532 lines
68 KiB
XML
532 lines
68 KiB
XML
<document id="281C6A91AB607FCAE34A4A41B82E3D2C" ID-CLB-Dataset="53922" ID-DOI="10.11646/zootaxa.5149.1.1" ID-GBIF-Dataset="caecca9d-72a3-4d9a-836d-9ac3e0ded512" ID-ISSN="1175-5326" ID-Zenodo-Dep="6605961" ID-ZooBank="BF98000F-04F9-4D22-9D93-4C3A2418D7EF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654152510384" checkinUser="plazi" docAuthor="Probst, Rodolfo Da Silva & Brandão, Carlos Roberto Ferreira" docDate="2022" docId="F62B8F24FFC4E311FF0FFF0C8494D9AF" docLanguage="en" docName="zootaxa.5149.1.1.pdf" docOrigin="Zootaxa 5149 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Basiceros tumucumaquensis Probst & Brandao 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="69" masterDocId="0A12F75CFF84E355FF98FF9B811BDB5E" masterDocTitle="A taxonomic revision of the dirt ants, Basiceros Schulz, 1906 (Hymenoptera, Formicidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="65" updateTime="1702002095603" updateUser="plazi">
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<mods:title id="4CE0ADD2E02D99486116BD7D9F8E2CF6">A taxonomic revision of the dirt ants, Basiceros Schulz, 1906 (Hymenoptera, Formicidae)</mods:title>
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<mods:name id="2D74CE193863B6822F98DFF9AE26B5D6" type="personal">
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<mods:namePart id="97ED395BB54FB51170926246443F9010">Probst, Rodolfo Da Silva</mods:namePart>
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<mods:nameIdentifier id="8455FD18654ACF0204BC391613FCD62C" type="email">probstrodolfo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4472ED832536EB395A53ED4622FE3378">Brandão, Carlos Roberto Ferreira</mods:namePart>
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<mods:nameIdentifier id="E7309AF7665AF8D5F0F3B74A20DB5A70" type="email">probstrodolfo@gmail.com</mods:nameIdentifier>
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<mods:title id="6D704AF987784B4E13D330C28F1C8420">Zootaxa</mods:title>
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<mods:part id="D512632249AD34BE79D967C2FEC0F723">
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<mods:date id="FB2FE0376168A1B6C6C6778F1352417C">2022</mods:date>
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<mods:number id="6A8DAB98BE4EFC336F465C7B3F2135CC">2022-06-02</mods:number>
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<mods:number id="9850064B3854BD31437F402149197181">5149</mods:number>
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<treatment id="F62B8F24FFC4E311FF0FFF0C8494D9AF" ID-DOI="http://doi.org/10.5281/zenodo.7636279" ID-GBIF-Taxon="209289995" ID-Zenodo-Dep="7636279" LSID="urn:lsid:plazi:treatment:F62B8F24FFC4E311FF0FFF0C8494D9AF" httpUri="http://treatment.plazi.org/id/F62B8F24FFC4E311FF0FFF0C8494D9AF" lastPageId="68" lastPageNumber="69" pageId="64" pageNumber="65">
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<subSubSection id="36986DB9FFC4E315FF0FFF0C825EDBEF" box="[151,837,151,177]" pageId="64" pageNumber="65" type="nomenclature">
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<paragraph id="7E3D3E32FFC4E315FF0FFF0C825EDBEF" blockId="64.[151,837,151,213]" box="[151,837,151,177]" pageId="64" pageNumber="65">
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<heading id="2575895EFFC4E315FF0FFF0C825EDBEF" bold="true" box="[151,837,151,177]" fontSize="11" level="2" pageId="64" pageNumber="65" reason="4">
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<emphasis id="4CF6E220FFC4E315FF0FFF0C825EDBEF" bold="true" box="[151,837,151,177]" pageId="64" pageNumber="65">
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<taxonomicName id="B98245B1FFC4E315FF0FFF0C83FBDBEF" authority="Probst & Brandao" authorityName="Probst & Brandao" authorityYear="2022" box="[151,736,151,177]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="tumucumaquensis" status="sp. nov.">
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<emphasis id="4CF6E220FFC4E315FF0FFF0C80E9DBEF" bold="true" box="[151,498,151,177]" italics="true" pageId="64" pageNumber="65">Basiceros tumucumaquensis</emphasis>
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Probst & Brandão
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</taxonomicName>
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<taxonomicNameLabel id="57C55F5BFFC4E315FD7FFF0C825EDBEF" box="[743,837,151,177]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="36986DB9FFC4E315FF0FFF20804ADB8B" box="[151,337,187,213]" pageId="64" pageNumber="65" type="description">
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<paragraph id="7E3D3E32FFC4E315FF0FFF20804ADB8B" blockId="64.[151,837,151,213]" box="[151,337,187,213]" pageId="64" pageNumber="65">
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(
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<figureCitation id="E6B922B7FFC4E315FF07FF20803BDB8B" box="[159,288,187,213]" captionStart-0="FIGURE 26" captionStart-1="FIGURE 27" captionStart-2="FIGURE 28" captionStartId-0="65.[151,250,1148,1172]" captionStartId-1="66.[151,250,1712,1736]" captionStartId-2="67.[151,250,1649,1673]" captionTargetBox-0="[151,1436,345,1125]" captionTargetBox-1="[155,1433,489,1689]" captionTargetBox-2="[167,1421,525,1626]" captionTargetId-0="figure-81@65.[151,1436,345,1125]" captionTargetId-1="figure-129@66.[155,1433,489,1689]" captionTargetId-2="figure-173@67.[167,1421,525,1626]" captionTargetPageId-0="65" captionTargetPageId-1="66" captionTargetPageId-2="67" captionText-0="FIGURE 26. Basiceros tumucumaquensis sp. nov. holotype worker (MZSP, CASENT0647157, Brazil: Amapá); mandibles in A. dorsal view, B. oblique view, C. lateral view, D. ventral view, highlighting ventrolabrum, E. dorsal view, F. lateral view, G. full-face view, H. oblique view of head and pronotum. Scale bars: A–D = 0.2 mm, E–F = 1 mm, G–H = 0.5 mm." captionText-1="FIGURE 27. Basiceros tumucumaquensis sp. nov. paratype worker (MZSP, CASENT0647158, Brazil: Amapá); A. full-face view, D. dorsal view, E. lateral view. Labrum from non-type worker (MZSP, Brazil: Rondônia); B. dorsal view and C. ventral view; lo: lateral lobe, black arrow: specialized seta on anterior margin. Crossed arrows give the orientation: a, anterior; p, posterior. Scale bars: A = 0.5 mm, D–E =1 mm, B–C = 0.1 mm." captionText-2="FIGURE 28. Basiceros tumucumaquensis sp. nov. paratype male (MZSP, CASENT0647167, Brazil: Amapá); A. head and antenna, B. lateral view, C. full-face view, D. dorsal view. Scale bars: A–B, D =1 mm, C = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6606025" figureDoi-1="http://doi.org/10.5281/zenodo.6606027" figureDoi-2="http://doi.org/10.5281/zenodo.6606031" httpUri-0="https://zenodo.org/record/6606025/files/figure.png" httpUri-1="https://zenodo.org/record/6606027/files/figure.png" httpUri-2="https://zenodo.org/record/6606031/files/figure.png" pageId="64" pageNumber="65">Figs 26–28</figureCitation>
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,
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<figureCitation id="E6B922B7FFC4E315FEB4FF208052DB8B" box="[300,329,187,213]" captionStart="FIGURE 32" captionStartId="71.[151,250,2006,2030]" captionTargetBox="[261,1327,1118,1982]" captionTargetId="figure-37@71.[261,1327,1118,1982]" captionTargetPageId="71" captionText="FIGURE 32. Distribution map of Basiceros scambognathus and B. tumucumaquensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6606041" httpUri="https://zenodo.org/record/6606041/files/figure.png" pageId="64" pageNumber="65">32</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="36986DB9FFC4E315FF0FFE98806CDA8F" pageId="64" pageNumber="65" type="materials_examined">
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<paragraph id="7E3D3E32FFC4E315FF0FFE98806CDA8F" blockId="64.[151,1437,259,2014]" pageId="64" pageNumber="65">
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<emphasis id="4CF6E220FFC4E315FF0FFE988053DA43" bold="true" box="[151,328,259,285]" pageId="64" pageNumber="65">Type material.</emphasis>
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<materialsCitation id="CEEA346FFFC4E315FEC9FE988503DA1F" ID-GBIF-Occurrence="4029332323" collectingDate="2018-12-25" collectionCode="PNMT, R, MZSP" collectorName="R. Probst & O. Guilherme" country="Brazil" elevation="0" latitude="1.242333" location="Amapa" longLatPrecision="14" longitude="-52.419777" pageId="64" pageNumber="65" specimenCode="CASENT0647157" specimenCount="1" stateProvince="Amapa" typeStatus="holotype">
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<emphasis id="4CF6E220FFC4E315FEC9FE98830BDA43" box="[337,528,259,285]" italics="true" pageId="64" pageNumber="65">
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<typeStatus id="A1398090FFC4E315FEC9FE9880A3DA43" box="[337,440,259,285]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
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worker
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</emphasis>
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:
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<collectingCountry id="06957EA2FFC4E315FDBAFE988397DA43" box="[546,652,259,285]" name="Brazil" pageId="64" pageNumber="65">BRAZIL</collectingCountry>
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:
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<collectingRegion id="BC46F0D0FFC4E315FD0FFE9883F0DA43" box="[663,747,259,285]" country="Brazil" name="Amapa" pageId="64" pageNumber="65">Amapá</collectingRegion>
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,
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<collectionCode id="1893A6F7FFC4E315FD60FE9F825CDA40" box="[760,839,260,286]" pageId="64" pageNumber="65">PNMT</collectionCode>
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,
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±
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<quantity id="B97A93D7FFC4E315FBF9FE98858FDA43" box="[1121,1172,259,285]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="64" pageNumber="65" unit="m" value="10.0">10m</quantity>
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,
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<quantity id="B97A93D7FFC4E315FB3AFE9885FEDA43" box="[1186,1253,259,285]" metricMagnitude="2" metricUnit="m" metricValue="1.3199999999999998" pageId="64" pageNumber="65" unit="m" value="132.0">132m</quantity>
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</elevation>
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,
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<date id="0A3C18F2FFC4E315FB6AFE98846EDA40" box="[1266,1397,259,286]" pageId="64" pageNumber="65" value="2018-12-25">
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<collectingDate id="1A78E11AFFC4E315FB6AFE98846EDA40" box="[1266,1397,259,286]" pageId="64" pageNumber="65" value="2018-12-25">25.xii.2018</collectingDate>
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</date>
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,
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<collectionCode id="1893A6F7FFC4E315FA1BFE9F8482DA40" box="[1411,1433,260,286]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="64" pageNumber="65">R</collectionCode>
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.
