates, 1891 imaged from NHMUK collection; (c) Chlaenius (Lissauchenius) variipes Chaudoir, 1856 imaged from NHMUK collection; (d) Chlaenius (Lithochlaenius) agilis Chaudoir, 1856." figureDoi="http://doi.org/10.5281/zenodo.6361571" httpUri="https://zenodo.org/record/6361571/files/figure.png" pageId="19" pageNumber="470">Fig. 6d</figureCitation>
)
</paragraph>
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<subSubSection id="E9AF31BC5304DC6EFF6DFA388399FA68" pageId="19" pageNumber="470" type="reference_group">
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<treatmentCitationGroup id="81A545195304DC6EFF6DFA388399FA68" pageId="19" pageNumber="470">
<treatmentCitation id="201444265304DC6EFF6DFA38814CFA4B" author="Chaudoir, M." box="[151,545,1474,1498]" page="246" pageId="19" pageNumber="470" year="1856">
<taxonomicName id="66B519B45304DC6EFF6DFA38814CFA4B" ID-CoL="5Y7LJ" authority="Chaudoir, 1856: 246" authorityName="Chaudoir" authorityPageNumber="246" authorityYear="1856" box="[151,545,1474,1498]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="93C1BE255304DC6EFF6DFA388252FA4B" box="[151,319,1474,1498]" italics="true" pageId="19" pageNumber="470">Chlaenius agilis</emphasis>
<bibRefCitation id="C5241FC65304DC6EFEB2FA38814CFA4B" author="Chaudoir, M." box="[328,545,1474,1498]" pageId="19" pageNumber="470" pagination="187 - 291" refId="ref26044" refString="Chaudoir, M. (1856) Memoire sur la famille des carabiques. 6 - e partie. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 29 (3), 187 - 291." type="journal article" year="1856">Chaudoir, 1856: 246</bibRefCitation>
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;
<taxonomicName id="66B519B45304DC6EFDD4FA388769FA4B" ID-CoL="5XVBL" authority="Bates, 1889 b: 212" authorityName="Bates, H. E. Andrewes" authorityPageNumber="212" authorityYear="1889" box="[558,1028,1474,1498]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis id="93C1BE255304DC6EFDD4FA388056FA4B" box="[558,827,1474,1498]" italics="true" pageId="19" pageNumber="470">Chlaenius anchomenoides</emphasis>
Bates, 1889b: 212
</taxonomicName>
;
<taxonomicName id="66B519B45304DC6EFBEBFA38839DFA68" ID-CoL="69V6K" authority="Jedlicka, 1956 a: 194" authorityName="Jedlicka" authorityPageNumber="194" authorityYear="1956" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis id="93C1BE255304DC6EFBEBFA388781FA4B" box="[1041,1260,1474,1498]" italics="true" pageId="19" pageNumber="470">Chlaenius nuristanus</emphasis>
Jedlicka, 1956a: 194
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5304DC6EFF6DF9DD81BAF8AF" pageId="19" pageNumber="470" type="materials_examined">
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<emphasis id="93C1BE255304DC6EFF6DF9DD82EDF9D0" bold="true" box="[151,384,1575,1601]" pageId="19" pageNumber="470">Material examined.</emphasis>
<materialsCitation id="11DD686A5304DC6EFE7FF9DD8385F9F7" ID-GBIF-Occurrence="3709719374" collectingDate="1887-05" collectingDateMax="2015-08-29" collectingDateMin="1887-05" collectionCode="PMNH, NHMUK" collectorName="Mishkat " country="Pakistan" county="Swat" elevation="1219" location="Kannai" municipality="Kalam" pageId="19" pageNumber="470" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Khyber Pakhtunkhwa">
