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<document id="B791DDD8452947EE35526A361003981F" ID-CLB-Dataset="53922" ID-DOI="10.11646/zootaxa.5149.1.1" ID-GBIF-Dataset="caecca9d-72a3-4d9a-836d-9ac3e0ded512" ID-ISSN="1175-5326" ID-Zenodo-Dep="6605961" ID-ZooBank="BF98000F-04F9-4D22-9D93-4C3A2418D7EF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654152510384" checkinUser="plazi" docAuthor="Probst, Rodolfo Da Silva &amp; Brandão, Carlos Roberto Ferreira" docDate="2022" docId="F62B8F24FF8BE343FF0FF936826CD803" docLanguage="en" docName="zootaxa.5149.1.1.pdf" docOrigin="Zootaxa 5149 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Basiceros conjugans Brown 1974" docType="treatment" docVersion="7" lastPageNumber="22" masterDocId="0A12F75CFF84E355FF98FF9B811BDB5E" masterDocTitle="A taxonomic revision of the dirt ants, Basiceros Schulz, 1906 (Hymenoptera, Formicidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="16" updateTime="1699338740759" updateUser="ExternalLinkService">
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<mods:title id="01EBB0246139B2258C2F71FEEC6074E2">A taxonomic revision of the dirt ants, Basiceros Schulz, 1906 (Hymenoptera, Formicidae)</mods:title>
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<mods:namePart id="6810C2D8AB666561CEDB6AB7D739609D">Probst, Rodolfo Da Silva</mods:namePart>
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<mods:namePart id="FA6456FBCEC966C28E4209E9D89D8DB2">Brandão, Carlos Roberto Ferreira</mods:namePart>
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<treatment id="F62B8F24FF8BE343FF0FF936826CD803" ID-DOI="http://doi.org/10.5281/zenodo.7636259" ID-GBIF-Taxon="209289994" ID-Zenodo-Dep="7636259" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803" httpUri="http://treatment.plazi.org/id/F62B8F24FF8BE343FF0FF936826CD803" lastPageId="22" lastPageNumber="22" pageId="15" pageNumber="16">
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<paragraph id="7E3D3E32FF8BE35AFF0FF936832CDD99" blockId="15.[151,567,1709,1772]" box="[151,567,1709,1735]" pageId="15" pageNumber="16">
<heading id="2575895EFF8BE35AFF0FF936832CDD99" bold="true" box="[151,567,1709,1735]" fontSize="11" level="2" pageId="15" pageNumber="16" reason="4">
<taxonomicName id="B98245B1FF8BE35AFF0FF936832CDD99" ID-CoL="5WD7M" ID-ENA="604459" authority="Brown, 1974" authorityName="Brown" authorityYear="1974" box="[151,567,1709,1735]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF8BE35AFF0FF936832CDD99" bold="true" box="[151,567,1709,1735]" pageId="15" pageNumber="16">
<emphasis id="4CF6E220FF8BE35AFF0FF9368089DD99" bold="true" box="[151,402,1709,1735]" italics="true" pageId="15" pageNumber="16">Basiceros conjugans</emphasis>
<bibRefCitation id="1A1343C3FF8BE35AFE01F936832CDD99" author="Brown, W. L. Jr." box="[409,567,1709,1735]" pageId="15" pageNumber="16" pagination="131 - 140" refId="ref61175" refString="Brown, W. L. Jr. (1974) A supplement to the revision of the ant genus Basiceros. Journal of the New York Entomological Society, 82, 131 - 140." type="journal article" year="1974">Brown, 1974</bibRefCitation>
</emphasis>
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<paragraph id="7E3D3E32FF8BE35AFF0FF94A809DDDB5" blockId="15.[151,567,1709,1772]" box="[151,390,1745,1772]" pageId="15" pageNumber="16">
(
<figureCitation id="E6B922B7FF8BE35AFF07F94A8006DDB5" box="[159,285,1745,1771]" captionStart="FIGURE 2" captionStartId="8.[151,250,1710,1734]" captionTargetBox="[384,1203,181,1686]" captionTargetId="figure-17@8.[384,1203,181,1686]" captionTargetPageId="8" captionText="FIGURE 2. Morphological measurements of Basiceros species; A. lateral view of worker and B. dorsal view of alate gyne of B. conjugans (MCZ546461, Peru: 15 km NE Puerto Maldonado), EL: Eye Length, PDL: Pedicel Length, AFL: Apical Flagellomere Length, ML: Mesosoma Length (= Webers Length), MTL: Metatibia Length, PTH: Petiole Height, PTL: Petiole Length, PPL: Postpetiole Length, MSL: Mesoscutum Length, MSW: Mesoscutum Width, MLL: Mesoscutellum Length, MLW: Mesoscutellum Width, MFL: Metafemur Length, PTW: Petiole Width, PPW: Postpetiole Width, GW: Width of gaster. C. full-face view of Basiceros browni sp. nov., paratype worker (MZSP, Ecuador: Pichincha, Otongachi), HL: Head Length, HL2: Head Length 2, HW1: Head Width 1, MdL: Mandible length, SL1: Scape Length 1, SL2: Scape Length 2, FuL: Funiculus Length. D. full-face view of male of B. manni (MCZ, Costa Rica: Heredia), HW2: Head Width 2, LOD: Lateral Ocellus Length, MOD: Median Ocellus Length, OOD: Oculo-ocellar distance, A3L: Antennomere 3 Length. Figures not to scale." figureDoi="http://doi.org/10.5281/zenodo.6605967" httpUri="https://zenodo.org/record/6605967/files/figure.png" pageId="15" pageNumber="16">Figs 2AB</figureCitation>
,
<figureCitation id="E6B922B7FF8BE35AFEB1F94A804EDDB2" box="[297,341,1745,1772]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="17.[151,250,1530,1554]" captionStartId-1="18.[151,250,1544,1568]" captionStartId-2="20.[151,250,1218,1242]" captionTargetBox-0="[155,1432,777,1506]" captionTargetBox-1="[151,1435,561,1520]" captionTargetBox-2="[154,1433,181,1195]" captionTargetId-0="figure-265@17.[155,1432,777,1506]" captionTargetId-1="figure-199@18.[151,1435,561,1520]" captionTargetId-2="figure-17@20.[154,1433,181,1195]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="20" captionText-0="FIGURE 5. Basiceros conjugans, holotype worker (MCZ 32179, Ecuador: Limoncocha); A. full-face view, B. lateral view, and C. dorsal view. Scale bars: A = 0.5 mm, BC = 1 mm." captionText-1="FIGURE 6. Basiceros conjugans, paratypes gyne and male (MCZ 546463, Ecuador: Limoncocha), Ecuador: Limoncocha); A and D. full-face view, B and E. dorsal view, C and F. lateral view. Scale bars: A, D = 0.5 mm, B, C, E, F =1 mm." captionText-2="FIGURE 7. Basiceros conjugans, larva (MZSP, Peru: Madre de Dios); A. head in anteroventral view; B. head in oblique view; C. lateral view; D. dorsal pilosity; E. anus. Scale bars: A, B, E = 0.1 mm; C = 0.5 mm; E = 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6605973" figureDoi-1="http://doi.org/10.5281/zenodo.6605977" figureDoi-2="http://doi.org/10.5281/zenodo.6605979" httpUri-0="https://zenodo.org/record/6605973/files/figure.png" httpUri-1="https://zenodo.org/record/6605977/files/figure.png" httpUri-2="https://zenodo.org/record/6605979/files/figure.png" pageId="15" pageNumber="16">57</figureCitation>
,
<figureCitation id="E6B922B7FF8BE35AFEF9F94A8065DDB5" box="[353,382,1745,1771]" captionStart="FIGURE 31" captionStartId="71.[151,250,1069,1093]" captionTargetBox="[261,1327,181,1045]" captionTargetId="figure-17@71.[261,1327,181,1045]" captionTargetPageId="71" captionText="FIGURE 31. Distribution map of Basiceros species from the disciger clade (in part)." figureDoi="http://doi.org/10.5281/zenodo.6606039" httpUri="https://zenodo.org/record/6606039/files/figure.png" pageId="15" pageNumber="16">31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FF8BE35AFF0FF88D82EADC73" box="[151,1009,1813,1838]" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="7E3D3E32FF8BE35AFF0FF88D82EADC73" blockId="15.[151,1009,1813,1838]" box="[151,1009,1813,1838]" pageId="15" pageNumber="16">
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<treatmentCitation id="FF231823FF8BE35AFF0FF88D8339DC73" author="Brown, W. L. Jr." box="[151,546,1813,1837]" page="134" pageId="15" pageNumber="16" year="1974">
<taxonomicName id="B98245B1FF8BE35AFF0FF88D8339DC73" ID-CoL="5WD7M" authority="Brown 1974: 134" authorityName="Brown" authorityPageNumber="134" authorityYear="1974" box="[151,546,1813,1837]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF8BE35AFF0FF88D8073DC73" box="[151,360,1814,1837]" italics="true" pageId="15" pageNumber="16">Basiceros conjugans</emphasis>
<bibRefCitation id="1A1343C3FF8BE35AFEF7F88D8339DC73" author="Brown, W. L. Jr." box="[367,546,1813,1837]" pageId="15" pageNumber="16" pagination="131 - 140" refId="ref61175" refString="Brown, W. L. Jr. (1974) A supplement to the revision of the ant genus Basiceros. Journal of the New York Entomological Society, 82, 131 - 140." type="journal article" year="1974">Brown 1974: 134</bibRefCitation>
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,
<figureCitation id="E6B922B7FF8BE35AFDB5F88E839EDC73" box="[557,645,1813,1837]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="7.[151,250,1879,1903]" captionStartId-1="8.[151,250,1710,1734]" captionTargetBox-0="[149,1439,181,1854]" captionTargetBox-1="[384,1203,181,1686]" captionTargetId-0="figure-17@7.[149,1439,181,1854]" captionTargetId-1="figure-17@8.[384,1203,181,1686]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 1. Wing venation of Basiceros males. A. forewing and B. hindwing of B. conjugans (RSPPC, Peru). C. forewing of B. manni male (MCZ, Costa Rica: Heredia). Fused veins are indicated with a plus sign and crossveins by a dash. Cell names (in white) are indicated in italics. Scale bars: 1 mm." captionText-1="FIGURE 2. Morphological measurements of Basiceros species; A. lateral view of worker and B. dorsal view of alate gyne of B. conjugans (MCZ546461, Peru: 15 km NE Puerto Maldonado), EL: Eye Length, PDL: Pedicel Length, AFL: Apical Flagellomere Length, ML: Mesosoma Length (= Webers Length), MTL: Metatibia Length, PTH: Petiole Height, PTL: Petiole Length, PPL: Postpetiole Length, MSL: Mesoscutum Length, MSW: Mesoscutum Width, MLL: Mesoscutellum Length, MLW: Mesoscutellum Width, MFL: Metafemur Length, PTW: Petiole Width, PPW: Postpetiole Width, GW: Width of gaster. C. full-face view of Basiceros browni sp. nov., paratype worker (MZSP, Ecuador: Pichincha, Otongachi), HL: Head Length, HL2: Head Length 2, HW1: Head Width 1, MdL: Mandible length, SL1: Scape Length 1, SL2: Scape Length 2, FuL: Funiculus Length. D. full-face view of male of B. manni (MCZ, Costa Rica: Heredia), HW2: Head Width 2, LOD: Lateral Ocellus Length, MOD: Median Ocellus Length, OOD: Oculo-ocellar distance, A3L: Antennomere 3 Length. Figures not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.6605965" figureDoi-1="http://doi.org/10.5281/zenodo.6605967" httpUri-0="https://zenodo.org/record/6605965/files/figure.png" httpUri-1="https://zenodo.org/record/6605967/files/figure.png" pageId="15" pageNumber="16">Figs 12</figureCitation>
(worker, gyne, and male),
<collectingCountry id="06957EA2FF8BE35AFC0EF88E82F7DC73" box="[918,1004,1813,1837]" name="Ecuador" pageId="15" pageNumber="16">Ecuador</collectingCountry>
.
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</paragraph>
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<emphasis id="4CF6E220FF8BE35AFF0FF8C7805CDC2B" bold="true" box="[151,327,1883,1910]" pageId="15" pageNumber="16">Type material.</emphasis>
<materialsCitation id="CEEA346FFF8BE35AFEC8F8C781ECDCE0" ID-GBIF-Occurrence="4029332356" accessionNumber="MCZ 32179" collectingDate="1964-09" collectingDateMax="1964-11-31" collectingDateMin="1964-09" collectionCode="R, MCZ, NHMB" collectorName="Hermann" country="Ecuador" location="Limoncocha" pageId="15" pageNumber="16" specimenCount="8" specimenCount-male="3" specimenCount-worker="5" typeStatus="holotype">
<collectingCountry id="06957EA2FF8BE35AFEC8F8C780C7DC28" box="[336,476,1884,1910]" name="Ecuador" pageId="15" pageNumber="16">ECUADOR</collectingCountry>
:
<location id="7B5D68E9FF8BE35AFE70F8C0836FDC2B" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF8BE35AFE70F8C0836FDC2B" box="[488,628,1883,1909]" country="Ecuador" name="Limoncocha" pageId="15" pageNumber="16">Limoncocha</location>
&amp; vic.:
<date id="0A3C18F2FF8BE35AFD4EF8C0827ADC2B" box="[726,865,1883,1909]" pageId="15" pageNumber="16" value="1964-09" valueMax="1964-11-31" valueMin="1964-09-00">
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</date>
, H.
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.
