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<document id="5E82D04000CC778B115FE1961A11620E" ID-CLB-Dataset="3997" ID-DOI="10.11646/zootaxa.5052.3.6" ID-GBIF-Dataset="9e958680-584d-4057-bc60-8ae20cb9f210" ID-ISSN="1175-5326" ID-Zenodo-Dep="5572202" ID-ZooBank="9682EB53-E37B-4FB1-8208-17AF1E8B2571" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634301788653" checkinUser="plazi" docAuthor="Díaz, Luis M., Fong, Ansel, Salas, Diego &amp; Hedges, S. Blair" docDate="2021" docId="DD4987C1FF96247CFF3DFC385F30FE6F" docLanguage="en" docName="zootaxa.5052.3.6.pdf" docOrigin="Zootaxa 5052 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arrhyton albicollum Díaz &amp; Fong &amp; Salas &amp; Hedges 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="414" masterDocId="2170FFB9FF942474FFAAFFEA5C7DFFC2" masterDocTitle="A new semifossorial snake of the genus Arrhyton (Squamata: Dipsadidae) from eastern Cuba, with taxonomic comments on other species" masterLastPageNumber="418" masterPageNumber="406" pageNumber="408" updateTime="1699460104163" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B0B96C53B6BF5621F3B0A4BCEAB98A03">A new semifossorial snake of the genus Arrhyton (Squamata: Dipsadidae) from eastern Cuba, with taxonomic comments on other species</mods:title>
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<mods:namePart id="9776EF6D1F298F31C2F3882F995F676D">Díaz, Luis M.</mods:namePart>
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<mods:affiliation id="4BBF5458FC49BB8E00742BE4EBC54E81">Museo Nacional de Historia Natural de Cuba. Obispo # 61, esquina Oficios, Plaza de Armas, Habana Vieja, CP 10100. luismdiazbeltran @ gmail. com; https: // orcid. org / 0000 - 0002 - 7213 - 7239</mods:affiliation>
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<mods:namePart id="EBDA8DA298D945535AA9D8D9F7912384">Fong, Ansel</mods:namePart>
<mods:affiliation id="754AD8F4223F79C769534F7B930AADE4">Centro Oriental de Ecosistemas y Biodiversidad (BIOECO), Museo de Historia Natural “ Tomás Romay ”, Enramadas # 601, Santiago de Cuba 90100, Cuba.</mods:affiliation>
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<mods:namePart id="2C4ECD9FA781DC7B97A444E4F97FCB7F">Salas, Diego</mods:namePart>
<mods:affiliation id="2D4962BDD8D85496F4DB2E117DF5D5E7">Calle Máximo Gómez # 4 entre H y Final, Guanabacoa, La Habana 12500, Cuba.</mods:affiliation>
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<mods:namePart id="597E507EBD84DEF07C9ABFC55B288FBA">Hedges, S. Blair</mods:namePart>
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<mods:affiliation id="D0A152F6391CD06A35A450C1183D6738">Department of Biology, Center for Biodiversity, Temple University, 1925 N 12 th St Suite 502, Philadelphia, PA 19122, USA. sbh @ temple. edu; https: // orcid. org / 0000 - 0002 - 0652 - 2411</mods:affiliation>
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<taxonomicName id="92E04D54FF962476FF3DFC385DF3FC2F" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[151,398,978,1005]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF962476FF3DFC385DF3FC2F" bold="true" box="[151,398,978,1005]" italics="true" pageId="2" pageNumber="408">Arrhyton albicollum</emphasis>
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<taxonomicNameLabel id="7CA757BEFF962476FE3FFC395D8EFC2F" box="[405,499,979,1005]" pageId="2" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="1DFA655CFF962476FF3DFC1D5D6FFBD3" box="[151,274,1015,1041]" pageId="2" pageNumber="408" type="description">
<paragraph id="555F36D7FF962476FF3DFC1D5D6FFBD3" blockId="2.[151,499,978,1041]" box="[151,274,1015,1041]" pageId="2" pageNumber="408">
(
<figureCitation id="CDDB2A52FF962476FF35FC1D5D74FBD3" box="[159,265,1015,1041]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="4.[151,246,1853,1877]" captionStartId-1="5.[151,246,1755,1779]" captionStartId-2="6.[151,246,769,793]" captionStartId-3="6.[151,246,1980,2004]" captionStartId-4="7.[151,246,1041,1065]" captionStartId-5="7.[151,246,1978,2002]" captionStartId-6="9.[151,246,1944,1968]" captionTargetBox-0="[164,1423,179,1827]" captionTargetBox-1="[234,1345,193,1732]" captionTargetBox-2="[248,1336,184,738]" captionTargetBox-3="[271,1318,929,1955]" captionTargetBox-4="[206,1377,636,1011]" captionTargetBox-5="[202,1386,1162,1955]" captionTargetBox-6="[184,1404,181,1920]" captionTargetId-0="figure-17@4.[164,1423,179,1827]" captionTargetId-1="figure-19@5.[213,1375,181,1732]" captionTargetId-2="figure-17@6.[248,1340,181,745]" captionTargetId-3="figure-87@6.[261,1326,925,1956]" captionTargetId-4="figure-229@7.[202,1386,633,1018]" captionTargetId-5="figure-293@7.[202,1386,1162,1955]" captionTargetId-6="figure-19@9.[184,1404,181,1920]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="7" captionTargetPageId-6="9" captionText-0="FIGURE 1. Diversity of snakes of the genus Arrhyton. (A) A. albicollum sp. nov., holotype (MNHNCu 6000) in life, from Gibara, Holguín; (B) A. redimitum, from Macambo, Guantánamo; (C) A. dolichura (DSP 8), from Rincón de Guanabo, La Habana; (D) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (E) A. supernum from La Melba, Guantánamo; (F) A. tanyplectum (MNHNCu 6002), from Mogote del Valle, Viñales, Pinar del Río; (G) A. vittatum, from Soroa,Artemisa; (H) A. taeniatum (MNHNNCu 6008), Río Máximo, Camagüey. Photos: José Luis Ponce de León (D) and Luis M. Díaz (all the others)." captionText-1="FIGURE 2. Head patterns of preserved snakes of the genus Arrhyton. (A) A. albicollum sp. nov. (holotype MNHNCu 6000), Gibara, Holguín; (B) A. redimitum (BSC.H 2670), El Cobre, Santiago de Cuba; (C) A. redimitum (BSC.H 3151), La Pimienta, Baconao, Santiago de Cuba; (D) A. redimitum (BSC.H 3963), Playa Blanca, Baracoa, Guantánamo; (E) Arrhyton aff. ainictum (BSC.H 4154), El Cucán, near Jiguaní, Granma; (F) A. dolichura (CMST 27), Bosque de La Habana, La Habana; (G) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (H) A. supernum (MNHNCu 2704) from Yunque de Baracoa, Guantánamo (holotype); (I) A. taeniatum (MNHNCu 4622), Loma de Canasí, Mayabeque; (J) A. tanyplectum (MNHNCu 6002), Mogote del Valle, Viñales, Pinar del Río; (K) A. tanyplectum (MNHNCu 6003), Idem.; (L) A. vittatum (MNHNCu 6004), Soroa, Artemisa. Photos: Ansel Fong (BE) and Luis M. Díaz (all the others)." captionText-2="FIGURE 3. Maximum-likelihood phylogeny of snakes of the genus Arrhyton from sequences of two mitochondrial genes (cytochrome b and ND4). Each species name is followed by GenBank accession numbers of the cytochrome b and ND4 genes, respectively. Voucher specimens are listed in Appendix 1. Numbers at nodes are bootstrap support values. The scale bar indicates 2% sequence divergence on a branch." captionText-3="FIGURE 4. Line-drawing of A. albicollum sp. nov. holotype (MNHNCu 6000) head in dorsal (A), ventral (B), frontal (C), and lateral (D) views. Scale bars = 2 mm." captionText-4="FIGURE 5. Distribution of Arrhyton in Cuba. Localities, other than the type-locality of A. albicollum sp. nov., are based on Rodríguez et al. (2013), Cajigas &amp; Velazco (2020), the authors visual records, and specimens listed in Appendix 1. See the Discussion for Arrhyton aff. ainictum (with question mark) and A. aff. dolichura." captionText-5="FIGURE 6. Type locality of Arrhyton albicollum sp. nov. Cueva de los Panaderos at Gibara, Holguín Province. The degraded habitat comprised of grasses and spiny bushes (dominated by the invasive Dichrostachys cinerea). Photo: Antonio Cádiz." captionText-6="FIGURE 7. Arrhyton albicollum sp. nov. unvouchered adult individual from the type locality. The mucronate rostral scale is evident in the lower photograph. Photos: Diego Salas Pantoja." figureDoi-0="http://doi.org/10.5281/zenodo.5572204" figureDoi-1="http://doi.org/10.5281/zenodo.5572206" figureDoi-2="http://doi.org/10.5281/zenodo.5572208" figureDoi-3="http://doi.org/10.5281/zenodo.5572210" figureDoi-4="http://doi.org/10.5281/zenodo.5572212" figureDoi-5="http://doi.org/10.5281/zenodo.5572214" figureDoi-6="http://doi.org/10.5281/zenodo.5572216" httpUri-0="https://zenodo.org/record/5572204/files/figure.png" httpUri-1="https://zenodo.org/record/5572206/files/figure.png" httpUri-2="https://zenodo.org/record/5572208/files/figure.png" httpUri-3="https://zenodo.org/record/5572210/files/figure.png" httpUri-4="https://zenodo.org/record/5572212/files/figure.png" httpUri-5="https://zenodo.org/record/5572214/files/figure.png" httpUri-6="https://zenodo.org/record/5572216/files/figure.png" pageId="2" pageNumber="408">Figs. 17</figureCitation>
)
</paragraph>
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<subSubSection id="1DFA655CFF962476FF37FBD2593AFBB4" pageId="2" pageNumber="408" type="materials_examined">
<paragraph id="555F36D7FF962476FF37FBD2593AFBB4" blockId="2.[151,1437,1080,2043]" pageId="2" pageNumber="408">
<materialsCitation id="E5883C8AFF962476FF37FBD2593AFBB4" ID-GBIF-Occurrence="3387243301" collectingDate="2010-01" collectionCode="WGS" collectorName="Alexis Silva Garcia &amp; Jose Raul Suarez Bauza" county="Juvenile" latitude="21.105862" location="Cueva de los Panaderos" longLatPrecision="1" longitude="-76.13822" municipality="Gibara" pageId="2" pageNumber="408" specimenCode="MHNHCu 6000" specimenCount="1" stateProvince="Holguin" typeStatus="holotype">
<emphasis id="6794EAC5FF962476FF37FBD25D77FB90" bold="true" box="[157,266,1080,1106]" pageId="2" pageNumber="408">
<typeStatus id="8A5B8875FF962476FF37FBD25D77FB90" box="[157,266,1080,1106]" pageId="2" pageNumber="408" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCode id="05469EACFF962476FEB2FBD25DADFB90" box="[280,464,1080,1106]" pageId="2" pageNumber="408">MHNHCu 6000</specimenCode>
.
<specimenCount id="43E6FD5EFF962476FE73FBD25E0AFB90" box="[473,631,1080,1106]" pageId="2" pageNumber="408" type="juvenile">Juvenile male</specimenCount>
. Entrance of
<location id="503F600CFF962476FCA2FBD2586CFB90" LSID="urn:lsid:plazi:treatment:DD4987C1FF96247CFF3DFC385F30FE6F:503F600CFF962476FCA2FBD2586CFB90" box="[776,1041,1080,1106]" county="Juvenile" latitude="21.105862" longLatPrecision="1" longitude="-76.13822" municipality="Gibara" name="Cueva de los Panaderos" pageId="2" pageNumber="408" stateProvince="Holguin">Cueva de los Panaderos</location>
(
<geoCoordinate id="30D45010FF962476FBB5FBD258E7FB91" box="[1055,1178,1080,1107]" degrees="21.105861" direction="north" orientation="latitude" pageId="2" pageNumber="408" precision="1" value="21.105862">21.105861</geoCoordinate>
,
<geoCoordinate id="30D45010FF962476FB0FFBD25955FB90" box="[1189,1320,1080,1106]" degrees="76.138222" direction="west" orientation="longitude" pageId="2" pageNumber="408" precision="1" value="-76.13822">-76.138222</geoCoordinate>
, WGS84),
<collectingMunicipality id="B53BACADFF962476FF3DFBB65C9BFBB4" box="[151,230,1116,1142]" pageId="2" pageNumber="408">Gibara</collectingMunicipality>
,
<collectingRegion id="9724F835FF962476FF5BFBB65D2DFBB4" box="[241,336,1116,1142]" country="Cuba" name="Holguin" pageId="2" pageNumber="408">Holguín</collectingRegion>
. Collected by
<collectorName id="F8155301FF962476FE45FBB65EB6FBB4" box="[495,715,1116,1142]" pageId="2" pageNumber="408">Alexis Silva García</collectorName>
and
<collectorName id="F8155301FF962476FCABFBB65870FBB4" box="[769,1037,1116,1142]" pageId="2" pageNumber="408">José Raúl Suárez Bauzá</collectorName>
on the
<collectingDate id="311AE9FFFF962476FBCAFBB6593EFBB4" box="[1120,1347,1116,1143]" pageId="2" pageNumber="408">
7th of
<date id="215E1017FF962476FB03FBB7593EFBB4" box="[1193,1347,1116,1143]" pageId="2" pageNumber="408" value="2010-01">January 2010</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1DFA655CFF962477FF6DFB6A5D97FCD7" lastPageId="3" lastPageNumber="409" pageId="2" pageNumber="408" type="diagnosis">
<paragraph id="555F36D7FF962476FF6DFB6A5977FA8C" blockId="2.[151,1437,1080,2043]" pageId="2" pageNumber="408">
<emphasis id="6794EAC5FF962476FF6DFB6A5D47FB58" bold="true" box="[199,314,1152,1178]" pageId="2" pageNumber="408">Diagnosis</emphasis>
. A species of the genus
<taxonomicName id="92E04D54FF962476FDF0FB6A5EC2FB58" authorityName=", Maglio" authorityYear="1974" box="[602,703,1152,1178]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="genus">
<emphasis id="6794EAC5FF962476FDF0FB6A5EC2FB58" box="[602,703,1152,1178]" italics="true" pageId="2" pageNumber="408">Arrhyton</emphasis>
</taxonomicName>
, related to
<taxonomicName id="92E04D54FF962476FCECFB6A5FA7FB58" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[838,986,1152,1178]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF962476FCECFB6A5FA7FB58" box="[838,986,1152,1178]" italics="true" pageId="2" pageNumber="408">A. redimitum</emphasis>
</taxonomicName>
, as shown in the molecular phylogeny of
<figureCitation id="CDDB2A52FF962476FF12FB4E5C83FB7C" box="[184,254,1188,1214]" captionStart="FIGURE 3" captionStartId="6.[151,246,769,793]" captionTargetBox="[248,1336,184,738]" captionTargetId="figure-17@6.[248,1340,181,745]" captionTargetPageId="6" captionText="FIGURE 3. Maximum-likelihood phylogeny of snakes of the genus Arrhyton from sequences of two mitochondrial genes (cytochrome b and ND4). Each species name is followed by GenBank accession numbers of the cytochrome b and ND4 genes, respectively. Voucher specimens are listed in Appendix 1. Numbers at nodes are bootstrap support values. The scale bar indicates 2% sequence divergence on a branch." figureDoi="http://doi.org/10.5281/zenodo.5572208" httpUri="https://zenodo.org/record/5572208/files/figure.png" pageId="2" pageNumber="408">Fig. 3</figureCitation>
. It can be distinguished from other species of the genus by the combination of (1) a wide immaculate white neck band or collar; (2) faint body stripes on a gray to grayish-tan background; (3) a solidly black and sharply defined cephalic pattern (“cap”,
<emphasis id="6794EAC5FF962476FDF7FB075EE6FAC4" box="[605,667,1261,1286]" italics="true" pageId="2" pageNumber="408">sensu</emphasis>
<bibRefCitation id="31714B26FF962476FD09FB065F30FAC5" author="Schwartz, A." box="[675,845,1260,1287]" pageId="2" pageNumber="408" pagination="99 - 114" refId="ref7833" refString="Schwartz, A. (1965) A review of the colubrid snake genus Arrhyton with a description of a new subspecies from southern Oriente province, Cuba. Proceedings of the Biological Society of Washington, 78, 99 - 114." type="journal article" year="1965">Schwartz 1965</bibRefCitation>
), surrounded by whitish gray; (4) a short and solidly pigmented dark stripe crossing the eye, interrupted posteriorly by the white neck band, (5) a mucronate rostral scale (see Discussion), (6) high number of ventral scales (145), and high number of subcaudal scales (132).