<collectingCountry id="D9A222A75304DC6EFE7FF9DD8166F9D0" box="[389,523,1575,1601]" name="Pakistan" pageId="19" pageNumber="470">PAKISTAN</collectingCountry>
—
<collectingRegion id="6371ACD55304DC6EFDD7F9DD8072F9D0" box="[557,799,1575,1601]" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="19" pageNumber="470">Khyber Pakhtunkhwa</collectingRegion>
•
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;
<collectingCounty id="486B1ABB5304DC6EFC91F9DD80CBF9D0" box="[875,934,1575,1601]" pageId="19" pageNumber="470">Swat</collectingCounty>
,
<collectingMunicipality id="416EF84D5304DC6EFC4AF9DD8090F9D0" box="[944,1021,1575,1601]" pageId="19" pageNumber="470">Kalam</collectingMunicipality>
,
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;
<date id="D50B44F75304DC6EFB9CF9DD8792F9D3" box="[1126,1279,1575,1602]" pageId="19" pageNumber="470" value="2015-08-29">
<collectingDate id="C54FBD1F5304DC6EFB9CF9DD8792F9D3" box="[1126,1279,1575,1602]" pageId="19" pageNumber="470" value="2015-08-29">29 Aug. 2015</collectingDate>
</date>
;
<collectorName id="0C4007E15304DC6EFAF0F9DD860BF9D0" box="[1290,1382,1575,1601]" pageId="19" pageNumber="470">Mishkat</collectorName>
leg.;
<collectionCode id="C7A4FAF25304DC6EFF6DF9B68385F9F7" box="[151,232,1612,1638]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="19" pageNumber="470" type="Museum">PMNH</collectionCode>
</materialsCitation>
<materialsCitation id="11DD686A5304DC6EFF15F9B687CFF9F7" ID-GBIF-Occurrence="3709719398" box="[239,1186,1611,1638]" collectingDate="1887-05" collectingDateMax="2015-08-29" collectingDateMin="1887-05" collectionCode="PMNH, NHMUK" collectorName="Mishkat " country="Pakistan" county="Swat" elevation="1219" location="Kannai" municipality="Kalam" pageId="19" pageNumber="470" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Khyber Pakhtunkhwa">
•
<specimenCount id="B7B3A9BE5304DC6EFF05F9B18246F9F7" box="[255,299,1611,1638]" pageId="19" pageNumber="470" type="male">3 ♂</specimenCount>
,
<specimenCount id="B7B3A9BE5304DC6EFECCF9B1820CF9F7" box="[310,353,1611,1638]" pageId="19" pageNumber="470" type="female">1 ♀</specimenCount>
;
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,
<location id="A46A34EC5304DC6EFE30F9B181A4F9F4" LSID="urn:lsid:plazi:treatment:291CD3215304DC69FF6DFAA38791FE65:A46A34EC5304DC6EFE30F9B181A4F9F4" box="[458,713,1611,1637]" country="Pakistan" county="Swat" municipality="Kalam" name="Garam Chashma Road" pageId="19" pageNumber="470" stateProvince="Khyber Pakhtunkhwa">Garam Chashma Road</location>
,
<location id="A46A34EC5304DC6EFD2EF9B18065F9F4" LSID="urn:lsid:plazi:treatment:291CD3215304DC69FF6DFAA38791FE65:A46A34EC5304DC6EFD2EF9B18065F9F4" box="[724,776,1611,1637]" country="Pakistan" county="Swat" municipality="Kalam" name="Ruji" pageId="19" pageNumber="470" stateProvince="Khyber Pakhtunkhwa">Ruji</location>
;
<date id="D50B44F75304DC6EFCE9F9B180C0F9F7" box="[787,941,1611,1638]" pageId="19" pageNumber="470" value="2015-08-22">
