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<document id="C568DC5E097BFBCC0530AED4EB2F28FF" ID-CLB-Dataset="301504" ID-DOI="10.1093/zooldnnean/zlad146" ID-GBIF-Dataset="5e66aeaf-44c5-4be9-924d-cc13680ace6a" ID-ISSN="0024-4082" ID-Zenodo-Dep="13315269" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722845279467" checkinUser="plazi" docAuthor="Lumen, Ryan &amp; Kamdńskd, Marcdn J." docDate="2024" docId="C27387A7FFE0FFCF9A050201FB0A7162" docLanguage="en" docName="zlad146.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (201)" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Biolus asperipennis" docType="treatment" docVersion="7" lastPageNumber="376" masterDocId="3E4AFFDFFFF2FFDD9A72053DFF83770F" masterDocTitle="evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae" masterLastPageNumber="386" masterPageNumber="358" pageNumber="376" updateTime="1732206558056" updateUser="felipe">
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<mods:title id="86A21361C7CE843EB66C796E718244B5">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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<mods:namePart id="14E70C9106E646448BE0F4D328ECBF7F">Lumen, Ryan</mods:namePart>
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<mods:namePart id="215B95E4DB252A1D64107A128CB504B9">Kamdńskd, Marcdn J.</mods:namePart>
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<mods:title id="B7A96C3C052A8043B305867A7D7EBCED">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="39AFD428D887050514927BD8C3244C53">2024</mods:date>
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<mods:identifier id="2CE6BAC644652B6B20E34B9BC033B5BC" type="DOI">10.1093/zooldnnean/zlad146</mods:identifier>
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<treatment id="C27387A7FFE0FFCF9A050201FB0A7162" ID-DOI="http://doi.org/10.5281/zenodo.13315095" ID-GBIF-Taxon="238560668" ID-Zenodo-Dep="13315095" LSID="urn:lsid:plazi:treatment:C27387A7FFE0FFCF9A050201FB0A7162" httpUri="http://treatment.plazi.org/id/C27387A7FFE0FFCF9A050201FB0A7162" lastPageNumber="376" pageId="18" pageNumber="376">
<subSubSection id="02C0653AFFE0FFCF9A050201FD767058" box="[119,757,1852,1879]" pageId="18" pageNumber="376" type="nomenclature">
<paragraph id="4A6536B1FFE0FFCF9A050201FD767058" blockId="18.[113,762,1852,2009]" box="[119,757,1852,1879]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9A050201FD767058" box="[119,757,1852,1879]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="4">
<emphasis id="78AEEAA3FFE0FFCF9A050201FD767058" bold="true" box="[119,757,1852,1879]" pageId="18" pageNumber="376">
2.
<taxonomicName id="8DDA4D32FFE0FFCF9AFC0201FDAB7059" authority="(Mulsant and Rey)" baseAuthorityName="Mulsant and Rey" box="[142,552,1852,1879]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9AFC0201FEE17059" bold="true" box="[142,354,1852,1879]" italics="true" pageId="18" pageNumber="376">Biolus asperipennis</emphasis>
(Mulsant and Rey)
</taxonomicName>
stat. res.,comb. nov.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE0FFCF9B290246FB0A7162" pageId="18" pageNumber="376" type="description">
<paragraph id="4A6536B1FFE0FFCF9B290246FD92709A" blockId="18.[113,762,1852,2009]" box="[347,529,1915,1941]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9B290246FD92709A" box="[347,529,1915,1941]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="3">
<emphasis id="78AEEAA3FFE0FFCF9B290246FD92709A" bold="true" box="[347,529,1915,1941]" pageId="18" pageNumber="376">
(
<figureCitation id="D2E12A34FFE0FFCF9B140246FE30709A" box="[358,435,1915,1941]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.species.A, Biolus analisetosus (Koch) comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey) stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey) comb. nov. Lectotype; E, B. graafi (Koch) comb. nov. Holotype; F,B. granulatus (Fabricius) comb. nov. Lectotype; G, B. omeri (Koch) comb.nov.Holotype; H, B. sauroides (Koch) comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." figureDoi="http://doi.org/10.5281/zenodo.13315277" httpUri="https://zenodo.org/record/13315277/files/figure.png" pageId="18" pageNumber="376">Figs 2B</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9BCF0246FE5C709A" box="[445,479,1915,1941]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used in the cladistic analysis.Arrows indicate particular states of given characters from the morphological matrix [e.g." figureDoi="http://doi.org/10.5281/zenodo.13315289" httpUri="https://zenodo.org/record/13315289/files/figure.png" pageId="18" pageNumber="376">6G</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9B9B0246FD85709A" box="[489,518,1915,1941]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotina species.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres highlighted red. Scale = 1mm." figureDoi="http://doi.org/10.5281/zenodo.13315291" httpUri="https://zenodo.org/record/13315291/files/figure.png" pageId="18" pageNumber="376">8B</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A03029FFD7970D6" blockId="18.[113,762,1852,2009]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A03029FFED670B6" box="[113,341,1954,1977]" italics="true" pageId="18" pageNumber="376">Eurynotus asperipennis</emphasis>