</paragraph>
<paragraph id="555F36D7FF962476FF67FAB35870F885" blockId="2.[151,1437,1080,2043]" pageId="2" pageNumber="408">
Most
<taxonomicName id="92E04D54FF962476FEA6FAB25DBAFAB0" box="[268,455,1368,1394]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="6794EAC5FF962476FEA6FAB25D0CFAB0" box="[268,369,1368,1394]" italics="true" pageId="2" pageNumber="408">Arrhyton</emphasis>
species
</taxonomicName>
have basically three well-defined body stripes on a tan or brown background, but there is variation in this character and two additional paravertebral stripes may be present depending on the species, giving some taxa a quinquelineate pattern (
<figureCitation id="CDDB2A52FF962476FD9FFA4A5ECFFA78" box="[565,690,1440,1466]" captionStart="FIGURE 1" captionStartId="4.[151,246,1853,1877]" captionTargetBox="[164,1423,179,1827]" captionTargetId="figure-17@4.[164,1423,179,1827]" captionTargetPageId="4" captionText="FIGURE 1. Diversity of snakes of the genus Arrhyton. (A) A. albicollum sp. nov., holotype (MNHNCu 6000) in life, from Gibara, Holguín; (B) A. redimitum, from Macambo, Guantánamo; (C) A. dolichura (DSP 8), from Rincón de Guanabo, La Habana; (D) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (E) A. supernum from La Melba, Guantánamo; (F) A. tanyplectum (MNHNCu 6002), from Mogote del Valle, Viñales, Pinar del Río; (G) A. vittatum, from Soroa,Artemisa; (H) A. taeniatum (MNHNNCu 6008), Río Máximo, Camagüey. Photos: José Luis Ponce de León (D) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572204" httpUri="https://zenodo.org/record/5572204/files/figure.png" pageId="2" pageNumber="408">Fig. 1BH</figureCitation>
).
<taxonomicName id="92E04D54FF962476FD62FA4A5FD0FA78" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[712,941,1440,1466]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF962476FD62FA4A5FD0FA78" box="[712,941,1440,1466]" italics="true" pageId="2" pageNumber="408">Arrhyton albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF962476FC1CFA4A586CFA78" bold="true" box="[950,1041,1440,1466]" pageId="2" pageNumber="408">
<taxonomicNameLabel id="7CA757BEFF962476FC1CFA4A586CFA78" box="[950,1041,1440,1466]" pageId="2" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is closely related to
<taxonomicName id="92E04D54FF962476FAA8FA4A59E8FA78" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[1282,1429,1440,1466]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF962476FAA8FA4A59E8FA78" box="[1282,1429,1440,1466]" italics="true" pageId="2" pageNumber="408">A. redimitum</emphasis>
</taxonomicName>
. Juveniles of
<taxonomicName id="92E04D54FF962476FE81FA2E5DC2FA1C" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[299,447,1476,1502]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF962476FE81FA2E5DC2FA1C" box="[299,447,1476,1502]" italics="true" pageId="2" pageNumber="408">A. redimitum</emphasis>
</taxonomicName>
may have a pair of small occipital spots interrupted by the middorsal stripe or show narrow post-occipital indentations of the paler colored areas on either side of the middorsal stripe that abruptly disconnect the paravertebral stripes from the cephalic pattern (
<figureCitation id="CDDB2A52FF962476FCC0F9E65FB3F9E4" box="[874,974,1548,1574]" captionStart="FIGURE 1" captionStartId="4.[151,246,1853,1877]" captionTargetBox="[164,1423,179,1827]" captionTargetId="figure-17@4.[164,1423,179,1827]" captionTargetPageId="4" captionText="FIGURE 1. Diversity of snakes of the genus Arrhyton. (A) A. albicollum sp. nov., holotype (MNHNCu 6000) in life, from Gibara, Holguín; (B) A. redimitum, from Macambo, Guantánamo; (C) A. dolichura (DSP 8), from Rincón de Guanabo, La Habana; (D) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (E) A. supernum from La Melba, Guantánamo; (F) A. tanyplectum (MNHNCu 6002), from Mogote del Valle, Viñales, Pinar del Río; (G) A. vittatum, from Soroa,Artemisa; (H) A. taeniatum (MNHNNCu 6008), Río Máximo, Camagüey. Photos: José Luis Ponce de León (D) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572204" httpUri="https://zenodo.org/record/5572204/files/figure.png" pageId="2" pageNumber="408">Figs. 1B</figureCitation>
and
<figureCitation id="CDDB2A52FF962476FBA0F9E6582FF9E4" box="[1034,1106,1548,1574]" captionStart="FIGURE 2" captionStartId="5.[151,246,1755,1779]" captionTargetBox="[234,1345,193,1732]" captionTargetId="figure-19@5.[213,1375,181,1732]" captionTargetPageId="5" captionText="FIGURE 2. Head patterns of preserved snakes of the genus Arrhyton. (A) A. albicollum sp. nov. (holotype MNHNCu 6000), Gibara, Holguín; (B) A. redimitum (BSC.H 2670), El Cobre, Santiago de Cuba; (C) A. redimitum (BSC.H 3151), La Pimienta, Baconao, Santiago de Cuba; (D) A. redimitum (BSC.H 3963), Playa Blanca, Baracoa, Guantánamo; (E) Arrhyton aff. ainictum (BSC.H 4154), El Cucán, near Jiguaní, Granma; (F) A. dolichura (CMST 27), Bosque de La Habana, La Habana; (G) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (H) A. supernum (MNHNCu 2704) from Yunque de Baracoa, Guantánamo (holotype); (I) A. taeniatum (MNHNCu 4622), Loma de Canasí, Mayabeque; (J) A. tanyplectum (MNHNCu 6002), Mogote del Valle, Viñales, Pinar del Río; (K) A. tanyplectum (MNHNCu 6003), Idem.; (L) A. vittatum (MNHNCu 6004), Soroa, Artemisa. Photos: Ansel Fong (BE) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572206" httpUri="https://zenodo.org/record/5572206/files/figure.png" pageId="2" pageNumber="408">2BD</figureCitation>
). The cephalic “cap” is dark brown, generally paler in the middle and outlined dark in adults. The dark stripe across the eye from the sides of snout is continuous with the lateral stripe on each side, running along the body to the tail. None of the available specimens of
<taxonomicName id="92E04D54FF962476FE9EF9925DBAF950" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[308,455,1656,1682]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF962476FE9EF9925DBAF950" box="[308,455,1656,1682]" italics="true" pageId="2" pageNumber="408">A. redimitum</emphasis>
</taxonomicName>
has a conical, abrupt, dark projection on the rostral scale. Like most
<taxonomicName id="92E04D54FF962476FB48F9925909F950" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[1250,1396,1656,1682]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF962476FB48F9925909F950" box="[1250,1396,1656,1682]" italics="true" pageId="2" pageNumber="408">A. redimitum</emphasis>
</taxonomicName>
,
<taxonomicName id="92E04D54FF962476FA2EF9925D72F974" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF962476FA2EF9925D72F974" italics="true" pageId="2" pageNumber="408">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF962476FEBFF9765D12F974" bold="true" box="[277,367,1692,1718]" pageId="2" pageNumber="408">
<taxonomicNameLabel id="7CA757BEFF962476FEBFF9765D12F974" box="[277,367,1692,1718]" pageId="2" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a dark spot on each of the internasal scales and stippled first labials. The ventral scale count of
<taxonomicName id="92E04D54FF962476FF1DF92A5D32F918" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[183,335,1728,1754]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF962476FF1DF92A5D32F918" box="[183,335,1728,1754]" italics="true" pageId="2" pageNumber="408">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF962476FEFDF92A5DCFF918" bold="true" box="[343,434,1728,1754]" pageId="2" pageNumber="408">
<taxonomicNameLabel id="7CA757BEFF962476FEFDF92A5DCFF918" box="[343,434,1728,1754]" pageId="2" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(145) is within the range of
<taxonomicName id="92E04D54FF962476FD50F92A5FF1F918" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[762,908,1728,1754]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF962476FD50F92A5FF1F918" box="[762,908,1728,1754]" italics="true" pageId="2" pageNumber="408">A. redimitum</emphasis>
</taxonomicName>
. However, only the
<typeStatus id="8A5B8875FF962476FBDFF92A58AAF918" box="[1141,1239,1728,1754]" pageId="2" pageNumber="408" type="holotype">holotype</typeStatus>
(MCZ 42505) of
<taxonomicName id="92E04D54FF962476FF3DF90E5C83F93C" box="[151,254,1764,1790]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="landoi">
<emphasis id="6794EAC5FF962476FF3DF90E5C83F93C" box="[151,254,1764,1790]" italics="true" pageId="2" pageNumber="408">A. landoi</emphasis>
</taxonomicName>
(=
<taxonomicName id="92E04D54FF962476FE8EF90E5DC9F93C" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[292,436,1764,1790]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF962476FE8EF90E5DC9F93C" box="[292,436,1764,1790]" italics="true" pageId="2" pageNumber="408">A. redimitum</emphasis>
</taxonomicName>
) has 150 ventrals, the remaining sampled individuals of this species have 110144 (mean 128, n = 54) ventrals. The number of subcaudal scales of
<taxonomicName id="92E04D54FF962476FCB6F8E25FCEF8E0" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[796,947,1800,1826]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF962476FCB6F8E25FCEF8E0" box="[796,947,1800,1826]" italics="true" pageId="2" pageNumber="408">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF962476FC10F8E25869F8E0" bold="true" box="[954,1044,1800,1826]" pageId="2" pageNumber="408">
<taxonomicNameLabel id="7CA757BEFF962476FC10F8E25869F8E0" box="[954,1044,1800,1826]" pageId="2" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is higher than that of
<taxonomicName id="92E04D54FF962476FAA1F8E259E1F8E0" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[1291,1436,1800,1826]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF962476FAA1F8E259E1F8E0" box="[1291,1436,1800,1826]" italics="true" pageId="2" pageNumber="408">A. redimitum</emphasis>
</taxonomicName>
(63125, mean 89, n = 44; specimens with missing tail tips were not included).