<collectorName id="D3775BE4FF8BE35AFC34F8C7850FDC28" box="[940,1044,1884,1910]" pageId="15" pageNumber="16">Hermann</collectorName>
col., n. (
<specimenCount id="6884F5BBFF8BE35AFBE5F8C7841ADC2B" box="[1149,1281,1883,1910]" pageId="15" pageNumber="16" type="worker">one worker</specimenCount>
<typeStatus id="A1398090FF8BE35AFA86F8C0849ADC2B" box="[1310,1409,1883,1909]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
;
<specimenCount id="6884F5BBFF8BE35AFA16F8C781EFDCC7" pageId="15" pageNumber="16" type="worker">4 workers</specimenCount>
,
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<typeStatus id="A1398090FF8BE35AFEE2F8E480FDDCC4" box="[378,486,1919,1946]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
) [
<collectionCode id="1893A6F7FF8BE35AFE67F81B8324DCC4" box="[511,575,1920,1946]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="15" pageNumber="16" type="Museum">MCZ</collectionCode>
32179] (examined); (one alate gyne,
<specimenCount id="6884F5BBFF8BE35AFC61F81B8578DCC4" box="[1017,1123,1919,1946]" pageId="15" pageNumber="16" type="male">one male</specimenCount>
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) [
<collectionCode id="1893A6F7FF8BE35AFA9CF8E48445DCC4" box="[1284,1374,1919,1946]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="15" pageNumber="16">NHMB</collectionCode>
] (examined)
</materialsCitation>
.
<materialsCitation id="CEEA346FFF8BE35AFE9AF83F834EDCBC" ID-GBIF-Occurrence="4029332439" collectingDate="1972-02-10" collectingDateMax="1972-02-25" collectingDateMin="1972-02-10" collectionCode="ANIC" collectorName="Stewart B. Peck &amp; Jarmila Peck" country="Colombia" location="Leticia" pageId="15" pageNumber="16" specimenCount="1" specimenCount-worker="1" typeStatus="paratype">
<collectingCountry id="06957EA2FF8BE35AFE9AF83F8087DCE0" box="[258,412,1956,1982]" name="Colombia" pageId="15" pageNumber="16">COLOMBIA</collectingCountry>
:
<location id="7B5D68E9FF8BE35AFE3FF83880E2DCE3" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF8BE35AFE3FF83880E2DCE3" box="[423,505,1955,1981]" country="Colombia" name="Leticia" pageId="15" pageNumber="16">Leticia</location>
:
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<quantity id="B97A93D7FF8BE35AFD9CF83F8325DCE3" box="[516,574,1955,1982]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="16" unit="km" value="7.0">7 km</quantity>
North of Leticia
</locationDeviation>
,
<date id="0A3C18F2FF8BE35AFC92F83882B3DCE0" box="[778,936,1955,1982]" pageId="15" pageNumber="16" value="1972-02-10" valueMax="1972-02-25" valueMin="1972-02-10">
<collectingDate id="1A78E11AFF8BE35AFC92F83882B3DCE0" box="[778,936,1955,1982]" pageId="15" pageNumber="16" value="1972-02-10" valueMax="1972-02-25" valueMin="1972-02-10">1025.ii.1972</collectingDate>
</date>
,
<collectorName id="D3775BE4FF8BE35AFC2CF83F8572DCE3" box="[948,1129,1955,1982]" pageId="15" pageNumber="16">Stewart B. Peck</collectorName>
&amp;
<collectorName id="D3775BE4FF8BE35AFB17F8388404DCE3" box="[1167,1311,1955,1981]" pageId="15" pageNumber="16">Jarmila Peck</collectorName>
cols., (
<specimenCount id="6884F5BBFF8BE35AFAEAF83F81FCDCBC" pageId="15" pageNumber="16" type="worker">one worker</specimenCount>
<typeStatus id="A1398090FF8BE35AFE9CF85C807EDCBC" box="[260,357,1991,2018]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
) [
<collectionCode id="1893A6F7FF8BE35AFEE2F85C80DFDCBC" box="[378,452,1991,2018]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="15" pageNumber="16">ANIC</collectionCode>
] (examined)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FF94E345FF5FFF0D856DDBA6" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="7E3D3E32FF94E345FF5FFF0D856DDBA6" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">
<emphasis id="4CF6E220FF94E345FF5FFF0D8059DBEE" bold="true" box="[199,322,150,176]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
Dark amber to black; integument punctuate-foveate; clypeomandibular space narrow; corners of vertexal margin rounded, sulcus present medially and separating posterofrontal tumosity; petiolar node truncate anteriorly; postpetiolar and petiolar nodes usually densely covered by squamiform hairs.
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FF94E346FF5FFE998209DCE7" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17" type="description">
<paragraph id="7E3D3E32FF94E345FF5FFE9983C7DAF2" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">
<emphasis id="4CF6E220FF94E345FF5FFE998042DA42" bold="true" box="[199,345,258,284]" pageId="16" pageNumber="17">Description.</emphasis>
<emphasis id="4CF6E220FF94E345FEF9FE9980A9DA42" box="[353,434,258,284]" italics="true" pageId="16" pageNumber="17">Worker</emphasis>
(n=4). HL 1.251.28, HL2 1.251.30, HW1 1.281.31, MdL 0.720.78, SL1 0.840.89, SL2 0.890.94, PDL 0.110.13, A3L 0.030.05, AFL 0.360.38, FuL 0.971.00, EL 0.160.17, EW 0.140.16, ML 1.691.72, MfL 1.331.34, MtL 1.001.06,
<collectingCountry id="06957EA2FF94E345FDE5FED083BADA3B" box="[637,673,331,357]" name="Philippines" pageId="16" pageNumber="17">PH</collectingCountry>
0.38, PL 0.77.081, PW 0.360.38, PPL 0.470.50, PPW 0.530.61, GL 1.591.63, GW 1.131.19, TL 6.586.61, CI 100105, CS 1.271.28, MCI 5662, SI 6973, ESI 1719, SAI2 247252, EI1 0.230.26, MFI 9597, PTI 204216.
</paragraph>
<paragraph id="7E3D3E32FF94E345FF5FFE2D80FFDE26" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">Body yellowish-amber to dark brown; appendages slightly lighter, yellowish to dark brown. Mandibles covered by tiny piligerous punctures, apex with short yellowish setae; interdental setae present, yellowish and filiform, subequal to teeth length. Basimandibular seta present, narrow and erect, slightly clavate. Suberect and clavate hairs in the median portion of the stipe dorsa. Frontoclypeal margin covered by spaced piligerous punctuation; lateroclypeal region with short and squamiform, decumbent yellow hairs. Occipital region predominantly covered by coarse (foveolar) piligerous punctuations; pilosity close to the posterolateral region of the head and the vertexal margin composed of whitish to yellowish, short and subdecumbent, squamiform hair. Lateral and vertexal margins of head covered by erect to suberect clavate hairs, yellowish and in the following conformation: one hair above the eyes, at the anterior limit of the antennal scrobe; four hairs on the posterolateral margin of the head, surrounding the meeting point between the posterior limit of the scrobe and the vertexal margin; one hair near the posterolateral corner of the head; four or five hairs on each side of the vertexal margin, separated by the median groove. A pair of yellowish erect and clavate hairs in the frontal region, close to the vertexal margin. Lateroventral head margin covered by whitish and squamiform hairs, subdecumbent; ventral surface of head with curved suberect and subdecumbent filiform setae, length variable. Occipital carina surrounded by four to six long and filiform hairs. Yellowish to whitish plumose pilosity, usually subdecumbent, on the mesosoma and metasoma in the following conformation: surrounding the anterolateral margin of pronotum; on the mesonotum and propodeum dorsa; marginating the sloping face of propodeum; densely on petiolar and postpetiolar node dorsa; three pairs on each side of the anterolateral region of the postpetiolar sternite; on the anterolateral region of procoxae and meso and metacoxae dorsa; on legs, from trochanters to basitarsi dorsum. Clavate hairs in the following conformation: one pair of erect hairs present on mesonotum dorsum close to promesonotal suture; erect to suberect hairs close to gastral sternite and tergite margins. Mesosternum shelf (surrounded by the epicnemial fossa) with short, filiform setae along its length. Long and filiform setae present on the anterior portion of procoxae and median portion of first gastral sternite. Thick and suberect setae present on the ventral margin of basitarsus to apex of distal tarsomere. Antennae pilosity: dorsal surface of the scape primarily covered by short, subdecumbent and squamiform hairs; external margin of scape with long, erect, and squamiform (on its apical half) hairs; funiculus densely covered with short yellowish setae; ventral margin of scape with longitudinal rows of median, curved and subdecumbent setae. Remaining of integument with thick piligerous punctuations.</paragraph>
<paragraph id="7E3D3E32FF94E345FF5FFA198073DDBE" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="17">Body mostly smooth and shiny on glabrous regions. Mandible and clypeus dorsa sparsely covered with punctuations. Anteroventral portion of mandibles and ventral margin of scapes finely alveolate. Head punctuate-foveate, posterolateral surface of antennal scrobe punctuate-rugose. Pronotum sparsely foveate. Dorsum of mesonotum, anterodorsal region of the propodeum, dorsal surface of the meso- and metacoxae, and dorsum of petiolar and postpetiolar node punctuate-rugose. Mesopleuron and lateral of the propodeum smooth and shiny or subopaque. Surface of propodeal declivity punctuate-rugose. Gaster densely punctuate-reticulate; tergite of abdominal segments V, VI, and VII finely and densely punctuate, slightly opaque, tergal margins smooth and shiny; sculpture sparser in the first gastral sternite, especially in the longitudinal axis of median region.Antennal scapes smooth or finely rugulose, usually shiny. Funiculi densely and finely punctuate, usually opaque. Legs smooth or superficially rugose; procoxae punctuate-foveate.</paragraph>
<paragraph id="7E3D3E32FF94E344FF5FF97185B3DA43" blockId="16.[151,1437,150,2013]" lastBlockId="17.[151,1437,151,753]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">Head subtrapezoidal, sides delimited by raised margin extending from eye height to posterolateral region. Vertexal margin with gently convex corners slightly projected posterad, and with median groove dividing tumosity on the posterofrontal region of the head. Cervical margin carinate. Palp formula 2,2; palps strongly fused, giving the impression of being unsegmented; maxillary palp slightly larger in size and slightly wider than labial; labial palp apically clavate. Stipes subrectangular. Labrum cuneiform; distal margin bilobed, lobes separated by a narrow cleft, basal region canaliculate. Mandibles triangular; in full-face view, external margins slightly concave; basal margin lamellar; basal angle obtuse, followed by a masticatory margin with ten triangular teeth, apical tooth slightly curved; in lateral view mandibular apex slightly curved ventrally. Clypeomandibular space narrow. Anterior clypeal margin lamellate; anterolateral portion gently convex; anterior margin slightly concave in its median portion. Scape with slightly obtuse basal angle, followed by translucent and crenulated lamellar portion. Antennal fossa deeply impressed. Antennal scrobe comparatively deep in its lower half, posterior limit faintly distinct.</paragraph>
<paragraph id="7E3D3E32FF95E344FF5FFEBC85DAD9DB" blockId="17.[151,1437,151,753]" pageId="17" pageNumber="18">Lateral profile of mesosoma with promesonotal complex subglobular. In dorsal view, promesonotal suture practically indistinct; metanotal suture broad and strongly impressed, longitudinally costulate. Mesopleuron anteriorly marginate, interrupted at the meeting of a conspicuous epicnemial fossa. In dorsal view, propodeum subrectangular. In lateral view, anterior portion of propodeum slightly oblique posteriorly, abruptly followed by the sloping face. Propodeal slope laterally carinate and with transverse carina connecting to short, triangular and acute projections. Opening of propodeal spiracle rounded. Metapleural gland bulla prominent, protruding; opening transversal and covered by cuticular lamella. Petiolar peduncle longitudinally carinate on dorsal surface. In dorsal view, propodeal spiracle projected laterally. In lateral view, petiolar node with anterior face truncate, posterior face covered by pilosity; postpetiole slightly longer than petiolar node. Subpetiolar process highly variable: anterior process, followed by spiniform lamella or anterior process bifid, followed by spines and/or angular lamellar process.</paragraph>
<paragraph id="7E3D3E32FF95E344FF5FFD148579D9AF" blockId="17.[151,1437,151,753]" pageId="17" pageNumber="18">In dorsal view, petiolar node longitudinally subrectangular; postpetiole (pilosity excluded) slightly wider than long; posterior margin convex and widely inserted in the anterior gastral cavity. Calcar of strigil pectinate. Pro-, meso-, and metabasitarsi longer than the sum of other tarsomeres. Tarsal claws simple.</paragraph>
<caption id="2AFD6EBAFF95E344FF0FFA6183A1DD68" ID-DOI="http://doi.org/10.5281/zenodo.6605973" ID-Zenodo-Dep="6605973" httpUri="https://zenodo.org/record/6605973/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1530,1554]" targetBox="[155,1432,777,1506]" targetPageId="17">
<paragraph id="7E3D3E32FF95E344FF0FFA6183A1DD68" blockId="17.[151,1437,1530,1590]" pageId="17" pageNumber="18">
<emphasis id="4CF6E220FF95E344FF0FFA618009DD4C" bold="true" box="[151,274,1530,1554]" pageId="17" pageNumber="18">FIGURE 5.</emphasis>
<taxonomicName id="B98245B1FF95E344FE80FA6080F2DD4C" authorityName="Brown" authorityYear="1974" box="[280,489,1531,1554]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF95E344FE80FA6080F2DD4C" box="[280,489,1531,1554]" italics="true" pageId="17" pageNumber="18">Basiceros conjugans</emphasis>
</taxonomicName>
, holotype worker (MCZ 32179, Ecuador: Limoncocha); A. full-face view, B. lateral view, and C. dorsal view. Scale bars: A = 0.5 mm, BC = 1 mm.
