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<mods:titleInfo id="EEB7956A76AD6F45302A9EE8D9514C68">
<mods:title id="E0E1E3A8B6C08A4BDB38B3172B306EE4">Resin bees of genus Megachile, subgenera Callomegachile and Carinula (Hymenoptera, Megachilidae) from Thailand with description of a new species</mods:title>
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<mods:namePart id="AFD319F60083BB8C7E45FBA3B139B49E">Chatthanabun, Nontawat</mods:namePart>
<mods:affiliation id="7819E7D1545DBE74460D3506E0A3C43C">Center of Excellence in Entomology and Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="306A1DDD77686D3968AE4193CB3F20D2">Ascher, John S.</mods:namePart>
<mods:affiliation id="96554CA21F84795A48FC208A816F662B">Insect Diversity Lab, Department of Biological Sciences, National University of Singapore, 16 Science Drive 4 S 3 Level 4, 117558, Singapore</mods:affiliation>
</mods:name>
<mods:name id="37A8C58D75531200E2A6147E419FA7B1" type="personal">
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<mods:roleTerm id="AE3F44ADD002AF465005B686F45B8A25">Author</mods:roleTerm>
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<mods:namePart id="DCA4857013D70BF6949BC05837C459B0">Pinkaew, Nantasak</mods:namePart>
<mods:affiliation id="465518B1749384C1474A486256935290">Department of Entomology, Faculty of Agriculture at Kamphaengsaen, Kasetsart University, Kamphaengsaen Campus, Nakhon Pathom, 73140, Thailand</mods:affiliation>
</mods:name>
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<mods:roleTerm id="2EBC1131C467DA1D1AB2CDBA8B9B6DBC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2A6ECEB2FF94CF8132680865A7F9600D">Thanoosing, Chawatat</mods:namePart>
<mods:affiliation id="39918FE9CD9D7B2996F0DAA5C15579BC">Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK &amp; Department of Life Sciences, Imperial College London, South Kensington Campus, Exhibition Road, London SW 7 2 AZ, UK</mods:affiliation>
</mods:name>
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<mods:namePart id="DABE6DC2F43608DF141FFB216C1AA191">Traiyasut, Prapun</mods:namePart>
<mods:affiliation id="E1A5644EC8DBC6C23B8A6F849EC5912A">Program in Biology, Faculty of Science, Ubon Ratchathani Rajabhat University, Ubon Ratchathani, 34000, Thailand</mods:affiliation>
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<mods:namePart id="684B6A07F9E4B4FE23DB1B66DBFD2691">Warrit, Natapot</mods:namePart>
<mods:affiliation id="C4C1E74040EC045D1D49123E46CAF116">Center of Excellence in Entomology and Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:date id="79C2797DC9C13F4AF096497EE4BB8EC4">2020</mods:date>
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<mods:number id="13FD0309373C277486D1ABEC00BFE718">997</mods:number>
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<treatment id="DD1DE8D984D452E783D19EBD4C5E1414" ID-GBIF-Taxon="170531172" LSID="urn:lsid:plazi:treatment:DD1DE8D984D452E783D19EBD4C5E1414" httpUri="http://treatment.plazi.org/id/DD1DE8D984D452E783D19EBD4C5E1414" lastPageNumber="95" pageId="0" pageNumber="95" scope_family="Megachilidae" scope_order="Hymenoptera">
<subSubSection id="2236EFC092E6B69D9EB74ACDA7298865" pageId="0" pageNumber="95" type="nomenclature">
<paragraph id="C1A61270F3BDA51DD9EE5FC7AE3BD69B" pageId="0" pageNumber="95">
<taxonomicName id="6353554577F85FE29A75D267E3937390" ID-CoL="8SPMM" LSID="DD1DE8D9-84D4-52E7-83D1-9EBD4C5E1414" authority="Meade-Waldo, 1914" authorityName="Meade-Waldo" authorityYear="1914" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile (Callomegachile) atratiformis" order="Hymenoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="atratiformis" subGenus="Callomegachile">Megachile (Callomegachile) atratiformis Meade-Waldo, 1914</taxonomicName>