</paragraph>
<paragraph id="555F36D7FF962477FF6DF8BA5EA2FF17" blockId="2.[151,1437,1080,2043]" lastBlockId="3.[151,1437,150,789]" lastPageId="3" lastPageNumber="409" pageId="2" pageNumber="408">
<taxonomicName id="92E04D54FF962476FF6DF8BA5DE2F8A8" authorityName="Werner" authorityYear="1909" box="[199,415,1872,1898]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="dolichura">
<emphasis id="6794EAC5FF962476FF6DF8BA5DE2F8A8" box="[199,415,1872,1898]" italics="true" pageId="2" pageNumber="408">Arrhyton dolichura</emphasis>
</taxonomicName>
(
<figureCitation id="CDDB2A52FF962476FE07F8BA5E71F8A8" box="[429,524,1872,1898]" captionStart="FIGURE 1" captionStartId="4.[151,246,1853,1877]" captionTargetBox="[164,1423,179,1827]" captionTargetId="figure-17@4.[164,1423,179,1827]" captionTargetPageId="4" captionText="FIGURE 1. Diversity of snakes of the genus Arrhyton. (A) A. albicollum sp. nov., holotype (MNHNCu 6000) in life, from Gibara, Holguín; (B) A. redimitum, from Macambo, Guantánamo; (C) A. dolichura (DSP 8), from Rincón de Guanabo, La Habana; (D) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (E) A. supernum from La Melba, Guantánamo; (F) A. tanyplectum (MNHNCu 6002), from Mogote del Valle, Viñales, Pinar del Río; (G) A. vittatum, from Soroa,Artemisa; (H) A. taeniatum (MNHNNCu 6008), Río Máximo, Camagüey. Photos: José Luis Ponce de León (D) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572204" httpUri="https://zenodo.org/record/5572204/files/figure.png" pageId="2" pageNumber="408">Figs. 1C</figureCitation>
and
<figureCitation id="CDDB2A52FF962476FDEBF8BA5E1FF8A8" box="[577,610,1872,1898]" captionStart="FIGURE 2" captionStartId="5.[151,246,1755,1779]" captionTargetBox="[234,1345,193,1732]" captionTargetId="figure-19@5.[213,1375,181,1732]" captionTargetPageId="5" captionText="FIGURE 2. Head patterns of preserved snakes of the genus Arrhyton. (A) A. albicollum sp. nov. (holotype MNHNCu 6000), Gibara, Holguín; (B) A. redimitum (BSC.H 2670), El Cobre, Santiago de Cuba; (C) A. redimitum (BSC.H 3151), La Pimienta, Baconao, Santiago de Cuba; (D) A. redimitum (BSC.H 3963), Playa Blanca, Baracoa, Guantánamo; (E) Arrhyton aff. ainictum (BSC.H 4154), El Cucán, near Jiguaní, Granma; (F) A. dolichura (CMST 27), Bosque de La Habana, La Habana; (G) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (H) A. supernum (MNHNCu 2704) from Yunque de Baracoa, Guantánamo (holotype); (I) A. taeniatum (MNHNCu 4622), Loma de Canasí, Mayabeque; (J) A. tanyplectum (MNHNCu 6002), Mogote del Valle, Viñales, Pinar del Río; (K) A. tanyplectum (MNHNCu 6003), Idem.; (L) A. vittatum (MNHNCu 6004), Soroa, Artemisa. Photos: Ansel Fong (BE) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572206" httpUri="https://zenodo.org/record/5572206/files/figure.png" pageId="2" pageNumber="408">2F</figureCitation>
),
<taxonomicName id="92E04D54FF962476FDDFF8BA5F7DF8A8" authorityName="Hedges &amp; Garrido" authorityYear="1992" box="[629,768,1872,1898]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="procerum">
<emphasis id="6794EAC5FF962476FDDFF8BA5F7DF8A8" box="[629,768,1872,1898]" italics="true" pageId="2" pageNumber="408">A. procerum</emphasis>
</taxonomicName>
(
<figureCitation id="CDDB2A52FF962476FCA4F8BA5F12F8A8" box="[782,879,1872,1898]" captionStart="FIGURE 1" captionStartId="4.[151,246,1853,1877]" captionTargetBox="[164,1423,179,1827]" captionTargetId="figure-17@4.[164,1423,179,1827]" captionTargetPageId="4" captionText="FIGURE 1. Diversity of snakes of the genus Arrhyton. (A) A. albicollum sp. nov., holotype (MNHNCu 6000) in life, from Gibara, Holguín; (B) A. redimitum, from Macambo, Guantánamo; (C) A. dolichura (DSP 8), from Rincón de Guanabo, La Habana; (D) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (E) A. supernum from La Melba, Guantánamo; (F) A. tanyplectum (MNHNCu 6002), from Mogote del Valle, Viñales, Pinar del Río; (G) A. vittatum, from Soroa,Artemisa; (H) A. taeniatum (MNHNNCu 6008), Río Máximo, Camagüey. Photos: José Luis Ponce de León (D) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572204" httpUri="https://zenodo.org/record/5572204/files/figure.png" pageId="2" pageNumber="408">Figs. 1D</figureCitation>
and
<figureCitation id="CDDB2A52FF962476FC0EF8BA5FB4F8A8" box="[932,969,1872,1898]" captionStart="FIGURE 2" captionStartId="5.[151,246,1755,1779]" captionTargetBox="[234,1345,193,1732]" captionTargetId="figure-19@5.[213,1375,181,1732]" captionTargetPageId="5" captionText="FIGURE 2. Head patterns of preserved snakes of the genus Arrhyton. (A) A. albicollum sp. nov. (holotype MNHNCu 6000), Gibara, Holguín; (B) A. redimitum (BSC.H 2670), El Cobre, Santiago de Cuba; (C) A. redimitum (BSC.H 3151), La Pimienta, Baconao, Santiago de Cuba; (D) A. redimitum (BSC.H 3963), Playa Blanca, Baracoa, Guantánamo; (E) Arrhyton aff. ainictum (BSC.H 4154), El Cucán, near Jiguaní, Granma; (F) A. dolichura (CMST 27), Bosque de La Habana, La Habana; (G) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (H) A. supernum (MNHNCu 2704) from Yunque de Baracoa, Guantánamo (holotype); (I) A. taeniatum (MNHNCu 4622), Loma de Canasí, Mayabeque; (J) A. tanyplectum (MNHNCu 6002), Mogote del Valle, Viñales, Pinar del Río; (K) A. tanyplectum (MNHNCu 6003), Idem.; (L) A. vittatum (MNHNCu 6004), Soroa, Artemisa. Photos: Ansel Fong (BE) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572206" httpUri="https://zenodo.org/record/5572206/files/figure.png" pageId="2" pageNumber="408">2G</figureCitation>
), and
<taxonomicName id="92E04D54FF962476FBA1F8BA58CFF8A8" authorityName="Schwartz &amp; Garrido" authorityYear="1981" box="[1035,1202,1872,1898]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="tanyplectum">
<emphasis id="6794EAC5FF962476FBA1F8BA58CFF8A8" box="[1035,1202,1872,1898]" italics="true" pageId="2" pageNumber="408">A. tanyplectum</emphasis>
</taxonomicName>
(
<figureCitation id="CDDB2A52FF962476FB6BF8BA5961F8A8" box="[1217,1308,1872,1898]" captionStart="FIGURE 1" captionStartId="4.[151,246,1853,1877]" captionTargetBox="[164,1423,179,1827]" captionTargetId="figure-17@4.[164,1423,179,1827]" captionTargetPageId="4" captionText="FIGURE 1. Diversity of snakes of the genus Arrhyton. (A) A. albicollum sp. nov., holotype (MNHNCu 6000) in life, from Gibara, Holguín; (B) A. redimitum, from Macambo, Guantánamo; (C) A. dolichura (DSP 8), from Rincón de Guanabo, La Habana; (D) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (E) A. supernum from La Melba, Guantánamo; (F) A. tanyplectum (MNHNCu 6002), from Mogote del Valle, Viñales, Pinar del Río; (G) A. vittatum, from Soroa,Artemisa; (H) A. taeniatum (MNHNNCu 6008), Río Máximo, Camagüey. Photos: José Luis Ponce de León (D) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572204" httpUri="https://zenodo.org/record/5572204/files/figure.png" pageId="2" pageNumber="408">Figs. 1F</figureCitation>
and
<figureCitation id="CDDB2A52FF962476FAF8F8BA59F2F8A9" box="[1362,1423,1872,1899]" captionStart="FIGURE 2" captionStartId="5.[151,246,1755,1779]" captionTargetBox="[234,1345,193,1732]" captionTargetId="figure-19@5.[213,1375,181,1732]" captionTargetPageId="5" captionText="FIGURE 2. Head patterns of preserved snakes of the genus Arrhyton. (A) A. albicollum sp. nov. (holotype MNHNCu 6000), Gibara, Holguín; (B) A. redimitum (BSC.H 2670), El Cobre, Santiago de Cuba; (C) A. redimitum (BSC.H 3151), La Pimienta, Baconao, Santiago de Cuba; (D) A. redimitum (BSC.H 3963), Playa Blanca, Baracoa, Guantánamo; (E) Arrhyton aff. ainictum (BSC.H 4154), El Cucán, near Jiguaní, Granma; (F) A. dolichura (CMST 27), Bosque de La Habana, La Habana; (G) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (H) A. supernum (MNHNCu 2704) from Yunque de Baracoa, Guantánamo (holotype); (I) A. taeniatum (MNHNCu 4622), Loma de Canasí, Mayabeque; (J) A. tanyplectum (MNHNCu 6002), Mogote del Valle, Viñales, Pinar del Río; (K) A. tanyplectum (MNHNCu 6003), Idem.; (L) A. vittatum (MNHNCu 6004), Soroa, Artemisa. Photos: Ansel Fong (BE) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572206" httpUri="https://zenodo.org/record/5572206/files/figure.png" pageId="2" pageNumber="408">2J, K</figureCitation>
), have long tails which are 4046% of the TL versus 2540% of the TL of other species (39% in
<taxonomicName id="92E04D54FF962476FB70F89E590EF84C" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[1242,1395,1908,1934]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF962476FB70F89E590EF84C" box="[1242,1395,1908,1934]" italics="true" pageId="2" pageNumber="408">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF962476FAD1F89E5CB5F870" bold="true" pageId="2" pageNumber="408">
<taxonomicNameLabel id="7CA757BEFF962476FAD1F89E5CB5F870" pageId="2" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). These snakes have well defined stripes, and usually a vivid yellow or cream coloration on the lower flanks and the belly. In none of them is the cephalic dorsal pattern or “cap” solid black (as in
<taxonomicName id="92E04D54FF962476FBE3F85658A2F814" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[1097,1247,1980,2006]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF962476FBE3F85658A2F814" box="[1097,1247,1980,2006]" italics="true" pageId="2" pageNumber="408">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF962476FB4EF8565940F814" bold="true" box="[1252,1341,1980,2006]" pageId="2" pageNumber="408">
<taxonomicNameLabel id="7CA757BEFF962476FB4EF8565940F814" box="[1252,1341,1980,2006]" pageId="2" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and the number of subcaudal scales of
<taxonomicName id="92E04D54FF962476FE45F80A5EFBF838" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[495,646,2016,2042]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF962476FE45F80A5EFBF838" box="[495,646,2016,2042]" italics="true" pageId="2" pageNumber="408">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF962476FD27F80A5E9BF838" bold="true" box="[653,742,2016,2042]" pageId="2" pageNumber="408">
<taxonomicNameLabel id="7CA757BEFF962476FD27F80A5E9BF838" box="[653,742,2016,2042]" pageId="2" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is in the range of
<taxonomicName id="92E04D54FF962476FC04F80A582BF838" authorityName="Schwartz &amp; Garrido" authorityYear="1981" box="[942,1110,2016,2042]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="tanyplectum">
<emphasis id="6794EAC5FF962476FC04F80A582BF838" box="[942,1110,2016,2042]" italics="true" pageId="2" pageNumber="408">A. tanyplectum</emphasis>
</taxonomicName>
(
<tableCitation id="1862036CFF962476FBCEF80A58CAF838" box="[1124,1207,2016,2042]" captionStart="TABLE 1" captionStartId="3.[151,239,836,860]" captionTargetPageId="3" captionText="TABLE 1. Summarized comparative measurements and scale counts of Arrhyton. An asterisk indicates a subcaudal count that corresponds to a tail with a missing tip (see Hedges &amp; Garrido 1992)." httpUri="http://table.plazi.org/id/019F665FFF972477FF3DFCAE5FD1FC43" pageId="2" pageNumber="408" tableUuid="019F665FFF972477FF3DFCAE5FD1FC43">Table 1</tableCitation>
).
<taxonomicName id="92E04D54FF962476FB65F80A59E1F838" authorityName="Schwartz &amp; Garrido" authorityYear="1981" box="[1231,1436,2016,2042]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="408" phylum="Chordata" rank="species" species="ainictum">
<emphasis id="6794EAC5FF962476FB65F80A59E1F838" box="[1231,1436,2016,2042]" italics="true" pageId="2" pageNumber="408">Arrhyton ainictum</emphasis>
</taxonomicName>
has a well-defined quinquelineate pattern, 137 ventrals (
<emphasis id="6794EAC5FF972477FCBBFF7D5F55FF72" box="[785,808,151,176]" italics="true" pageId="3" pageNumber="409">vs</emphasis>
.
<quantity id="92189B32FF972477FC9DFF7C5F02FF73" box="[823,895,150,177]" metricMagnitude="0" metricUnit="m" metricValue="3.683" pageId="3" pageNumber="409" unit="in" value="145.0">145 in</quantity>
<taxonomicName id="92E04D54FF972477FC2CFF7D5863FF73" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[902,1054,151,177]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF972477FC2CFF7D5863FF73" box="[902,1054,151,177]" italics="true" pageId="3" pageNumber="409">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF972477FB8FFF7D5802FF73" bold="true" box="[1061,1151,151,177]" pageId="3" pageNumber="409">
<taxonomicNameLabel id="7CA757BEFF972477FB8FFF7D5802FF73" box="[1061,1151,151,177]" pageId="3" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), 108 subcaudals (
<emphasis id="6794EAC5FF972477FAFFFF7D59E0FF72" box="[1365,1437,151,176]" italics="true" pageId="3" pageNumber="409">versus</emphasis>
132), and shorter tail which is 33% of TL (
<emphasis id="6794EAC5FF972477FDDCFF515EF0FF16" box="[630,653,187,212]" italics="true" pageId="3" pageNumber="409">vs</emphasis>
. 39%).