</paragraph>
</caption>
<paragraph id="7E3D3E32FF95E344FF5FF9FE82ECDC6C" blockId="17.[151,1436,1636,2023]" pageId="17" pageNumber="18">
<emphasis id="4CF6E220FF95E344FF5FF9FE8019DD21" box="[199,258,1637,1663]" italics="true" pageId="17" pageNumber="18">Gyne</emphasis>
(n=3). HL 1.311.35, HL2 1.341.39, HW1 1.141.19, MdL 0.720.75, SL1 0.840.88, SL2 0.850.94, PDL 0.130.14, A3L 0.05, AFL 0.380.41, FuL 0.971.00, EL 0.200.23, EW 0.160.19, LOD 0.070.08, MOD 0.080.10, OOD 0.370.43, ML 1.701.84, MSL 0.750.81, MSW 0.820.87, MLL 0.300.31, MLW 0.450.50, MfL 1.231.25, MtL 0.971.00,
<collectingCountry id="06957EA2FF95E344FD92F94A8335DDB5" box="[522,558,1745,1771]" name="Philippines" pageId="17" pageNumber="18">PH</collectingCountry>
0.410.44, PL 0.750.81, PPL 0.530.55, PW 0.210.34, PPW 0.50.63, GL 1.882.03, GW 1.251.36, TL 6.917.30, CI 8488, CS 1.231.27, MCI 5455, SI 7479, ESI 2226, SAI2 226 232, EI1 0.310.32, MTI 105116, MLI 145160, MFI 9295, PTI 184188.
</paragraph>
<paragraph id="7E3D3E32FF95E347FF5FF8A78441D947" blockId="17.[151,1436,1636,2023]" lastBlockId="18.[151,1437,151,537]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
Color and sculpture similar to the worker; size slightly larger. Cephalic dorsum with three ocelli: median ocelli inserted slightly below and lateral ocelli inserted just above a pair of clavate and erect hairs. Head and waist pilosity as in workers. Pilosity of anterolateral margin of pronotum denser and longer than in workers. Whitish and subdecumbent, squamiform hairs close to humeral angles and surrounding the posterior limit of pronotum; short and sparse on the dorsum of mesoscutum; on the scutoscutellar sulcus axillae; on the dorsum of mesoscutellum; a suberect pair on the metanotal flange. Whitish erect to suberect and clavate hairs on each side of pronotum, one pair close to the pronotal suture; a pair on the metanotal flange; narrower and arranged in eight pairs on the dorsum of mesoscutum; one pair in each parapside; one on each lateral portion of axilla, one on the lateral margin of mesoscutellum. Hairs on gaster more abundant than on workers. In dorsal view, mesoscutum anteriorly rounded, slightly cuneiform, posterior margin slightly convex in the meeting with the scutellar suture; notauli indistinct; parapsidal lines narrow and inconspicuous, involved by the sculpture; shallow, parapsis rudimentary; tegula narrow, apical margin rounded. Prescutellum narrow; axillae projected posteriorly, rounded and slightly depressed. Scutellar sulcus well marked. Mesoscutellum transversely subrectangular, posterior limit concave. Dorsal face of propodeum strongly inclined. In lateral view, anapleural sulcus broader anteriorly at the connection with the epicnemial fossa, narrowing posteriorly. First gastral sternite with median region slightly projecting on its basal half. Forewing mostly conforming to
<typeStatus id="A1398090FF96E347FEA6FE648075D947" box="[318,366,511,537]" pageId="18" pageNumber="19">type</typeStatus>
2, transversal vein 1m-cu present as an appendage; hindwing with fix submedian hamuli.
</paragraph>
<caption id="2AFD6EBAFF96E347FF0FF9938411DD1D" ID-DOI="http://doi.org/10.5281/zenodo.6605977" ID-Zenodo-Dep="6605977" httpUri="https://zenodo.org/record/6605977/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1544,1568]" targetBox="[151,1435,561,1520]" targetPageId="18">
<paragraph id="7E3D3E32FF96E347FF0FF9938411DD1D" blockId="18.[151,1436,1544,1604]" pageId="18" pageNumber="19">
<emphasis id="4CF6E220FF96E347FF0FF993800FDD7E" bold="true" box="[151,276,1544,1568]" pageId="18" pageNumber="19">FIGURE 6.</emphasis>
<taxonomicName id="B98245B1FF96E347FE83F99280F6DD7E" authorityName="Brown" authorityYear="1974" box="[283,493,1545,1568]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF96E347FE83F99280F6DD7E" box="[283,493,1545,1568]" italics="true" pageId="18" pageNumber="19">Basiceros conjugans</emphasis>
</taxonomicName>
, paratypes gyne and male (MCZ 546463, Ecuador: Limoncocha), Ecuador: Limoncocha); A and D. full-face view, B and E. dorsal view, C and F. lateral view. Scale bars: A, D = 0.5 mm, B, C, E, F =1 mm.
</paragraph>
</caption>
<paragraph id="7E3D3E32FF96E347FF5FF9E98365DC3B" blockId="18.[151,1437,1650,2037]" pageId="18" pageNumber="19">
<emphasis id="4CF6E220FF96E347FF5FF9E9801ADDD2" box="[199,257,1650,1676]" italics="true" pageId="18" pageNumber="19">Male</emphasis>
(n=2). HL 0.850.93, HW1 0.770.81, HW2
<emphasis id="4CF6E220FF96E347FD67F9E9822BDDD2" bold="true" box="[767,816,1650,1676]" pageId="18" pageNumber="19">0.63</emphasis>
0.95, MdL 0.400.43, SL2 0.150.21, PDL 0.100.13, A3L 0.280.31, AFL 0.440.48, EL 0.300.33, EW 0.230.26, LOD 0.09, MOD 0.080.09, OOD 0.310.34, ML 1.431.59, MSL 0.750.85, MSW 0.700.78, MLL 0.300.34, MLW 0.430.48, MfL 1.151.25, MtL 0.830.88,
<collectingCountry id="06957EA2FF96E347FF0FF94481A0DDA7" box="[151,187,1759,1785]" name="Philippines" pageId="18" pageNumber="19">PH</collectingCountry>
0.250.30, PL 0.630.70, PW 0.230.33, PPL 0.310.36, PPW 0.340.40, GL 1.361.50, GW 0.981.10, TL 5.015.41, CI 8491, CS 0.810.87, MCI 4447, SI 1928, ESI 150200, SAI 5070, SAI2 3450, EI1 0.650.71, EI2 80
<emphasis id="4CF6E220FF96E347FF6AF8BD8007DC1E" bold="true" box="[242,284,1830,1856]" pageId="18" pageNumber="19">124</emphasis>
, MTI 8798, MLI 136141, MFI 6167, PTI 231250. (In bold: measurements suffered influence of prolegs partially covering compound eyes).
</paragraph>
<paragraph id="7E3D3E32FF96E346FF5FF8F581CCDA1F" blockId="18.[151,1437,1650,2037]" lastBlockId="19.[151,1437,151,1977]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Size slightly smaller than the conspecific gyne. Color black; postpetiole and first gastral tergite disc brown; appendages yellowish to light brown. Apical portion of mandibles smooth and yellowish. Wings yellowish to brown. Mandible dorsa and apex with long and fine yellow hairs, semi-erect to subdecumbent, slightly longer on apical portion of mandibles. Head with two main
<typeStatus id="A1398090FF96E347FDE1F84083AFDCAB" box="[633,692,2011,2037]" pageId="18" pageNumber="19">types</typeStatus>
of hair: medium and fine, yellow and subdecumbent, primarily on the frontal disc of clypeus; long and whitish to yellowish (sometimes with a curved apex), present along the genal carina, on the vertexal margin, and the ventral surface. The second hair
<typeStatus id="A1398090FF97E346FC29FF2082FADB8B" box="[945,993,187,213]" pageId="19" pageNumber="20">type</typeStatus>
widely present throughout the body: on mesosomal dorsum, waist, and gaster, exceptionally long on the ocellar region, on procoxae and dorsum of petiole. Antennomeres with short and yellowish appressed setae. Legs with medium and yellowish decumbent to decumbent setae.
</paragraph>
<paragraph id="7E3D3E32FF97E346FF5FFED081CED947" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">Body uniformly punctuate-reticulate, sculpture varying on diameter and degree of impression. Apical portion of mandibles smooth and shiny. Irregular longitudinal rugae present in the neck to faintly on the vertex margin, close to occipital carina. Irregular transverse rugae on the dorsum of mesoscutellum, on the metakatepisternum and propodeum. Mesoanepisternum slightly darker, smooth and shiny over a little more than half its length; posteromedial portion of mesokatepisternum smooth and shiny. Dorsolateral rugae present on the anterior portion of petiolar node.</paragraph>
<paragraph id="7E3D3E32FF97E346FF5FFDB8808EDE27" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">
Head subpiriform; occipital margin wide and lamellar, medially concave. Palp formula 1,1; palpi slightly swollen and flattened on apical half; maxillary palp appears to be slightly longer and wider than labial. Mentum narrow and triangular. Stipes subrectangular. Labrum elongated, distal margin bilobed; approximately ten long setae present. Mandibles triangular, curved towards their apexes; masticatory margin with nine teeth of similar size. Clypeus with central disc convex, slightly elevated; lateral regions depressed; anterior margin lamellar and slightly concave. Postgenal carina present, shaped like a longitudinal line, smooth and shiny just after the supraclypeal region and extending posteriorly on the head until close to the upper limits of the antennal scrobes. Antennal arch medially expanded as swollen posterolateral lobe, completely hiding the antennal bulb in frontal view. Antenna pedicel longer than wide, third antennomere about three times longer than the pedicel. Eyes large and globular, protruding from the cephalic capsule. Ocelli caramel-colored, projected. Mesoscutum cuneiform in dorsal view, elongated anteriorly. Smooth and shiny carina present on the anteromedial region of mesoscutum, extending as a line near the dorsal margin. Notauli V-shaped, converging at the center of mesoscutellum and extending as a median longitudinal sulcus to the line of the scutellar suture. Parapsidal lines shiny; curved on its anterior portion, subparallel and directed anterolaterally to the parascutal flange. Parapsis oval. Transcutellar sulcus slightly angled medially. Axillae protruded, strongly curved posteriorly; hook-shaped. Anapleural suture broad, strongly impressed and scrobiculate—mesoanepisternum conspicuously elevated relative to mesokatepisternum. Scutoscutellar sutured narrower compared to other
<taxonomicName id="B98245B1FF97E346FE4FFBF8835FDF22" authorityName="Schulz" authorityYear="1906" box="[471,580,1123,1148]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="4CF6E220FF97E346FE4FFBF8835FDF22" box="[471,580,1123,1148]" italics="true" pageId="19" pageNumber="20">Basiceros</emphasis>
</taxonomicName>
males; smooth and with a median transverse carina. Mesoscutellum subrectangular; posteromedially depressed; posterior margin strongly concave and depressed. Metanotum with posterior margin lamellar. Propodeum armed with triangular, laminar projections. Propodeal lobes auricular. Calcar of strigil short and pectinate. Tarsal claws simple; short arolia present. Petiole claviform; in lateral view petiolar node low; petiolar spiracle projected in dorsal view. Subpetiolar process with anteriorly curved projection followed by 14 teeth or with lamella or both (as
<bibRefCitation id="1A1343C3FF97E346FD94FA8C83B0DE6F" author="Brown, W. L. Jr." box="[524,683,1303,1329]" pageId="19" pageNumber="20" pagination="131 - 140" refId="ref61175" refString="Brown, W. L. Jr. (1974) A supplement to the revision of the ant genus Basiceros. Journal of the New York Entomological Society, 82, 131 - 140." type="journal article" year="1974">Brown (1974)</bibRefCitation>
mentioned: “very inconstant”). Postpetiole approximately half the length of petiole in lateral view. In dorsal view, petiolar node rounded. Forewing
<typeStatus id="A1398090FF97E346FBA8FAA0857BDE0B" box="[1072,1120,1339,1365]" pageId="19" pageNumber="20">type</typeStatus>
1 or
<typeStatus id="A1398090FF97E346FB05FAA085D6DE0B" box="[1181,1229,1339,1365]" pageId="19" pageNumber="20">type</typeStatus>
2. Hindwing with six submedian hamuli.
</paragraph>
<paragraph id="7E3D3E32FF97E346FF5FFA188209DCE7" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">
<emphasis id="4CF6E220FF97E346FF5FFA188011DEC2" box="[199,266,1411,1436]" italics="true" pageId="19" pageNumber="20">Larva</emphasis>
(first description, based on
<specimenCount id="6884F5BBFF97E346FDCBFA188214DEC0" box="[595,783,1411,1438]" pageId="19" pageNumber="20" type="generic">three specimens</specimenCount>
). Approximate length through spiracles:
<quantity id="B97A93D7FF97E346FB7FFA188458DEC3" box="[1255,1347,1411,1437]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="19" pageNumber="20" unit="mm" value="3.7">3.7 mm</quantity>
; profile pogonomyrmecoid: largest diameter near middle of abdominal region, thoracic region slender than abdominal region; curved ventrally. Anus ventral, anal opening weakly convex. Spiracles small. Integument of ventral region densely covered by spinules arranged in transverse rows. Pilosity moderately dense, denser near abdominal apex, sparse on the rest of the body; hairs long (
<quantity id="B97A93D7FF97E346FDEAF9888200DD73" box="[626,795,1555,1581]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="19" pageNumber="20" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.100.30 mm</quantity>
), flexible and indented; alveolus and articular membrane present or not. Cranium suboctagonal, antennae very small. Clypeus protruded. Labrum bilobed, slightly wider than twice its length; ventral face densely spinulose; anteroventral margin of each lobe with about 45 rounded sensilla; anterodorsal border with two isolated short setae resembling trichoid sensilla. Mandibles pogonomyrmecoid, long and narrow, medially curved; apical tooth slightly curved and with rounded apex, separated from other teeth by conspicuous diastema; two protruding teeth projected from the basal margin; anteroventral margin with spiculae in short and arcuate subtransversal lines. Maxillae parabolic, dorsum with 4 setae, apex spinulose; maxillary palps digitiform, elongated and narrow, with three apical encapsulated sensilla and one spinulose lateral sensillum; galea digitiform, with two apical encapsulated sensilla. Labium covered by short transverse rows of spinules; those developed, capillary (about
<quantity id="B97A93D7FF97E346FE2EF8CC8300DC2F" box="[438,539,1879,1905]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="mm" value="0.02">0.02 mm</quantity>
) and densely covering the rest of labium surface; labial palp papillary, with five apical sensilla (two encapsulated and three with spinules); opening of sericeous gland transversal, bearing a short isolated sensillum on each side. Hypopharynx spinulose.