<figureCitation id="48669A090DA6AFAEE44306FE628BC166" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Thai distribution of Megachile (Callomegachile) atratiformis." figureDoi="10.3897/zookeys.997.34935.figure1" httpUri="https://binary.pensoft.net/fig/479630" pageId="0" pageNumber="95">Figs 1</figureCitation>
<figureCitation id="69423B3753339220DABF45D72943B9E6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megachile (Callomegachile) atratiformis Meade-Waldo, 1914, female a dorsal view b lateral view c frontal view d frontal view of mandible and labrum e dorsal view of mesoscutum f dorsal view of metasoma g lateral view of metasoma and scopa." figureDoi="10.3897/zookeys.997.34935.figure2" httpUri="https://binary.pensoft.net/fig/479631" pageId="0" pageNumber="95">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F306DB341A5346B57804703033C484C5" pageId="0" pageNumber="95" type="reference_group">
<paragraph id="CAA357638B1049A1CAAFB97E6E776592" pageId="0" pageNumber="95">
<taxonomicName id="299D6595402489714D4FC2CE747E633F" authorityName="Meade-Waldo" authorityYear="1914" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile atratiformis" order="Hymenoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="atratiformis">Megachile atratiformis</taxonomicName>
Meade-Waldo, 1914: 456; Female syntype (NHMUK, examined) Mergui (Low Tenasserim), Myanmar.
</paragraph>
<paragraph id="3A5A9457685EAB58B2E730E3B83036D5" pageId="0" pageNumber="95">
<taxonomicName id="C89609E7A63D226FB5A0CAFBF32B0D75" class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma (Alocanthedon) atratiforme" order="Hymenoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="atratiforme" subGenus="Alocanthedon">Chalicodoma (Alocanthedon) atratiforme</taxonomicName>
Engel &amp; Gonzalez, 2011: 68-70.
</paragraph>
</subSubSection>
<subSubSection id="4CEED10534ED6F1D464AA5727743D604" pageId="0" pageNumber="95" type="diagnosis">
<paragraph id="792143DE882B2232E1B40CA7E8C337F0" pageId="0" pageNumber="95">Diagnosis.</paragraph>
<paragraph id="C7EEE497075485F53999995A1C9E03D5" pageId="0" pageNumber="95">
Female can be easily recognized by its large body size (18-22 mm); black body covered with black hairs throughout (Fig.
<figureCitation id="5950F077FD9D39DB83F5CE9C0A602915" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megachile (Callomegachile) atratiformis Meade-Waldo, 1914, female a dorsal view b lateral view c frontal view d frontal view of mandible and labrum e dorsal view of mesoscutum f dorsal view of metasoma g lateral view of metasoma and scopa." figureDoi="10.3897/zookeys.997.34935.figure2" httpUri="https://binary.pensoft.net/fig/479631" pageId="0" pageNumber="95">2a</figureCitation>
); clypeus without median tubercle; mandibles four teeth; apical margin of labrum truncate without tooth (Fig.
<figureCitation id="1FCE3793BC67C348EE18F97D6FC1CB44" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megachile (Callomegachile) atratiformis Meade-Waldo, 1914, female a dorsal view b lateral view c frontal view d frontal view of mandible and labrum e dorsal view of mesoscutum f dorsal view of metasoma g lateral view of metasoma and scopa." figureDoi="10.3897/zookeys.997.34935.figure2" httpUri="https://binary.pensoft.net/fig/479631" pageId="0" pageNumber="95">2d</figureCitation>
); mesoscutum with weak transverse wrinkle pattern on disc, posteriorly also with weakly transverse wrinkle pattern (Fig.