<collectingDate id="C54FBD1F5304DC6EFCE9F9B180C0F9F7" box="[787,941,1611,1638]" pageId="19" pageNumber="470" value="2015-08-22">22 Aug. 2015</collectingDate>
</date>
,
<collectorName id="0C4007E15304DC6EFC4DF9B1877EF9F4" box="[951,1043,1611,1637]" pageId="19" pageNumber="470">Mishkat</collectorName>
leg.;
<collectionCode id="C7A4FAF25304DC6EFBABF9B687CFF9F7" box="[1105,1186,1612,1638]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="19" pageNumber="470" type="Museum">PMNH</collectionCode>
</materialsCitation>
<materialsCitation id="11DD686A5304DC6EFB53F9B68049F918" ID-GBIF-Occurrence="3709719428" collectingDate="1887-05" collectingDateMax="2015-08-29" collectingDateMin="1887-05" collectionCode="PMNH, NHMUK" collectorName="Mishkat " country="Pakistan" county="Swat" elevation="1219" location="Kannai" municipality="Kalam" pageId="19" pageNumber="470" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Khyber Pakhtunkhwa">
•
<specimenCount id="B7B3A9BE5304DC6EFB43F9B1878BF9F7" box="[1209,1254,1611,1638]" pageId="19" pageNumber="470" type="male">2 ♂</specimenCount>
;
<location id="A46A34EC5304DC6EFB0BF9B1862FF9F4" LSID="urn:lsid:plazi:treatment:291CD3215304DC69FF6DFAA38791FE65:A46A34EC5304DC6EFB0BF9B1862FF9F4" box="[1265,1346,1611,1637]" country="Pakistan" county="Swat" municipality="Kalam" name="Chitral" pageId="19" pageNumber="470" stateProvince="Khyber Pakhtunkhwa">Chitral</location>
,
<location id="A46A34EC5304DC6EFAB7F9B183D6F918" LSID="urn:lsid:plazi:treatment:291CD3215304DC69FF6DFAA38791FE65:A46A34EC5304DC6EFAB7F9B183D6F918" country="Pakistan" county="Swat" municipality="Kalam" name="Khanis Fir" pageId="19" pageNumber="470" stateProvince="Khyber Pakhtunkhwa">Khanis Fir</location>
;
<date id="D50B44F75304DC6EFF3DF995820EF91B" box="[199,355,1647,1674]" pageId="19" pageNumber="470" value="2015-08-23">
<collectingDate id="C54FBD1F5304DC6EFF3DF995820EF91B" box="[199,355,1647,1674]" pageId="19" pageNumber="470" value="2015-08-23">23 Aug. 2015</collectingDate>
</date>
;
<collectorName id="0C4007E15304DC6EFE95F99582A6F918" box="[367,459,1647,1673]" pageId="19" pageNumber="470">Mishkat</collectorName>
leg.;
<collectionCode id="C7A4FAF25304DC6EFDF1F9958130F918" box="[523,605,1647,1673]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="19" pageNumber="470" type="Museum">PMNH</collectionCode>
—
<location id="A46A34EC5304DC6EFD83F9958049F918" LSID="urn:lsid:plazi:treatment:291CD3215304DC69FF6DFAA38791FE65:A46A34EC5304DC6EFD83F9958049F918" box="[633,804,1647,1673]" country="Pakistan" county="Swat" municipality="Kalam" name="Gilgit Baltistan" pageId="19" pageNumber="470" stateProvince="Khyber Pakhtunkhwa">Gilgit Baltistan</location>
</materialsCitation>
<materialsCitation id="11DD686A5304DC6EFCD1F98A82CFF93F" ID-GBIF-Occurrence="3709719417" collectingDate="1887-05" collectingDateMax="2015-08-29" collectingDateMin="1887-05" collectionCode="PMNH, NHMUK" collectorName="Mishkat &amp; Ibad " country="Pakistan" county="Swat" elevation="1219" location="Kannai" municipality="Kalam" pageId="19" pageNumber="470" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Khyber Pakhtunkhwa">