</paragraph>
<paragraph id="555F36D7FF972477FF6DFF3559E9FE13" blockId="3.[151,1437,150,789]" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF6DFF355DEDFF3A" baseAuthorityName="Gundlach" baseAuthorityYear="1861" box="[199,400,223,249]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="vittatum">
<emphasis id="6794EAC5FF972477FF6DFF355DEDFF3A" box="[199,400,223,249]" italics="true" pageId="3" pageNumber="409">Arrhyton vittatum</emphasis>
</taxonomicName>
(
<figureCitation id="CDDB2A52FF972477FE0AFF355E7EFF3B" box="[416,515,223,249]" captionStart="FIGURE 1" captionStartId="4.[151,246,1853,1877]" captionTargetBox="[164,1423,179,1827]" captionTargetId="figure-17@4.[164,1423,179,1827]" captionTargetPageId="4" captionText="FIGURE 1. Diversity of snakes of the genus Arrhyton. (A) A. albicollum sp. nov., holotype (MNHNCu 6000) in life, from Gibara, Holguín; (B) A. redimitum, from Macambo, Guantánamo; (C) A. dolichura (DSP 8), from Rincón de Guanabo, La Habana; (D) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (E) A. supernum from La Melba, Guantánamo; (F) A. tanyplectum (MNHNCu 6002), from Mogote del Valle, Viñales, Pinar del Río; (G) A. vittatum, from Soroa,Artemisa; (H) A. taeniatum (MNHNNCu 6008), Río Máximo, Camagüey. Photos: José Luis Ponce de León (D) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572204" httpUri="https://zenodo.org/record/5572204/files/figure.png" pageId="3" pageNumber="409">Figs. 1G</figureCitation>
and
<figureCitation id="CDDB2A52FF972477FD91FF355E26FF3B" box="[571,603,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,246,1755,1779]" captionTargetBox="[234,1345,193,1732]" captionTargetId="figure-19@5.[213,1375,181,1732]" captionTargetPageId="5" captionText="FIGURE 2. Head patterns of preserved snakes of the genus Arrhyton. (A) A. albicollum sp. nov. (holotype MNHNCu 6000), Gibara, Holguín; (B) A. redimitum (BSC.H 2670), El Cobre, Santiago de Cuba; (C) A. redimitum (BSC.H 3151), La Pimienta, Baconao, Santiago de Cuba; (D) A. redimitum (BSC.H 3963), Playa Blanca, Baracoa, Guantánamo; (E) Arrhyton aff. ainictum (BSC.H 4154), El Cucán, near Jiguaní, Granma; (F) A. dolichura (CMST 27), Bosque de La Habana, La Habana; (G) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (H) A. supernum (MNHNCu 2704) from Yunque de Baracoa, Guantánamo (holotype); (I) A. taeniatum (MNHNCu 4622), Loma de Canasí, Mayabeque; (J) A. tanyplectum (MNHNCu 6002), Mogote del Valle, Viñales, Pinar del Río; (K) A. tanyplectum (MNHNCu 6003), Idem.; (L) A. vittatum (MNHNCu 6004), Soroa, Artemisa. Photos: Ansel Fong (BE) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572206" httpUri="https://zenodo.org/record/5572206/files/figure.png" pageId="3" pageNumber="409">2L</figureCitation>
) is easily distinguished from
<taxonomicName id="92E04D54FF972477FC06FF355839FF3B" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[940,1092,223,249]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF972477FC06FF355839FF3B" box="[940,1092,223,249]" italics="true" pageId="3" pageNumber="409">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF972477FBE6FF3558DAFF3B" bold="true" box="[1100,1191,223,249]" pageId="3" pageNumber="409">
<taxonomicNameLabel id="7CA757BEFF972477FBE6FF3558DAFF3B" box="[1100,1191,223,249]" pageId="3" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and the other species of
<taxonomicName id="92E04D54FF972477FF1FFEE95D67FEDF" authorityName=", Maglio" authorityYear="1974" box="[181,282,259,285]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="genus">
<emphasis id="6794EAC5FF972477FF1FFEE95D67FEDF" box="[181,282,259,285]" italics="true" pageId="3" pageNumber="409">Arrhyton</emphasis>
</taxonomicName>
in that it has a reddish brown coloration. The middorsal stripe may be absent, paler, or wider than those stripes on the flanks. Intercalated paravertebral stripes are sometimes slightly evident or generally absent in this species. The ventral coloration is pearl-white to iridescent grayish-pink. Juveniles may have some evidence of small occipital pale spots; the dark cephalic dorsal pattern is diffuse in adults (“soot-like”), not conspicuously surrounded by a distinctive pale coloration, and the snout is mostly pale brown. The number of ventral (107123) and subcaudal (5281) scales is notably lower than
<taxonomicName id="92E04D54FF972477FD99FE5D5EB7FE13" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[563,714,439,465]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF972477FD99FE5D5EB7FE13" box="[563,714,439,465]" italics="true" pageId="3" pageNumber="409">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF972477FD7BFE5D5F56FE13" bold="true" box="[721,811,439,465]" pageId="3" pageNumber="409">
<taxonomicNameLabel id="7CA757BEFF972477FD7BFE5D5F56FE13" box="[721,811,439,465]" pageId="3" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<tableCitation id="1862036CFF972477FC93FE5D5FF1FE13" box="[825,908,439,465]" captionStart="TABLE 1" captionStartId="3.[151,239,836,860]" captionTargetPageId="3" captionText="TABLE 1. Summarized comparative measurements and scale counts of Arrhyton. An asterisk indicates a subcaudal count that corresponds to a tail with a missing tip (see Hedges &amp; Garrido 1992)." httpUri="http://table.plazi.org/id/019F665FFF972477FF3DFCAE5FD1FC43" pageId="3" pageNumber="409" tableUuid="019F665FFF972477FF3DFCAE5FD1FC43">Table 1</tableCitation>
) and the tail is shorter, 2933% TL (
<emphasis id="6794EAC5FF972477FA81FE5D593FFE12" box="[1323,1346,439,464]" italics="true" pageId="3" pageNumber="409">vs</emphasis>
. 39%).
</paragraph>
<paragraph id="555F36D7FF972477FF0AFE3159E8FDFF" blockId="3.[151,1437,150,789]" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF0AFE315D0FFE36" authorityName="Hedges &amp; Garrido" authorityYear="1992" box="[160,370,475,501]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="supernum">
<emphasis id="6794EAC5FF972477FF0AFE315D0FFE36" box="[160,370,475,501]" italics="true" pageId="3" pageNumber="409">Arrhyton supernum</emphasis>
</taxonomicName>
(
<figureCitation id="CDDB2A52FF972477FE2BFE315D9FFE37" box="[385,482,475,501]" captionStart="FIGURE 1" captionStartId="4.[151,246,1853,1877]" captionTargetBox="[164,1423,179,1827]" captionTargetId="figure-17@4.[164,1423,179,1827]" captionTargetPageId="4" captionText="FIGURE 1. Diversity of snakes of the genus Arrhyton. (A) A. albicollum sp. nov., holotype (MNHNCu 6000) in life, from Gibara, Holguín; (B) A. redimitum, from Macambo, Guantánamo; (C) A. dolichura (DSP 8), from Rincón de Guanabo, La Habana; (D) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (E) A. supernum from La Melba, Guantánamo; (F) A. tanyplectum (MNHNCu 6002), from Mogote del Valle, Viñales, Pinar del Río; (G) A. vittatum, from Soroa,Artemisa; (H) A. taeniatum (MNHNNCu 6008), Río Máximo, Camagüey. Photos: José Luis Ponce de León (D) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572204" httpUri="https://zenodo.org/record/5572204/files/figure.png" pageId="3" pageNumber="409">Figs. 1E</figureCitation>
and
<figureCitation id="CDDB2A52FF972477FDB7FE315E3DFE37" box="[541,576,475,501]" captionStart="FIGURE 2" captionStartId="5.[151,246,1755,1779]" captionTargetBox="[234,1345,193,1732]" captionTargetId="figure-19@5.[213,1375,181,1732]" captionTargetPageId="5" captionText="FIGURE 2. Head patterns of preserved snakes of the genus Arrhyton. (A) A. albicollum sp. nov. (holotype MNHNCu 6000), Gibara, Holguín; (B) A. redimitum (BSC.H 2670), El Cobre, Santiago de Cuba; (C) A. redimitum (BSC.H 3151), La Pimienta, Baconao, Santiago de Cuba; (D) A. redimitum (BSC.H 3963), Playa Blanca, Baracoa, Guantánamo; (E) Arrhyton aff. ainictum (BSC.H 4154), El Cucán, near Jiguaní, Granma; (F) A. dolichura (CMST 27), Bosque de La Habana, La Habana; (G) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (H) A. supernum (MNHNCu 2704) from Yunque de Baracoa, Guantánamo (holotype); (I) A. taeniatum (MNHNCu 4622), Loma de Canasí, Mayabeque; (J) A. tanyplectum (MNHNCu 6002), Mogote del Valle, Viñales, Pinar del Río; (K) A. tanyplectum (MNHNCu 6003), Idem.; (L) A. vittatum (MNHNCu 6004), Soroa, Artemisa. Photos: Ansel Fong (BE) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572206" httpUri="https://zenodo.org/record/5572206/files/figure.png" pageId="3" pageNumber="409">2H</figureCitation>
) has two occipital pale spots, instead of a broad white band, interrupted by the middorsal stripe; the overall coloration is dark brown, compared with the grayish color of
<taxonomicName id="92E04D54FF972477FBD3FE155976FDDB" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[1145,1291,511,537]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF972477FBD3FE155976FDDB" box="[1145,1291,511,537]" italics="true" pageId="3" pageNumber="409">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF972477FABEFE155911FDDB" bold="true" box="[1300,1388,511,537]" pageId="3" pageNumber="409">
<taxonomicNameLabel id="7CA757BEFF972477FABEFE155911FDDB" box="[1300,1388,511,537]" pageId="3" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the number of ventral (124128) and subcaudal (107108) scales is much lower (
<tableCitation id="1862036CFF972477FC7EFDC9585EFDFF" box="[980,1059,547,573]" captionStart="TABLE 1" captionStartId="3.[151,239,836,860]" captionTargetPageId="3" captionText="TABLE 1. Summarized comparative measurements and scale counts of Arrhyton. An asterisk indicates a subcaudal count that corresponds to a tail with a missing tip (see Hedges &amp; Garrido 1992)." httpUri="http://table.plazi.org/id/019F665FFF972477FF3DFCAE5FD1FC43" pageId="3" pageNumber="409" tableUuid="019F665FFF972477FF3DFCAE5FD1FC43">Table 1</tableCitation>
), and the tail is 26% TL (
<emphasis id="6794EAC5FF972477FA9BFDC9593AFDFE" box="[1329,1351,547,572]" italics="true" pageId="3" pageNumber="409">vs</emphasis>
. 39%).
</paragraph>
<paragraph id="555F36D7FF972477FF6DFDAD5D97FCD7" blockId="3.[151,1437,150,789]" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF6DFDAD5DE4FDA2" authorityName="Gunther" authorityYear="1858" box="[199,409,583,609]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="taeniatum">
<emphasis id="6794EAC5FF972477FF6DFDAD5DE4FDA2" box="[199,409,583,609]" italics="true" pageId="3" pageNumber="409">Arrhyton taeniatum</emphasis>
</taxonomicName>
(
<figureCitation id="CDDB2A52FF972477FE01FDAD5E72FDA3" box="[427,527,583,609]" captionStart="FIGURE 1" captionStartId="4.[151,246,1853,1877]" captionTargetBox="[164,1423,179,1827]" captionTargetId="figure-17@4.[164,1423,179,1827]" captionTargetPageId="4" captionText="FIGURE 1. Diversity of snakes of the genus Arrhyton. (A) A. albicollum sp. nov., holotype (MNHNCu 6000) in life, from Gibara, Holguín; (B) A. redimitum, from Macambo, Guantánamo; (C) A. dolichura (DSP 8), from Rincón de Guanabo, La Habana; (D) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (E) A. supernum from La Melba, Guantánamo; (F) A. tanyplectum (MNHNCu 6002), from Mogote del Valle, Viñales, Pinar del Río; (G) A. vittatum, from Soroa,Artemisa; (H) A. taeniatum (MNHNNCu 6008), Río Máximo, Camagüey. Photos: José Luis Ponce de León (D) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572204" httpUri="https://zenodo.org/record/5572204/files/figure.png" pageId="3" pageNumber="409">Figs. 1H</figureCitation>
and
<figureCitation id="CDDB2A52FF972477FDE1FDAD5E19FDA3" box="[587,612,583,609]" captionStart="FIGURE 2" captionStartId="5.[151,246,1755,1779]" captionTargetBox="[234,1345,193,1732]" captionTargetId="figure-19@5.[213,1375,181,1732]" captionTargetPageId="5" captionText="FIGURE 2. Head patterns of preserved snakes of the genus Arrhyton. (A) A. albicollum sp. nov. (holotype MNHNCu 6000), Gibara, Holguín; (B) A. redimitum (BSC.H 2670), El Cobre, Santiago de Cuba; (C) A. redimitum (BSC.H 3151), La Pimienta, Baconao, Santiago de Cuba; (D) A. redimitum (BSC.H 3963), Playa Blanca, Baracoa, Guantánamo; (E) Arrhyton aff. ainictum (BSC.H 4154), El Cucán, near Jiguaní, Granma; (F) A. dolichura (CMST 27), Bosque de La Habana, La Habana; (G) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (H) A. supernum (MNHNCu 2704) from Yunque de Baracoa, Guantánamo (holotype); (I) A. taeniatum (MNHNCu 4622), Loma de Canasí, Mayabeque; (J) A. tanyplectum (MNHNCu 6002), Mogote del Valle, Viñales, Pinar del Río; (K) A. tanyplectum (MNHNCu 6003), Idem.; (L) A. vittatum (MNHNCu 6004), Soroa, Artemisa. Photos: Ansel Fong (BE) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572206" httpUri="https://zenodo.org/record/5572206/files/figure.png" pageId="3" pageNumber="409">2I</figureCitation>
) is the largest species (up to
<quantity id="92189B32FF972477FC0DFDAC587FFDA3" box="[935,1026,582,609]" metricMagnitude="-1" metricUnit="m" metricValue="4.57" pageId="3" pageNumber="409" unit="mm" value="457.0">457 mm</quantity>
SVL, MNHNCu 4622) and it occurs syntopically with
<taxonomicName id="92E04D54FF972477FEF8FD815DA2FD47" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[338,479,619,645]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF972477FEF8FD815DA2FD47" box="[338,479,619,645]" italics="true" pageId="3" pageNumber="409">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF972477FE4EFD815E45FD47" bold="true" box="[484,568,619,645]" pageId="3" pageNumber="409">
<taxonomicNameLabel id="7CA757BEFF972477FE4EFD815E45FD47" box="[484,568,619,645]" pageId="3" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
It is the only species which always lacks a loreal scale. It has wide body stripes that in some individuals are indistinct from the brown background, vivid yellow flanks and belly, the highest number of ventral scales (168189,
<emphasis id="6794EAC5FF972477FEFFFD595D11FD0E" box="[341,364,691,716]" italics="true" pageId="3" pageNumber="409">vs</emphasis>
.