<figureCitation id="6330602ACC497FE31F664C0497A0564E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megachile (Callomegachile) atratiformis Meade-Waldo, 1914, female a dorsal view b lateral view c frontal view d frontal view of mandible and labrum e dorsal view of mesoscutum f dorsal view of metasoma g lateral view of metasoma and scopa." figureDoi="10.3897/zookeys.997.34935.figure2" httpUri="https://binary.pensoft.net/fig/479631" pageId="0" pageNumber="95">2e</figureCitation>
); yellow wings; black scopa. Male can be easily recognized by the presence of black setae patch on medial cell of forewings; apical of T6 with shallow concavity; front tibia modified (
<bibRefCitation id="7CD6FB06CFF05C08F21EFE174A541E1A" DOI="https://doi.org/10.3897/zookeys.101.1182" author="Engel, MS" journalOrPublisher="ZooKeys" pageId="0" pageNumber="95" pagination="51 - 80" refId="B20" refString="Engel, MS, Gonzalez, VH, 2011. Alocanthedon, a new subgenus of Chalicodoma from Southeast Asia (Hymenoptera, Megachilidae). ZooKeys 101: 51 - 80, DOI: https://doi.org/10.3897/zookeys.101.1182" title="Alocanthedon, a new subgenus of Chalicodoma from Southeast Asia (Hymenoptera, Megachilidae)." url="https://doi.org/10.3897/zookeys.101.1182" volume="101" year="2011">Engel and Gonzalez 2011</bibRefCitation>
).
</paragraph>
<caption id="2784A9655E48DDF87CC9437C90B49BE4" doi="10.3897/zookeys.997.34935.figure1" httpUri="https://binary.pensoft.net/fig/479630" pageId="0" pageNumber="95" start="Figure 1" startId="F1">
<paragraph id="07B981E8F17D499AE3DBCCA86071AA27" pageId="0" pageNumber="95">
<emphasis id="DB5887AE283CEFE011C29A51D8C474C9" bold="true" pageId="0" pageNumber="95">Figure 1.</emphasis>
Thai distribution of
<taxonomicName id="8A08F226DF228869D2E1924A4FDE5F70" authorityName="Meade-Waldo" authorityYear="1914" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile (Callomegachile) atratiformis" order="Hymenoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="atratiformis" subGenus="Callomegachile">Megachile (Callomegachile) atratiformis</taxonomicName>
.
</paragraph>
</caption>
<caption id="0E03FF7991131223687D2D005B028A40" doi="10.3897/zookeys.997.34935.figure2" httpUri="https://binary.pensoft.net/fig/479631" pageId="0" pageNumber="95" start="Figure 2" startId="F2">
<paragraph id="9A3E9C52489E62F2280BF40D4A17CEC5" pageId="0" pageNumber="95">
<emphasis id="990ACC73DEAB718A18EDD157C9673AD9" bold="true" pageId="0" pageNumber="95">Figure 2.</emphasis>
<taxonomicName id="FCEB1B39BE833B5E71185E442697898E" authorityName="Meade-Waldo" authorityYear="1914" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile (Callomegachile) atratiformis" order="Hymenoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="atratiformis" subGenus="Callomegachile">Megachile (Callomegachile) atratiformis</taxonomicName>
Meade-Waldo, 1914, female
<emphasis id="2245D88F84725F453F123B3A57673902" bold="true" pageId="0" pageNumber="95">a</emphasis>
dorsal view
<emphasis id="49D204334B5D6CE7E76725435EF5C955" bold="true" pageId="0" pageNumber="95">b</emphasis>
lateral view
<emphasis id="A6891E0857788B4A0FE7ACFD4285EE84" bold="true" pageId="0" pageNumber="95">c</emphasis>
frontal view