Mulsant and Rey, 1854a: 23.– Mulsant and Rey 1854b: 169; Lacordaire 1859: 239; Gebien 1938: 414;
</paragraph>
<paragraph id="4A6536B1FFE0FFCF995805ADFC1B77C8" blockId="18.[810,1459,144,199]" pageId="18" pageNumber="376">
Koch 1954a: 280; 1954b: 2;
<bibRefCitation id="2E4B4B40FFE0FFCF9E2205ADFAC477A7" author="Banaszkdewdcz M" box="[1104,1351,144,168]" pageId="18" pageNumber="376" pagination="59 - 77" refId="ref25294" refString="Banaszkdewdcz M. Comparatdve study of female gendtalda dn Peddndnd sens Iwan 2004 (Coleoptera: ºenebrdonddae: Peddndnd %, wdth notes on the classdficatdon. Annales Zoologici 2006; 56: 59 - 77." type="journal article" year="2006">Banaszkiewicz 2006: 63</bibRefCitation>
; Kamiński 2016: 235.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF995805D4FB0E7577" blockId="18.[810,1460,233,632]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF995805D4FC13760F" box="[810,912,233,256]" italics="true" pageId="18" pageNumber="376">Diagnosis:</emphasis>
Lacks sclerite in bursa copulatrix (present in
<taxonomicName id="8DDA4D32FFE0FFCF9FED05D4FC1E762F" authorityName="Koch" authorityYear="1954" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF9FED05D4FC1E762F" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE0FFCF99920434FBE9762F" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[992,1130,264,288]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF99920434FBE9762F" box="[992,1130,264,288]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
).
<taxonomicName id="8DDA4D32FFE0FFCF9EF80435FACF762F" box="[1162,1356,264,288]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9EF80435FACF762F" box="[1162,1356,264,288]" italics="true" pageId="18" pageNumber="376">Biolus asperipennis</emphasis>
</taxonomicName>
has two puncture
<typeStatus id="95618813FFE0FFCF99E30415FC4B764F" box="[913,968,296,320]" pageId="18" pageNumber="376">types</typeStatus>
on ventrite V (single
<typeStatus id="95618813FFE0FFCF9EC80415FB6B764F" box="[1210,1256,296,320]" pageId="18" pageNumber="376">type</typeStatus>
in
<taxonomicName id="8DDA4D32FFE0FFCF9F7D0415FAE77630" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[1295,1380,296,319]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9F7D0415FAE77630" box="[1295,1380,296,319]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE0FFCF9FED0415FC127650" baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9FED0415FC127650" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
), and golden setae on the tarsi (dark/black in
<taxonomicName id="8DDA4D32FFE0FFCF9F2F047AFA2E7651" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[1373,1453,327,350]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9F2F047AFA2E7651" box="[1373,1453,327,350]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9958045AFC1C7671" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[810,927,358,382]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE0FFCF9958045AFC1C7671" box="[810,927,358,382]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
</taxonomicName>
, and
<taxonomicName id="8DDA4D32FFE0FFCF99A5045AFBB07671" box="[983,1075,358,382]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF99A5045AFBB07671" box="[983,1075,358,382]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
). Without close inspection,
<taxonomicName id="8DDA4D32FFE0FFCF9F2A045AFA307671" box="[1368,1459,358,382]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9F2A045AFA307671" box="[1368,1459,358,382]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
(length
<quantity id="8D229B54FFE0FFCF990C04B8FC707692" box="[894,1011,389,413]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="18" pageNumber="376" unit="mm" value="16.5" valueMax="18.0" valueMin="15.0">15–18 mm</quantity>
; width
<quantity id="8D229B54FFE0FFCF9E3004B8FB307692" box="[1090,1203,389,413]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="10.0" metricValueMin="7.5" pageId="18" pageNumber="376" unit="mm" value="8.75" valueMax="10.0" valueMin="7.5">7.5–10 mm</quantity>
) specimens may appear similar to smaller specimens of
<taxonomicName id="8DDA4D32FFE0FFCF9E0F0498FA9176B3" box="[1149,1298,421,444]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9E0F0498FA9176B3" box="[1149,1298,421,444]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
(length
<quantity id="8D229B54FFE0FFCF9F030498FCD576D4" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="18" pageNumber="376" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">19–22 mm</quantity>
; width
<quantity id="8D229B54FFE0FFCF99ED04F9FC7D76D3" box="[927,1022,452,476]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="18" pageNumber="376" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
); however,
<taxonomicName id="8DDA4D32FFE0FFCF9E0504F8FA8476D3" box="[1143,1287,453,476]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9E0504F8FA8476D3" box="[1143,1287,453,476]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
has golden tarsal setae (black or dark in