</paragraph>
</subSubSection>
<caption id="2AFD6EBAFF90E341FF0FFB59858FDFA0" ID-DOI="http://doi.org/10.5281/zenodo.6605979" ID-Zenodo-Dep="6605979" httpUri="https://zenodo.org/record/6605979/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1218,1242]" targetBox="[154,1433,181,1195]" targetPageId="20">
<paragraph id="7E3D3E32FF90E341FF0FFB59858FDFA0" blockId="20.[151,1436,1218,1279]" pageId="20" pageNumber="21">
<emphasis id="4CF6E220FF90E341FF0FFB598008DF85" bold="true" box="[151,275,1218,1243]" pageId="20" pageNumber="21">FIGURE 7.</emphasis>
<taxonomicName id="B98245B1FF90E341FE81FB5880F1DF84" authorityName="Brown" authorityYear="1974" box="[281,490,1219,1242]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF90E341FE81FB5880F1DF84" box="[281,490,1219,1242]" italics="true" pageId="20" pageNumber="21">Basiceros conjugans</emphasis>
</taxonomicName>
, larva (MZSP, Peru: Madre de Dios); A. head in anteroventral view; B. head in oblique view; C. lateral view; D. dorsal pilosity; E. anus. Scale bars: A, B, E = 0.1 mm; C = 0.5 mm; E = 0.05 mm.
</paragraph>
</caption>
<subSubSection id="36986DB9FF90E341FF5FFAB68017DE89" pageId="20" pageNumber="21" type="etymology">
<paragraph id="7E3D3E32FF90E341FF5FFAB68017DE89" blockId="20.[151,1436,1325,2036]" pageId="20" pageNumber="21">
<emphasis id="4CF6E220FF90E341FF5FFAB68055DE19" bold="true" box="[199,334,1325,1351]" pageId="20" pageNumber="21">Etymology.</emphasis>
from the Latin
<emphasis id="4CF6E220FF90E341FD9FFAB5836CDE19" box="[519,631,1326,1351]" italics="true" pageId="20" pageNumber="21">conjugant</emphasis>
- (
<taxonomicName id="B98245B1FF90E341FD0BFAB5821DDE19" authorityName="Brown" authorityYear="1974" box="[659,774,1326,1351]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF90E341FD0BFAB5821DDE19" box="[659,774,1326,1351]" italics="true" pageId="20" pageNumber="21">conjugans</emphasis>
</taxonomicName>
), present participle of the verb
<emphasis id="4CF6E220FF90E341FBECFAB585FFDE19" box="[1140,1252,1326,1351]" italics="true" pageId="20" pageNumber="21">conjugare</emphasis>
, meaning “connecting”, “uniting”. Such an epithet is undoubtedly a reference to the intermediate morphology of this species among the other taxa of the then considered separate genera
<taxonomicName id="B98245B1FF90E341FCAEFAEE82B2DED1" authorityName=", Weber" authorityYear="1950" box="[822,937,1397,1423]" class="Insecta" family="Formicidae" genus="Aspididris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="4CF6E220FF90E341FCAEFAEE82B2DED1" box="[822,937,1397,1423]" italics="true" pageId="20" pageNumber="21">Aspididris</emphasis>
</taxonomicName>
and
<taxonomicName id="B98245B1FF90E341FC46FAED8550DED1" authorityName="Schulz" authorityYear="1906" box="[990,1099,1398,1423]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="4CF6E220FF90E341FC46FAED8550DED1" box="[990,1099,1398,1423]" italics="true" pageId="20" pageNumber="21">Basiceros</emphasis>
</taxonomicName>
. When describing the species,
<bibRefCitation id="1A1343C3FF90E341FF0FFA02802DDEED" author="Brown, W. L. Jr." box="[151,310,1433,1459]" pageId="20" pageNumber="21" pagination="131 - 140" refId="ref61175" refString="Brown, W. L. Jr. (1974) A supplement to the revision of the ant genus Basiceros. Journal of the New York Entomological Society, 82, 131 - 140." type="journal article" year="1974">Brown (1974)</bibRefCitation>
refers to
<taxonomicName id="B98245B1FF90E341FE3BFA01832DDEED" authorityName="Brown" authorityYear="1974" box="[419,566,1434,1459]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF90E341FE3BFA01832DDEED" box="[419,566,1434,1459]" italics="true" pageId="20" pageNumber="21">B. conjugans</emphasis>
</taxonomicName>
as the “additional intergradient” and formally synonymizes
<taxonomicName id="B98245B1FF90E341FB79FA02844FDEED" authorityName=", Weber" authorityYear="1950" box="[1249,1364,1433,1459]" class="Insecta" family="Formicidae" genus="Aspididris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="4CF6E220FF90E341FB79FA02844FDEED" box="[1249,1364,1433,1459]" italics="true" pageId="20" pageNumber="21">Aspididris</emphasis>
</taxonomicName>
under
<taxonomicName id="B98245B1FF90E341FF0FFA25801FDE89" authorityName="Schulz" authorityYear="1906" box="[151,260,1470,1495]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="4CF6E220FF90E341FF0FFA25801FDE89" box="[151,260,1470,1495]" italics="true" pageId="20" pageNumber="21">Basiceros</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FF90E340FF5FFA7A8209D84B" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="discussion">
<paragraph id="7E3D3E32FF90E341FF5FFA7A827EDCD9" blockId="20.[151,1436,1325,2036]" pageId="20" pageNumber="21">
<emphasis id="4CF6E220FF90E341FF5FFA7A8054DEA5" bold="true" box="[199,335,1505,1531]" pageId="20" pageNumber="21">Comments.</emphasis>
<bibRefCitation id="1A1343C3FF90E341FECEFA7A80EEDEA5" author="Brown, W. L. Jr." box="[342,501,1505,1531]" pageId="20" pageNumber="21" pagination="131 - 140" refId="ref61175" refString="Brown, W. L. Jr. (1974) A supplement to the revision of the ant genus Basiceros. Journal of the New York Entomological Society, 82, 131 - 140." type="journal article" year="1974">Brown (1974)</bibRefCitation>
, based on the shape of the vertexal margin and the clavate hairs distributed along its length, speculates on the intermediate state of
<taxonomicName id="B98245B1FF90E341FD38F99D8229DD41" authorityName="Brown" authorityYear="1974" box="[672,818,1542,1567]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF90E341FD38F99D8229DD41" box="[672,818,1542,1567]" italics="true" pageId="20" pageNumber="21">B. conjugans</emphasis>
</taxonomicName>
within the genus when comparing it to
<taxonomicName id="B98245B1FF90E341FB6BF99D8477DD41" baseAuthorityName="Mayr" baseAuthorityYear="1887" box="[1267,1388,1541,1567]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="disciger">
<emphasis id="4CF6E220FF90E341FB6BF99D8477DD41" box="[1267,1388,1541,1567]" italics="true" pageId="20" pageNumber="21">B. disciger</emphasis>
</taxonomicName>
and
<taxonomicName id="B98245B1FF90E341FF0FF9B181E0DD1D" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[151,251,1578,1603]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FF90E341FF0FF9B181E0DD1D" box="[151,251,1578,1603]" italics="true" pageId="20" pageNumber="21">B. manni</emphasis>
</taxonomicName>
.
<taxonomicName id="B98245B1FF90E341FE90F9B180F4DD1D" authorityName="Brown" authorityYear="1974" box="[264,495,1578,1603]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF90E341FE90F9B180F4DD1D" box="[264,495,1578,1603]" italics="true" pageId="20" pageNumber="21">Basiceros conjugans</emphasis>
</taxonomicName>
presents what Brown categorizes as “an intermediate pattern of transversal ridges on the vertexal margin”; the same for the concentration of clavate hairs in this region. The following could be added to the list of “intermediate” characters: head shape, more elongated than in
<taxonomicName id="B98245B1FF90E341FC24F9E9852EDDD5" baseAuthorityName="Mayr" baseAuthorityYear="1887" box="[956,1077,1649,1675]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="disciger">
<emphasis id="4CF6E220FF90E341FC24F9E9852EDDD5" box="[956,1077,1649,1675]" italics="true" pageId="20" pageNumber="21">B. disciger</emphasis>
</taxonomicName>
and
<taxonomicName id="B98245B1FF90E341FBF1F9E985FCDDD5" baseAuthorityName="Weber" baseAuthorityYear="1950" box="[1129,1255,1649,1675]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="militaris">
<emphasis id="4CF6E220FF90E341FBF1F9E985FCDDD5" box="[1129,1255,1649,1675]" italics="true" pageId="20" pageNumber="21">B. militaris</emphasis>
</taxonomicName>
; the presence of a narrow clypeomandibular space, separating the clypeus from the basal region of the mandibles—absent in
<taxonomicName id="B98245B1FF90E341FACBF90D81CADD8D" baseAuthorityName="Mayr" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="disciger">
<emphasis id="4CF6E220FF90E341FACBF90D81CADD8D" italics="true" pageId="20" pageNumber="21">B. disciger</emphasis>
</taxonomicName>
and
<taxonomicName id="B98245B1FF90E341FE9EF921809FDD8D" baseAuthorityName="Weber" baseAuthorityYear="1950" box="[262,388,1721,1747]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="militaris">
<emphasis id="4CF6E220FF90E341FE9EF921809FDD8D" box="[262,388,1721,1747]" italics="true" pageId="20" pageNumber="21">B. militaris</emphasis>
</taxonomicName>
and broad in
<taxonomicName id="B98245B1FF90E341FD84F921839ADD8D" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[540,641,1722,1747]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FF90E341FD84F921839ADD8D" box="[540,641,1722,1747]" italics="true" pageId="20" pageNumber="21">B. manni</emphasis>
</taxonomicName>
and
<taxonomicName id="B98245B1FF90E341FD2FF921825EDD8D" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[695,837,1721,1747]" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="4CF6E220FF90E341FD2FF921825EDD8D" box="[695,837,1721,1747]" italics="true" pageId="20" pageNumber="21">B. singularis</emphasis>
</taxonomicName>
; and the specialized pilosity (clavate or subplumose), denser than observed for
<taxonomicName id="B98245B1FF90E341FE2BF9458337DDA9" baseAuthorityName="Mayr" baseAuthorityYear="1887" box="[435,556,1757,1783]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="disciger">
<emphasis id="4CF6E220FF90E341FE2BF9458337DDA9" box="[435,556,1757,1783]" italics="true" pageId="20" pageNumber="21">B. disciger</emphasis>
</taxonomicName>
and
<taxonomicName id="B98245B1FF90E341FDFAF94583FADDA9" baseAuthorityName="Weber" baseAuthorityYear="1950" box="[610,737,1757,1783]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="militaris">
<emphasis id="4CF6E220FF90E341FDFAF94583FADDA9" box="[610,737,1757,1783]" italics="true" pageId="20" pageNumber="21">B. militaris</emphasis>
</taxonomicName>
—what could explain the greater tendency for soil and vegetation particles to adhere to females of
<taxonomicName id="B98245B1FF90E341FDB7F89983DBDC45" authorityName="Brown" authorityYear="1974" box="[559,704,1794,1819]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF90E341FDB7F89983DBDC45" box="[559,704,1794,1819]" italics="true" pageId="20" pageNumber="21">B. conjugans</emphasis>
</taxonomicName>
compared to other taxa from the
<taxonomicName id="B98245B1FF90E341FBB5F89A859CDC45" baseAuthorityName="Mayr" baseAuthorityYear="1887" box="[1069,1159,1793,1819]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="disciger">
<emphasis id="4CF6E220FF90E341FBB5F89A859CDC45" box="[1069,1159,1793,1819]" italics="true" pageId="20" pageNumber="21">disciger</emphasis>
</taxonomicName>
clade. In addition, males of
<taxonomicName id="B98245B1FF90E341FF2DF8BD805DDC61" authorityName="Brown" authorityYear="1974" box="[181,326,1830,1855]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF90E341FF2DF8BD805DDC61" box="[181,326,1830,1855]" italics="true" pageId="20" pageNumber="21">B. conjugans</emphasis>
</taxonomicName>
may present a complete 1m-cu vein, closing the discal cell. This characteristic is found in
<taxonomicName id="B98245B1FF90E341FAB7F8BD8487DC61" authorityName="Schulz" authorityYear="1906" box="[1327,1436,1830,1855]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="4CF6E220FF90E341FAB7F8BD8487DC61" box="[1327,1436,1830,1855]" italics="true" pageId="20" pageNumber="21">Basiceros</emphasis>
</taxonomicName>
males from the
<taxonomicName id="B98245B1FF90E341FEDEF8D280ADDC3D" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[326,438,1865,1891]" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="4CF6E220FF90E341FEDEF8D280ADDC3D" box="[326,438,1865,1891]" italics="true" pageId="20" pageNumber="21">singularis</emphasis>
</taxonomicName>
clade, but it is absent in other males from the
<taxonomicName id="B98245B1FF90E341FC25F8D2850CDC3D" baseAuthorityName="Mayr" baseAuthorityYear="1887" box="[957,1047,1865,1891]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="disciger">
<emphasis id="4CF6E220FF90E341FC25F8D2850CDC3D" box="[957,1047,1865,1891]" italics="true" pageId="20" pageNumber="21">disciger</emphasis>
</taxonomicName>
clade, confirming that this species might have a combination of characteristics of these two clades.