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<collectorName id="D3775BE4FFC4E315FF0FFEBC81C4DA1F" box="[151,223,295,321]" pageId="64" pageNumber="65">Probst</collectorName>
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&
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<collectorName id="D3775BE4FFC4E315FE91FEBC80BCDA1F" box="[265,423,295,321]" pageId="64" pageNumber="65">O. Guilherme</collectorName>
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cols. (
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<emphasis id="4CF6E220FFC4E315FE61FEB3839EDA1F" box="[505,645,295,321]" italics="true" pageId="64" pageNumber="65">
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<collectionCode id="1893A6F7FFC4E315FE61FEB38316DA1F" box="[505,525,296,321]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="64" pageNumber="65">R</collectionCode>
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.
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<collectorName id="D3775BE4FFC4E315FD84FEBC839EDA1F" box="[540,645,295,321]" pageId="64" pageNumber="65">S. Probst</collectorName>
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</emphasis>
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#678) [
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<collectionCode id="1893A6F7FFC4E315FD79FEBC8236DA1F" box="[737,813,295,321]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="64" pageNumber="65" type="Museum">MZSP</collectionCode>
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,
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<specimenCode id="2E249649FFC4E315FCA4FEBD850ADA1F" box="[828,1041,294,321]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT0647157</specimenCode>
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]
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</materialsCitation>
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.
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<materialsCitation id="CEEA346FFFC4E315FBBEFEB3820DDAF3" ID-GBIF-Occurrence="4029332313" collectingDate="2018-12-25" collectionCode="MZSP, MPEG, DZUP, MCZ, JTL" country="Brazil" elevation="0" latitude="1.242333" location="Amapa" longLatPrecision="14" longitude="-52.419777" pageId="64" pageNumber="65" specimenCode="CASENT0647158, CASENT0647159, CASENT0647160, CASENT0647161, CASENT0647162, CASENT0647163, CASENT0647164, CASENT0647165, CASENT0647166" specimenCount="9" specimenCount-worker="9" stateProvince="Amapa" typeStatus="paratype">
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<emphasis id="4CF6E220FFC4E315FBBEFEB385EADA1F" box="[1062,1265,295,321]" italics="true" pageId="64" pageNumber="65">
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<typeStatus id="A1398090FFC4E315FBBEFEB38596DA1F" box="[1062,1165,296,321]" pageId="64" pageNumber="65" type="paratype">Paratype</typeStatus>
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workers
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</emphasis>
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: Same data as holotype [
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<collectionCode id="1893A6F7FFC4E315FE9DFED0804ADA3B" box="[261,337,331,357]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="64" pageNumber="65" type="Museum">MZSP</collectionCode>
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,
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<specimenCount id="6884F5BBFFC4E315FEC4FED080D6DA3B" box="[348,461,331,357]" pageId="64" pageNumber="65" type="worker">4 workers</specimenCount>
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,
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<specimenCode id="2E249649FFC4E315FE4FFED083B3DA38" box="[471,680,331,358]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT0647158</specimenCode>
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,
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<specimenCode id="2E249649FFC4E315FD2BFED0829FDA38" box="[691,900,331,358]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT0647159</specimenCode>
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,
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<specimenCode id="2E249649FFC4E315FC17FED0857ADA3B" box="[911,1121,331,357]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT0647160</specimenCode>
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,
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<specimenCode id="2E249649FFC4E315FBF3FED08426DA3B" box="[1131,1341,331,357]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT0647161</specimenCode>
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;
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<collectionCode id="1893A6F7FFC4E315FADFFED08483DA3B" box="[1351,1432,331,357]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="64" pageNumber="65">MPEG</collectionCode>
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,
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<specimenCount id="6884F5BBFFC4E315FF0FFEF4800BDAD7" box="[151,272,367,393]" pageId="64" pageNumber="65" type="worker">2 workers</specimenCount>
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,
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<specimenCode id="2E249649FFC4E315FEB9FEF480E8DAD7" box="[289,499,367,393]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT0647162</specimenCode>
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,
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<specimenCode id="2E249649FFC4E315FD9DFEF483CCDAD7" box="[517,727,367,393]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT0647163</specimenCode>
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;
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<collectionCode id="1893A6F7FFC4E315FD71FEEB8229DAD4" box="[745,818,368,394]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="64" pageNumber="65" type="University or college">DZUP</collectionCode>
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,
|
||
<specimenCount id="6884F5BBFFC4E315FCDBFEEB82D1DAD7" box="[835,970,367,394]" pageId="64" pageNumber="65" type="worker">one worker</specimenCount>
|
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,
|
||
<specimenCode id="2E249649FFC4E315FC44FEF485B5DAD7" box="[988,1198,367,393]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT0647164</specimenCode>
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;
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<collectionCode id="1893A6F7FFC4E315FB58FEF4841BDAD7" box="[1216,1280,367,393]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="64" pageNumber="65" type="Museum">MCZ</collectionCode>
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,
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<specimenCount id="6884F5BBFFC4E315FA89FEEB8483DAD7" box="[1297,1432,367,394]" pageId="64" pageNumber="65" type="worker">one worker</specimenCount>
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,
|
||
<specimenCode id="2E249649FFC4E315FF0FFE088071DAF0" box="[151,362,403,430]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT0647165</specimenCode>
|
||
;
|
||
<collectionCode id="1893A6F7FFC4E315FEECFE0F80BFDAF0" box="[372,420,404,430]" pageId="64" pageNumber="65">JTL</collectionCode>
|
||
,
|
||
<specimenCount id="6884F5BBFFC4E315FE37FE0F8334DAF3" box="[431,559,403,430]" pageId="64" pageNumber="65" type="worker">one worker</specimenCount>
|
||
,
|
||
<specimenCode id="2E249649FFC4E315FDA2FE098214DAF3" box="[570,783,402,429]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT0647166</specimenCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="CEEA346FFFC4E315FCB8FE0F8068DA8F" ID-GBIF-Occurrence="4029332437" collectingDate="2018-12-25" collectionCode="MZSP" country="Brazil" elevation="0" latitude="1.242333" location="Amapa" longLatPrecision="14" longitude="-52.419777" pageId="64" pageNumber="65" specimenCode="CASENT0647167" specimenCount="1" specimenCount-male="1" stateProvince="Amapa" typeStatus="paratype">
|
||
<emphasis id="4CF6E220FFC4E315FCB8FE0F82D8DAF3" box="[800,963,403,429]" italics="true" pageId="64" pageNumber="65">
|
||
<typeStatus id="A1398090FFC4E315FCB8FE0F829CDAF3" box="[800,903,404,429]" pageId="64" pageNumber="65" type="paratype">Paratype</typeStatus>
|
||
male
|
||
</emphasis>
|
||
: Same data as holotype [
|
||
<specimenCount id="6884F5BBFFC4E315FB41FE0F845FDAF3" box="[1241,1348,403,430]" pageId="64" pageNumber="65" type="male">one male</specimenCount>
|
||
,
|
||
<collectionCode id="1893A6F7FFC4E315FAD7FE088483DAF3" box="[1359,1432,403,429]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="64" pageNumber="65" type="Museum">MZSP</collectionCode>
|
||
,
|
||
<specimenCode id="2E249649FFC4E315FF0FFE2D8076DA8F" box="[151,365,438,465]" collectionCode="CASENT" pageId="64" pageNumber="65">CASENT0647167</specimenCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36986DB9FFC4E317FF5FFE4080E8DA1F" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65" type="diagnosis">
|
||
<paragraph id="7E3D3E32FFC4E315FF5FFE4083B2D963" blockId="64.[151,1437,259,2014]" pageId="64" pageNumber="65">
|
||
<emphasis id="4CF6E220FFC4E315FF5FFE408059DAAB" bold="true" box="[199,322,475,501]" pageId="64" pageNumber="65">Diagnosis.</emphasis>
|
||
Mandibles bizarre, apical portion strongly curved ventrally, basidorsal sulcus present, slightly transversal to mandibular dorsa, basal angle translucent. Clypeus densely covered with decumbent squamiform hairs. Head dorsum rugulose, with long clavate hairs.