•
<specimenCount id="B7B3A9BE5304DC6EFCC7F9958005F91B" box="[829,872,1647,1674]" pageId="19" pageNumber="470" type="female">1 ♀</specimenCount>
;
<location id="A46A34EC5304DC6EFC8CF99580A9F918" LSID="urn:lsid:plazi:treatment:291CD3215304DC69FF6DFAA38791FE65:A46A34EC5304DC6EFC8CF99580A9F918" box="[886,964,1647,1673]" country="Pakistan" county="Swat" municipality="Kalam" name="Ghizer" pageId="19" pageNumber="470" stateProvince="Khyber Pakhtunkhwa">Ghizer</location>
,
<location id="A46A34EC5304DC6EFC2AF995877BF918" LSID="urn:lsid:plazi:treatment:291CD3215304DC69FF6DFAA38791FE65:A46A34EC5304DC6EFC2AF995877BF918" box="[976,1046,1647,1673]" country="Pakistan" county="Swat" municipality="Kalam" name="Gupis" pageId="19" pageNumber="470" stateProvince="Khyber Pakhtunkhwa">Gupis</location>
,
<location id="A46A34EC5304DC6EFBDBF99587C6F918" LSID="urn:lsid:plazi:treatment:291CD3215304DC69FF6DFAA38791FE65:A46A34EC5304DC6EFBDBF99587C6F918" box="[1057,1195,1647,1673]" country="Pakistan" county="Swat" municipality="Kalam" name="Khalti Lake" pageId="19" pageNumber="470" stateProvince="Khyber Pakhtunkhwa">Khalti Lake</location>
;
<date id="D50B44F75304DC6EFB4CF9958623F918" box="[1206,1358,1647,1673]" pageId="19" pageNumber="470" value="2014-09-06">
<collectingDate id="C54FBD1F5304DC6EFB4CF9958623F918" box="[1206,1358,1647,1673]" pageId="19" pageNumber="470" value="2014-09-06">06 Sep. 2014</collectingDate>
</date>
; Mishkat &amp;
<collectorName id="0C4007E15304DC6EFF25F969827CF93C" box="[223,273,1683,1709]" pageId="19" pageNumber="470">Ibad</collectorName>
leg.;
<collectionCode id="C7A4FAF25304DC6EFEABF96E82CFF93F" box="[337,418,1684,1710]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="19" pageNumber="470" type="Museum">PMNH</collectionCode>
</materialsCitation>
<materialsCitation id="11DD686A5304DC6EFE50F96E8784F943" ID-GBIF-Occurrence="3709719336" collectingDate="1887-05" collectingDateMax="2015-08-29" collectingDateMin="1887-05" collectionCode="PMNH, NHMUK" collectorName=" H. G. Champion " country="Pakistan" county="Swat" elevation="1219" location="Kannai" municipality="Kalam" pageId="19" pageNumber="470" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Khyber Pakhtunkhwa">
•
<specimenCount id="B7B3A9BE5304DC6EFE46F9698285F93F" box="[444,488,1683,1710]" pageId="19" pageNumber="470" type="female">2 ♀</specimenCount>
; “
<collectorName id="0C4007E15304DC6EFDFBF96981DBF93C" box="[513,694,1683,1710]" pageId="19" pageNumber="470">N.W. Pro. India</collectorName>
” [
<collectingRegion id="6371ACD55304DC6EFD34F96980A6F93C" box="[718,971,1682,1709]" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="19" pageNumber="470">Khyber Pakhtunkhwa</collectingRegion>
],
<location id="A46A34EC5304DC6EFC24F96E8600F93C" LSID="urn:lsid:plazi:treatment:291CD3215304DC69FF6DFAA38791FE65:A46A34EC5304DC6EFC24F96E8600F93C" box="[990,1389,1683,1710]" country="Pakistan" county="Swat" municipality="Kalam" name="Hazara Dist. Lower Kaghan Valley" pageId="19" pageNumber="470" stateProvince="Khyber Pakhtunkhwa">Hazara Dist. Lower Kaghan Valley</location>
; alt.