<quantity id="92189B32FF972477FED1FD585DBEFD0F" box="[379,451,690,717]" metricMagnitude="0" metricUnit="m" metricValue="3.683" pageId="3" pageNumber="409" unit="in" value="145.0">145 in</quantity>
<taxonomicName id="92E04D54FF972477FE60FD595E25FD0F" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[458,600,691,717]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF972477FE60FD595E25FD0F" box="[458,600,691,717]" italics="true" pageId="3" pageNumber="409">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF972477FDF4FD595ECEFD0F" bold="true" box="[606,691,691,717]" pageId="3" pageNumber="409">
<taxonomicNameLabel id="7CA757BEFF972477FDF4FD595ECEFD0F" box="[606,691,691,717]" pageId="3" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), a projected and upturned rostral scale, a laterally expanded head and smaller eyes which are far from the head outline when seen dorsally
<emphasis id="6794EAC5FF972477FCDDFD3D5FF0FD32" box="[887,909,727,752]" italics="true" pageId="3" pageNumber="409">vs</emphasis>
. larger and slightly protruded eyes from the head outline in
<taxonomicName id="92E04D54FF972477FEABFD115DF2FCD7" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[257,399,763,789]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF972477FEABFD115DF2FCD7" box="[257,399,763,789]" italics="true" pageId="3" pageNumber="409">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF972477FE3FFD115D97FCD7" bold="true" box="[405,490,763,789]" pageId="3" pageNumber="409">
<taxonomicNameLabel id="7CA757BEFF972477FE3FFD115D97FCD7" box="[405,490,763,789]" pageId="3" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<caption id="019F665FFF972477FF3DFCAE5FD1FC43" ID-Table-UUID="019F665FFF972477FF3DFCAE5FD1FC43" httpUri="http://table.plazi.org/id/019F665FFF972477FF3DFCAE5FD1FC43" pageId="3" pageNumber="409" startId="3.[151,239,836,860]" targetBox="[163,1363,921,1348]" targetIsTable="true" targetPageId="3">
<paragraph id="555F36D7FF972477FF3DFCAE5FD1FC43" blockId="3.[151,1436,836,897]" pageId="3" pageNumber="409">
<emphasis id="6794EAC5FF972477FF3DFCAE5D74FC9E" bold="true" box="[151,265,836,860]" pageId="3" pageNumber="409">TABLE 1.</emphasis>
Summarized comparative measurements and scale counts of
<taxonomicName id="92E04D54FF972477FC38FCAE5F8FFC9E" authorityName=", Maglio" authorityYear="1974" box="[914,1010,836,860]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="genus">
<emphasis id="6794EAC5FF972477FC38FCAE5F8FFC9E" box="[914,1010,836,860]" italics="true" pageId="3" pageNumber="409">Arrhyton</emphasis>
</taxonomicName>
. An asterisk indicates a subcaudal count that corresponds to a tail with a missing tip (see
<bibRefCitation id="31714B26FF972477FD31FC825FDDFC43" author="Hedges, S. B. &amp; Garrido, O. H." box="[667,928,872,897]" pageId="3" pageNumber="409" pagination="168 - 177" refId="ref7460" refString="Hedges, S. B. &amp; Garrido, O. H. (1992) Cuban snakes of the genus Arrhyton: two new species and a reconsideration of A. redimitum Cope. Herpetologica, 48 (2), 168 - 177." type="journal article" year="1992">Hedges &amp; Garrido 1992</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="555F36D7FF972477FEB7FC42595CFA81" pageId="3" pageNumber="409">
<table id="27E0C477FF97DB8BFF09FC73592EFA86" box="[163,1363,921,1348]" gridcols="5" gridrows="10" pageId="3" pageNumber="409">
<tr id="EBD03495FF97DB8BFF09FC73592EFC0C" box="[163,1363,921,974]" gridrow="0" pageId="3" pageNumber="409">
<th id="A8015DE9FF97DB8BFF09FC735D0CFC0C" box="[163,369,921,974]" gridcol="0" gridrow="0" pageId="3" pageNumber="409">Species</th>
<th id="A8015DE9FF97DB8BFDACFC735EE3FC0C" box="[518,670,921,974]" gridcol="1" gridrow="0" pageId="3" pageNumber="409">Maximum SVL (mm)</th>
<th id="A8015DE9FF97DB8BFD73FC735F21FC0C" box="[729,860,921,974]" gridcol="2" gridrow="0" pageId="3" pageNumber="409">Maximum Ta (mm)</th>
<th id="A8015DE9FF97DB8BFC1BFC735848FC0C" box="[945,1077,921,974]" gridcol="3" gridrow="0" pageId="3" pageNumber="409">Ventral scales</th>
<th id="A8015DE9FF97DB8BFB1AFC73592EFC0C" box="[1200,1363,921,974]" gridcol="4" gridrow="0" pageId="3" pageNumber="409">Subcaudal scales</th>
</tr>
<tr id="EBD03495FF97DB8BFF09FC0D592EFC3D" box="[163,1363,999,1023]" gridrow="1" pageId="3" pageNumber="409">
<th id="A8015DE9FF97DB8BFF09FC0D5D0CFC3D" box="[163,369,999,1023]" gridcol="0" gridrow="1" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF09FC0D5D5DFC3D" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[163,288,999,1023]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF972477FF09FC0D5D5DFC3D" box="[163,288,999,1023]" italics="true" pageId="3" pageNumber="409">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF972477FE8CFC0D5D0CFC3D" bold="true" box="[294,369,999,1023]" pageId="3" pageNumber="409">
<taxonomicNameLabel id="7CA757BEFF972477FE8CFC0D5D0CFC3D" box="[294,369,999,1023]" pageId="3" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="A8015DE9FF97DB8BFDACFC0D5EE3FC3D" box="[518,670,999,1023]" gridcol="1" gridrow="1" pageId="3" pageNumber="409">143.5</td>
<td id="A8015DE9FF97DB8BFD73FC0D5F21FC3D" box="[729,860,999,1023]" gridcol="2" gridrow="1" pageId="3" pageNumber="409">90.5</td>
<td id="A8015DE9FF97DB8BFC1BFC0D5848FC3D" box="[945,1077,999,1023]" gridcol="3" gridrow="1" pageId="3" pageNumber="409">145</td>
<td id="A8015DE9FF97DB8BFB1AFC0D592EFC3D" box="[1200,1363,999,1023]" gridcol="4" gridrow="1" pageId="3" pageNumber="409">132</td>
</tr>
<tr id="EBD03495FF97DB8BFF09FBE5592EFBE5" box="[163,1363,1039,1063]" gridrow="2" pageId="3" pageNumber="409">
<th id="A8015DE9FF97DB8BFF09FBE55D0CFBE5" box="[163,369,1039,1063]" gridcol="0" gridrow="2" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF09FBFA5D62FBE5" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[163,287,1039,1063]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF972477FF09FBFA5D62FBE5" box="[163,287,1039,1063]" italics="true" pageId="3" pageNumber="409">A. redimitum</emphasis>
</taxonomicName>
</th>
<td id="A8015DE9FF97DB8BFDACFBE55EE3FBE5" box="[518,670,1039,1063]" gridcol="1" gridrow="2" pageId="3" pageNumber="409">250</td>
<td id="A8015DE9FF97DB8BFD73FBE55F21FBE5" box="[729,860,1039,1063]" gridcol="2" gridrow="2" pageId="3" pageNumber="409">158</td>
<td id="A8015DE9FF97DB8BFC1BFBE55848FBE5" box="[945,1077,1039,1063]" gridcol="3" gridrow="2" pageId="3" pageNumber="409">115150</td>
<td id="A8015DE9FF97DB8BFB1AFBE5592EFBE5" box="[1200,1363,1039,1063]" gridcol="4" gridrow="2" pageId="3" pageNumber="409">69125</td>
</tr>
<tr id="EBD03495FF97DB8BFF09FBDD592EFB92" box="[163,1363,1079,1104]" gridrow="3" pageId="3" pageNumber="409">
<th id="A8015DE9FF97DB8BFF09FBDD5D0CFB92" box="[163,369,1079,1104]" gridcol="0" gridrow="3" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF09FBD25D6CFB8D" authorityName="Schwartz &amp; Garrido" authorityYear="1981" box="[163,273,1080,1103]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="ainictum">
<emphasis id="6794EAC5FF972477FF09FBD25D6CFB8D" box="[163,273,1080,1103]" italics="true" pageId="3" pageNumber="409">A. ainictum</emphasis>
</taxonomicName>
</th>
<td id="A8015DE9FF97DB8BFDACFBDD5EE3FB92" box="[518,670,1079,1104]" gridcol="1" gridrow="3" pageId="3" pageNumber="409">363</td>
<td id="A8015DE9FF97DB8BFD73FBDD5F21FB92" box="[729,860,1079,1104]" gridcol="2" gridrow="3" pageId="3" pageNumber="409">176</td>
<td id="A8015DE9FF97DB8BFC1BFBDD5848FB92" box="[945,1077,1079,1104]" gridcol="3" gridrow="3" pageId="3" pageNumber="409">137</td>
<td id="A8015DE9FF97DB8BFB1AFBDD592EFB92" box="[1200,1363,1079,1104]" gridcol="4" gridrow="3" pageId="3" pageNumber="409">108</td>
</tr>
<tr id="EBD03495FF97DB8BFF09FB8A592EFBBA" box="[163,1363,1120,1144]" gridrow="4" pageId="3" pageNumber="409">
<th id="A8015DE9FF97DB8BFF09FB8A5D0CFBBA" box="[163,369,1120,1144]" gridcol="0" gridrow="4" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF09FB8A5D67FBBA" authorityName="Werner" authorityYear="1909" box="[163,282,1120,1144]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="dolichura">
<emphasis id="6794EAC5FF972477FF09FB8A5D67FBBA" box="[163,282,1120,1144]" italics="true" pageId="3" pageNumber="409">A. dolichura</emphasis>
</taxonomicName>
</th>
<td id="A8015DE9FF97DB8BFDACFB8A5EE3FBBA" box="[518,670,1120,1144]" gridcol="1" gridrow="4" pageId="3" pageNumber="409">282</td>
<td id="A8015DE9FF97DB8BFD73FB8A5F21FBBA" box="[729,860,1120,1144]" gridcol="2" gridrow="4" pageId="3" pageNumber="409">200</td>
<td id="A8015DE9FF97DB8BFC1BFB8A5848FBBA" box="[945,1077,1120,1144]" gridcol="3" gridrow="4" pageId="3" pageNumber="409">123132</td>
<td id="A8015DE9FF97DB8BFB1AFB8A592EFBBA" box="[1200,1363,1120,1144]" gridcol="4" gridrow="4" pageId="3" pageNumber="409">101127</td>
</tr>
<tr id="EBD03495FF97DB8BFF09FB6D592EFB63" box="[163,1363,1159,1185]" gridrow="5" pageId="3" pageNumber="409">
<th id="A8015DE9FF97DB8BFF09FB6D5D0CFB63" box="[163,369,1159,1185]" gridcol="0" gridrow="5" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF09FB635D67FB62" authorityName="Hedges &amp; Garrido" authorityYear="1992" box="[163,282,1161,1184]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="procerum">
<emphasis id="6794EAC5FF972477FF09FB635D67FB62" box="[163,282,1161,1184]" italics="true" pageId="3" pageNumber="409">A. procerum</emphasis>
</taxonomicName>
</th>
<td id="A8015DE9FF97DB8BFDACFB6D5EE3FB63" box="[518,670,1159,1185]" gridcol="1" gridrow="5" pageId="3" pageNumber="409">267</td>
<td id="A8015DE9FF97DB8BFD73FB6D5F21FB63" box="[729,860,1159,1185]" gridcol="2" gridrow="5" pageId="3" pageNumber="409">211</td>
<td id="A8015DE9FF97DB8BFC1BFB6D5848FB63" box="[945,1077,1159,1185]" gridcol="3" gridrow="5" pageId="3" pageNumber="409">142145</td>
<td id="A8015DE9FF97DB8BFB1AFB6D592EFB63" box="[1200,1363,1159,1185]" gridcol="4" gridrow="5" pageId="3" pageNumber="409">137140*</td>
</tr>
<tr id="EBD03495FF97DB8BFF09FB5B592EFB0B" box="[163,1363,1201,1225]" gridrow="6" pageId="3" pageNumber="409">
<th id="A8015DE9FF97DB8BFF09FB5B5D0CFB0B" box="[163,369,1201,1225]" gridcol="0" gridrow="6" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF09FB5B5D61FB0B" authorityName="Hedges &amp; Garrido" authorityYear="1992" box="[163,284,1201,1225]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="supernum">
<emphasis id="6794EAC5FF972477FF09FB5B5D61FB0B" box="[163,284,1201,1225]" italics="true" pageId="3" pageNumber="409">A. supernum</emphasis>
</taxonomicName>
</th>
<td id="A8015DE9FF97DB8BFDACFB5B5EE3FB0B" box="[518,670,1201,1225]" gridcol="1" gridrow="6" pageId="3" pageNumber="409">255</td>
<td id="A8015DE9FF97DB8BFD73FB5B5F21FB0B" box="[729,860,1201,1225]" gridcol="2" gridrow="6" pageId="3" pageNumber="409">171</td>
<td id="A8015DE9FF97DB8BFC1BFB5B5848FB0B" box="[945,1077,1201,1225]" gridcol="3" gridrow="6" pageId="3" pageNumber="409">124128</td>
<td id="A8015DE9FF97DB8BFB1AFB5B592EFB0B" box="[1200,1363,1201,1225]" gridcol="4" gridrow="6" pageId="3" pageNumber="409">107108</td>
</tr>
<tr id="EBD03495FF97DB8BFF09FB33592EFB33" box="[163,1363,1241,1265]" gridrow="7" pageId="3" pageNumber="409">
<th id="A8015DE9FF97DB8BFF09FB335D0CFB33" box="[163,369,1241,1265]" gridcol="0" gridrow="7" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF09FB305D60FB33" authorityName="Gunther" authorityYear="1858" box="[163,285,1242,1265]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="taeniatum">
<emphasis id="6794EAC5FF972477FF09FB305D60FB33" box="[163,285,1242,1265]" italics="true" pageId="3" pageNumber="409">A. taeniatum</emphasis>
</taxonomicName>
</th>
<td id="A8015DE9FF97DB8BFDACFB335EE3FB33" box="[518,670,1241,1265]" gridcol="1" gridrow="7" pageId="3" pageNumber="409">457</td>
<td id="A8015DE9FF97DB8BFD73FB335F21FB33" box="[729,860,1241,1265]" gridcol="2" gridrow="7" pageId="3" pageNumber="409">150</td>
<td id="A8015DE9FF97DB8BFC1BFB335848FB33" box="[945,1077,1241,1265]" gridcol="3" gridrow="7" pageId="3" pageNumber="409">168189</td>
<td id="A8015DE9FF97DB8BFB1AFB33592EFB33" box="[1200,1363,1241,1265]" gridcol="4" gridrow="7" pageId="3" pageNumber="409">7399</td>
</tr>
<tr id="EBD03495FF97DB8BFF09FAE8592EFAD9" box="[163,1363,1282,1307]" gridrow="8" pageId="3" pageNumber="409">
<th id="A8015DE9FF97DB8BFF09FAE85D0CFAD9" box="[163,369,1282,1307]" gridcol="0" gridrow="8" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF09FAE95D4FFAD8" authorityName="Schwartz &amp; Garrido" authorityYear="1981" box="[163,306,1282,1306]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="tanyplectum">
<emphasis id="6794EAC5FF972477FF09FAE95D4FFAD8" box="[163,306,1282,1306]" italics="true" pageId="3" pageNumber="409">A. tanyplectum</emphasis>
</taxonomicName>
</th>
<td id="A8015DE9FF97DB8BFDACFAE85EE3FAD9" box="[518,670,1282,1307]" gridcol="1" gridrow="8" pageId="3" pageNumber="409">321</td>
<td id="A8015DE9FF97DB8BFD73FAE85F21FAD9" box="[729,860,1282,1307]" gridcol="2" gridrow="8" pageId="3" pageNumber="409">236</td>
<td id="A8015DE9FF97DB8BFC1BFAE85848FAD9" box="[945,1077,1282,1307]" gridcol="3" gridrow="8" pageId="3" pageNumber="409">141146</td>
<td id="A8015DE9FF97DB8BFB1AFAE8592EFAD9" box="[1200,1363,1282,1307]" gridcol="4" gridrow="8" pageId="3" pageNumber="409">121133</td>
</tr>
<tr id="EBD03495FF97DB8BFF09FAC1592EFA86" box="[163,1363,1323,1348]" gridrow="9" pageId="3" pageNumber="409">