|
||
</paragraph>
|
||
<paragraph id="7E3D3E32FFC4E315FF5FFDDC8556D84B" blockId="64.[151,1437,259,2014]" pageId="64" pageNumber="65">
|
||
<emphasis id="4CF6E220FFC4E315FF5FFDDC807FD93F" bold="true" box="[199,356,583,609]" pageId="64" pageNumber="65">Descriptions.</emphasis>
|
||
<emphasis id="4CF6E220FFC4E315FEF3FDDC80A7D93F" box="[363,444,583,609]" italics="true" pageId="64" pageNumber="65">Worker</emphasis>
|
||
(n=5, range downward from size of smallest to biggest individual. Includes
|
||
<typeStatus id="A1398090FFC4E315FA85FDDC8499D93F" box="[1309,1410,583,609]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
,
|
||
<specimenCount id="6884F5BBFFC4E315FA16FDDC801ED9D8" pageId="64" pageNumber="65" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
|
||
, and a non-type each from Amapá and Rondônia). HL 1.28–1.44, HL2 1.22–1.47, HW1 1.04–1.24, MdL 0.48–0.58, SL1 0.97–0.99, SL2 1.02–1.08, PDL 0.09–0.11, A3L 0.03–0.04, AFL
|
||
<geoCoordinate id="1BB658F5FFC4E315FBB4FD148577D9F4" box="[1068,1132,655,682]" degrees="0.385" direction="north" orientation="latitude" pageId="64" pageNumber="65" precision="55" value="0.385">0.385</geoCoordinate>
|
||
<geoCoordinate id="1BB658F5FFC4E315FBF4FD1485A1D9F4" box="[1132,1210,655,682]" degrees="0.409" direction="west" orientation="longitude" pageId="64" pageNumber="65" precision="55" value="-0.409">–0.409</geoCoordinate>
|
||
, FuL 0.95–1.02, EL 0.09–0.12, EW 0.09, ML 1.556–1.7, MfL 1.28–1.35, MtL 1.0–1.06,
|
||
<collectingCountry id="06957EA2FFC4E315FC38FD2F82DFD990" box="[928,964,692,718]" name="Philippines" pageId="64" pageNumber="65">PH</collectingCountry>
|
||
0.32–0.35, PL 0.74–0.76, PW 0.29–0.33, PPL 0.51–0.54, PPW 0.51–0.538, GL 1.54–1.71, GW 1.04–1.12, TL 6.2–6.63, CI 81–85, CS 1.16–1.31, MCI 40– 42, SI 84–97, ESI 9–11, SAI2 3011–3347, EI1 0.15–0.16, MFI 81–91, PTI 213–234.
|
||
</paragraph>
|
||
<paragraph id="7E3D3E32FFC4E315FF5FFC848064DE50" blockId="64.[151,1437,259,2014]" pageId="64" pageNumber="65">Medium size comparatively. Coloration reddish-brown to brown; appendages lighter. Mandible dorsa covered by minute sparse piligerous punctuation, bearing erect to suberect setae; basal portion with clavate pilosity suberect to subdecumbent; mandibular apex with medium yellowish setae. Basimandibular setae present, narrow and erect. Clavate suberect hairs on the dorsum of each stipe. Anterior margin of labrum with long filiform hairs. Clypeal dorsum densely covered with decumbent squamiform hairs. Head with basal layer of hairs with similar morphology as clypeal pilosity (although longer), sparse and located at margin of sculpture; erect and slightly clavate pilosity as follows: one pair on frons, close to the median head depression, one pair on the temple, around the sulcus of vertexal margin, a pair present anterior to and a pair near the posterior limit of eyes; several on vertexal corners and superficial emargination projected above the eyes until the vertexal margin. Mesosoma, petiole, postpetiole, and first gastral sternite with basal pilosity suberect to sudecumbent, slightly clavate to clavate. Erect to suberect pilosity on mesosomal dorsum, petiole and postpetiole, coxae, and gaster; relatively abundant, clavate, yellowish; pygidium with short, filiform, suberect hairs. Antennal scapes mainly with yellowish suberect clavate hairs; funiculus covered by short, appressed, and yellowish setae. Tibiae and femora with abundant subdecumbent clavate pilosity, tarsi with subdecumbent setae.</paragraph>
|
||
<paragraph id="7E3D3E32FFC4E315FF5FFA8C840FDE27" blockId="64.[151,1437,259,2014]" pageId="64" pageNumber="65">Surface of antennal scrobes matte, finely strigulate; head dorsum rugulose, median temple punctate. Pronotum laterally punctuate-reticulate; anterior portion rugulose. Surface of mesopleuron and propodeum matte, rugulae sparse. Petiole and postpetiole rugulose. Procoxae areolate-rugulose. Gaster densely areolate-punctate.</paragraph>
|
||
<paragraph id="7E3D3E32FFC4E315FF5FFA18822EDC74" blockId="64.[151,1437,259,2014]" pageId="64" pageNumber="65">
|
||
Head oblong; posterolateral corners round; vertexal margin slightly raised medially. Vertexal crest present, shallow; sulcus around head visible; temple medially depressed. Mandibles aberrant, trapezoidal; external margin slightly concave; masticatory margin straight bearing 10–11 teeth, apical tooth slightly broader and longer; apical portion of mandibles strongly curved ventrally to the midpoint of their length; basal angle strongly convex, portion translucent; in full-face view, sulcus present on basal portion, extending from the masticatory margin to at least half of the lamina, slightly transversal. Labrum short and triangular, overall shape similar to
|
||
<taxonomicName id="B98245B1FFC4E315FBFEF9A38429DD0F" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[1126,1330,1591,1617]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC4E315FBFEF9A38429DD0F" box="[1126,1330,1591,1617]" italics="true" pageId="64" pageNumber="65">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="E6B922B7FFC4E315FAF4F9AC81ACDD2B" captionStart="FIGURE 2" captionStartId="8.[151,250,1710,1734]" captionTargetBox="[384,1203,181,1686]" captionTargetId="figure-17@8.[384,1203,181,1686]" captionTargetPageId="8" captionText="FIGURE 2. Morphological measurements of Basiceros species; A. lateral view of worker and B. dorsal view of alate gyne of B. conjugans (MCZ546461, Peru: 15 km NE Puerto Maldonado), EL: Eye Length, PDL: Pedicel Length, AFL: Apical Flagellomere Length, ML: Mesosoma Length (= Weber’s Length), MTL: Metatibia Length, PTH: Petiole Height, PTL: Petiole Length, PPL: Postpetiole Length, MSL: Mesoscutum Length, MSW: Mesoscutum Width, MLL: Mesoscutellum Length, MLW: Mesoscutellum Width, MFL: Metafemur Length, PTW: Petiole Width, PPW: Postpetiole Width, GW: Width of gaster. C. full-face view of Basiceros browni sp. nov., paratype worker (MZSP, Ecuador: Pichincha, Otongachi), HL: Head Length, HL2: Head Length 2, HW1: Head Width 1, MdL: Mandible length, SL1: Scape Length 1, SL2: Scape Length 2, FuL: Funiculus Length. D. full-face view of male of B. manni (MCZ, Costa Rica: Heredia), HW2: Head Width 2, LOD: Lateral Ocellus Length, MOD: Median Ocellus Length, OOD: Oculo-ocellar distance, A3L: Antennomere 3 Length. Figures not to scale." figureDoi="http://doi.org/10.5281/zenodo.6605967" httpUri="https://zenodo.org/record/6605967/files/figure.png" pageId="64" pageNumber="65">Figs 2E</figureCitation>
|
||
and
|
||