<quantity id="664DCFD25304DC6EFF6DF94D8242F940" box="[151,303,1719,1746]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" metricValueMax="1.524" metricValueMin="0.9144" pageId="19" pageNumber="470" unit="ft" value="4000.0" valueMax="5000.0" valueMin="3000.0">
<elevation id="2A9885045304DC6EFF6DF94D8242F940" box="[151,303,1719,1746]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" metricValueMax="1.524" metricValueMin="0.9144" pageId="19" pageNumber="470" unit="ft" value="4000.0" valueMax="5000.0" valueMin="3000.0">3000–5000 ft</elevation>
</quantity>
;
<date id="D50B44F75304DC6EFEC0F94D82BCF940" box="[314,465,1719,1746]" pageId="19" pageNumber="470" value="1927-05-26">
<collectingDate id="C54FBD1F5304DC6EFEC0F94D82BCF940" box="[314,465,1719,1746]" pageId="19" pageNumber="470" value="1927-05-26">26 May 1927</collectingDate>
</date>
;
<collectorName id="0C4007E15304DC6EFE21F94D81E0F940" box="[475,653,1719,1745]" pageId="19" pageNumber="470">H.G. Champion</collectorName>
leg.;
<collectionCode id="C7A4FAF25304DC6EFD30F9428059F943" box="[714,820,1720,1746]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="470" type="Museum">NHMUK</collectionCode>
(
<collectorName id="0C4007E15304DC6EFCB8F94D8702F940" box="[834,1135,1719,1745]" pageId="19" pageNumber="470">H.G. Champion Collection</collectorName>
1953-156)
</materialsCitation>
<materialsCitation id="11DD686A5304DC6EFB15F94281BEF8AF" ID-GBIF-Occurrence="3709719400" collectingDate="1887-05" collectingDateMax="2015-08-29" collectingDateMin="1887-05" collectionCode="PMNH, NHMUK" collectorName="Oburthur" country="Pakistan" county="Swat" elevation="1219" location="Kannai" municipality="Kalam" pageId="19" pageNumber="470" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Khyber Pakhtunkhwa">
•
<specimenCount id="B7B3A9BE5304DC6EFB05F94D8646F943" box="[1279,1323,1719,1746]" pageId="19" pageNumber="470" type="male">2 ♂</specimenCount>
; Kashmir, Goorais; alt.
<quantity id="664DCFD25304DC6EFED2F9218216F964" box="[296,379,1755,1781]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="19" pageNumber="470" unit="ft" value="7000.0">
<elevation id="2A9885045304DC6EFED2F9218216F964" box="[296,379,1755,1781]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="19" pageNumber="470" unit="ft" value="7000.0">7000 ft</elevation>
</quantity>
;
<date id="D50B44F75304DC6EFE7DF9268291F964" box="[391,508,1755,1782]" pageId="19" pageNumber="470" value="1887-05">
<collectingDate id="C54FBD1F5304DC6EFE7DF9268291F964" box="[391,508,1755,1782]" pageId="19" pageNumber="470" value="1887-05">May 1887</collectingDate>
</date>
; [genitalia extacted and attached to label]; ex coll.
<collectorName id="0C4007E15304DC6EFBCEF92187F3F964" box="[1076,1182,1755,1781]" pageId="19" pageNumber="470">Oburthür</collectorName>
; Co-type green circled
<typeStatus id="7E0EDC955304DC6EFF6DF8FA83AAF88B" box="[151,199,1792,1818]" pageId="19" pageNumber="470">type</typeStatus>
label;
<taxonomicName id="66B519B45304DC6EFEEEF9058057F888" authority="Bates, H. E. Andrewes" authorityName="Bates, H. E. Andrewes" authorityYear="1889" box="[276,826,1791,1818]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis id="93C1BE255304DC6EFEEEF9058156F888" box="[276,571,1791,1817]" italics="true" pageId="19" pageNumber="470">Chlaenius anchomenoides</emphasis>
Bates, H.E. Andrewes
</taxonomicName>
det.