</paragraph>
<paragraph id="7E3D3E32FF90E340FF5FF80A82AFDA1F" blockId="20.[151,1436,1325,2036]" lastBlockId="21.[151,1437,151,2014]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
In terms of variation, workers of
<taxonomicName id="B98245B1FF90E341FDDBF80983CDDCF5" authorityName="Brown" authorityYear="1974" box="[579,726,1938,1963]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF90E341FDDBF80983CDDCF5" box="[579,726,1938,1963]" italics="true" pageId="20" pageNumber="21">B. conjugans</emphasis>
</taxonomicName>
present variable integumentary and pilosity colorations, depth of the metanotal suture, shape of the petiolar node and subpetiolar process, integumentary sculpture (degree of impression and brightness), and size. However, together with
<taxonomicName id="B98245B1FF90E341FCD3F84182EADCAD" baseAuthorityName="Mayr" baseAuthorityYear="1887" box="[843,1009,2010,2035]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="convexiceps">
<emphasis id="4CF6E220FF90E341FCD3F84182EADCAD" box="[843,1009,2010,2035]" italics="true" pageId="20" pageNumber="21">B. convexiceps</emphasis>
</taxonomicName>
, it is one of the most morphologically “cohesive” species. The specialized pilosity pattern (with emphasis on the vertexal margin, back of propodeum, petiolar node, and dorsal margin of the postpetiole) and head and mesosoma morphologies are present similarly on all examined specimens. It is important to mention that the pilosity is present as described for specimens in good conditions, but some specimens—probably resultant of abrasion—lack several special hairs, such as those on the vertexal margin and the back of the mesoscutum (in the case of gynes).
</paragraph>
<paragraph id="7E3D3E32FF91E340FF5FFED080F1D9DB" blockId="21.[151,1437,151,2014]" pageId="21" pageNumber="22">
The
<typeStatus id="A1398090FF91E340FF63FED08046DA3B" box="[251,349,331,357]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
has a reddish-brown color and is probably a young worker, as other examined specimens from
<collectingCountry id="06957EA2FF91E340FF0FFEF481EFDAD7" box="[151,244,367,393]" name="Ecuador" pageId="21" pageNumber="22">Ecuador</collectingCountry>
are darker colored. Other Ecuadorian specimens have a similar reddish-brown color, usually within series containing darker specimens. The most extraordinary integumental coloration comes from Peruvian specimens collected in
<collectingRegion id="BC46F0D0FF91E340FE98FE2C8024DA8F" box="[256,319,439,465]" country="Ecuador" name="Napo" pageId="21" pageNumber="22">Napo</collectingRegion>
, close to the border with
<collectingCountry id="06957EA2FF91E340FDF9FE2C83D8DA8F" box="[609,707,439,465]" name="Ecuador" pageId="21" pageNumber="22">Ecuador</collectingCountry>
: a clear yellowish-amber and very bright integument. Regarding the morphology of the subpetiolar process, in the case of gynes and workers, it varies from: a set of two spines, composed of an angulate anteroventral “boot”-shaped or spine, followed by a lamella; a process with an anteroventral projection followed by 56 spines; or by lamellar processes and/or fused spines. The male subpetiolar process varies from an anterior curved process to a set of several short spines followed by a lamellar process at the end of the peduncles ventral margin.
</paragraph>
<paragraph id="7E3D3E32FF91E340FF5FFD148209D84B" blockId="21.[151,1437,151,2014]" pageId="21" pageNumber="22">In addition, some specimens were partially covered by litter and soil particles, making it difficult to see certain structures, such as the ocelli on the gynes. Males may show variation in the forewing venation, with the length of transversal 1m-cu vein present differentially in the same specimen: it may be absent or present as a short appendix in one wing while present as complete in the other wing.</paragraph>
</subSubSection>
<subSubSection id="36986DB9FF91E340FF5FFC8585EFDE9F" pageId="21" pageNumber="22" type="distribution">
<paragraph id="7E3D3E32FF91E340FF5FFC85800EDF07" blockId="21.[151,1437,151,2014]" pageId="21" pageNumber="22">
<emphasis id="4CF6E220FF91E340FF5FFC85807AD866" bold="true" box="[199,353,798,824]" pageId="21" pageNumber="22">Distribution.</emphasis>
This species is known for the Eastern Amazonian rainforests in South America and from secondary forests in
<collectingCountry id="06957EA2FF91E340FEAEFCD8808DD803" box="[310,406,835,861]" name="Trinidad and Tobago" pageId="21" pageNumber="22">Trinidad</collectingCountry>
—more precisely, at the Arena Forest Reserve, place of a sustainable management system of a tropical forest (Shelterwood System), developed in the 1950s. For the present study,
<taxonomicName id="B98245B1FF91E340FB19FCFC840FD8DE" authorityName="Brown" authorityYear="1974" box="[1153,1300,871,896]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF91E340FB19FCFC840FD8DE" box="[1153,1300,871,896]" italics="true" pageId="21" pageNumber="22">B. conjugans</emphasis>
</taxonomicName>
distribution is considerably expanded based on new records for the Brazilian states of Pará and Rondônia,
<collectingCountry id="06957EA2FF91E340FB53FC10841AD8FB" box="[1227,1281,907,933]" name="Peru" pageId="21" pageNumber="22">Peru</collectingCountry>
, and
<collectingCountry id="06957EA2FF91E340FAA6FC108482D8FB" box="[1342,1433,907,933]" name="Guyana" pageId="21" pageNumber="22">Guyana</collectingCountry>
.
<taxonomicName id="B98245B1FF91E340FF0FFC34809AD896" authorityName="Brown" authorityYear="1974" box="[151,385,943,968]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF91E340FF0FFC34809AD896" box="[151,385,943,968]" italics="true" pageId="21" pageNumber="22">Basiceros conjugans</emphasis>
</taxonomicName>
was known to
<collectingCountry id="06957EA2FF91E340FDADFC348362D897" box="[565,633,943,969]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
based on a single worker collected in Porto Walter (
<bibRefCitation id="1A1343C3FF91E340FB44FC348461D897" author="Delabie, J. H. C." box="[1244,1402,943,969]" pageId="21" pageNumber="22" pagination="272" refId="ref61422" refString="Delabie, J. H. C. (2000) First record of Basiceros conjugans (Hymenoptera: Formicidae: Basicerotini) in Brazil. Revista de Biologia Tropical, 48, 272." type="journal article" year="2000">Delabie 2000</bibRefCitation>
), a municipality in the state of
<collectingRegion id="BC46F0D0FF91E340FE49FC48831CD8B3" box="[465,519,979,1005]" country="Brazil" name="Acre" pageId="21" pageNumber="22">Acre</collectingRegion>
close to the Peruvian border. That worker was found in the stomach contents of the amphibian
<taxonomicName id="B98245B1FF91E340FEDAFC6C83ACDF4F" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[322,695,1014,1041]" class="Amphibia" family="Bufonidae" genus="Bufo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="typhonius">
<emphasis id="4CF6E220FF91E340FEDAFC6C80F6DF4F" box="[322,493,1015,1041]" italics="true" pageId="21" pageNumber="22">Bufo typhonius</emphasis>
(Linnaeus, 1758)
</taxonomicName>
(=
<taxonomicName id="B98245B1FF91E340FD7AFC6C85AADF4C" authority="(Laurenti, 1768))" authorityName="" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[738,1201,1015,1042]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="margaritifera">
<emphasis id="4CF6E220FF91E340FD7AFC6C82FADF4F" box="[738,993,1015,1041]" italics="true" pageId="21" pageNumber="22">Rhinella margaritifera</emphasis>
(Laurenti, 1768))
</taxonomicName>
. The material examined here indicates that this species possibly occurs in southern
<collectingCountry id="06957EA2FF91E340FCFDFB8082C3DF6B" box="[869,984,1051,1077]" name="Venezuela" pageId="21" pageNumber="22">Venezuela</collectingCountry>
and the Brazilian states of Roraima and
<collectingRegion id="BC46F0D0FF91E340FF0FFBA4800ADF07" box="[151,273,1087,1113]" country="Venezuela" name="Amazonas" pageId="21" pageNumber="22">Amazonas</collectingRegion>
.
</paragraph>
<paragraph id="7E3D3E32FF91E340FF5FFBF885EFDE9F" blockId="21.[151,1437,151,2014]" pageId="21" pageNumber="22">
<emphasis id="4CF6E220FF91E340FF5FFBF8809CDF22" bold="true" box="[199,391,1122,1149]" pageId="21" pageNumber="22">Natural history.</emphasis>
Little is known about the biology of this dirt ant. Specimens were manually collected from rotten logs or leaf litter samples, with most of the examined material collected via Winkler extractors. In 2012, as a student of the Neotropical Ant Course at the Sachavacayoc Center (
<collectingCountry id="06957EA2FF91E340FC35FB3082FEDF9B" box="[941,997,1195,1221]" name="Peru" pageId="21" pageNumber="22">Peru</collectingCountry>
,
<collectingRegion id="BC46F0D0FF91E340FC69FB308585DF9B" box="[1009,1182,1195,1221]" country="Peru" name="Madre de Dios" pageId="21" pageNumber="22">Madre de Díos</collectingRegion>
), one of us (RSP) collected part of a colony nesting in a rotten trunk. Next to the collected material (larvae, a pupa, few workers) was an unidentified gastropod shell, the body of a turtle ant worker (
<taxonomicName id="B98245B1FF91E340FCDEFB6882F6DE53" box="[838,1005,1267,1293]" class="Insecta" family="Formicidae" genus="Cephalotes" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4CF6E220FF91E340FCDEFB6882DFDE53" box="[838,964,1267,1293]" italics="true" pageId="21" pageNumber="22">Cephalotes</emphasis>
sp.
</taxonomicName>
), and two cephalic capsules: one from an unidentified trap-jaw ant worker (
<taxonomicName id="B98245B1FF91E340FDADFA8C821DDE6F" box="[565,774,1303,1329]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4CF6E220FF91E340FDADFA8C83C4DE6F" box="[565,735,1303,1329]" italics="true" pageId="21" pageNumber="22">Odontomachus</emphasis>
sp.
</taxonomicName>
) and one from the termite
<taxonomicName id="B98245B1FF91E340FBABFA8C8417DE6E" baseAuthorityName="Emerson" baseAuthorityYear="1925" box="[1075,1292,1303,1328]" class="Insecta" family="Termitidae" genus="Uncitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="teevani">
<emphasis id="4CF6E220FF91E340FBABFA8C8417DE6E" box="[1075,1292,1303,1328]" italics="true" pageId="21" pageNumber="22">Uncitermes teevani</emphasis>
</taxonomicName>
(
<taxonomicName id="B98245B1FF91E340FA82FA8C8483DE6F" authorityName="Latreille" authorityYear="1802" box="[1306,1432,1303,1329]" class="Insecta" family="Termitidae" kingdom="Animalia" order="Blattodea" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">Termitidae</taxonomicName>
,
<taxonomicName id="B98245B1FF91E340FF0FFAA08021DE08" authorityName="Engel &amp; Krishna" authorityYear="2004" box="[151,314,1339,1366]" class="Insecta" family="Termitidae" kingdom="Animalia" order="Blattodea" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subFamily" subFamily="Syntermitinae">Syntermitinae</taxonomicName>
). These observations, associated with a cryptic life habit, suggest that
<taxonomicName id="B98245B1FF91E340FBF8FAA085EFDE0A" authorityName="Brown" authorityYear="1974" box="[1120,1268,1339,1364]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="conjugans">
<emphasis id="4CF6E220FF91E340FBF8FAA085EFDE0A" box="[1120,1268,1339,1364]" italics="true" pageId="21" pageNumber="22">B. conjugans</emphasis>
</taxonomicName>
might feed on carcasses of other arthropods. It cannot be ruled out that this species might prey on termites, which has occasionally been observed for other
<taxonomicName id="B98245B1FF91E340FE3EFA188308DEC2" authorityName="Schulz" authorityYear="1906" box="[422,531,1411,1436]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="4CF6E220FF91E340FE3EFA188308DEC2" box="[422,531,1411,1436]" italics="true" pageId="21" pageNumber="22">Basiceros</emphasis>
</taxonomicName>
(see natural history accounts for
<taxonomicName id="B98245B1FF91E340FC1EFA18855BDEC2" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[902,1088,1411,1436]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FF91E340FC1EFA18855BDEC2" box="[902,1088,1411,1436]" italics="true" pageId="21" pageNumber="22">Basiceros manni</emphasis>
</taxonomicName>
and
<taxonomicName id="B98245B1FF91E340FBEDFA18841FDEC3" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[1141,1284,1411,1437]" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="4CF6E220FF91E340FBEDFA18841FDEC3" box="[1141,1284,1411,1437]" italics="true" pageId="21" pageNumber="22">B. singularis</emphasis>
</taxonomicName>
).
<taxonomicName id="B98245B1FF91E340FA83FA1881F3DE9E" baseAuthorityName="Emerson" baseAuthorityYear="1925" class="Insecta" family="Termitidae" genus="Uncitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="teevani">
<emphasis id="4CF6E220FF91E340FA83FA1881F3DE9E" italics="true" pageId="21" pageNumber="22">Uncitermes teevani</emphasis>
</taxonomicName>
does not build a nest and lives in the interstices of litter (Dr.
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Moura, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FF91E343FF5FFA51826CD803" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="7E3D3E32FF91E343FF5FFA51826CD803" blockId="21.[151,1437,151,2014]" lastBlockId="22.[151,1437,150,862]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="4CF6E220FF91E340FF5FFA5180AADEBA" bold="true" box="[199,433,1482,1508]" pageId="21" pageNumber="22">Material examined.</emphasis>
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<collectingCountry id="06957EA2FF91E340FE20FA508339DEBB" box="[440,546,1483,1509]" name="Brazil" pageId="21" pageNumber="22">BRAZIL</collectingCountry>
:
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:
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,
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,
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,
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col. (
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) [
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];
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<collectingMunicipality id="9E59A448FF91E340FE21FA7480F4DD57" box="[441,495,1519,1545]" pageId="21" pageNumber="22">Pará</collectingMunicipality>
:
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,
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,
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, C.A.
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.