<figureCitation id="E6B922B7FFC4E315FF77F9C08015DD2B" box="[239,270,1627,1653]" captionStart="FIGURE 3" captionStartId="14.[151,250,1673,1697]" captionTargetBox="[156,1432,921,1649]" captionTargetId="figure-348@14.[156,1432,921,1649]" captionTargetPageId="14" captionText="FIGURE 3. Basiceros browni sp. nov., holotype worker (MCZ, Ecuador: Los Ríos); A. full-face view, B. lateral view, and C. dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6605969" httpUri="https://zenodo.org/record/6605969/files/figure.png" pageId="64" pageNumber="65">3E</figureCitation>
|
||
from
|
||
<bibRefCitation id="1A1343C3FFC4E315FECCF9C08306DD2B" author="Probst, R. S. & Wray, B. D. & Moreau, C. S. & Brandao, C. R. F." box="[340,541,1627,1654]" pageId="64" pageNumber="65" pagination="1 - 12" refId="ref62641" refString="Probst, R. S., Wray, B. D., Moreau, C. S. & Brandao, C. R. F. (2019) A phylogenetic analysis of the dirt ants, Basiceros (Formicidae: Myrmicinae): inferring life histories through morphological convergence. Insect Systematics and Diversity, 3 (4), 1 - 12. https: // doi. org / 10.1093 / isd / ixz 013" type="journal article" year="2019">
|
||
Probst
|
||
<emphasis id="4CF6E220FFC4E315FE3CF9C780CDDD2B" box="[420,470,1627,1653]" italics="true" pageId="64" pageNumber="65">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
and
|
||
<figureCitation id="E6B922B7FFC4E315FDCDF9C083CDDD2B" box="[597,726,1627,1653]" captionStart="FIGURE 27" captionStartId="66.[151,250,1712,1736]" captionTargetBox="[155,1433,489,1689]" captionTargetId="figure-129@66.[155,1433,489,1689]" captionTargetPageId="66" captionText="FIGURE 27. Basiceros tumucumaquensis sp. nov. paratype worker (MZSP, CASENT0647158, Brazil: Amapá); A. full-face view, D. dorsal view, E. lateral view. Labrum from non-type worker (MZSP, Brazil: Rondônia); B. dorsal view and C. ventral view; lo: lateral lobe, black arrow: specialized seta on anterior margin. Crossed arrows give the orientation: a, anterior; p, posterior. Scale bars: A = 0.5 mm, D–E =1 mm, B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6606027" httpUri="https://zenodo.org/record/6606027/files/figure.png" pageId="64" pageNumber="65">Fig 27B–C</figureCitation>
|
||
); lateral lobes short, distal margin with cleft extending slightly longer and more slit-like than on
|
||
<taxonomicName id="B98245B1FFC4E315FD8AF91B83C4DDC7" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[530,735,1663,1689]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC4E315FD8AF91B83C4DDC7" box="[530,735,1663,1689]" italics="true" pageId="64" pageNumber="65">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
. Frontoclypeal surface convex; anterior margin medially concave. Tumosity present on frons, circular-shaped and with central concavity. Eyes inserted on the dorsal margin of antennal scrobes, these deep and extending posteriorly to the vertexal margin. Scape flattened and with a biangulate basal lobe, margin between angles concave; external margin lamellar and crenulated. Antennal apical segment slightly longer than the sum of three anterior antennomeres.
|
||
</paragraph>
|
||
<paragraph id="7E3D3E32FFC4E314FF5FF8AF81F2DA1F" blockId="64.[151,1437,259,2014]" lastBlockId="65.[151,1436,151,321]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">Mesosoma robust; in lateral view, promesonotal profile slightly convex dorsally; metanotal suture strongly impressed, extending as the anapleural sulcus, transversely rugulose. Propodeum posteriorly oblique; propodeal projections short, triangular and slightly upwards, connected by strong transverse carina; propodeal spiracle elliptic, spiracle directed posteriorly. Petiole nodiform, peduncle short; node anteriorly oblique and strongly tapered posterad. Subpetiolar process consisting of series of long sharp denticles. In lateral view, postpetiole slightly longer than length of petiolar node; slightly dorsoventrally flattened, dorsal face slightly projected on its posterior half, posterior margin bulged at the meeting with gaster. In dorsal view, petiolar node slightly oblong, ogive-shaped; anterior margin of postpetiole strongly concave, posterior margin convex. Gaster oval; anterior margin concave; first gastral sternite visibly longer than tergite. Metabasitarsus conspicuously long, longer than metatibia. Tarsal claws simple.</paragraph>
|
||
<caption id="2AFD6EBAFFC5E314FF0FFBE78410DF82" ID-DOI="http://doi.org/10.5281/zenodo.6606025" ID-Zenodo-Dep="6606025" httpUri="https://zenodo.org/record/6606025/files/figure.png" pageId="65" pageNumber="66" startId="65.[151,250,1148,1172]" targetBox="[151,1436,345,1125]" targetPageId="65">
|
||
<paragraph id="7E3D3E32FFC5E314FF0FFBE78410DF82" blockId="65.[151,1437,1148,1245]" pageId="65" pageNumber="66">
|
||
<emphasis id="4CF6E220FFC5E314FF0FFBE7803BDFCA" bold="true" box="[151,288,1148,1172]" pageId="65" pageNumber="66">FIGURE 26.</emphasis>
|
||
<taxonomicName id="B98245B1FFC5E314FEBFFBE6835CDFCA" authority="Probst & Brandão, 2022" authorityName="Probst & Brandão" authorityYear="2022" box="[295,583,1149,1172]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="tumucumaquensis" status="sp. nov.">
|
||
<emphasis id="4CF6E220FFC5E314FEBFFBE6835CDFCA" box="[295,583,1149,1172]" italics="true" pageId="65" pageNumber="66">Basiceros tumucumaquensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4CF6E220FFC5E314FDD5FBE68385DFCB" bold="true" box="[589,670,1149,1173]" pageId="65" pageNumber="66">
|
||
<taxonomicNameLabel id="57C55F5BFFC5E314FDD5FBE68385DFCB" box="[589,670,1149,1173]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype worker (MZSP, CASENT0647157, Brazil: Amapá); mandibles in A. dorsal view, B. oblique view, C. lateral view, D. ventral view, highlighting ventrolabrum, E. dorsal view, F. lateral view, G. full-face view, H. oblique view of head and pronotum. Scale bars: A–D = 0.2 mm, E–F = 1 mm, G–H = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E3D3E32FFC5E314FF5FFA9085B9DECF" blockId="65.[151,1437,1291,2037]" pageId="65" pageNumber="66">
|
||
<emphasis id="4CF6E220FFC5E314FF5FFA90801ADE7B" box="[199,257,1291,1317]" italics="true" pageId="65" pageNumber="66">Male</emphasis>
|
||
. (n=1) HL 1.072, HW1 0.954, HW2 1.113, MdL 0.431, SL2 0.21, PDL 0.118, A3L 0.387, AFL 0.63, EL 0.347, LOD 0.11, MOD 0.083, OOD 0.396, ML 1.78, MSL 1.009, MSW 0.85, MLL 0.32, MLW 0.552, MfL 1.469, MtL 1.023,
|
||
<collectingCountry id="06957EA2FFC5E314FE83FAC88024DE33" box="[283,319,1363,1389]" name="Philippines" pageId="65" pageNumber="66">PH</collectingCountry>
|
||
0.36, PL 0.8, PW 0.317, PPL 0.474, PPW 0.462, GL 1.627, GW 1.17, TL 6.18, CI 88, CS 1.01, MCI 40, SI 22, ESI 165, SAI 54, SAI2 33, EI1 0.63, EI2 85, MTI 84, MLI 172, MFI 64, PTI 222.