<emphasis id="93C1BE255304DC6EFC8BF8FA80E0F886" bold="true" box="[881,909,1792,1815]" pageId="19" pageNumber="470">①</emphasis>
;
<taxonomicName id="66B519B45304DC6EFC66F9058601F888" authority="Mishkat Ullah 2017" authorityName="Mishkat Ullah" authorityYear="2017" box="[924,1388,1791,1818]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="agilis" subGenus="Lithochlaenius">
<emphasis id="93C1BE255304DC6EFC66F9058763F888" box="[924,1038,1791,1817]" italics="true" pageId="19" pageNumber="470">Chlaenius</emphasis>
(
<emphasis id="93C1BE255304DC6EFBE4F8FA8758F888" box="[1054,1077,1792,1817]" italics="true" pageId="19" pageNumber="470">L.</emphasis>
)
<emphasis id="93C1BE255304DC6EFBBCF90587E9F888" box="[1094,1156,1791,1817]" italics="true" pageId="19" pageNumber="470">agilis</emphasis>
Mishkat Ullah 2017
</taxonomicName>
det.
<emphasis id="93C1BE255304DC6EFF6DF8DE83DEF8AA" bold="true" box="[151,179,1828,1851]" pageId="19" pageNumber="470">②</emphasis>
;
<collectionCode id="C7A4FAF25304DC6EFF38F8DE8241F8AF" box="[194,300,1828,1854]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="470" type="Museum">NHMUK</collectionCode>
(
<collectorName id="0C4007E15304DC6EFEC6F8D9810FF8AC" box="[316,610,1827,1853]" pageId="19" pageNumber="470">H.E. Andrewes Collection</collectorName>
1945-97)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5304DC6EFF3DF8BD8251F85C" pageId="19" pageNumber="470" type="description">
<paragraph id="A10A62375304DC6EFF3DF8BD8251F85C" blockId="19.[151,1437,1575,2033]" pageId="19" pageNumber="470">
<emphasis id="93C1BE255304DC6EFF3DF8BD8213F8F0" bold="true" box="[199,382,1863,1889]" pageId="19" pageNumber="470">Measurements.</emphasis>
Male (n=5): TL=
<quantity id="664DCFD25304DC6EFDABF8BD81AAF8F3" box="[593,711,1863,1890]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="19" pageNumber="470" unit="mm" value="15.4">15.40 mm</quantity>
, HW=
<quantity id="664DCFD25304DC6EFCE1F8BD80EEF8F3" box="[795,899,1863,1890]" metricMagnitude="-3" metricUnit="m" metricValue="2.79" pageId="19" pageNumber="470" unit="mm" value="2.79">2.79 mm</quantity>
, PL=
<quantity id="664DCFD25304DC6EFC33F8BD875CF8F3" box="[969,1073,1863,1890]" metricMagnitude="-3" metricUnit="m" metricValue="2.87" pageId="19" pageNumber="470" unit="mm" value="2.87">2.87 mm</quantity>
, PW=
<quantity id="664DCFD25304DC6EFB7BF8BD8785F8F3" box="[1153,1256,1863,1890]" metricMagnitude="-3" metricUnit="m" metricValue="3.26" pageId="19" pageNumber="470" unit="mm" value="3.26">3.26 mm</quantity>
, EL=
<quantity id="664DCFD25304DC6EFACAF8BD86F5F8F3" box="[1328,1432,1863,1890]" metricMagnitude="-3" metricUnit="m" metricValue="9.61" pageId="19" pageNumber="470" unit="mm" value="9.61">9.61 mm</quantity>
, EW=