<th id="A8015DE9FF97DB8BFF09FAC15D0CFA86" box="[163,369,1323,1348]" gridcol="0" gridrow="9" pageId="3" pageNumber="409">
<taxonomicName id="92E04D54FF972477FF09FAC65D71FA81" baseAuthorityName="Gundlach" baseAuthorityYear="1861" box="[163,268,1324,1347]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="vittatum">
<emphasis id="6794EAC5FF972477FF09FAC65D71FA81" box="[163,268,1324,1347]" italics="true" pageId="3" pageNumber="409">A. vittatum</emphasis>
</taxonomicName>
</th>
<td id="A8015DE9FF97DB8BFDACFAC15EE3FA86" box="[518,670,1323,1348]" gridcol="1" gridrow="9" pageId="3" pageNumber="409">207</td>
<td id="A8015DE9FF97DB8BFD73FAC15F21FA86" box="[729,860,1323,1348]" gridcol="2" gridrow="9" pageId="3" pageNumber="409">87</td>
<td id="A8015DE9FF97DB8BFC1BFAC15848FA86" box="[945,1077,1323,1348]" gridcol="3" gridrow="9" pageId="3" pageNumber="409">107123</td>
<td id="A8015DE9FF97DB8BFB1AFAC1592EFA86" box="[1200,1363,1323,1348]" gridcol="4" gridrow="9" pageId="3" pageNumber="409">5281</td>
</tr>
</table>
</paragraph>
<subSubSection id="1DFA655CFF972473FF6DFA995F1CFE37" lastPageId="7" lastPageNumber="413" pageId="3" pageNumber="409" type="description">
<paragraph id="555F36D7FF972477FF6DFA995D35F833" blockId="3.[151,1437,1395,2033]" pageId="3" pageNumber="409">
<emphasis id="6794EAC5FF972477FF6DFA995DA1FA4F" bold="true" box="[199,476,1395,1421]" pageId="3" pageNumber="409">
Description of
<typeStatus id="8A5B8875FF972477FEDFFA995DA1FA4F" box="[373,476,1395,1421]" pageId="3" pageNumber="409" type="holotype">holotype</typeStatus>
</emphasis>
. See
<figureCitation id="CDDB2A52FF972477FDB2FA995E24FA4F" box="[536,601,1395,1421]" captionStart="FIGURE 4" captionStartId="6.[151,246,1980,2004]" captionTargetBox="[271,1318,929,1955]" captionTargetId="figure-87@6.[261,1326,925,1956]" captionTargetPageId="6" captionText="FIGURE 4. Line-drawing of A. albicollum sp. nov. holotype (MNHNCu 6000) head in dorsal (A), ventral (B), frontal (C), and lateral (D) views. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5572210" httpUri="https://zenodo.org/record/5572210/files/figure.png" pageId="3" pageNumber="409">Fig. 4</figureCitation>
for head scales. SVL
<quantity id="92189B32FF972477FCE1FA995FBDFA4C" box="[843,960,1395,1422]" metricMagnitude="-1" metricUnit="m" metricValue="1.435" pageId="3" pageNumber="409" unit="mm" value="143.5">143.5 mm</quantity>
; Ta
<quantity id="92189B32FF972477FC46FA995832FA4C" box="[1004,1103,1395,1422]" metricMagnitude="-2" metricUnit="m" metricValue="9.05" pageId="3" pageNumber="409" unit="mm" value="90.5">90.5 mm</quantity>
(39% TL),
<quantity id="92189B32FF972477FB7AFA995951FA4C" box="[1232,1324,1395,1422]" metricMagnitude="-1" metricUnit="m" metricValue="2.34" pageId="3" pageNumber="409" unit="mm" value="234.0">234 mm</quantity>
TL. Head distinct from body; head length
<quantity id="92189B32FF972477FDA3FA7D5E22FA70" box="[521,607,1431,1458]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="3" pageNumber="409" unit="mm" value="8.6">8.6 mm</quantity>
(6% SVL), head width
<quantity id="92189B32FF972477FCD9FA7D5FB4FA70" box="[883,969,1431,1458]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="409" unit="mm" value="5.5">5.5 mm</quantity>
(64% head length) at widest point, head height
<quantity id="92189B32FF972477FF4DFA515D40FA14" box="[231,317,1467,1494]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="409" unit="mm" value="3.5">3.5 mm</quantity>
at tallest point; interocular distance
<quantity id="92189B32FF972477FD4FFA515F46FA14" box="[741,827,1467,1494]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="409" unit="mm" value="3.7">3.7 mm</quantity>
(67% head width); snout-eye distance
<quantity id="92189B32FF972477FAAAFA51592BFA14" box="[1280,1366,1467,1494]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="409" unit="mm" value="2.2">2.2 mm</quantity>
(25% head length, HL); nostril-eye distance
<quantity id="92189B32FF972477FDE5FA355ED7FA38" box="[591,682,1503,1530]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="409" unit="mm" value="1.5">1.5 mm</quantity>
; rostral subtriangular,
<quantity id="92189B32FF972477FC00FA35587DFA38" box="[938,1024,1503,1530]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="409" unit="mm" value="1.9">1.9 mm</quantity>
wide and
<quantity id="92189B32FF972477FBD3FA3558B2FA38" box="[1145,1231,1503,1530]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="409" unit="mm" value="1.2">1.2 mm</quantity>
high, well visible in dorsal view; prefrontal divided,
<quantity id="92189B32FF972477FD80F9E95EFEF9DC" box="[554,643,1539,1566]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="409" unit="mm" value="1.6">1.6 mm</quantity>
long (17% HL), 2.3 times longer than internasals; prefrontals not in contact with orbit, but instead contacting supraocular and upper half of the preocular; frontal subtriangular
<quantity id="92189B32FF972477FAEDF9CD59E0F980" box="[1351,1437,1575,1602]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="409" unit="mm" value="2.5">2.5 mm</quantity>
long (29% HL) and
<quantity id="92189B32FF972477FED0F9A15DADF9A4" box="[378,464,1611,1638]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="409" unit="mm" value="2.2">2.2 mm</quantity>
wide; supraocular
<quantity id="92189B32FF972477FD0CF9A15E81F9A4" box="[678,764,1611,1638]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="409" unit="mm" value="1.8">1.8 mm</quantity>
long (21% HL) and
<quantity id="92189B32FF972477FC4CF9A15841F9A4" box="[998,1084,1611,1638]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="409" unit="mm" value="0.9">0.9 mm</quantity>
wide, twice wider distally than proximally; parietals
<quantity id="92189B32FF972477FE20F9855D9FF948" box="[394,482,1647,1674]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="409" unit="mm" value="3.4">3.4 mm</quantity>
long (39% HL),
<quantity id="92189B32FF972477FD02F9855F7DF948" box="[680,768,1647,1674]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="409" unit="mm" value="2.4">2.4 mm</quantity>
wide; nasal divided; nostril horizontally oval, its posterior border lying on the nasal suture, but most of the opening is on the prenasal scale; prenasal scale
<quantity id="92189B32FF972477FB78F9795954F96C" box="[1234,1321,1683,1710]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="409" unit="mm" value="0.7">0.7 mm</quantity>
(8% HL); postnasal scale
<quantity id="92189B32FF972477FEE7F95D5DDBF910" box="[333,422,1719,1746]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="409" unit="mm" value="0.6">0.6 mm</quantity>
(7% HL); loreal single, quadrangular,
<quantity id="92189B32FF972477FCC4F95D5FB5F910" box="[878,968,1719,1746]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="409" unit="mm" value="0.6">0.6 mm</quantity>
long (7% HL) and
<quantity id="92189B32FF972477FB1EF95D5973F910" box="[1204,1294,1719,1746]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="409" unit="mm" value="0.4">0.4 mm</quantity>
high; loreal contacting second supralabial; preocular single, two times higher than wide, entering orbit posteriorly; eye diameter
<quantity id="92189B32FF972477FF3DF9155C92F8D8" box="[151,239,1791,1818]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="409" unit="mm" value="1.9">1.9 mm</quantity>
(22% HL), pupil round; postoculars 2/2, upper postocular twice higher than lower postocular; temporals 1+2+3 on both sides of head; supralabials 7/7, of which 3rd and 4th are in contact with the eye; 6th supralabial slightly tallest than 5th, both in contact with first temporal; infralabials 8/8, first pair in contact with mental, 1st to 3rd contacting first chinshields, 4th and 5th contacting second chinshields; dorsals smooth, rhomb shaped, with no pits externally visible; dorsal scales row formula 171717; cloacal plate divided; number of ventral and subcaudal scales were already mentioned in the diagnosis; a terminal conical projection about five times longer than the two last subcaudals.
</paragraph>
<caption id="019F665FFF902470FF3DF8D75880F827" ID-DOI="http://doi.org/10.5281/zenodo.5572204" ID-Zenodo-Dep="5572204" httpUri="https://zenodo.org/record/5572204/files/figure.png" pageId="4" pageNumber="410" startId="4.[151,246,1853,1877]" targetBox="[164,1423,179,1827]" targetPageId="4">
<paragraph id="555F36D7FF902470FF3DF8D75880F827" blockId="4.[151,1437,1853,2021]" pageId="4" pageNumber="410">
<emphasis id="6794EAC5FF902470FF3DF8D75D74F897" bold="true" box="[151,265,1853,1877]" pageId="4" pageNumber="410">FIGURE 1</emphasis>
. Diversity of snakes of the genus
<taxonomicName id="92E04D54FF902470FDFEF8D75ED7F897" authorityName=", Maglio" authorityYear="1974" box="[596,682,1853,1877]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="genus">
<emphasis id="6794EAC5FF902470FDFEF8D75ED7F897" box="[596,682,1853,1877]" italics="true" pageId="4" pageNumber="410">Arrhyton</emphasis>
</taxonomicName>
. (
<emphasis id="6794EAC5FF902470FD15F8D75EACF897" bold="true" box="[703,721,1853,1877]" pageId="4" pageNumber="410">A</emphasis>
)
<taxonomicName id="92E04D54FF902470FD75F8D75F1FF897" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[735,866,1853,1877]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF902470FD75F8D75F1FF897" box="[735,866,1853,1877]" italics="true" pageId="4" pageNumber="410">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF902470FCC2F8D75FC8F897" bold="true" box="[872,949,1853,1877]" pageId="4" pageNumber="410">
<taxonomicNameLabel id="7CA757BEFF902470FCC2F8D75FC8F897" box="[872,949,1853,1877]" pageId="4" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (MNHNCu 6000) in life, from Gibara, Holguín; (
<emphasis id="6794EAC5FF902470FF54F88B5D73F8BB" bold="true" box="[254,270,1889,1913]" pageId="4" pageNumber="410">B</emphasis>
)
<taxonomicName id="92E04D54FF902470FEB5F88B5DE3F8BB" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[287,414,1889,1913]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF902470FEB5F88B5DE3F8BB" box="[287,414,1889,1913]" italics="true" pageId="4" pageNumber="410">A. redimitum</emphasis>
</taxonomicName>
, from Macambo, Guantánamo; (
<emphasis id="6794EAC5FF902470FD43F88B5E86F8BB" bold="true" box="[745,763,1889,1913]" pageId="4" pageNumber="410">C</emphasis>
)
<taxonomicName id="92E04D54FF902470FCA6F88B5FFAF8BB" authorityName="Werner" authorityYear="1909" box="[780,903,1889,1913]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="dolichura">
<emphasis id="6794EAC5FF902470FCA6F88B5FFAF8BB" box="[780,903,1889,1913]" italics="true" pageId="4" pageNumber="410">A. dolichura</emphasis>
</taxonomicName>
(DSP 8), from Rincón de Guanabo, La Habana; (
<emphasis id="6794EAC5FF902470FA28F88B59E9F8BB" bold="true" box="[1410,1428,1889,1913]" pageId="4" pageNumber="410">D</emphasis>
)
<taxonomicName id="92E04D54FF902470FF3DF86F5D6FF85F" authorityName="Hedges &amp; Garrido" authorityYear="1992" box="[151,274,1925,1949]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="procerum">
<emphasis id="6794EAC5FF902470FF3DF86F5D6FF85F" box="[151,274,1925,1949]" italics="true" pageId="4" pageNumber="410">A. procerum</emphasis>
</taxonomicName>
(MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (
<emphasis id="6794EAC5FF902470FC20F86F5FE7F85F" bold="true" box="[906,922,1925,1949]" pageId="4" pageNumber="410">E</emphasis>
)
<taxonomicName id="92E04D54FF902470FC01F86F585AF85F" authorityName="Hedges &amp; Garrido" authorityYear="1992" box="[939,1063,1925,1949]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="supernum">
<emphasis id="6794EAC5FF902470FC01F86F585AF85F" box="[939,1063,1925,1949]" italics="true" pageId="4" pageNumber="410">A. supernum</emphasis>
</taxonomicName>
from La Melba, Guantánamo; (
<emphasis id="6794EAC5FF902470FAC2F86F590AF85F" bold="true" box="[1384,1399,1925,1949]" pageId="4" pageNumber="410">F</emphasis>
)
<taxonomicName id="92E04D54FF902470FA2DF86F5D71F803" authorityName="Schwartz &amp; Garrido" authorityYear="1981" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="tanyplectum">
<emphasis id="6794EAC5FF902470FA2DF86F5D71F803" italics="true" pageId="4" pageNumber="410">A. tanyplectum</emphasis>
</taxonomicName>
(MNHNCu 6002), from Mogote del Valle, Viñales, Pinar del Río; (
<emphasis id="6794EAC5FF902470FC24F8435FDCF803" bold="true" box="[910,929,1961,1985]" pageId="4" pageNumber="410">G</emphasis>
)
<taxonomicName id="92E04D54FF902470FC04F843586BF803" baseAuthorityName="Gundlach" baseAuthorityYear="1861" box="[942,1046,1961,1985]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="vittatum">
<emphasis id="6794EAC5FF902470FC04F843586BF803" box="[942,1046,1961,1985]" italics="true" pageId="4" pageNumber="410">A. vittatum</emphasis>
</taxonomicName>
, from Soroa, Artemisa; (
<emphasis id="6794EAC5FF902470FAA8F8435968F803" bold="true" box="[1282,1301,1961,1985]" pageId="4" pageNumber="410">H</emphasis>
)
<taxonomicName id="92E04D54FF902470FA88F84359E1F803" authorityName="Gunther" authorityYear="1858" box="[1314,1436,1961,1985]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="taeniatum">
<emphasis id="6794EAC5FF902470FA88F84359E1F803" box="[1314,1436,1961,1985]" italics="true" pageId="4" pageNumber="410">A. taeniatum</emphasis>
</taxonomicName>
(MNHNNCu 6008), Río Máximo, Camagüey. Photos: José Luis Ponce de León (D) and Luis M. Díaz (all the others).