|
||
</paragraph>
|
||
<paragraph id="7E3D3E32FFC5E314FF5FFA008036DD7F" blockId="65.[151,1437,1291,2037]" pageId="65" pageNumber="66">Coloration black, with areas slightly ferruginous (e.g., propodeal lobes and scutoscutellar sulcus). Coxae and central portion of femora dark, rest of appendages lighter, yellowish. Pilosity primarily composed of long filiform whitish hairs covering the body, mainly head, dorsum of promesonotum, and sparser on gaster. Wigs brown, microtrichia dense.</paragraph>
|
||
<paragraph id="7E3D3E32FFC5E314FF5FF9B085F5DDD3" blockId="65.[151,1437,1291,2037]" pageId="65" pageNumber="66">Integument matte, with a grainy aspect. Head covered with irregular rugulae, punctuate in between; mesosoma dorsally rugulose. Mesokatepisternum rugulose, mesoanepisternum rugulose-punctate, other integument covered by dense rugulose-punctate sculpture that resemble reticulate punctuations on lower magnification.</paragraph>
|
||
<paragraph id="7E3D3E32FFC5E317FF5FF90C8386DA43" blockId="65.[151,1437,1291,2037]" lastBlockId="66.[151,1437,151,465]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
|
||
Mandibles sinuous, elongated; rugulose. Masticatory margin with 10 short triangular teeth, second and third teeth from the base slightly broader than others; apical tooth falcate; external margins of mandibles convex. Head subpiriform, cervical flange conspicuous. Antennal torsion conspicuous. Ocelli large and positioned on cephalic crest; median ocellus inserted after a transversely rugose cleft. As for other
|
||
<taxonomicName id="B98245B1FFC5E314FC7AF89F8554DC43" authorityName=": Wheeler" authorityYear="1916" box="[994,1103,1796,1821]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4CF6E220FFC5E314FC7AF89F8554DC43" box="[994,1103,1796,1821]" italics="true" pageId="65" pageNumber="66">Basiceros</emphasis>
|
||
</taxonomicName>
|
||
males, a deep concave sulcus is present along the frontoclypeal region. Central disc of clypeus convex, central portion raised; anterior margin and lateral portions slightly depressed. Anterior clypeal margin slightly concave medially, lateral margin convex. Eyes globular, large, and shiny. Antennal fossa deep. In dorsal view, mesoscutum slightly cuneiform anteriorly, as wide as long; anterior margin with smooth and shiny median carina, extending along dorsum of mesoscutum but not reaching notauli, these lightly impressed. Scutoscutellar suture broadly concave. Parapsides oval, slightly deep. Parapsidal lines smooth and shiny. Scutoscutellar sulcus broad and deep, transversally costulate. In dorsal view, mesoscutellum transversely rectangular, longitudinally divided by a median posterior sulcus. Propodeal projections round, propodeal lobes auricular and slightly narrow. Petiole and gaster similar to conspecific females. Subpetiolar projection absent, only obsolete spine on peduncle present; petiolar spiracle slightly projected laterally. Forewing
|
||
<typeStatus id="A1398090FFC6E317FF0FFE9881DCDA43" box="[151,199,259,285]" pageId="66" pageNumber="67">type</typeStatus>
|
||
2; hindwing with five submedian hamuli.
|
||
</paragraph>
|
||
<paragraph id="7E3D3E32FFC6E317FF5FFEBC80E8DA1F" blockId="66.[151,1437,151,465]" box="[199,499,295,321]" pageId="66" pageNumber="67">
|
||
<emphasis id="4CF6E220FFC6E317FF5FFEBC806EDA1F" box="[199,373,295,321]" italics="true" pageId="66" pageNumber="67">Gyne and larva</emphasis>
|
||
. unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36986DB9FFC6E317FF5FFED08213DA8F" pageId="66" pageNumber="67" type="etymology">
|
||
<paragraph id="7E3D3E32FFC6E317FF5FFED08213DA8F" blockId="66.[151,1437,151,465]" pageId="66" pageNumber="67">
|
||
<emphasis id="4CF6E220FFC6E317FF5FFED08055DA3B" bold="true" box="[199,334,331,357]" pageId="66" pageNumber="67">Etymology.</emphasis>
|
||
The species name
|
||
<taxonomicName id="B98245B1FFC6E317FD87FED083F3DA3A" authorityName="Probst & Brandão" authorityYear="2022" box="[543,744,331,356]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="tumucumaquensis">
|
||
<emphasis id="4CF6E220FFC6E317FD87FED083F3DA3A" box="[543,744,331,356]" italics="true" pageId="66" pageNumber="67">tumucumaquensis</emphasis>
|
||
</taxonomicName>
|
||
is masculine and a Latin singular adjective in the Nominative, paying homage to the Tumucumaque Mountains National Park (PNMT),
|
||
<typeStatus id="A1398090FFC6E317FC42FEF48511DAD7" box="[986,1034,367,393]" pageId="66" pageNumber="67">type</typeStatus>
|
||
locality and protection area named after the Tumuc-Humac mountain range in the states of
|
||
<collectingRegion id="BC46F0D0FFC6E317FD67FE08822ADAF3" box="[767,817,403,429]" country="Suriname" name="Para" pageId="66" pageNumber="67">Pará</collectingRegion>
|
||
and
|
||
<collectingRegion id="BC46F0D0FFC6E317FCFBFE0882A3DAF3" box="[867,952,403,429]" country="Brazil" name="Amapa" pageId="66" pageNumber="67">Amapá</collectingRegion>
|
||
. The PNMT is Brazil’s largest national park and one of the world’s largest protected tropical forests.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="2AFD6EBAFFC6E317FF0FF92B83E0DC6A" ID-DOI="http://doi.org/10.5281/zenodo.6606027" ID-Zenodo-Dep="6606027" httpUri="https://zenodo.org/record/6606027/files/figure.png" pageId="66" pageNumber="67" startId="66.[151,250,1712,1736]" targetBox="[155,1433,489,1689]" targetPageId="66">
|
||
<paragraph id="7E3D3E32FFC6E317FF0FF92B83E0DC6A" blockId="66.[151,1437,1712,1845]" pageId="66" pageNumber="67">
|
||
<emphasis id="4CF6E220FFC6E317FF0FF92B8039DD96" bold="true" box="[151,290,1712,1736]" pageId="66" pageNumber="67">FIGURE 27.</emphasis>
|
||
<taxonomicName id="B98245B1FFC6E317FEB1F92A8351DD96" authority="Probst & Brandão, 2022" authorityName="Probst & Brandão" authorityYear="2022" box="[297,586,1713,1736]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="tumucumaquensis" status="sp. nov.">
|
||
<emphasis id="4CF6E220FFC6E317FEB1F92A8351DD96" box="[297,586,1713,1736]" italics="true" pageId="66" pageNumber="67">Basiceros tumucumaquensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4CF6E220FFC6E317FDCAF92A83BFDD97" bold="true" box="[594,676,1713,1737]" pageId="66" pageNumber="67">
|
||
<taxonomicNameLabel id="57C55F5BFFC6E317FDCAF92A83BFDD97" box="[594,676,1713,1737]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
paratype worker (MZSP, CASENT0647158, Brazil: Amapá); A. full-face view, D. dorsal view, E. lateral view. Labrum from non-type worker (MZSP, Brazil: Rondônia); B. dorsal view and C. ventral view; lo: lateral lobe, black arrow: specialized seta on anterior margin. Crossed arrows give the orientation: a, anterior; p, posterior. Scale bars: A = 0.5 mm, D–E =1 mm, B–C = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="36986DB9FFC6E316FF5FF8F883F7DAAB" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67" type="discussion">
|
||
<paragraph id="7E3D3E32FFC6E316FF5FF8F88022DA1F" blockId="66.[151,1436,1891,2025]" lastBlockId="67.[151,1437,151,501]" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67">
|
||
<emphasis id="4CF6E220FFC6E317FF5FF8F88054DC23" bold="true" box="[199,335,1891,1917]" pageId="66" pageNumber="67">Comments.</emphasis>
|
||
<taxonomicName id="B98245B1FFC6E317FECEF8FF8388DC23" authority="Probst & Brandão, 2022" authorityName="Probst & Brandão" authorityYear="2022" box="[342,659,1892,1917]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="tumucumaquensis" status="sp. nov.">
|
||
<emphasis id="4CF6E220FFC6E317FECEF8FF8388DC23" box="[342,659,1892,1917]" italics="true" pageId="66" pageNumber="67">Basiceros tumucumaquensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4CF6E220FFC6E317FD01F8F883E9DC23" bold="true" box="[665,754,1891,1917]" pageId="66" pageNumber="67">
|
||
<taxonomicNameLabel id="57C55F5BFFC6E317FD01F8F883E9DC23" box="[665,754,1891,1917]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be separated from other
|
||
<taxonomicName id="B98245B1FFC6E317FBABF8FF85BBDC23" authorityName=": Wheeler" authorityYear="1916" box="[1075,1184,1892,1917]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4CF6E220FFC6E317FBABF8FF85BBDC23" box="[1075,1184,1892,1917]" italics="true" pageId="66" pageNumber="67">Basiceros</emphasis>
|
||
</taxonomicName>
|
||
by the combination of mandibular and labrum morphologies and general pilosity on the worker, and by head and mandibular morphologies, pilosity, and mesosomal sculpture on the male. This new species is very similar to
|
||
<taxonomicName id="B98245B1FFC6E317FB1FF837844DDC9B" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[1159,1366,1963,1989]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC6E317FB1FF837844DDC9B" box="[1159,1366,1963,1989]" italics="true" pageId="66" pageNumber="67">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
, from which the worker caste can be differentiated based on head shape and pilosity, body pilosity, mesosomal and gastral sculpture, and mandibular shape. The male of this new species looks like a chimera between males of
|
||
<taxonomicName id="B98245B1FFC7E316FA9CFF0C848EDBEE" authorityName="Brown" authorityYear="1974" box="[1284,1429,151,176]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="conjugans">
|
||
<emphasis id="4CF6E220FFC7E316FA9CFF0C848EDBEE" box="[1284,1429,151,176]" italics="true" pageId="67" pageNumber="68">B. conjugans</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B98245B1FFC7E316FF0FFF2081E6DB8A" authorityName="Brown & Kempf" authorityYear="1960" box="[151,253,187,212]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="manni">
|
||
<emphasis id="4CF6E220FFC7E316FF0FFF2081E6DB8A" box="[151,253,187,212]" italics="true" pageId="67" pageNumber="68">B. manni</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="B98245B1FFC7E316FEA3FF208313DB8B" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[315,520,187,213]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC7E316FEA3FF208313DB8B" box="[315,520,187,213]" italics="true" pageId="67" pageNumber="68">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
. Body coloration is almost uniformly dark; mandibles are sinuous and not subtriangular like in
|
||
<taxonomicName id="B98245B1FFC7E316FEA5FF448312DBA7" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[317,521,223,249]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC7E316FEA5FF448312DBA7" box="[317,521,223,249]" italics="true" pageId="67" pageNumber="68">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
; the apical tooth follows the profile of the masticatory margin, not being projected as in
|
||
<taxonomicName id="B98245B1FFC7E316FF4BFE9880BADA43" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[211,417,259,285]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC7E316FF4BFE9880BADA43" box="[211,417,259,285]" italics="true" pageId="67" pageNumber="68">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
; the head is subpiriform, resembling other males of
|
||
<taxonomicName id="B98245B1FFC7E316FC68FE988546DA42" authorityName=": Wheeler" authorityYear="1916" box="[1008,1117,259,284]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4CF6E220FFC7E316FC68FE988546DA42" box="[1008,1117,259,284]" italics="true" pageId="67" pageNumber="68">Basiceros</emphasis>
|
||
</taxonomicName>
|
||
—in
|
||
<taxonomicName id="B98245B1FFC7E316FB0EFE98847FDA43" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[1174,1380,259,285]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC7E316FB0EFE98847FDA43" box="[1174,1380,259,285]" italics="true" pageId="67" pageNumber="68">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
, it is subhexagonal.