<emphasis id="5ECD55110C2E6D5CD14F82DA2C8836C5" bold="true" pageId="0" pageNumber="95">d</emphasis>
frontal view of mandible and labrum
<emphasis id="2A47745F873D818577AE0CA5F9D2A66F" bold="true" pageId="0" pageNumber="95">e</emphasis>
dorsal view of mesoscutum
<emphasis id="EE2E90CDFE1B0A277C48D41FE421F82D" bold="true" pageId="0" pageNumber="95">f</emphasis>
dorsal view of metasoma
<emphasis id="FDCE7AB0DD9666BFE836E439BB7DD3D1" bold="true" pageId="0" pageNumber="95">g</emphasis>
lateral view of metasoma and scopa.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="2BA4B1D460ABD13D77C3265F87EC978A" pageId="0" pageNumber="95" type="literature records">
<paragraph id="48AB526146BAE57D1BC7CD7D545A4588" pageId="0" pageNumber="95">Literature records.</paragraph>
<paragraph id="08BAF1646C7062FD236420BD729FA641" pageId="0" pageNumber="95">
Malaysia. Negeri Sembilan, Pahang (
<bibRefCitation id="83FEA8244E31BB376DFA8424C013B5E1" DOI="https://doi.org/10.1080/00222931408693601" author="Meade-Waldo, G" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="95" pagination="450 - 464" refId="B39" refString="Meade-Waldo, G, 1914. Note on the Hymenoptera in the collection of the British Museum, with descriptions of new species. Annals and Magazine of Natural History 14: 450 - 464, DOI: https://doi.org/10.1080/00222931408693601" title="Note on the Hymenoptera in the collection of the British Museum, with descriptions of new species." url="https://doi.org/10.1080/00222931408693601" volume="14" year="1914">Meade-Waldo 1914</bibRefCitation>
; Gonzalez and Engel 2011;
<bibRefCitation id="B6F99D8CE5CA293C823C7189511C01AE" author="Ascher, JS" journalOrPublisher="Raffles Bulletin of Zoology Supplement" pageId="0" pageNumber="95" publicationUrl="www.discoverlife.org/mp/20q?guide=Apoidea_species" refId="B3" refString="Ascher, JS, Pickering, J, 2020. Bee Species Guide (Hymenoptera: Apoidea: Anthophila). http:// www.discoverlife.org/mp/20q?guide=Apoidea_species" title="Bee Species Guide (Hymenoptera: Apoidea: Anthophila). http: //" url="www.discoverlife.org/mp/20q?guide=Apoidea_species" year="2020">Ascher and Pickering 2020</bibRefCitation>
); Myanmar. Mergui Archipelago in Tanintharyi Region (
<bibRefCitation id="739F3534E8AF41754C9C308D718AB04E" DOI="https://doi.org/10.1080/00222931408693601" author="Meade-Waldo, G" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="95" pagination="450 - 464" refId="B39" refString="Meade-Waldo, G, 1914. Note on the Hymenoptera in the collection of the British Museum, with descriptions of new species. Annals and Magazine of Natural History 14: 450 - 464, DOI: https://doi.org/10.1080/00222931408693601" title="Note on the Hymenoptera in the collection of the British Museum, with descriptions of new species." url="https://doi.org/10.1080/00222931408693601" volume="14" year="1914">Meade-Waldo 1914</bibRefCitation>
; Gonzalez and Engel 2011;
<bibRefCitation id="2F57ABEE272022AF949C809A16D58E4D" author="Ascher, JS" journalOrPublisher="Raffles Bulletin of Zoology Supplement" pageId="0" pageNumber="95" publicationUrl="www.discoverlife.org/mp/20q?guide=Apoidea_species" refId="B3" refString="Ascher, JS, Pickering, J, 2020. Bee Species Guide (Hymenoptera: Apoidea: Anthophila). http:// www.discoverlife.org/mp/20q?guide=Apoidea_species" title="Bee Species Guide (Hymenoptera: Apoidea: Anthophila). http: //" url="www.discoverlife.org/mp/20q?guide=Apoidea_species" year="2020">Ascher and Pickering 2020</bibRefCitation>