<materialsCitation id="CEEA346FFF91E340FB7DFA748329DD73" ID-GBIF-Occurrence="4029332398" collectionCode="MZSP" collectorName="Souza" country="Brazil" pageId="21" pageNumber="22" specimenCount="1" specimenCount-worker="1" stateProvince="Acre">
<collectorName id="D3775BE4FF91E340FB7DFA748432DD57" box="[1253,1321,1519,1545]" pageId="21" pageNumber="22">Souza</collectorName>
<emphasis id="4CF6E220FF91E340FAA8FA74847BDD57" box="[1328,1376,1519,1545]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
. col., CARSTE 6040 (
<specimenCount id="6884F5BBFF91E340FEC1F98880DADD70" box="[345,449,1555,1582]" pageId="21" pageNumber="22" type="worker">1 worker</specimenCount>
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;
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<collectingRegion id="BC46F0D0FF91E340FDD8F98883AADD73" box="[576,689,1555,1581]" country="Brazil" name="Rondonia" pageId="21" pageNumber="22">Rondônia</collectingRegion>
:
<collectingMunicipality id="9E59A448FF91E340FD26F9888252DD73" box="[702,841,1555,1581]" pageId="21" pageNumber="22">Porto Velho</collectingMunicipality>
,
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,
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,
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.
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.
<collectorName id="D3775BE4FF91E340FECBF9AC8096DD0F" box="[339,397,1591,1617]" pageId="21" pageNumber="22">Silva</collectorName>
&amp;
<collectorName id="D3775BE4FF91E340FE36F9AC8365DD0F" box="[430,638,1591,1617]" pageId="21" pageNumber="22">Albuquerque, E.Z.</collectorName>
cols. (
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],
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,
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&amp;
<collectorName id="D3775BE4FF91E340FA84F9AC847BDD0F" box="[1308,1376,1591,1617]" pageId="21" pageNumber="22">Prado</collectorName>
, L. P. cols. (
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<collectingMunicipality id="9E59A448FF91E340FE2FF9C08326DD2B" box="[439,573,1627,1653]" pageId="21" pageNumber="22">Porto Velho</collectingMunicipality>
,
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,
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,
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,
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.
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.
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&amp;
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cols. (
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<collectionCode id="1893A6F7FF91E340FE61F9E58351DDC7" box="[505,586,1662,1689]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="22" type="Museum">MZSP</collectionCode>
],
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<collectingDate id="1A78E11AFF91E340FC30F9E48573DDC7" box="[936,1128,1663,1689]" pageId="21" pageNumber="22" value="2011-03-27" valueMax="2011-04-09" valueMin="2011-03-27">27.iii09.iv.2011</collectingDate>
</date>
,
<collectionCode id="1893A6F7FF91E340FBEAF9E48593DDC7" box="[1138,1160,1663,1689]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="22">R</collectionCode>
.
<collectionCode id="1893A6F7FF91E340FB13F9E485BADDC7" box="[1163,1185,1663,1689]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="22">R</collectionCode>
.
<collectorName id="D3775BE4FF91E340FB32F9E485FFDDC7" box="[1194,1252,1663,1689]" pageId="21" pageNumber="22">Silva</collectorName>
&amp;
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cols. (
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) [
<collectionCode id="1893A6F7FF91E340FE0CF93980FDDDE3" box="[404,486,1698,1725]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="22" type="Museum">MZSP</collectionCode>
]
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.
<materialsCitation id="CEEA346FFF91E340FE60F9388039DDBF" ID-GBIF-Occurrence="4029332338" collectionCode="IAVH" collectorName="Orito &amp; Territorio Kofan" country="Colombia" elevation="1000" latitude="0.5" location="Narino" longLatPrecision="1308" longitude="-77.21667" pageId="21" pageNumber="22" specimenCount="4" specimenCount-worker="4" stateProvince="Narino">
<collectingCountry id="06957EA2FF91E340FE60F9388389DDE3" box="[504,658,1699,1725]" name="Colombia" pageId="21" pageNumber="22">COLOMBIA</collectingCountry>
:
<collectingRegion id="BC46F0D0FF91E340FD06F93883F7DDE3" box="[670,748,1699,1725]" country="Colombia" name="Narino" pageId="21" pageNumber="22">Nariño</collectingRegion>
<collectorName id="D3775BE4FF91E340FD6CF9388229DDE3" box="[756,818,1699,1725]" pageId="21" pageNumber="22">Orito</collectorName>
,
<collectorName id="D3775BE4FF91E340FCA6F93882E1DDE3" box="[830,1018,1699,1725]" pageId="21" pageNumber="22">Territorio Kofan</collectorName>
,
<geoCoordinate id="1BB658F5FF91E340FB9FF938857CDDE3" box="[1031,1127,1699,1725]" degrees="00" direction="north" minutes="30" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="0.5">00°30N</geoCoordinate>
<geoCoordinate id="1BB658F5FF91E340FBE8F93885CCDDE3" box="[1136,1239,1699,1725]" degrees="77" direction="west" minutes="13" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="-77.21667">77°13W</geoCoordinate>
,
<quantity id="B97A93D7FF91E340FB7BF9388429DDE3" box="[1251,1330,1699,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="m" value="1000.0">
<elevation id="F5AFD901FF91E340FB7BF9388429DDE3" box="[1251,1330,1699,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="m" value="1000.0">1000m</elevation>
</quantity>
(
<specimenCount id="6884F5BBFF91E340FADAF93881A2DDBF" pageId="21" pageNumber="22" type="worker">4 workers</specimenCount>
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<collectionCode id="1893A6F7FF91E340FF48F95D8000DDBF" box="[208,283,1734,1761]" country="Colombia" httpUri="http://grbio.org/cool/6phm-mi1y" name="Instituto de Ivestigacion de los Recursos Biologicos Alexander von Humboldt" pageId="21" pageNumber="22">IAVH</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="CEEA346FFF91E340FEB5F95C852CDDBF" ID-GBIF-Occurrence="4029332368" box="[301,1079,1734,1762]" collectingDate="1965-05" collectionCode="MCZ" collectorName="La Pena" country="Ecuador" location="Coca" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Narino">
<collectingCountry id="06957EA2FF91E340FEB5F95C80A2DDBF" box="[301,441,1735,1761]" name="Ecuador" pageId="21" pageNumber="22">ECUADOR</collectingCountry>
:
<location id="7B5D68E9FF91E340FE5CF95C8319DDBF" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF91E340FE5CF95C8319DDBF" box="[452,514,1735,1761]" country="Ecuador" name="Coca" pageId="21" pageNumber="22" stateProvince="Narino">Coca</location>
:
<date id="0A3C18F2FF91E340FD95F95D8340DDBF" box="[525,603,1734,1761]" pageId="21" pageNumber="22" value="1965-05">
<collectingDate id="1A78E11AFF91E340FD95F95D8340DDBF" box="[525,603,1734,1761]" pageId="21" pageNumber="22" value="1965-05">v.1965</collectingDate>
</date>
,
<collectorName id="D3775BE4FF91E340FDF0F95C83D8DDBF" box="[616,707,1735,1761]" pageId="21" pageNumber="22">La Peña</collectorName>
col., n. 546612 (
<specimenCount id="6884F5BBFF91E340FC10F95C82CFDDBC" box="[904,980,1735,1762]" pageId="21" pageNumber="22" type="male">1 male</specimenCount>
) [
<collectionCode id="1893A6F7FF91E340FC73F95D852BDDBF" box="[1003,1072,1734,1761]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="21" pageNumber="22" type="Museum">MCZ</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="CEEA346FFF91E340FBDBF95C82A3DC58" ID-GBIF-Occurrence="4029332378" collectionCode="LACM" collectorName="Stewart &amp; Peck" country="Ecuador" elevation="600" location="20 km S de Tena" pageId="21" pageNumber="22" specimenCode="LACM 326569" specimenCount="1" specimenCount-worker="1" stateProvince="Napo">
<collectingRegion id="BC46F0D0FF91E340FBDBF95C859FDDBF" box="[1091,1156,1735,1761]" country="Ecuador" name="Napo" pageId="21" pageNumber="22">Napo</collectingRegion>
:
<location id="7B5D68E9FF91E340FB17F95C8452DDBF" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF91E340FB17F95C8452DDBF" box="[1167,1353,1735,1761]" country="Ecuador" name="20 km S de Tena" pageId="21" pageNumber="22" stateProvince="Napo">
<quantity id="B97A93D7FF91E340FB17F95C85D4DDBF" box="[1167,1231,1735,1761]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="22" unit="km" value="20.0">20km</quantity>
S de Tena
</location>
,
<quantity id="B97A93D7FF91E340FACDF95C8483DDBF" box="[1365,1432,1735,1761]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="22" unit="m" value="600.0">
<elevation id="F5AFD901FF91E340FACDF95C8483DDBF" box="[1365,1432,1735,1761]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="22" unit="m" value="600.0">600m</elevation>
</quantity>
, 11.vii,
<collectorName id="D3775BE4FF91E340FF7CF9708021DC5B" box="[228,314,1771,1797]" pageId="21" pageNumber="22">Stewart</collectorName>
B.
<collectorName id="D3775BE4FF91E340FEF8F970808DDC5B" box="[352,406,1771,1797]" pageId="21" pageNumber="22">Peck</collectorName>
&amp; Jarmila Peck cols.,
<specimenCode id="2E249649FF91E340FD08F9708227DC5B" box="[656,828,1770,1797]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="21" pageNumber="22" type="Museum">LACM 326569</specimenCode>
(
<specimenCount id="6884F5BBFF91E340FCD5F97082AADC58" box="[845,945,1771,1798]" pageId="21" pageNumber="22" type="worker">1 worker</specimenCount>
)
</materialsCitation>
<materialsCitation id="CEEA346FFF91E340FC27F971828ADC77" ID-GBIF-Occurrence="4029332349" accessionNumber="PSW11365" collectingDate="1976-06-18" collectingDateMax="1991-08-05" collectingDateMin="1976-06-18" collectionCode="LACM, MCZ, UCDC, GUPC, MZSP, MNH, RSPPC" collectorName="Stewart B. Peck &amp; Jarmila Peck" country="Ecuador" county="Limoncocha" elevation="250" location="Long." municipality="Jatun Sacha" pageId="21" pageNumber="22" specimenCount="7" specimenCount-worker="7" stateProvince="Napo">
[
<collectionCode id="1893A6F7FF91E340FC5DF9718504DC5B" box="[965,1055,1770,1797]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="21" pageNumber="22" type="Museum">LACM</collectionCode>
];
<collectingCounty id="975C46BEFF91E340FBA9F97085DBDC5B" box="[1073,1216,1771,1797]" pageId="21" pageNumber="22">Limoncocha</collectingCounty>
,
<quantity id="B97A93D7FF91E340FB52F9718409DC5B" box="[1226,1298,1770,1797]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="22" unit="m" value="250.0">
<elevation id="F5AFD901FF91E340FB52F9718409DC5B" box="[1226,1298,1770,1797]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="22" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<date id="0A3C18F2FF91E340FA86F970848CDC5B" box="[1310,1431,1771,1797]" pageId="21" pageNumber="22" value="1976-06-18">
<collectingDate id="1A78E11AFF91E340FA86F970848CDC5B" box="[1310,1431,1771,1797]" pageId="21" pageNumber="22" value="1976-06-18">18.vi.1976</collectingDate>
</date>
,
<collectorName id="D3775BE4FF91E340FF0FF8948051DC77" box="[151,330,1807,1833]" pageId="21" pageNumber="22">Stewart B. Peck</collectorName>
&amp;
<collectorName id="D3775BE4FF91E340FEF5F89480E6DC77" box="[365,509,1807,1833]" pageId="21" pageNumber="22">Jarmila Peck</collectorName>
cols., n. 546454 (
<specimenCount id="6884F5BBFF91E340FD5FF8948230DC74" box="[711,811,1807,1834]" pageId="21" pageNumber="22" type="worker">1 worker</specimenCount>
) [
<collectionCode id="1893A6F7FF91E340FCD8F895829EDC77" box="[832,901,1806,1833]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="21" pageNumber="22" type="Museum">MCZ</collectionCode>
];
</materialsCitation>
<materialsCitation id="CEEA346FFF91E340FC0FF89485E6DC13" ID-GBIF-Occurrence="4029332326" accessionNumber="PSW11365" collectingDate="1976-06-18" collectingDateMax="1991-08-05" collectingDateMin="1976-06-18" collectionCode="LACM, MCZ, UCDC, GUPC, MZSP, MNH, RSPPC" collectorName="P. S. Ward" country="Ecuador" county="Limoncocha" elevation="250" location="Long." municipality="Jatun Sacha" pageId="21" pageNumber="22" specimenCount="7" specimenCount-worker="7" stateProvince="Napo">