<quantity id="664DCFD25304DC6EFF27F891822AF817" box="[221,327,1899,1926]" metricMagnitude="-3" metricUnit="m" metricValue="2.77" pageId="19" pageNumber="470" unit="mm" value="2.77">2.77 mm</quantity>
. Ratios PL/PW= 0.87, HW/PW= 0.85 and EL/EW= 3.46. Female (n=5): TL=
<quantity id="664DCFD25304DC6EFB22F8918622F817" box="[1240,1359,1899,1926]" metricMagnitude="-2" metricUnit="m" metricValue="1.612" pageId="19" pageNumber="470" unit="mm" value="16.12">16.12 mm</quantity>
, HW=
<quantity id="664DCFD25304DC6EFF6DF8758393F83B" box="[151,254,1935,1962]" metricMagnitude="-3" metricUnit="m" metricValue="2.92" pageId="19" pageNumber="470" unit="mm" value="2.92">2.92 mm</quantity>
, PL=
<quantity id="664DCFD25304DC6EFEB8F87582C4F83B" box="[322,425,1935,1962]" metricMagnitude="-3" metricUnit="m" metricValue="2.99" pageId="19" pageNumber="470" unit="mm" value="2.99">2.99 mm</quantity>
, PW=
<quantity id="664DCFD25304DC6EFE0CF8758130F83B" box="[502,605,1935,1962]" metricMagnitude="-3" metricUnit="m" metricValue="3.39" pageId="19" pageNumber="470" unit="mm" value="3.39">3.39 mm</quantity>
, EL=
<quantity id="664DCFD25304DC6EFD58F8758064F83B" box="[674,777,1935,1962]" metricMagnitude="-3" metricUnit="m" metricValue="9.99" pageId="19" pageNumber="470" unit="mm" value="9.99">9.99 mm</quantity>
, EW=
<quantity id="664DCFD25304DC6EFCA2F87580ADF83B" box="[856,960,1935,1962]" metricMagnitude="-3" metricUnit="m" metricValue="2.99" pageId="19" pageNumber="470" unit="mm" value="2.99">2.99 mm</quantity>
. Ratios PL/PW= 0.88, HW/PW= 0.85 and EL/EW= 3.33.
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5304DC69FF3DF82D8277FEF5" lastPageId="20" lastPageNumber="471" pageId="19" pageNumber="470" type="diagnosis">
<paragraph id="A10A62375304DC69FF3DF82D8277FEF5" blockId="19.[151,1437,1575,2033]" lastBlockId="20.[151,1437,150,501]" lastPageId="20" lastPageNumber="471" pageId="19" pageNumber="470">
<emphasis id="93C1BE255304DC6EFF3DF82D822FF860" bold="true" box="[199,322,2007,2033]" pageId="19" pageNumber="470">Diagnosis.</emphasis>
Body elongated. Head black with slight metallic violet greenish tinge, smooth moderately covered with very minute punctures. Antennal article Imore ovid, labrum slightly emarginate. Mandibles, antenna article III and legs black; palpi, antennalarticles I–II and IV–IX reddish brown. Pronotum black with metallic violet greenish tinge smooth, slightly transverse rugose, basal fovea deep, sparsely setiferous punctate. Elytra black, humeral angle obtuse, striae deep and punctate, intervals convex raised with wide setiferous punctures throughout; pubescence dense from interval VI–VIII toward apical 1/3
<superScript id="56C0CF7F5303DC69FD18FEDC819DFEA5" attach="none" box="[738,752,294,308]" fontSize="6" pageId="20" pageNumber="471">rd</superScript>
. Legs black and slender. Ventral surface black and pubescent throughout.