</paragraph>
</caption>
<caption id="019F665FFF912471FF3DF9315811F832" ID-DOI="http://doi.org/10.5281/zenodo.5572206" ID-Zenodo-Dep="5572206" httpUri="https://zenodo.org/record/5572206/files/figure.png" pageId="5" pageNumber="411" startId="5.[151,246,1755,1779]" targetBox="[234,1345,193,1732]" targetPageId="5">
<paragraph id="555F36D7FF912471FF3DF9315811F832" blockId="5.[151,1437,1755,2032]" pageId="5" pageNumber="411">
<emphasis id="6794EAC5FF912471FF3DF9315D77F931" bold="true" box="[151,266,1755,1779]" pageId="5" pageNumber="411">FIGURE 2</emphasis>
. Head patterns of preserved snakes of the genus
<taxonomicName id="92E04D54FF912471FCA8F9365F20F936" authorityName=", Maglio" authorityYear="1974" box="[770,861,1756,1780]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="genus">
<emphasis id="6794EAC5FF912471FCA8F9365F20F936" box="[770,861,1756,1780]" italics="true" pageId="5" pageNumber="411">Arrhyton</emphasis>
</taxonomicName>
. (
<emphasis id="6794EAC5FF912471FCD9F9315FF8F931" bold="true" box="[883,901,1755,1779]" pageId="5" pageNumber="411">A</emphasis>
)
<taxonomicName id="92E04D54FF912471FC3CF9365862F936" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[918,1055,1756,1780]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF912471FC3CF9365862F936" box="[918,1055,1756,1780]" italics="true" pageId="5" pageNumber="411">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF912471FB8CF9365805F936" bold="true" box="[1062,1144,1756,1780]" pageId="5" pageNumber="411">
<taxonomicNameLabel id="7CA757BEFF912471FB8CF9365805F936" box="[1062,1144,1756,1780]" pageId="5" pageNumber="411" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype MNHNCu 6000), Gibara, Holguín; (
<emphasis id="6794EAC5FF912471FEF8F8EA5D1EF8DA" bold="true" box="[338,355,1792,1816]" pageId="5" pageNumber="411">B</emphasis>
)
<taxonomicName id="92E04D54FF912471FED8F8EA5EEFF8DA" authority="(BSC. H 2670)" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[370,658,1792,1816]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF912471FED8F8EA5D8BF8DA" box="[370,502,1792,1816]" italics="true" pageId="5" pageNumber="411">A. redimitum</emphasis>
(BSC.H 2670)
</taxonomicName>
, El Cobre, Santiago de Cuba; (
<emphasis id="6794EAC5FF912471FC65F9155F9CF8D5" bold="true" box="[975,993,1791,1815]" pageId="5" pageNumber="411">C</emphasis>
)
<taxonomicName id="92E04D54FF912471FC5AF8EA5809F8DA" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[1008,1140,1792,1816]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF912471FC5AF8EA5809F8DA" box="[1008,1140,1792,1816]" italics="true" pageId="5" pageNumber="411">A. redimitum</emphasis>
</taxonomicName>
(BSC.H 3151), La Pimienta, Baconao, Santiago de Cuba; (
<emphasis id="6794EAC5FF912471FE6CF8CE5DA5F8FE" bold="true" box="[454,472,1828,1852]" pageId="5" pageNumber="411">D</emphasis>
)
<taxonomicName id="92E04D54FF912471FE4DF8CE5E17F8FE" baseAuthorityName="BSC. H" baseAuthorityYear="2670" box="[487,618,1828,1852]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="redimitum">
<emphasis id="6794EAC5FF912471FE4DF8CE5E17F8FE" box="[487,618,1828,1852]" italics="true" pageId="5" pageNumber="411">A. redimitum</emphasis>
</taxonomicName>
(BSC.H 3963), Playa Blanca, Baracoa, Guantánamo; (
<emphasis id="6794EAC5FF912471FB30F8CE58D6F8FE" bold="true" box="[1178,1195,1828,1852]" pageId="5" pageNumber="411">E</emphasis>
)
<taxonomicName id="92E04D54FF912471FB10F8CE59E0F8F9" authorityName="Schwartz &amp; Garrido" authorityYear="1981" box="[1210,1437,1828,1852]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="ainictum">
<emphasis id="6794EAC5FF912471FB10F8CE5968F8FE" box="[1210,1301,1828,1852]" italics="true" pageId="5" pageNumber="411">Arrhyton</emphasis>
aff.
<emphasis id="6794EAC5FF912471FAEEF8CE59E0F8F9" box="[1348,1437,1828,1851]" italics="true" pageId="5" pageNumber="411">ainictum</emphasis>
</taxonomicName>
(BSC.H 4154), El Cucán, near Jiguaní, Granma; (
<emphasis id="6794EAC5FF912471FD0DF8A25ECBF8A2" bold="true" box="[679,694,1864,1888]" pageId="5" pageNumber="411">F</emphasis>
)
<taxonomicName id="92E04D54FF912471FD63F8A25F36F8A2" authorityName="Werner" authorityYear="1909" box="[713,843,1864,1888]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="dolichura">
<emphasis id="6794EAC5FF912471FD63F8A25F36F8A2" box="[713,843,1864,1888]" italics="true" pageId="5" pageNumber="411">A. dolichura</emphasis>
</taxonomicName>
(CMST 27), Bosque de La Habana, La Habana; (
<emphasis id="6794EAC5FF912471FACBF8AD5908F89D" bold="true" box="[1377,1397,1863,1887]" pageId="5" pageNumber="411">G</emphasis>
)
<taxonomicName id="92E04D54FF912471FA2CF8A25C84F841" authorityName="Hedges &amp; Garrido" authorityYear="1992" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="procerum">
<emphasis id="6794EAC5FF912471FA2CF8A25C84F841" italics="true" pageId="5" pageNumber="411">A. procerum</emphasis>
</taxonomicName>
(MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (
<emphasis id="6794EAC5FF912471FC37F8865FCCF846" bold="true" box="[925,945,1900,1924]" pageId="5" pageNumber="411">H</emphasis>
)
<taxonomicName id="92E04D54FF912471FC69F886583AF841" authorityName="Hedges &amp; Garrido" authorityYear="1992" box="[963,1095,1900,1923]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="supernum">
<emphasis id="6794EAC5FF912471FC69F886583AF841" box="[963,1095,1900,1923]" italics="true" pageId="5" pageNumber="411">A. supernum</emphasis>
</taxonomicName>
(MNHNCu 2704) from Yunque de Baracoa, Guantánamo (holotype); (
<emphasis id="6794EAC5FF912471FD82F87A5E4FF86A" bold="true" box="[552,562,1936,1960]" pageId="5" pageNumber="411">I</emphasis>
)
<taxonomicName id="92E04D54FF912471FDEEF87A5EB5F865" authorityName="Gunther" authorityYear="1858" box="[580,712,1936,1959]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="taeniatum">
<emphasis id="6794EAC5FF912471FDEEF87A5EB5F865" box="[580,712,1936,1959]" italics="true" pageId="5" pageNumber="411">A. taeniatum</emphasis>
</taxonomicName>
(MNHNCu 4622), Loma de Canasí, Mayabeque; (
<emphasis id="6794EAC5FF912471FB49F87A588DF86A" bold="true" box="[1251,1264,1936,1960]" pageId="5" pageNumber="411">J</emphasis>
)
<taxonomicName id="92E04D54FF912471FAABF87A59E1F86A" authorityName="Schwartz &amp; Garrido" authorityYear="1981" box="[1281,1436,1936,1960]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="tanyplectum">
<emphasis id="6794EAC5FF912471FAABF87A59E1F86A" box="[1281,1436,1936,1960]" italics="true" pageId="5" pageNumber="411">A. tanyplectum</emphasis>
</taxonomicName>
(MNHNCu 6002), Mogote del Valle, Viñales, Pinar del Río; (
<emphasis id="6794EAC5FF912471FC88F85E5F4BF80E" bold="true" box="[802,822,1972,1996]" pageId="5" pageNumber="411">K</emphasis>
)
<taxonomicName id="92E04D54FF912471FCE2F85E5F9EF80E" authorityName="Schwartz &amp; Garrido" authorityYear="1981" box="[840,995,1972,1996]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="tanyplectum">
<emphasis id="6794EAC5FF912471FCE2F85E5F9EF80E" box="[840,995,1972,1996]" italics="true" pageId="5" pageNumber="411">A. tanyplectum</emphasis>
</taxonomicName>
(MNHNCu 6003), Idem.; (
<emphasis id="6794EAC5FF912471FAADF85E5965F80E" bold="true" box="[1287,1304,1972,1996]" pageId="5" pageNumber="411">L</emphasis>
)
<taxonomicName id="92E04D54FF912471FA80F85E59E1F809" baseAuthorityName="Gundlach" baseAuthorityYear="1861" box="[1322,1436,1972,1995]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="vittatum">
<emphasis id="6794EAC5FF912471FA80F85E59E1F809" box="[1322,1436,1972,1995]" italics="true" pageId="5" pageNumber="411">A. vittatum</emphasis>
</taxonomicName>
(MNHNCu 6004), Soroa, Artemisa. Photos: Ansel Fong (BE) and Luis M. Díaz (all the others).
</paragraph>
</caption>
<caption id="019F665FFF922472FF3DFCEB5E68FC47" ID-DOI="http://doi.org/10.5281/zenodo.5572208" ID-Zenodo-Dep="5572208" httpUri="https://zenodo.org/record/5572208/files/figure.png" pageId="6" pageNumber="412" startId="6.[151,246,769,793]" targetBox="[248,1336,184,738]" targetPageId="6">
<paragraph id="555F36D7FF922472FF3DFCEB5E68FC47" blockId="6.[151,1437,769,901]" pageId="6" pageNumber="412">
<emphasis id="6794EAC5FF922472FF3DFCEB5D71FCDB" bold="true" box="[151,268,769,793]" pageId="6" pageNumber="412">FIGURE 3</emphasis>
. Maximum-likelihood phylogeny of snakes of the genus
<taxonomicName id="92E04D54FF922472FCC6FCEB5FBAFCDB" authorityName=", Maglio" authorityYear="1974" box="[876,967,769,793]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="412" phylum="Chordata" rank="genus">
<emphasis id="6794EAC5FF922472FCC6FCEB5FBAFCDB" box="[876,967,769,793]" italics="true" pageId="6" pageNumber="412">Arrhyton</emphasis>
</taxonomicName>
from sequences of two mitochondrial genes (cytochrome
<emphasis id="6794EAC5FF922472FEB4FCCF5D56FCFF" box="[286,299,805,829]" italics="true" pageId="6" pageNumber="412">b</emphasis>
and
<emphasis id="6794EAC5FF922472FEF6FCCF5DF6FCFF" box="[348,395,805,829]" italics="true" pageId="6" pageNumber="412">ND4</emphasis>
). Each species name is followed by GenBank accession numbers of the cytochrome
<emphasis id="6794EAC5FF922472FB42FCCF5888FCFF" box="[1256,1269,805,829]" italics="true" pageId="6" pageNumber="412">b</emphasis>
and
<emphasis id="6794EAC5FF922472FA8CFCCF592BFCFF" box="[1318,1366,805,829]" italics="true" pageId="6" pageNumber="412">ND4</emphasis>
genes, respectively. Voucher specimens are listed in Appendix 1. Numbers at nodes are bootstrap support values. The scale bar indicates 2% sequence divergence on a branch.
</paragraph>
</caption>
<caption id="019F665FFF922472FF3DF8565E6FF835" ID-DOI="http://doi.org/10.5281/zenodo.5572210" ID-Zenodo-Dep="5572210" httpUri="https://zenodo.org/record/5572210/files/figure.png" pageId="6" pageNumber="412" startId="6.[151,246,1980,2004]" targetBox="[271,1318,929,1955]" targetPageId="6">
<paragraph id="555F36D7FF922472FF3DF8565E6FF835" blockId="6.[151,1437,1980,2040]" pageId="6" pageNumber="412">
<emphasis id="6794EAC5FF922472FF3DF8565D74F816" bold="true" box="[151,265,1980,2004]" pageId="6" pageNumber="412">FIGURE 4</emphasis>
. Line-drawing of
<taxonomicName id="92E04D54FF922472FE6AF8565E35F816" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[448,584,1980,2004]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="412" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF922472FE6AF8565E35F816" box="[448,584,1980,2004]" italics="true" pageId="6" pageNumber="412">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF922472FDE5F8565EDDF816" bold="true" box="[591,672,1980,2004]" pageId="6" pageNumber="412">
<taxonomicNameLabel id="7CA757BEFF922472FDE5F8565EDDF816" box="[591,672,1980,2004]" pageId="6" pageNumber="412" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype (MNHNCu 6000) head in dorsal (A), ventral (B), frontal (C), and lateral (D) views. Scale bars = 2 mm.
</paragraph>
</caption>
<paragraph id="555F36D7FF932473FF6DFF7D5EE6FE6F" blockId="7.[151,1437,151,610]" pageId="7" pageNumber="413">
Color in life (
<figureCitation id="CDDB2A52FF932473FEC7FF7D5DB6FF73" box="[365,459,151,177]" captionStart="FIGURE 1" captionStartId="4.[151,246,1853,1877]" captionTargetBox="[164,1423,179,1827]" captionTargetId="figure-17@4.[164,1423,179,1827]" captionTargetPageId="4" captionText="FIGURE 1. Diversity of snakes of the genus Arrhyton. (A) A. albicollum sp. nov., holotype (MNHNCu 6000) in life, from Gibara, Holguín; (B) A. redimitum, from Macambo, Guantánamo; (C) A. dolichura (DSP 8), from Rincón de Guanabo, La Habana; (D) A. procerum (MNHNCu 6001), Gironcito, Península de Zapata, Matanzas; (E) A. supernum from La Melba, Guantánamo; (F) A. tanyplectum (MNHNCu 6002), from Mogote del Valle, Viñales, Pinar del Río; (G) A. vittatum, from Soroa,Artemisa; (H) A. taeniatum (MNHNNCu 6008), Río Máximo, Camagüey. Photos: José Luis Ponce de León (D) and Luis M. Díaz (all the others)." figureDoi="http://doi.org/10.5281/zenodo.5572204" httpUri="https://zenodo.org/record/5572204/files/figure.png" pageId="7" pageNumber="413">Fig. 1A</figureCitation>
): Dorsum gray to grayish-tan, with faint stripes. Each scale with somewhat darker pigmentation on the edge giving the overall dorsal region a net-like appearance. The neck band is immaculate white, and behind it there is a slightly darker, smoky-gray pigmentation that transitions to the dorsal color. Internasal scales with a dark spot on each. Another dark spot is present on each prenasal scale. First three supralabials and four infralabials stippled with dark brown. Rostral conical projection dark. Eye bicolored: upper half grayish-white; inferior half very dark brown, almost black. Venter paler than dorsum. A faint lateral stripe on scale row four, just defined by slightly darker pigmentation of scale borders, giving a serrated outline and bordered at the top by a narrow zone paler than the rest of the dorsum.