<collectingMunicipality id="9E59A448FF91E340FC0FF8948504DC77" box="[919,1055,1807,1833]" pageId="21" pageNumber="22">Jatun Sacha</collectingMunicipality>
,
<locationDeviation id="6F8F2710FF91E340FBB2F894847EDC77" box="[1066,1381,1807,1833]" location="Puerto Misahualli" pageId="21" pageNumber="22">
<quantity id="B97A93D7FF91E340FBB2F8948540DC77" box="[1066,1115,1807,1833]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="21" pageNumber="22" unit="km" value="7.0">7km</quantity>
ESE
<locationDeviation id="6F8F2710FF91E340FB03F894847EDC77" box="[1179,1381,1807,1833]" pageId="21" pageNumber="22">Puerto Misahuallí</locationDeviation>
</locationDeviation>
,
<locationDeviation id="6F8F2710FF91E340FAE8F8948486DC77" box="[1392,1437,1807,1833]" pageId="21" pageNumber="22">Lat.</locationDeviation>
-1.0667
<location id="7B5D68E9FF91E340FF60F8A88021DC13" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF91E340FF60F8A88021DC13" box="[248,314,1843,1869]" country="Ecuador" county="Limoncocha" municipality="Jatun Sacha" name="Long." pageId="21" pageNumber="22" stateProvince="Napo">Long.</location>
-77.6167,
<quantity id="B97A93D7FF91E340FE22F8A880E6DC13" box="[442,509,1843,1869]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="22" unit="m" value="400.0">
<elevation id="F5AFD901FF91E340FE22F8A880E6DC13" box="[442,509,1843,1869]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="22" unit="m" value="400.0">400m</elevation>
</quantity>
,
<date id="0A3C18F2FF91E340FD93F8A9838EDC13" box="[523,661,1842,1869]" pageId="21" pageNumber="22" value="1991-08-05">
<collectingDate id="1A78E11AFF91E340FD93F8A9838EDC13" box="[523,661,1842,1869]" pageId="21" pageNumber="22" value="1991-08-05">05.viii.1991</collectingDate>
</date>
,
<collectorName id="D3775BE4FF91E340FD3BF8A88205DC13" box="[675,798,1843,1869]" pageId="21" pageNumber="22">P. S. Ward</collectorName>
col., PSW11365-31 (
<specimenCount id="6884F5BBFF91E340FB83F8A88598DC10" box="[1051,1155,1843,1870]" pageId="21" pageNumber="22" type="worker">1 worker</specimenCount>
) [
<collectionCode id="1893A6F7FF91E340FB04F8A985E9DC13" box="[1180,1266,1842,1869]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="21" pageNumber="22" type="Museum">UCDC</collectionCode>
];
</materialsCitation>
<materialsCitation id="CEEA346FFF91E340FA9FF8A883ACDCCB" ID-GBIF-Occurrence="4029332416" accessionNumber="PSW11365" collectingDate="1976-06-18" collectingDateMax="1991-08-05" collectingDateMin="1976-06-18" collectionCode="LACM, MCZ, UCDC, GUPC, MZSP, MNH, RSPPC" collectorName="G. J Umphrey" country="Ecuador" county="Limoncocha" elevation="250" location="Long." municipality="Jatun Sacha" pageId="21" pageNumber="22" specimenCount="7" specimenCount-worker="7" stateProvince="Napo">
<location id="7B5D68E9FF91E340FA9FF8A881C1DC2F" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF91E340FA9FF8A881C1DC2F" country="Ecuador" county="Limoncocha" municipality="Jatun Sacha" name="Puerto Misahualli" pageId="21" pageNumber="22" stateProvince="Napo">Puerto Misahuallí</location>
,
<date id="0A3C18F2FF91E340FF7CF8CD807DDC2F" box="[228,358,1878,1905]" pageId="21" pageNumber="22" value="1978-07-15">
<collectingDate id="1A78E11AFF91E340FF7CF8CD807DDC2F" box="[228,358,1878,1905]" pageId="21" pageNumber="22" value="1978-07-15">15.vii.1978</collectingDate>
</date>
,
<collectorName id="D3775BE4FF91E340FEE9F8CC831CDC2F" box="[369,519,1879,1905]" pageId="21" pageNumber="22">G.J Umphrey</collectorName>
col., GJU 0774, (
<specimenCount id="6884F5BBFF91E340FD53F8CC8227DC2C" box="[715,828,1879,1906]" pageId="21" pageNumber="22" type="worker">2 workers</specimenCount>
) [
<collectionCode id="1893A6F7FF91E340FCC9F8CD82BFDC2F" box="[849,932,1878,1905]" pageId="21" pageNumber="22">GUPC</collectionCode>
]; same data (
<specimenCount id="6884F5BBFF91E340FBADF8CC8582DC2C" box="[1077,1177,1879,1906]" pageId="21" pageNumber="22" type="worker">1 worker</specimenCount>
) [
<collectionCode id="1893A6F7FF91E340FB35F8CD85E4DC2F" box="[1197,1279,1878,1905]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="22" type="Museum">MZSP</collectionCode>
]; same data (
<specimenCount id="6884F5BBFF91E340FA08F8CC81F3DCC8" pageId="21" pageNumber="22" type="worker">1 worker</specimenCount>
) [
<collectionCode id="1893A6F7FF91E340FF65F8E1805CDCCB" box="[253,327,1914,1941]" country="Hungary" httpUri="http://grbio.org/cool/34nf-ps25" name="Musei Nacionalis Hungarici" pageId="21" pageNumber="22">MNH</collectionCode>
]; same data (
<specimenCount id="6884F5BBFF91E340FE42F8E08325DCC8" box="[474,574,1915,1942]" pageId="21" pageNumber="22" type="worker">1 worker</specimenCount>
) [
<collectionCode id="1893A6F7FF91E340FDCBF8E183ABDCCB" box="[595,688,1914,1941]" pageId="21" pageNumber="22">RSPPC</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="CEEA346FFF91E340FD59F8E085DFDC83" ID-GBIF-Occurrence="4029332364" collectingDate="1975-05-18" collectingDateMax="1976-07-08" collectingDateMin="1975-05-18" collectionCode="MCZ" collectorName="Stewart B. Peck &amp; Jarmila Peck" country="Ecuador" county="Santo Domingo" elevation="700" location="Rio Palenque Station" municipality="Colorados" pageId="21" pageNumber="22" specimenCount="2" specimenCount-worker="2" stateProvince="Pichincha">
<collectingRegion id="BC46F0D0FF91E340FD59F8E0822FDCCB" box="[705,820,1915,1941]" country="Ecuador" name="Pichincha" pageId="21" pageNumber="22">Pichincha</collectingRegion>
:
<quantity id="B97A93D7FF91E340FCA6F8E0826CDCCB" box="[830,887,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="22" unit="km" value="4.0">4 km</quantity>
E
<collectingCounty id="975C46BEFF91E340FC0DF8E0855DDCCB" box="[917,1094,1915,1941]" pageId="21" pageNumber="22">Santo Domingo</collectingCounty>
de los
<collectingMunicipality id="9E59A448FF91E340FB0DF8E08410DCCB" box="[1173,1291,1915,1941]" pageId="21" pageNumber="22">Colorados</collectingMunicipality>
,
<date id="0A3C18F2FF91E340FA8EF8E0848CDCCB" box="[1302,1431,1915,1941]" pageId="21" pageNumber="22" value="1976-07-08">
<collectingDate id="1A78E11AFF91E340FA8EF8E0848CDCCB" box="[1302,1431,1915,1941]" pageId="21" pageNumber="22" value="1976-07-08">08.vii.1976</collectingDate>
</date>
,
<collectorName id="D3775BE4FF91E340FF0FF8048053DCE7" box="[151,328,1951,1977]" pageId="21" pageNumber="22">Stewart B. Peck</collectorName>
&amp;
<collectorName id="D3775BE4FF91E340FEF2F80480E2DCE7" box="[362,505,1951,1977]" pageId="21" pageNumber="22">Jarmila Peck</collectorName>
cols., n. 546456 (
<specimenCount id="6884F5BBFF91E340FD58F8048238DCE4" box="[704,803,1951,1978]" pageId="21" pageNumber="22" type="worker">1 worker</specimenCount>
) [
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,
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,
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<collectingDate id="1A78E11AFF91E340FF7FF859809ADC83" box="[231,385,1986,2013]" pageId="21" pageNumber="22" value="1975-05-18" valueMax="1975-05-30" valueMin="1975-05-18">1830.v.1975</collectingDate>
</date>
,
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&amp;
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cols., n. 546462 (
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<collectionCode id="1893A6F7FF91E340FBCAF859858CDC83" box="[1106,1175,1986,2013]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="21" pageNumber="22" type="Museum">MCZ</collectionCode>
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.
<materialsCitation id="CEEA346FFF91E343FB49F8588316DB8B" ID-GBIF-Occurrence="4029332404" collectingDate="1976-08-03" collectingDateMax="2009-09-30" collectingDateMin="1976-08-03" collectionCode="MZSP, T" collectorName="Tjitte" country="Ecuador" county="Los Tayos" elevation="1030" lastPageId="22" lastPageNumber="23" location="Chichipe" municipality="Zamora" pageId="21" pageNumber="22" specimenCount="1" specimenCount-worker="1" stateProvince="Morona-Santiago">
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:
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,
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<collectingDate id="1A78E11AFF92E343FE8FFF0C8089DBEF" box="[279,402,151,177]" pageId="22" pageNumber="23" value="1976-08-03">3.viii.1976</collectingDate>
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,
<collectorName id="D3775BE4FF92E343FE05FF0C80C1DBEF" box="[413,474,151,177]" pageId="22" pageNumber="23">Tjitte</collectorName>
de Vries col. (
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) [
<collectionCode id="1893A6F7FF92E343FD6CFF0D825DDBEF" box="[756,838,150,177]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="22" pageNumber="23" type="Museum">MZSP</collectionCode>
];
<collectingMunicipality id="9E59A448FF92E343FCCFFF0C82A8DBEF" box="[855,947,151,177]" pageId="22" pageNumber="23">Zamora</collectingMunicipality>
:
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,
<location id="7B5D68E9FF92E343FBB7FF0C85BEDBEF" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF92E343FBB7FF0C85BEDBEF" box="[1071,1189,151,177]" country="Ecuador" county="Los Tayos" municipality="Zamora" name="Copalinga" pageId="22" pageNumber="23" stateProvince="Morona-Santiago">Copalinga</location>
,
<location id="7B5D68E9FF92E343FB37FF0C85C7DBEF" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF92E343FB37FF0C85C7DBEF" box="[1199,1244,151,177]" country="Ecuador" county="Los Tayos" municipality="Zamora" name="Lat." pageId="22" pageNumber="23" stateProvince="Morona-Santiago">Lat.</location>
-4.091222
<location id="7B5D68E9FF92E343FAC2FF0C8486DBEF" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF92E343FAC2FF0C8486DBEF" box="[1370,1437,151,177]" country="Ecuador" county="Los Tayos" municipality="Zamora" name="Long." pageId="22" pageNumber="23" stateProvince="Morona-Santiago">Long.</location>
-78.96069,
<quantity id="B97A93D7FF92E343FE80FF208073DB8B" box="[280,360,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="22" pageNumber="23" unit="m" value="1030.0">
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,
<date id="0A3C18F2FF92E343FEEDFF2080F4DB8B" box="[373,495,187,213]" pageId="22" pageNumber="23" value="2009-09-30">
<collectingDate id="1A78E11AFF92E343FEEDFF2080F4DB8B" box="[373,495,187,213]" pageId="22" pageNumber="23" value="2009-09-30">30.ix.2009</collectingDate>
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,
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</materialsCitation>
.
<materialsCitation id="CEEA346FFF92E343FD80FF208251DB8B" ID-GBIF-Occurrence="4029332315" box="[536,842,187,213]" collectionCode="T" collectorName="Delsinne &amp; Arias-Penna" country="Ecuador" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Morona Santiago">
<collectorName id="D3775BE4FF92E343FD80FF208367DB8B" box="[536,636,187,213]" pageId="22" pageNumber="23">Delsinne</collectorName>
&amp;
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,
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</materialsCitation>
.
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cols. (
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) [
<collectionCode id="1893A6F7FF92E343FB80FF218571DB8B" box="[1048,1130,186,213]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="22" pageNumber="23" type="Museum">MZSP</collectionCode>
];
<location id="7B5D68E9FF92E343FBE4FF20842BDB8B" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF92E343FBE4FF20842BDB8B" box="[1148,1328,187,213]" country="Ecuador" latitude="-3.74572" longLatPrecision="1" longitude="-78.61436" name="Jardin Botanico" pageId="22" pageNumber="23" stateProvince="Morona Santiago">Jardin Botánico</location>
,
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,
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NEE Yantzaza
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,
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,
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,
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<collectingDate id="1A78E11AFF92E343FC1DFF4582E4DBA7" box="[901,1023,222,249]" pageId="22" pageNumber="23" value="2014-06-15">15.vi.2014</collectingDate>
</date>
,
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</materialsCitation>
.