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5303DC69FF3DFE9480EDFE40" pageId="20" pageNumber="471" type="discussion">
<paragraph id="A10A62375303DC69FF3DFE9480EDFE40" blockId="20.[151,1437,150,501]" pageId="20" pageNumber="471">
<emphasis id="93C1BE255303DC69FF3DFE948222FE19" bold="true" box="[199,335,366,392]" pageId="20" pageNumber="471">Comments.</emphasis>
This species was collected from the same habitat and locality inhabited by
<taxonomicName id="66B519B45303DC69FB4CFE948104FE3C" authority="(Grundmann, 1955)" authorityName="J. Schmidt" authorityYear="2007" baseAuthorityName="Grundmann" baseAuthorityYear="1955" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="kashmiricus" subGenus="Stenochlaenius">
<emphasis id="93C1BE255303DC69FB4CFE948645FE19" box="[1206,1320,366,392]" italics="true" pageId="20" pageNumber="471">Chlaenius</emphasis>
(
<emphasis id="93C1BE255303DC69FAC0FE948385FE3D" italics="true" pageId="20" pageNumber="471">Stenochlaenius</emphasis>
)
<emphasis id="93C1BE255303DC69FF03FE6882EDFE3D" box="[249,384,402,428]" italics="true" pageId="20" pageNumber="471">kashmiricus</emphasis>
(
<bibRefCitation id="C5241FC65303DC69FE75FE68810DFE3C" author="Grundmann, E." box="[399,608,402,429]" pageId="20" pageNumber="471" pagination="53 - 55" refId="ref26317" refString="Grundmann, E. (1955) Ein neuer Stenochlaenius aus Kaschmir. Col., Carabidae, Chlaeniinae. Koleopterologische Rundschau, 33, 53 - 55." type="journal article" year="1955">Grundmann, 1955</bibRefCitation>
)
</taxonomicName>
. Detailed comments can be found in the work of
<bibRefCitation id="C5241FC65303DC69FB68FE688627FE3D" author="Liu, Y. &amp; Kavanaugh, D. H. &amp; Shi, H. &amp; Liang, H." box="[1170,1354,402,429]" pageId="20" pageNumber="471" pagination="15 - 52" refId="ref27203" refString="Liu, Y., Kavanaugh, D. H., Shi, H. &amp; Liang, H. (2011) A Key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species. ZooKeys, 128, 15 - 52. https: // doi. org / 10.3897 / zookeys. 128.1804" type="journal article" year="2011">Liu et al. (2011)</bibRefCitation>
. In this current study, it was reported from Swat, Chitral and Ghizer areas.
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5303DC69FF3DFE208791FE65" box="[199,1276,474,501]" pageId="20" pageNumber="471" type="distribution">
<paragraph id="A10A62375303DC69FF3DFE208791FE65" blockId="20.[151,1437,150,501]" box="[199,1276,474,501]" pageId="20" pageNumber="471">
<emphasis id="93C1BE255303DC69FF3DFE20820CFE65" bold="true" box="[199,353,474,500]" pageId="20" pageNumber="471">Distribution.</emphasis>
<collectingCountry id="D9A222A75303DC69FE9DFE20829EFE65" box="[359,499,474,500]" name="Afghanistan" pageId="20" pageNumber="471">Afghanistan</collectingCountry>
,
<collectingCountry id="D9A222A75303DC69FE04FE20812CFE65" box="[510,577,474,500]" name="China" pageId="20" pageNumber="471">China</collectingCountry>
(
<collectingRegion id="6371ACD55303DC69FDAAFE2081C2FE65" box="[592,687,474,500]" country="China" name="Sichuan" pageId="20" pageNumber="471">Sichuan</collectingRegion>
;
<collectingRegion id="6371ACD55303DC69FD43FE21807EFE64" box="[697,787,475,501]" country="China" name="Yunnan" pageId="20" pageNumber="471">Yunnan</collectingRegion>
),
<collectingCountry id="D9A222A75303DC69FCDCFE2080E8FE65" box="[806,901,474,500]" name="Pakistan" pageId="20" pageNumber="471">Pakistan</collectingCountry>
and
<collectingCountry id="D9A222A75303DC69FC46FE208092FE65" box="[956,1023,474,500]" name="Nepal" pageId="20" pageNumber="471">Nepal</collectingCountry>
(Anichtchenko 2017).
</paragraph>
</subSubSection>
</treatment>
</document>
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