|
||
</paragraph>
|
||
<paragraph id="7E3D3E32FFC7E316FF5FFED083F7DAAB" blockId="67.[151,1437,151,501]" pageId="67" pageNumber="68">
|
||
In terms of variation, workers of
|
||
<taxonomicName id="B98245B1FFC7E316FDDBFED08236DA3A" authorityName="Probst & Brandão" authorityYear="2022" box="[579,813,331,356]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="tumucumaquensis">
|
||
<emphasis id="4CF6E220FFC7E316FDDBFED08236DA3A" box="[579,813,331,356]" italics="true" pageId="67" pageNumber="68">B. tumucumaquensis</emphasis>
|
||
</taxonomicName>
|
||
present slightly variations in integumentary coloration and sculpture, pilosity density, and subpetiolar process, size, and slightly on labrum morphology—the cleft on distal margin can vary on extension and inner margin of lateral lobes can be more or less straight. The
|
||
<typeStatus id="A1398090FFC7E316FB51FE088430DAF3" box="[1225,1323,403,429]" pageId="67" pageNumber="68" type="holotype">holotype</typeStatus>
|
||
and some
|
||
<typeStatus id="A1398090FFC7E316FF0FFE2C8019DA8F" box="[151,258,439,465]" pageId="67" pageNumber="68" type="paratype">paratypes</typeStatus>
|
||
had oribatid mites on the ventral face of mandibles. It is not possible to confirm whether such arachnids are phoretic, commensal myrmecophiles or parasites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="2AFD6EBAFFC7E316FF0FF9EA859DDDF3" ID-DOI="http://doi.org/10.5281/zenodo.6606031" ID-Zenodo-Dep="6606031" httpUri="https://zenodo.org/record/6606031/files/figure.png" pageId="67" pageNumber="68" startId="67.[151,250,1649,1673]" targetBox="[167,1421,525,1626]" targetPageId="67">
|
||
<paragraph id="7E3D3E32FFC7E316FF0FF9EA859DDDF3" blockId="67.[151,1437,1649,1710]" pageId="67" pageNumber="68">
|
||
<emphasis id="4CF6E220FFC7E316FF0FF9EA8038DDD7" bold="true" box="[151,291,1649,1673]" pageId="67" pageNumber="68">FIGURE 28.</emphasis>
|
||
<taxonomicName id="B98245B1FFC7E316FEB4F9E98355DDD7" authority="Probst & Brandão, 2022" authorityName="Probst & Brandão" authorityYear="2022" box="[300,590,1650,1673]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="tumucumaquensis" status="sp. nov.">
|
||
<emphasis id="4CF6E220FFC7E316FEB4F9E98355DDD7" box="[300,590,1650,1673]" italics="true" pageId="67" pageNumber="68">Basiceros tumucumaquensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4CF6E220FFC7E316FDCEF9E983B1DDD4" bold="true" box="[598,682,1650,1674]" pageId="67" pageNumber="68">
|
||
<taxonomicNameLabel id="57C55F5BFFC7E316FDCEF9E983B1DDD4" box="[598,682,1650,1674]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
paratype male (MZSP, CASENT0647167, Brazil: Amapá); A. head and antenna, B. lateral view, C. full-face view, D. dorsal view. Scale bars: A–B, D =1 mm, C = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="36986DB9FFC7E311FF5FF94780F8D9DB" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68" type="distribution">
|
||
<paragraph id="7E3D3E32FFC7E311FF5FF9478556DA1F" blockId="67.[151,1437,1756,2035]" lastBlockId="68.[151,1437,151,755]" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68">
|
||
<emphasis id="4CF6E220FFC7E316FF5FF947807ADDA8" bold="true" box="[199,353,1756,1782]" pageId="67" pageNumber="68">Distribution.</emphasis>
|
||
North and NW South America. Currently known from the PNMT, the state of
|
||
<collectingRegion id="BC46F0D0FFC7E316FB6EF947847DDDA8" box="[1270,1382,1756,1782]" country="Brazil" name="Rondonia" pageId="67" pageNumber="68">Rondônia</collectingRegion>
|
||
, and potentially from a forest fragment close to Manaus, state of Amazonas (Itanna Oliveira, pers. comm., material not examined). This species occurs in sympatry with its sister taxon
|
||
<taxonomicName id="B98245B1FFC7E316FCEEF8BE855FDC60" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[886,1092,1828,1854]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC7E316FCEEF8BE855FDC60" box="[886,1092,1828,1854]" italics="true" pageId="67" pageNumber="68">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
, and hints of cryptic variation came from
|
||
<specimenCount id="6884F5BBFFC7E316FE8FF8D380B8DC3C" box="[279,419,1864,1890]" pageId="67" pageNumber="68" type="worker">two workers</specimenCount>
|
||
collected in
|
||
<collectingRegion id="BC46F0D0FFC7E316FDACF8D383BADC3C" box="[564,673,1864,1890]" country="Brazil" name="Rondonia" pageId="67" pageNumber="68">Rondônia</collectingRegion>
|
||
(included in the phylogenetic analyses from
|
||
<bibRefCitation id="1A1343C3FFC7E316FB04F8D38474DC3C" author="Probst, R. S. & Wray, B. D. & Moreau, C. S. & Brandao, C. R. F." box="[1180,1391,1864,1890]" pageId="67" pageNumber="68" pagination="1 - 12" refId="ref62641" refString="Probst, R. S., Wray, B. D., Moreau, C. S. & Brandao, C. R. F. (2019) A phylogenetic analysis of the dirt ants, Basiceros (Formicidae: Myrmicinae): inferring life histories through morphological convergence. Insect Systematics and Diversity, 3 (4), 1 - 12. https: // doi. org / 10.1093 / isd / ixz 013" type="journal article" year="2019">
|
||
Probst
|
||
<emphasis id="4CF6E220FFC7E316FB73F8D28406DC3C" box="[1259,1309,1864,1890]" italics="true" pageId="67" pageNumber="68">et al</emphasis>
|
||
., 2019
|
||
</bibRefCitation>
|
||
). At that time, those
|
||
<specimenCount id="6884F5BBFFC7E316FED0F8F780C9DCD8" box="[328,466,1900,1926]" pageId="67" pageNumber="68" type="worker">two workers</specimenCount>
|
||
(BSCJ1 and BSCJ2) were considered to be populational variations within
|
||
<taxonomicName id="B98245B1FFC7E316FA89F8F681F9DCF4" baseAuthorityName="Brown" baseAuthorityYear="1949" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC7E316FA89F8F681F9DCF4" italics="true" pageId="67" pageNumber="68">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
—so much that Probst (2015) included photographs of one of those workers under
|
||
<taxonomicName id="B98245B1FFC7E316FB0BF80A8479DCF4" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[1171,1378,1936,1962]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC7E316FB0BF80A8479DCF4" box="[1171,1378,1936,1962]" italics="true" pageId="67" pageNumber="68">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