<materialsCitation id="CEEA346FFF92E343FBB1FF448447DBA7" ID-GBIF-Occurrence="4029332332" box="[1065,1372,223,249]" collectionCode="T" collectorName="Delsinne &amp; Arias-Penna" country="Ecuador" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Morona Santiago">
<collectorName id="D3775BE4FF92E343FBB1FF448596DBA7" box="[1065,1165,223,249]" pageId="22" pageNumber="23">Delsinne</collectorName>
&amp;
<collectorName id="D3775BE4FF92E343FB29FF448425DBA7" box="[1201,1342,223,249]" pageId="22" pageNumber="23">Arias-Penna</collectorName>
,
<collectionCode id="1893A6F7FF92E343FAD2FF448447DBA7" box="[1354,1372,223,249]" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="CEEA346FFF92E343FAF0FF448065DA43" ID-GBIF-Occurrence="4029332400" collectionCode="MZSP" country="Ecuador" pageId="22" pageNumber="23" specimenCount="1" specimenCount-worker="1" stateProvince="Morona Santiago">
cols. (
<specimenCount id="6884F5BBFF92E343FF3AFE988010DA40" box="[162,267,259,286]" pageId="22" pageNumber="23" type="worker">1 worker</specimenCount>
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<collectionCode id="1893A6F7FF92E343FEBEFE998063DA43" box="[294,376,258,285]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="22" pageNumber="23" type="Museum">MZSP</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="CEEA346FFF92E343FE16FE988580DA1C" ID-GBIF-Occurrence="4029332403" collectingDate="2006-10-16" collectionCode="LS" collectorName="C. J. Marshall" country="Guyana" elevation="735" location="New Romeo Camp" municipality="Acarai Mountains" pageId="22" pageNumber="23" specimenCode="LS04" specimenCount="2" specimenCount-worker="2" stateProvince="Upper Takutu-Upper Essequibo">
<collectingCountry id="06957EA2FF92E343FE16FE988312DA43" box="[398,521,259,285]" name="Guyana" pageId="22" pageNumber="23">GUYANA</collectingCountry>
:
<collectingRegion id="BC46F0D0FF92E343FD80FE988291DA43" box="[536,906,259,285]" country="Guyana" name="Upper Takutu-Upper Essequibo" pageId="22" pageNumber="23">Upper Takutu-Upper Essequibo</collectingRegion>
:
<collectingMunicipality id="9E59A448FF92E343FC00FE988572DA43" box="[920,1129,259,285]" pageId="22" pageNumber="23">Acaraí Mountains</collectingMunicipality>
, near
<location id="7B5D68E9FF92E343FB2DFE988483DA43" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF92E343FB2DFE988483DA43" box="[1205,1432,259,285]" country="Guyana" municipality="Acarai Mountains" name="New Romeo Camp" pageId="22" pageNumber="23" stateProvince="Upper Takutu-Upper Essequibo">New Romeo Camp</location>
, 1°20842”N 58°57496”W,
<quantity id="B97A93D7FF92E343FE56FEBD830ADA1F" box="[462,529,294,321]" metricMagnitude="2" metricUnit="m" metricValue="7.35" pageId="22" pageNumber="23" unit="m" value="735.0">
<elevation id="F5AFD901FF92E343FE56FEBD830ADA1F" box="[462,529,294,321]" metricMagnitude="2" metricUnit="m" metricValue="7.35" pageId="22" pageNumber="23" unit="m" value="735.0">735m</elevation>
</quantity>
,
<date id="0A3C18F2FF92E343FD83FEBC8395DA1F" box="[539,654,295,321]" pageId="22" pageNumber="23" value="2006-10-16">
<collectingDate id="1A78E11AFF92E343FD83FEBC8395DA1F" box="[539,654,295,321]" pageId="22" pageNumber="23" value="2006-10-16">16.x.2006</collectingDate>
</date>
,
<collectorName id="D3775BE4FF92E343FD01FEBC822DDA1F" box="[665,822,295,321]" pageId="22" pageNumber="23">C. J. Marshall</collectorName>
, TRS 061016-
<specimenCode id="2E249649FF92E343FC40FEBC850FDA1F" box="[984,1044,295,321]" collectionCode="LS" country="United Kingdom" name="Linnean Society of London" pageId="22" pageNumber="23">LS04</specimenCode>
(
<specimenCount id="6884F5BBFF92E343FBBBFEBC858FDA1C" box="[1059,1172,295,322]" pageId="22" pageNumber="23" type="worker">2 workers</specimenCount>
)
</materialsCitation>
<materialsCitation id="CEEA346FFF92E343FB3AFEBD82AFDA38" ID-GBIF-Occurrence="4029332375" collectionCode="USNM" collectorName="M, J. Sosa-Calvo" country="Guyana" pageId="22" pageNumber="23" specimenCode="LS06" specimenCount="1" specimenCount-worker="1" stateProvince="Upper Takutu-Upper Essequibo">
[
<collectionCode id="1893A6F7FF92E343FB30FEBD8419DA1F" box="[1192,1282,294,321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="23" type="Museum">USNM</collectionCode>
]; 1°20896”N 58°57491”W, 753
<collectorName id="D3775BE4FF92E343FEF3FED18333DA3B" box="[363,552,330,357]" pageId="22" pageNumber="23">M, J. Sosa-Calvo</collectorName>
col., TRS 061016-
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(
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)
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[
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]; 1°2306”N 58°56789”W,
<quantity id="B97A93D7FF92E343FACDFED08483DA3B" box="[1365,1432,331,357]" metricMagnitude="2" metricUnit="m" metricValue="2.94" pageId="22" pageNumber="23" unit="m" value="294.0">
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</quantity>
,
<collectionCode id="1893A6F7FF92E343FF0FFEF481B2DAD7" box="[151,169,367,393]" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T</collectionCode>
.
<collectionCode id="1893A6F7FF92E343FF2DFEF481D0DAD7" box="[181,203,367,393]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="23">R</collectionCode>
.
<collectorName id="D3775BE4FF92E343FF4FFEF48030DAD7" box="[215,299,367,393]" pageId="22" pageNumber="23">Schultz</collectorName>
&amp; J.
<collectorName id="D3775BE4FF92E343FEF4FEF480F6DAD7" box="[364,493,367,393]" pageId="22" pageNumber="23">Sosa-Calvo</collectorName>
cols., JSC061010-
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(
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)
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[
<collectionCode id="1893A6F7FF92E343FC17FEF582F2DAD7" box="[911,1001,366,393]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="23" type="Museum">USNM</collectionCode>
]; 1°23191”N 58°56808”W,
<quantity id="B97A93D7FF92E343FAA0FEF48461DAD7" box="[1336,1402,367,393]" metricMagnitude="2" metricUnit="m" metricValue="3.03" pageId="22" pageNumber="23" unit="m" value="303.0">
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</quantity>
,
<collectionCode id="1893A6F7FF92E343FA1EFEF48483DAD7" box="[1414,1432,367,393]" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T</collectionCode>
.
<collectionCode id="1893A6F7FF92E343FF0FFE0881B6DAF3" box="[151,173,403,429]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="23">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="CEEA346FFF92E343FF21FE088244DAF0" ID-GBIF-Occurrence="4029332366" box="[185,863,403,430]" collectionCode="LS" collectorName="Schultz &amp; Sosa-Calvo" country="Guyana" pageId="22" pageNumber="23" specimenCode="LS03" specimenCount="1" specimenCount-worker="1" stateProvince="Upper Takutu-Upper Essequibo">
<collectorName id="D3775BE4FF92E343FF21FE088016DAF3" box="[185,269,403,429]" pageId="22" pageNumber="23">Schultz</collectorName>
&amp; J.
<collectorName id="D3775BE4FF92E343FED5FE0880D5DAF3" box="[333,462,403,429]" pageId="22" pageNumber="23">Sosa-Calvo</collectorName>
cols., JSC061010-
<specimenCode id="2E249649FF92E343FD3DFE0883FBDAF3" box="[677,736,403,429]" collectionCode="LS" country="United Kingdom" name="Linnean Society of London" pageId="22" pageNumber="23">LS03</specimenCode>
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].
<collectingCounty id="975C46BEFF92E343FCF2FD2885B1D993" box="[874,1194,691,717]" pageId="22" pageNumber="23">Tambopata Research Center</collectingCounty>
:
<collectingMunicipality id="9E59A448FF92E343FB2CFD2885FAD993" box="[1204,1249,691,717]" pageId="22" pageNumber="23">Lat.</collectingMunicipality>
-13.14048
<location id="7B5D68E9FF92E343FAF9FD288487D993" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF92E343FAF9FD288487D993" box="[1377,1436,691,717]" country="Peru" county="Tambopata Research Center" municipality="Lat." name="Long" pageId="22" pageNumber="23" stateProvince="Madre de Dios">Long</location>
-69.62108,
<quantity id="B97A93D7FF92E343FE83FD4C8045D9AF" box="[283,350,727,753]" metricMagnitude="2" metricUnit="m" metricValue="3.18" pageId="22" pageNumber="23" unit="m" value="318.0">
<elevation id="F5AFD901FF92E343FE83FD4C8045D9AF" box="[283,350,727,753]" metricMagnitude="2" metricUnit="m" metricValue="3.18" pageId="22" pageNumber="23" unit="m" value="318.0">318m</elevation>
</quantity>
,
<date id="0A3C18F2FF92E343FEF5FD4C80CFD9AF" box="[365,468,727,753]" pageId="22" pageNumber="23" value="2001-08">
<collectingDate id="1A78E11AFF92E343FEF5FD4C80CFD9AF" box="[365,468,727,753]" pageId="22" pageNumber="23" value="2001-08">viii.2001</collectingDate>
</date>
,
<collectorName id="D3775BE4FF92E343FE7BFD4C834DD9AF" box="[483,598,727,753]" pageId="22" pageNumber="23">D. Feener</collectorName>
col., TRC-S05-
<specimenCode id="2E249649FF92E343FC97FD4D8232D9AF" box="[783,809,726,753]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="23">R1</specimenCode>
</materialsCitation>
<materialsCitation id="CEEA346FFF92E343FCB2FD4D85B8D9AF" ID-GBIF-Occurrence="4029332432" box="[810,1187,726,753]" collectionCode="CASENT" country="Peru" pageId="22" pageNumber="23" specimenCode="CASENT0637530" specimenCount="1" stateProvince="Madre de Dios">
C09/
<specimenCode id="2E249649FF92E343FCF8FD4D852DD9AF" box="[864,1078,726,753]" collectionCode="CASENT" pageId="22" pageNumber="23">CASENT0637530</specimenCode>
(1 gyne)
</materialsCitation>
<materialsCitation id="CEEA346FFF92E343FB36FD4C81EFD867" ID-GBIF-Occurrence="4029332363" collectingDate="2001-09" collectionCode="JTL" collectorName="D. Feener" country="Peru" elevation="289" location="Long." municipality="Lat." pageId="22" pageNumber="23" specimenCode="R1, CASENT0637518" specimenCount="1" specimenCount-worker="1" stateProvince="Madre de Dios">
[
<collectionCode id="1893A6F7FF92E343FB2DFD4C85F2D9AF" box="[1205,1257,727,753]" pageId="22" pageNumber="23">JTL</collectionCode>
);
<collectingMunicipality id="9E59A448FF92E343FA99FD4C8436D9AF" box="[1281,1325,727,753]" pageId="22" pageNumber="23">Lat.</collectingMunicipality>
-13.1381
<location id="7B5D68E9FF92E343FF0FFD6081C1D84B" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF92E343FF0FFD6081C1D84B" box="[151,218,763,789]" country="Peru" municipality="Lat." name="Long." pageId="22" pageNumber="23" stateProvince="Madre de Dios">Long.</location>
-69.62767,
<quantity id="B97A93D7FF92E343FEFEFD6080B2D84B" box="[358,425,763,789]" metricMagnitude="2" metricUnit="m" metricValue="2.8899999999999997" pageId="22" pageNumber="23" unit="m" value="289.0">
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</quantity>
,
<date id="0A3C18F2FF92E343FE2FFD608315D84B" box="[439,526,763,789]" pageId="22" pageNumber="23" value="2001-09">
<collectingDate id="1A78E11AFF92E343FE2FFD608315D84B" box="[439,526,763,789]" pageId="22" pageNumber="23" value="2001-09">ix.2001</collectingDate>
</date>
,
<collectorName id="D3775BE4FF92E343FD84FD608396D84B" box="[540,653,763,789]" pageId="22" pageNumber="23">D. Feener</collectorName>
col., TRC-S10-
<specimenCode id="2E249649FF92E343FCDBFD618245D84B" box="[835,862,762,789]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="23">R1</specimenCode>
C01/
<specimenCode id="2E249649FF92E343FC0CFD618570D84B" box="[916,1131,762,789]" collectionCode="CASENT" pageId="22" pageNumber="23">CASENT0637518</specimenCode>
(
<specimenCount id="6884F5BBFF92E343FB18FD6085FCD848" box="[1152,1255,763,790]" pageId="22" pageNumber="23" type="worker">1 worker</specimenCount>
) [
<collectionCode id="1893A6F7FF92E343FB67FD608428D84B" box="[1279,1331,763,789]" pageId="22" pageNumber="23">JTL</collectionCode>
). TRINIDAD &amp;
</materialsCitation>
<materialsCitation id="CEEA346FFF92E343FF66FC84826CD803" ID-GBIF-Occurrence="4029332314" collectionCode="R, LACM" collectorName="Snelling" country="Trinidad and Tobago" latitude="10.95" location="Arena Forest Reserve" longLatPrecision="1296" longitude="-61.433334" pageId="22" pageNumber="23" specimenCode="LACM 326570, LACM 326571" specimenCount="2" specimenCount-worker="2" stateProvince="Madre de Dios">
<collectingCountry id="06957EA2FF92E343FF66FC84806CD867" box="[254,375,799,825]" name="Trinidad and Tobago" pageId="22" pageNumber="23">TOBAGO</collectingCountry>
:
<location id="7B5D68E9FF92E343FE1BFC84839AD867" LSID="urn:lsid:plazi:treatment:F62B8F24FF8BE343FF0FF936826CD803:7B5D68E9FF92E343FE1BFC84839AD867" box="[387,641,799,825]" country="Trinidad and Tobago" latitude="10.95" longLatPrecision="1296" longitude="-61.433334" name="Arena Forest Reserve" pageId="22" pageNumber="23" stateProvince="Madre de Dios">Arena Forest Reserve</location>
:
<locationDeviation id="6F8F2710FF92E343FD16FC8582D6D867" box="[654,973,798,825]" location="San Rafael" pageId="22" pageNumber="23">
<quantity id="B97A93D7FF92E343FD16FC8583F8D867" box="[654,739,798,825]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="22" pageNumber="23" unit="km" value="1.05">1.05km</quantity>
SSE de San Rafael
</locationDeviation>
,
<geoCoordinate id="1BB658F5FF92E343FC43FC858527D867" box="[987,1084,798,825]" degrees="10" direction="north" minutes="57" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="10.95">10°57N</geoCoordinate>
<geoCoordinate id="1BB658F5FF92E343FBDEFC8485B6D867" box="[1094,1197,799,825]" degrees="61" direction="west" minutes="26" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="-61.433334">61°26W</geoCoordinate>
,
<collectionCode id="1893A6F7FF92E343FB24FC8485C9D867" box="[1212,1234,799,825]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="23">R</collectionCode>
.
<collectionCode id="1893A6F7FF92E343FB78FC8485EDD867" box="[1248,1270,799,825]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="23">R</collectionCode>
.
<collectorName id="D3775BE4FF92E343FA9CFC84847AD867" box="[1284,1377,799,825]" pageId="22" pageNumber="23">Snelling</collectorName>
col.,
<specimenCode id="2E249649FF92E343FF0FFCD88058D803" box="[151,323,834,861]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="22" pageNumber="23" type="Museum">LACM 326570</specimenCode>
(
<specimenCount id="6884F5BBFF92E343FECDFCD880A2D800" box="[341,441,835,862]" pageId="22" pageNumber="23" type="worker">1 worker</specimenCount>
) e
<specimenCode id="2E249649FF92E343FE43FCD8839CD803" box="[475,647,834,861]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="22" pageNumber="23" type="Museum">LACM 326571</specimenCode>
(
<specimenCount id="6884F5BBFF92E343FD01FCD883E5D800" box="[665,766,835,862]" pageId="22" pageNumber="23" type="worker">1 worker</specimenCount>
) [
<collectionCode id="1893A6F7FF92E343FC8BFCD98276D803" box="[787,877,834,861]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="22" pageNumber="23" type="Museum">LACM</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>