</paragraph>
<paragraph id="555F36D7FF932473FF6DFE5D5F1CFE37" blockId="7.[151,1437,151,610]" pageId="7" pageNumber="413">Color in alcohol (ten years after collection): similar to coloration in life, except that yellowish pigments completely disappeared. The faint stripes are even less obvious.</paragraph>
</subSubSection>
<subSubSection id="1DFA655CFF93247CFF64FE145870FE83" lastPageId="8" lastPageNumber="414" pageId="7" pageNumber="413" type="distribution">
<paragraph id="555F36D7FF932473FF64FE145F89FDA0" blockId="7.[151,1437,151,610]" pageId="7" pageNumber="413">
<emphasis id="6794EAC5FF932473FF64FE145D1AFDDA" bold="true" box="[206,359,510,536]" pageId="7" pageNumber="413">Distribution.</emphasis>
<taxonomicName id="92E04D54FF932473FEC4FE155E2FFDDB" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[366,594,511,537]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="413" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF932473FEC4FE155E2FFDDB" box="[366,594,511,537]" italics="true" pageId="7" pageNumber="413">Arrhyton albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF932473FDF3FE155ECFFDDB" bold="true" box="[601,690,511,537]" pageId="7" pageNumber="413">
<taxonomicNameLabel id="7CA757BEFF932473FDF3FE155ECFFDDB" box="[601,690,511,537]" pageId="7" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from the
<typeStatus id="8A5B8875FF932473FC6CFE155F8BFDDB" box="[966,1014,511,537]" pageId="7" pageNumber="413">type</typeStatus>
locality near the town of Gibara, ~
<quantity id="92189B32FF932473FAD5FE155CC6FDFF" metricMagnitude="4" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="413" unit="km" value="36.0">36 km</quantity>
NNE from the city of
<collectingRegion id="9724F835FF932473FE19FDC95E72FDFF" box="[435,527,547,573]" country="Cuba" name="Holguin" pageId="7" pageNumber="413">Holguín</collectingRegion>
in eastern
<collectingCountry id="2DF77647FF932473FD2DFDC95EBFFDFF" box="[647,706,547,573]" name="Cuba" pageId="7" pageNumber="413">Cuba</collectingCountry>
(
<figureCitation id="CDDB2A52FF932473FD7AFDC95F6EFDFF" box="[720,787,546,573]" captionStart="FIGURE 5" captionStartId="7.[151,246,1041,1065]" captionTargetBox="[206,1377,636,1011]" captionTargetId="figure-229@7.[202,1386,633,1018]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of Arrhyton in Cuba. Localities, other than the type-locality of A. albicollum sp. nov., are based on Rodríguez et al. (2013), Cajigas &amp; Velazco (2020), the authors visual records, and specimens listed in Appendix 1. See the Discussion for Arrhyton aff. ainictum (with question mark) and A. aff. dolichura." figureDoi="http://doi.org/10.5281/zenodo.5572212" httpUri="https://zenodo.org/record/5572212/files/figure.png" pageId="7" pageNumber="413">Fig. 5</figureCitation>
), but it probably occurs in suitable habitats between Bahía de Puerto Padre and Banes (
<collectingRegion id="9724F835FF932473FE78FDAD5E3BFDA3" box="[466,582,583,609]" country="Cuba" name="Las Tunas" pageId="7" pageNumber="413">Las Tunas</collectingRegion>
and
<collectingRegion id="9724F835FF932473FDD6FDAD5EA5FDA3" box="[636,728,583,609]" country="Cuba" name="Holguin" pageId="7" pageNumber="413">Holguín</collectingRegion>
Provinces, respectively).
</paragraph>
<caption id="019F665FFF932473FF3DFBFB5FB1FBB3" ID-DOI="http://doi.org/10.5281/zenodo.5572212" ID-Zenodo-Dep="5572212" httpUri="https://zenodo.org/record/5572212/files/figure.png" pageId="7" pageNumber="413" startId="7.[151,246,1041,1065]" targetBox="[206,1377,636,1011]" targetPageId="7">
<paragraph id="555F36D7FF932473FF3DFBFB5FB1FBB3" blockId="7.[151,1436,1041,1138]" pageId="7" pageNumber="413">
<emphasis id="6794EAC5FF932473FF3DFBFB5D77FBE8" bold="true" box="[151,266,1041,1066]" pageId="7" pageNumber="413">FIGURE 5</emphasis>
. Distribution of
<taxonomicName id="92E04D54FF932473FE12FBF85E6EFBE8" authorityName=", Maglio" authorityYear="1974" box="[440,531,1042,1066]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="413" phylum="Chordata" rank="genus">
<emphasis id="6794EAC5FF932473FE12FBF85E6EFBE8" box="[440,531,1042,1066]" italics="true" pageId="7" pageNumber="413">Arrhyton</emphasis>
</taxonomicName>
in Cuba. Localities, other than the type-locality of
<taxonomicName id="92E04D54FF932473FB82FBF858CCFBE8" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[1064,1201,1042,1066]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="413" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF932473FB82FBF858CCFBE8" box="[1064,1201,1042,1066]" italics="true" pageId="7" pageNumber="413">A. albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF932473FB10FBF85971FBE8" bold="true" box="[1210,1292,1042,1066]" pageId="7" pageNumber="413">
<taxonomicNameLabel id="7CA757BEFF932473FB10FBF85971FBE8" box="[1210,1292,1042,1066]" pageId="7" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, are based on
<bibRefCitation id="31714B26FF932473FF3DFBDC5DF6FB8C" author="Rodriguez Schettino, L. &amp; Mancina, C. A. &amp; Rivalta Gonzalez, V." box="[151,395,1078,1102]" pageId="7" pageNumber="413" pagination="1 - 92" refId="ref7780" refString="Rodriguez Schettino, L., Mancina, C. A. &amp; Rivalta Gonzalez, V. (2013) Reptiles of Cuba: Checklist and geographic distributions. Smithsonian Herpetological Information Service, 144, 1 - 92. https: // doi. org / 10.5479 / si. 23317515.144.1" type="journal article" year="2013">
Rodríguez
<emphasis id="6794EAC5FF932473FEA3FBDC5D4AFB8C" box="[265,311,1078,1102]" italics="true" pageId="7" pageNumber="413">et al</emphasis>
. (2013)
</bibRefCitation>
, Cajigas &amp; Velazco (2020), the authors visual records, and specimens listed in Appendix 1. See the Discussion for
<taxonomicName id="92E04D54FF932473FE9BFBB05E6EFBB3" authorityName="Schwartz &amp; Garrido" authorityYear="1981" box="[305,531,1114,1138]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="413" phylum="Chordata" rank="species" species="ainictum">
<emphasis id="6794EAC5FF932473FE9BFBB05DF1FBB0" box="[305,396,1114,1138]" italics="true" pageId="7" pageNumber="413">Arrhyton</emphasis>
aff.
<emphasis id="6794EAC5FF932473FE10FBB05E6EFBB3" box="[442,531,1114,1137]" italics="true" pageId="7" pageNumber="413">ainictum</emphasis>
</taxonomicName>
(with question mark) and
<taxonomicName id="92E04D54FF932473FCB5FBB05FBBFBB0" authorityName="Werner" authorityYear="1909" box="[799,966,1114,1138]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="413" phylum="Chordata" rank="species" species="dolichura">
<emphasis id="6794EAC5FF932473FCB5FBB05F53FBB3" box="[799,814,1114,1137]" italics="true" pageId="7" pageNumber="413">A</emphasis>
. aff.
<emphasis id="6794EAC5FF932473FCC9FBB05FBBFBB0" box="[867,966,1114,1138]" italics="true" pageId="7" pageNumber="413">dolichura</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="019F665FFF932473FF3DF850591CF834" ID-DOI="http://doi.org/10.5281/zenodo.5572214" ID-Zenodo-Dep="5572214" httpUri="https://zenodo.org/record/5572214/files/figure.png" pageId="7" pageNumber="413" startId="7.[151,246,1978,2002]" targetBox="[202,1386,1162,1955]" targetPageId="7">
<paragraph id="555F36D7FF932473FF3DF850591CF834" blockId="7.[151,1436,1978,2039]" pageId="7" pageNumber="413">
<emphasis id="6794EAC5FF932473FF3DF8505D75F810" bold="true" box="[151,264,1978,2002]" pageId="7" pageNumber="413">FIGURE 6</emphasis>
. Type locality of
<taxonomicName id="92E04D54FF932473FE17F8505EF6F810" authority="Díaz &amp; Fong &amp; Salas &amp; Hedges, 2021" authorityName="Díaz &amp; Fong &amp; Salas &amp; Hedges" authorityYear="2021" box="[445,651,1978,2002]" class="Reptilia" family="Dipsadidae" genus="Arrhyton" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="413" phylum="Chordata" rank="species" species="albicollum" status="sp. nov.">
<emphasis id="6794EAC5FF932473FE17F8505EF6F810" box="[445,651,1978,2002]" italics="true" pageId="7" pageNumber="413">Arrhyton albicollum</emphasis>
</taxonomicName>
<emphasis id="6794EAC5FF932473FD38F8515E9EF811" bold="true" box="[658,739,1979,2003]" pageId="7" pageNumber="413">
<taxonomicNameLabel id="7CA757BEFF932473FD38F8515E9EF811" box="[658,739,1979,2003]" pageId="7" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Cueva de los Panaderos at Gibara, Holguín Province. The degraded habitat comprised of grasses and spiny bushes (dominated by the invasive
<taxonomicName id="92E04D54FF932473FC20F8345814F834" box="[906,1129,2014,2038]" class="Magnoliopsida" family="Fabaceae" genus="Dichrostachys" kingdom="Plantae" order="Fabales" pageId="7" pageNumber="413" phylum="Tracheophyta" rank="species" species="cinerea">
<emphasis id="6794EAC5FF932473FC20F8345814F834" box="[906,1129,2014,2038]" italics="true" pageId="7" pageNumber="413">Dichrostachys cinerea</emphasis>
</taxonomicName>
). Photo: Antonio Cádiz.
</paragraph>
</caption>
<paragraph id="555F36D7FF9C247CFF6DFF7C5870FE83" blockId="8.[151,1437,150,429]" pageId="8" pageNumber="414">
<emphasis id="6794EAC5FF9C247CFF6DFF7C5D5EFF72" bold="true" box="[199,291,150,176]" pageId="8" pageNumber="414">Habitat</emphasis>
. The
<typeStatus id="8A5B8875FF9C247CFECEFF7D5DE9FF73" box="[356,404,151,177]" pageId="8" pageNumber="414">type</typeStatus>
specimen was found under a pile of rocks at the entrance of Cueva de los Panaderos during the day (
<figureCitation id="CDDB2A52FF9C247CFF54FF515D38FF14" box="[254,325,187,214]" captionStart="FIGURE 6" captionStartId="7.[151,246,1978,2002]" captionTargetBox="[202,1386,1162,1955]" captionTargetId="figure-293@7.[202,1386,1162,1955]" captionTargetPageId="7" captionText="FIGURE 6. Type locality of Arrhyton albicollum sp. nov. Cueva de los Panaderos at Gibara, Holguín Province. The degraded habitat comprised of grasses and spiny bushes (dominated by the invasive Dichrostachys cinerea). Photo: Antonio Cádiz." figureDoi="http://doi.org/10.5281/zenodo.5572214" httpUri="https://zenodo.org/record/5572214/files/figure.png" pageId="8" pageNumber="414">Fig. 6</figureCitation>
). Caves and sinkholes are common structures in the area. The locality has patches of secondary semideciduous forest on limestone rock; however, the native vegetation is being displaced by the invasive spiny-bush
<taxonomicName id="92E04D54FF9C247CFF3DFEE95DEEFEDE" box="[151,403,259,285]" class="Magnoliopsida" family="Fabaceae" genus="Dichrostachys" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="414" phylum="Tracheophyta" rank="species" species="cinerea">
<emphasis id="6794EAC5FF9C247CFF3DFEE95DEEFEDE" box="[151,403,259,285]" italics="true" pageId="8" pageNumber="414">Dichrostachys cinerea</emphasis>
</taxonomicName>
and the whole area is threatened by human use of caves and the trash dumping. Associated herpetofauna and complementary description of habitat is in
<bibRefCitation id="31714B26FF9C247CFCEAFECD5874FE83" author="Diaz, L. M. &amp; Cadiz, A. &amp; Villar, S. &amp; Bermudez, F." box="[832,1033,295,321]" pageId="8" pageNumber="414" pagination="116 - 119" refId="ref7026" refString="Diaz, L. M., Cadiz, A., Villar, S. &amp; Bermudez, F. (2014) Notes on the ecology and morphology of the Cuban khaki trope, Tropidophis hendersoni Hedges and Garrido (Squamata: Tropidophiidae), with a new locality record. IRCF Reptiles &amp; Amphibians, 21, 116 - 119. https: // doi. org / 10.17161 / randa. v 21 i 4.14009" type="journal article" year="2014">
Díaz
<emphasis id="6794EAC5FF9C247CFCD6FECD5FD0FE83" box="[892,941,295,321]" italics="true" pageId="8" pageNumber="414">et al</emphasis>
. (2014)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1DFA655CFF9C247CFF6DFEA15F30FE6F" pageId="8" pageNumber="414" type="etymology">
<paragraph id="555F36D7FF9C247CFF6DFEA15E53FE4B" blockId="8.[151,1437,150,429]" pageId="8" pageNumber="414">
<emphasis id="6794EAC5FF9C247CFF6DFEA15D33FEA7" bold="true" box="[199,334,331,357]" pageId="8" pageNumber="414">Etymology.</emphasis>
The specific epithet is derived from the Latin words
<emphasis id="6794EAC5FF9C247CFC09FEA15F9DFEA7" box="[931,992,331,357]" italics="true" pageId="8" pageNumber="414">albus</emphasis>
(white) and
<emphasis id="6794EAC5FF9C247CFBC4FEA158C7FEA7" box="[1134,1210,331,357]" italics="true" pageId="8" pageNumber="414">collum</emphasis>
(neck), in reference to the conspicuous white neck band.
</paragraph>
<paragraph id="555F36D7FF9C247CFF6DFE795F30FE6F" blockId="8.[151,1437,150,429]" box="[199,845,402,429]" pageId="8" pageNumber="414">
<emphasis id="6794EAC5FF9C247CFF6DFE795D9DFE6F" bold="true" box="[199,480,402,429]" pageId="8" pageNumber="414">Proposed English name</emphasis>
. Gibara White-collared Racerlet
</paragraph>
</subSubSection>
</treatment>
</document>