. The boundaries of
|
||
<taxonomicName id="B98245B1FFC7E316FEAFF82E8318DC90" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[311,515,1972,1998]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC7E316FEAFF82E8318DC90" box="[311,515,1972,1998]" italics="true" pageId="67" pageNumber="68">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
started to make more sense after the phylogenetic study of
|
||
<taxonomicName id="B98245B1FFC7E316FB01F82E841DDC90" authorityName=": Wheeler" authorityYear="1916" box="[1177,1286,1973,1998]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4CF6E220FFC7E316FB01F82E841DDC90" box="[1177,1286,1973,1998]" italics="true" pageId="67" pageNumber="68">Basiceros</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="1A1343C3FFC7E316FA8DF82F81C9DCAC" author="Probst, R. S. & Wray, B. D. & Moreau, C. S. & Brandao, C. R. F." pageId="67" pageNumber="68" pagination="1 - 12" refId="ref62641" refString="Probst, R. S., Wray, B. D., Moreau, C. S. & Brandao, C. R. F. (2019) A phylogenetic analysis of the dirt ants, Basiceros (Formicidae: Myrmicinae): inferring life histories through morphological convergence. Insect Systematics and Diversity, 3 (4), 1 - 12. https: // doi. org / 10.1093 / isd / ixz 013" type="journal article" year="2019">
|
||
Probst
|
||
<emphasis id="4CF6E220FFC7E316FAFDF82E848EDC90" box="[1381,1429,1972,1998]" italics="true" pageId="67" pageNumber="68">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
; see
|
||
<figureCitation id="E6B922B7FFC7E316FE90F8438058DCAD" box="[264,323,2008,2035]" captionStart="FIGURE 5" captionStartId="17.[151,250,1530,1554]" captionTargetBox="[155,1432,777,1506]" captionTargetId="figure-265@17.[155,1432,777,1506]" captionTargetPageId="17" captionText="FIGURE 5. Basiceros conjugans, holotype worker (MCZ 32179, Ecuador: Limoncocha); A. full-face view, B. lateral view, and C. dorsal view. Scale bars: A = 0.5 mm, B–C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6605973" httpUri="https://zenodo.org/record/6605973/files/figure.png" pageId="67" pageNumber="68">Fig 5</figureCitation>
|
||
), which
|
||
<emphasis id="4CF6E220FFC7E316FE3DF84383FDDCAC" box="[421,742,2008,2034]" italics="true" pageId="67" pageNumber="68">
|
||
recovered
|
||
<taxonomicName id="B98245B1FFC7E316FD81F84283FDDCAC" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[537,742,2008,2034]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="scambognathus">B. scambognathus</taxonomicName>
|
||
</emphasis>
|
||
as two sister clades. One clade grouped
|
||
<specimenCount id="6884F5BBFFC7E316FB34F843843BDCAC" box="[1196,1312,2008,2034]" pageId="67" pageNumber="68" type="male">three male</specimenCount>
|
||
specimens from Amazonas (North of
|
||
<collectingCountry id="06957EA2FFC0E311FE58FF0C8312DBEC" box="[448,521,151,178]" name="Brazil" pageId="68" pageNumber="69">Brazil</collectingCountry>
|
||
),
|
||
<collectingRegion id="BC46F0D0FFC0E311FD84FF0C83A9DBEF" box="[540,690,151,177]" country="Brazil" name="Minas Gerais" pageId="68" pageNumber="69">Minas Gerais</collectingRegion>
|
||
(Southeast of
|
||
<collectingCountry id="06957EA2FFC0E311FCCCFF0C8286DBEC" box="[852,925,151,178]" name="Brazil" pageId="68" pageNumber="69">Brazil</collectingCountry>
|
||
), and
|
||
<collectingCountry id="06957EA2FFC0E311FC47FF0C850DDBEF" box="[991,1046,151,177]" name="Peru" pageId="68" pageNumber="69">Peru</collectingCountry>
|
||
; the other clade grouped those
|
||
<specimenCount id="6884F5BBFFC0E311FAEAFF0C81EADB8B" pageId="68" pageNumber="69" type="worker">two workers</specimenCount>
|
||
from Rondônia (west of the Amazonian biome). In the same study,
|
||
<bibRefCitation id="1A1343C3FFC0E311FB9FFF2085FDDB8B" author="Probst, R. S. & Wray, B. D. & Moreau, C. S. & Brandao, C. R. F." box="[1031,1254,187,213]" pageId="68" pageNumber="69" pagination="1 - 12" refId="ref62641" refString="Probst, R. S., Wray, B. D., Moreau, C. S. & Brandao, C. R. F. (2019) A phylogenetic analysis of the dirt ants, Basiceros (Formicidae: Myrmicinae): inferring life histories through morphological convergence. Insect Systematics and Diversity, 3 (4), 1 - 12. https: // doi. org / 10.1093 / isd / ixz 013" type="journal article" year="2019">
|
||
Probst
|
||
<emphasis id="4CF6E220FFC0E311FBC0FF208590DB8B" box="[1112,1163,187,213]" italics="true" pageId="68" pageNumber="69">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
pointed out the unique morphology of these
|
||
<specimenCount id="6884F5BBFFC0E311FE4CFF448345DBA7" box="[468,606,223,249]" pageId="68" pageNumber="69" type="worker">two workers</specimenCount>
|
||
in terms of pilosity patterns, head, and body shape, and suggested that they may correspond to a cryptic species within
|
||
<taxonomicName id="B98245B1FFC0E311FDE5FE988251DA43" baseAuthorityName="Brown" baseAuthorityYear="1949" box="[637,842,259,285]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="scambognathus">
|
||
<emphasis id="4CF6E220FFC0E311FDE5FE988251DA43" box="[637,842,259,285]" italics="true" pageId="68" pageNumber="69">B. scambognathus</emphasis>
|
||
</taxonomicName>
|
||
. Supported by phylogenetic information from
|
||
<bibRefCitation id="1A1343C3FFC0E311FACDFE98803ADA1F" author="Probst, R. S. & Wray, B. D. & Moreau, C. S. & Brandao, C. R. F." pageId="68" pageNumber="69" pagination="1 - 12" refId="ref62641" refString="Probst, R. S., Wray, B. D., Moreau, C. S. & Brandao, C. R. F. (2019) A phylogenetic analysis of the dirt ants, Basiceros (Formicidae: Myrmicinae): inferring life histories through morphological convergence. Insect Systematics and Diversity, 3 (4), 1 - 12. https: // doi. org / 10.1093 / isd / ixz 013" type="journal article" year="2019">
|
||
Probst
|
||
<emphasis id="4CF6E220FFC0E311FF0FFEBC81D3DA1F" box="[151,200,295,321]" italics="true" pageId="68" pageNumber="69">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
and by studying additional specimens, we raise it here as a new species.