); Thailand. Uthai Thani (
<bibRefCitation id="FB1F51D8AFE5B3CBEDA613FDB86F9E47" DOI="https://doi.org/10.3897/zookeys.101.1182" author="Engel, MS" journalOrPublisher="ZooKeys" pageId="0" pageNumber="95" pagination="51 - 80" refId="B20" refString="Engel, MS, Gonzalez, VH, 2011. Alocanthedon, a new subgenus of Chalicodoma from Southeast Asia (Hymenoptera, Megachilidae). ZooKeys 101: 51 - 80, DOI: https://doi.org/10.3897/zookeys.101.1182" title="Alocanthedon, a new subgenus of Chalicodoma from Southeast Asia (Hymenoptera, Megachilidae)." url="https://doi.org/10.3897/zookeys.101.1182" volume="101" year="2011">Engel and Gonzalez 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BC00C248678AA3BF0576AFB001D55A91" pageId="0" pageNumber="95" type="materials_examined">
<paragraph id="6B97722FF54533BFB74DFF64F9080C52" pageId="0" pageNumber="95">Material examined.</paragraph>
<paragraph id="C16EADB942F02DBC3859B1D9038B5A57" pageId="0" pageNumber="95">
Female syntype. Myanmar: Tannintharyi Region. &quot;Type H. T.; B.M. TYPE HYM. 2037. 17. a.;
<taxonomicName id="43DE81D7C89FBD7804F5C2D3233D2C01" authorityName="Smith" authorityYear="1853" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile atrata" order="Hymenoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="274EA63D50BFCB8E07253DA1AB327826" italics="true" pageId="0" pageNumber="95">Megachile atrata</emphasis>
</taxonomicName>
(Var), Smith, female; Lower Tenasserim. Mergui, 11-89. C. T. Bingham.; Col. C. T. Bingham 96-30;
<taxonomicName id="B61BB41C2E2B483470DD6DFDFA0452F3" authorityName="Meade-Waldo" authorityYear="1914" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile (Eumegachile) atratiformis" order="Hymenoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="atratiformis" subGenus="Eumegachile">Megachile (Eumegachile) atratiformis</taxonomicName>
M.W., G. Meade-Waldo det., Type, female; NHMUK 013379843&quot;; Thailand. Nong Khai Province: 1♀, Phon Phisai district, Cowboy Coffee,
<geoCoordinate id="72822607E595FA76E6E3D69F81D30256" degrees="18" direction="north" minutes="6" orientation="latitude" precision="1" seconds="9.95" value="18.102764">18°6'9.95&quot;N</geoCoordinate>
,
<geoCoordinate id="EF50E8A0F33BCC6E56FC6E51133FBA29" degrees="103" direction="east" minutes="6" orientation="longitude" precision="1" seconds="17.96" value="103.10499">103°6'17.96&quot;E</geoCoordinate>
, Alt. 147.65 m, 15-I-2017, coll. N. Warrit et al. (leg. NC and NW). Ubon Ratchathani Province: 1♀, Phu Jong Na Yoi Nat. P., Kaeng Ka Lao,
<geoCoordinate id="2A14D9D33B70C0950FB66248EF8BEDDD" degrees="14" direction="north" minutes="26" orientation="latitude" precision="1" seconds="10.98" value="14.436383">14°26'10.98&quot;N</geoCoordinate>
,
<geoCoordinate id="65020F6C0F3B47803AF285B0CE93DFF5" degrees="105" direction="east" minutes="16" orientation="longitude" precision="1" seconds="37.05" value="105.27696">105°16'37.05&quot;E</geoCoordinate>
, Alt. 322 m, 5-I-2019, coll. N. Warrit et al (leg. NC and NW).
</paragraph>
</subSubSection>
<subSubSection id="F181509BD08C302AA4E78D9F07C4B76C" pageId="0" pageNumber="95" type="floral records">
<paragraph id="70D55B222DD8A18DF9EA64679A3D2DA3" pageId="0" pageNumber="95">Floral records.</paragraph>
<paragraph id="201EBFF59BB9AACCB3B68F933FC0CE64" pageId="0" pageNumber="95">
<bibRefCitation id="D121F3B03D1D58E69D52231CF874A3F1" DOI="https://doi.org/10.3897/zookeys.101.1182" author="Engel, MS" journalOrPublisher="ZooKeys" pageId="0" pageNumber="95" pagination="51 - 80" refId="B20" refString="Engel, MS, Gonzalez, VH, 2011. Alocanthedon, a new subgenus of Chalicodoma from Southeast Asia (Hymenoptera, Megachilidae). ZooKeys 101: 51 - 80, DOI: https://doi.org/10.3897/zookeys.101.1182" title="Alocanthedon, a new subgenus of Chalicodoma from Southeast Asia (Hymenoptera, Megachilidae)." url="https://doi.org/10.3897/zookeys.101.1182" volume="101" year="2011">Engel and Gonzalez (2011)</bibRefCitation>
noted the species was captured on an indigenous tree,
<taxonomicName id="2033601A9A334A1157C135BD166F6A68" class="Magnoliopsida" family="Dipterocarpaceae" genus="Dipterocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dipterocarpus obtusifolius" order="Malvales" pageId="0" pageNumber="95" phylum="Tracheophyta" rank="species" species="obtusifolius">
<emphasis id="1AD18FD7A30C476E7172E5B28EE70773" italics="true" pageId="0" pageNumber="95">Dipterocarpus obtusifolius</emphasis>
</taxonomicName>
Teijsman &amp; Miquel.
</paragraph>
</subSubSection>
<subSubSection id="BEA2A8833CFA3EE59B447001B52D0013" pageId="0" pageNumber="95" type="comments">
<paragraph id="03AE194ECDD7F74EE1781EF880CF2B5D" pageId="0" pageNumber="95">Comments.</paragraph>
<paragraph id="44DCFB5A86FCD881D6B88D5B62BA82AD" pageId="0" pageNumber="95">
<normalizedToken id="823D9C37CCD49900D6A4E3971204AACD" originalValue="Meade-Waldo’s">Meade-Waldo's</normalizedToken>
original material was composite (
<bibRefCitation id="E759E13E9B6BD2C37F2366D7FB8DE24C" DOI="https://doi.org/10.3897/zookeys.101.1182" author="Engel, MS" journalOrPublisher="ZooKeys" pageId="0" pageNumber="95" pagination="51 - 80" refId="B20" refString="Engel, MS, Gonzalez, VH, 2011. Alocanthedon, a new subgenus of Chalicodoma from Southeast Asia (Hymenoptera, Megachilidae). ZooKeys 101: 51 - 80, DOI: https://doi.org/10.3897/zookeys.101.1182" title="Alocanthedon, a new subgenus of Chalicodoma from Southeast Asia (Hymenoptera, Megachilidae)." url="https://doi.org/10.3897/zookeys.101.1182" volume="101" year="2011">Engel and Gonzalez 2011</bibRefCitation>
) so we exclude from the distribution records from Middle Tenasserim: Haundraw [also known as Houngdarau] Valley in Myanmar (=
<taxonomicName id="6B0AE586E685E83369DD3551C0799FEC" lsidName="M. odontophorum" pageId="0" pageNumber="95" rank="species" species="odontophorum">
<emphasis id="980E5886416838ECFFBE8094768F52AF" italics="true" pageId="0" pageNumber="95">M. odontophorum</emphasis>
</taxonomicName>
) and also from Penang in Malaysia.
</paragraph>
</subSubSection>
</treatment>
</document>
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