<taxonomicName id="8DDA4D32FFE0FFCF9E7D04D9FBEA76F4" box="[1039,1129,483,507]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9E7D04D9FBEA76F4" box="[1039,1129,483,507]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
), a weaker circumocular depres - sion, the pronotal process is more pronounced, and the male tibiae are also at most only slightly modified in
<taxonomicName id="8DDA4D32FFE0FFCF9E8D071EFADA7535" box="[1279,1369,546,570]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9E8D071EFADA7535" box="[1279,1369,546,570]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
, while in
<taxonomicName id="8DDA4D32FFE0FFCF9958077FFC387556" box="[810,955,578,601]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9958077FFC387556" box="[810,955,578,601]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
they are more strongly curved. See the diagnosis of
<taxonomicName id="8DDA4D32FFE0FFCF9936075CFC1D7577" box="[836,926,608,632]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9936075CFC1D7577" box="[836,926,608,632]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
below for more details.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF995807A7FC6D720A" blockId="18.[810,1460,666,1285]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF995807A7FC3E75BD" box="[810,957,666,690]" italics="true" pageId="18" pageNumber="376">Redescription:</emphasis>
Length
<quantity id="8D229B54FFE0FFCF9E5B07A7FB1A75BD" box="[1065,1177,666,690]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="18" pageNumber="376" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">19–22mm</quantity>
, width
<quantity id="8D229B54FFE0FFCF9E8907A7FADC75BD" box="[1275,1375,666,690]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="18" pageNumber="376" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11mm</quantity>
. Head: Punctate; punctures closely spaced, separated by ≤1 punc - ture diameter. Mentum without apical notch. Eye with weakly impressed sulcus situated around perimeter of dorsal lobe. Prothorax: Pronotum punctate, punctures separated by ≥1 puncture diameter. Hypomeron lightly sculptured/ wrinkled with small punctation and microtuberculation. Pterothorax: Elytra not costate, without macrotubercles on disc. Elytral intervals with or without microtuberculation. Epipleuron with many small, faint microtubercles. Abdomen: Finely punctate, punctures separated by&gt;1 puncture diam - eter. Abdominal ventrite V punctures separated by ≥ 1 punc - ture diameter with larger, setigerous punctures intermixed, females with median, apical notch. Legs: Male hind tibiae slightly flattened/twisted, appearing bent. Female legs un - modified. Tarsi with golden setae. Males: pro/meso tarsi robust with large, ventral, tomentose pads. Male terminalia: Parameres ‘bottle -shaped’, strongly tapered and truncate ap - ically. Female terminalia: Bursa copulatrix without accessory pouch or sclerites.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9958001BFA267272" blockId="18.[810,1460,1318,1436]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9958001BFC767231" box="[810,1013,1318,1342]" italics="true" pageId="18" pageNumber="376">Material examined:</emphasis>
<typeStatus id="95618813FFE0FFCF9E75001AFBF27230" box="[1031,1137,1319,1343]" pageId="18" pageNumber="376" type="lectotype">Lectotype</typeStatus>
, here designated (MNHN): ‘
<taxonomicName id="8DDA4D32FFE0FFCF995C007BFC1F7251" box="[814,924,1350,1374]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
? sp. cap.?; blue circle label ‘
<taxonomicName id="8DDA4D32FFE0FFCF9ECF007BFAA87251" box="[1213,1323,1350,1374]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
asperipennis Muls cap. b. sp.?’ also with a red label with handwriting: ‘Type’.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF993700B9FADD7293" blockId="18.[810,1460,1318,1436]" box="[837,1374,1412,1436]" pageId="18" pageNumber="376">(See Appendix S1 for additional material examined.)</paragraph>
<paragraph id="4A6536B1FFE0FFCF99580083FA8D713B" blockId="18.[810,1459,1470,1588]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF99580083FCE272DA" box="[810,865,1470,1493]" italics="true" pageId="18" pageNumber="376">Note:</emphasis>
As the original description lacks information defining the number or specificity of specimens used, one from the presumed
<typeStatus id="95618813FFE0FFCF995800C0FCF8711A" box="[810,891,1533,1557]" pageId="18" pageNumber="376" type="syntype">syntype</typeStatus>
series is here designated as the
<typeStatus id="95618813FFE0FFCF9EC900C1FA98711B" box="[1211,1307,1532,1556]" pageId="18" pageNumber="376" type="lectotype">lectotype</typeStatus>
to fix the taxo - nomic status of the species and genus (Fig. 2B).
</paragraph>
<paragraph id="4A6536B1FFE0FFCF99580368FB0A7162" blockId="18.[810,1161,1621,1645]" box="[810,1161,1621,1645]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF99580368FC2B7162" box="[810,936,1621,1645]" italics="true" pageId="18" pageNumber="376">Distribution:</emphasis>
South Africa (Fig. 9).
</paragraph>
</subSubSection>
</treatment>
</document>
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