|
||
</paragraph>
|
||
<paragraph id="7E3D3E32FFC0E311FF5FFED080F8D9DB" blockId="68.[151,1437,151,755]" pageId="68" pageNumber="69">
|
||
<emphasis id="4CF6E220FFC0E311FF5FFED0809CDA3A" bold="true" box="[199,391,330,357]" pageId="68" pageNumber="69">Natural history.</emphasis>
|
||
Virtually nothing is known about the biology of this species. The type-series was collected below surface, close to tree roots, with the nest apparently scattered in indistinct chambers. RSP also collected
|
||
<specimenCount id="6884F5BBFFC0E311FF0FFE08803DDAF3" box="[151,294,403,429]" pageId="68" pageNumber="69" type="worker">two workers</specimenCount>
|
||
from a leaf litter sample taken around superficial roots of a
|
||
<collectingCountry id="06957EA2FFC0E311FC6EFE088521DAF3" box="[1014,1082,403,429]" name="Brazil" pageId="68" pageNumber="69">Brazil</collectingCountry>
|
||
nut tree (
|
||
<taxonomicName id="B98245B1FFC0E311FB2AFE08848EDAF3" box="[1202,1429,403,429]" class="Magnoliopsida" family="Lecythidaceae" genus="Bertholletia" kingdom="Plantae" order="Ericales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="excelsa">
|
||
<emphasis id="4CF6E220FFC0E311FB2AFE08848EDAF3" box="[1202,1429,403,429]" italics="true" pageId="68" pageNumber="69">Bertholletia excelsa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B98245B1FFC0E311FF0FFE2C8025DA8C" box="[151,318,439,466]" class="Magnoliopsida" family="Lecythidaceae" kingdom="Plantae" order="Ericales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="family">Lecythidaceae</taxonomicName>
|
||
). Those
|
||
<specimenCount id="6884F5BBFFC0E311FE05FE2C8332DA8F" box="[413,553,439,465]" pageId="68" pageNumber="69" type="worker">two workers</specimenCount>
|
||
were kept alive in laboratory conditions for three days, in which sparse interactions between specimens were observed. Workers moved slowly through the terrarium, with occasionally slowpaced antennation and showing thanatosis when disturbed. Unfortunately, all the conducted “cafeteria” experiments failed and specimens did not accept any items offered. Similarly, part of the type-series was kept alive in laboratory conditions for several weeks, consistently ignoring live prey dropped inside the terrarium, but sometimes accepting freshly-killed adult fruit flies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36986DB9FFC0E311FF5FFD158494D9AF" pageId="68" pageNumber="69" type="materials_examined">
|
||
<paragraph id="7E3D3E32FFC0E311FF5FFD158494D9AF" blockId="68.[151,1437,151,755]" pageId="68" pageNumber="69">
|
||
<emphasis id="4CF6E220FFC0E311FF5FFD1580ABD9F6" bold="true" box="[199,432,654,680]" pageId="68" pageNumber="69">Material examined.</emphasis>
|
||
<materialsCitation id="CEEA346FFFC0E311FE2EFD1483CED9F7" ID-GBIF-Occurrence="4029332441" box="[438,725,655,681]" collectionCode="PNMT" country="Brazil" location="Amapa" pageId="68" pageNumber="69" specimenCount="1" stateProvince="Amapa">
|
||
<collectingCountry id="06957EA2FFC0E311FE2EFD14833BD9F7" box="[438,544,655,681]" name="Brazil" pageId="68" pageNumber="69">BRAZIL</collectingCountry>
|
||
:
|
||
<collectingRegion id="BC46F0D0FFC0E311FDB0FD148367D9F7" box="[552,636,655,681]" country="Brazil" name="Amapa" pageId="68" pageNumber="69">Amapá</collectingRegion>
|
||
,
|
||
<collectionCode id="1893A6F7FFC0E311FD1EFD1483CED9F7" box="[646,725,655,681]" pageId="68" pageNumber="69">PNMT</collectionCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="CEEA346FFFC0E311FD47FD148416D9F7" ID-GBIF-Occurrence="4029332379" box="[735,1293,654,681]" collectingDate="2018-12-25" collectionCode="R" country="Brazil" elevation="0" latitude="1.242333" longLatPrecision="14" longitude="-52.419777" pageId="68" pageNumber="69" specimenCount="1" stateProvince="Amapa">
|
||
<geoCoordinate id="1BB658F5FFC0E311FD47FD148253D9F7" box="[735,840,655,681]" degrees="1.242333" direction="north" orientation="latitude" pageId="68" pageNumber="69" precision="10" value="1.242333">1.242333</geoCoordinate>
|
||
<geoCoordinate id="1BB658F5FFC0E311FCD6FD1582C8D9F7" box="[846,979,654,681]" degrees="52.419778" direction="west" orientation="longitude" pageId="68" pageNumber="69" precision="10" value="-52.419777">-52.419778</geoCoordinate>
|
||
±
|
||
<elevation id="F5AFD901FFC0E311FC7AFD15857BD9F7" box="[994,1120,654,681]" isEnumeration="true" pageId="68" pageNumber="69">
|
||
<quantity id="B97A93D7FFC0E311FC7AFD158508D9F7" box="[994,1043,654,681]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="68" pageNumber="69" unit="m" value="10.0">10m</quantity>
|
||
,
|
||
<quantity id="B97A93D7FFC0E311FB85FD14857BD9F7" box="[1053,1120,655,681]" metricMagnitude="2" metricUnit="m" metricValue="1.3199999999999998" pageId="68" pageNumber="69" unit="m" value="132.0">132m</quantity>
|
||
</elevation>
|
||
,
|
||
<date id="0A3C18F2FFC0E311FBF2FD1585F6D9F7" box="[1130,1261,654,681]" pageId="68" pageNumber="69" value="2018-12-25">
|
||
<collectingDate id="1A78E11AFFC0E311FBF2FD1585F6D9F7" box="[1130,1261,654,681]" pageId="68" pageNumber="69" value="2018-12-25">25.xii.2018</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="1893A6F7FFC0E311FB6FFD148416D9F7" box="[1271,1293,655,681]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="68" pageNumber="69">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="CEEA346FFFC0E311FA8FFD1485B8D993" ID-GBIF-Occurrence="4029332359" collectionCode="RSPPC" collectorName="Probst & O. Guilherme" country="Brazil" pageId="68" pageNumber="69" specimenCount="1" stateProvince="Amapa">
|
||
<collectorName id="D3775BE4FFC0E311FA8FFD148444D9F7" box="[1303,1375,655,681]" pageId="68" pageNumber="69">Probst</collectorName>
|
||
&
|
||
<collectorName id="D3775BE4FFC0E311FA19FD148014D993" pageId="68" pageNumber="69">O. Guilherme</collectorName>
|
||
cols. (additional 20 non-type workers, several broken and/or dissected) [
|
||
<collectionCode id="1893A6F7FFC0E311FBA7FD298587D993" box="[1087,1180,690,717]" pageId="68" pageNumber="69">RSPPC</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="CEEA346FFFC0E311FB36FD288255D9AF" ID-GBIF-Occurrence="4029332382" collectingDate="2013-07-15" collectingDateMax="2013-07-30" collectingDateMin="2013-07-15" collectionCode="R" collectorName="Porto Velho & Area Mutum" country="Brazil" latitude="-9.580277" location="Rondonia" longLatPrecision="21" longitude="-65.05361" pageId="68" pageNumber="69" specimenCount="1" stateProvince="Rondonia">
|
||
<collectingRegion id="BC46F0D0FFC0E311FB36FD288404D993" box="[1198,1311,691,717]" country="Brazil" name="Rondonia" pageId="68" pageNumber="69">Rondônia</collectingRegion>
|
||
:
|
||
<collectorName id="D3775BE4FFC0E311FAB2FD2881ADD9AF" pageId="68" pageNumber="69">Porto Velho</collectorName>
|
||
,
|
||
<collectorName id="D3775BE4FFC0E311FF59FD43804AD9AF" box="[193,337,727,755]" pageId="68" pageNumber="69">Área Mutum</collectorName>
|
||
<geoCoordinate id="1BB658F5FFC0E311FEC0FD4C80C6D9AF" box="[344,477,727,753]" degrees="09" direction="south" minutes="34" orientation="latitude" pageId="68" pageNumber="69" precision="15" seconds="49" value="-9.580277">09°34’49”S</geoCoordinate>
|
||
<geoCoordinate id="1BB658F5FFC0E311FE7CFD4D8368D9AF" box="[484,627,726,753]" degrees="65" direction="west" minutes="03" orientation="longitude" pageId="68" pageNumber="69" precision="15" seconds="13" value="-65.05361">65°03’13”W</geoCoordinate>
|
||
,
|
||
<date id="0A3C18F2FFC0E311FDE6FD4D8230D9AF" box="[638,811,726,753]" pageId="68" pageNumber="69" value="2013-07-15" valueMax="2013-07-30" valueMin="2013-07-15">
|
||
<collectingDate id="1A78E11AFFC0E311FDE6FD4D8230D9AF" box="[638,811,726,753]" pageId="68" pageNumber="69" value="2013-07-15" valueMax="2013-07-30" valueMin="2013-07-15">15–30.vii.2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="1893A6F7FFC0E311FCAEFD4C8255D9AF" box="[822,846,727,753]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="68" pageNumber="69">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="CEEA346FFFC0E311FCCAFD4C8511D9AF" ID-GBIF-Occurrence="4029332417" box="[850,1034,727,753]" collectionCode="R" collectorName="Probst" country="Brazil" pageId="68" pageNumber="69" specimenCount="1" stateProvince="Rondonia">
|
||
S.
|
||
<collectorName id="D3775BE4FFC0E311FCF5FD4C82AED9AF" box="[877,949,727,753]" pageId="68" pageNumber="69">Probst</collectorName>
|
||
& G.
|
||
<collectionCode id="1893A6F7FFC0E311FC6CFD4C8511D9AF" box="[1012,1034,727,753]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="68" pageNumber="69">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="CEEA346FFFC0E311FB8CFD4C8490D9AF" ID-GBIF-Occurrence="4029332307" box="[1044,1419,726,754]" collectionCode="MZSP" collectorName="Mazao" country="Brazil" pageId="68" pageNumber="69" specimenCount="2" specimenCount-worker="2" stateProvince="Rondonia">
|
||
<collectorName id="D3775BE4FFC0E311FB8CFD4C857BD9AF" box="[1044,1120,727,753]" pageId="68" pageNumber="69">Mazão</collectorName>
|
||
cols. (
|
||
<specimenCount id="6884F5BBFFC0E311FB33FD4C8406D9AC" box="[1195,1309,727,754]" pageId="68" pageNumber="69" type="worker">2 workers</specimenCount>
|
||
) [
|
||
<collectionCode id="1893A6F7FFC0E311FAAAFD4D849FD9AF" box="[1330,1412,726,753]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="68" pageNumber="69" type="Museum">MZSP</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |