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<document id="F954A9564F71D1096F994E7A968840EB" ID-DOI="10.11646/zootaxa.4816.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3954685" ID-ZooBank="5154FD08-0AED-4826-A275-37AA9FBF4874" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595340302449" checkinUser="plazi" docAuthor="Akhil, S. V., Divya, M. &amp; Sabu, K. Thomas" docDate="2020" docId="6B0687C5A660FFF1ADD3FEAEE3779CB3" docLanguage="en" docName="zootaxa.4816.4.7.pdf" docOrigin="Zootaxa 4816 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Brachinus (Metabrachinus) hexagrammus Chaudoir 1876" docType="treatment" docVersion="5" lastPageNumber="583" masterDocId="973FFFBDA667FFF6AD44FFAAE6649856" masterDocTitle="Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India" masterLastPageNumber="600" masterPageNumber="576" pageNumber="583" updateTime="1698783441694" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="E97110988728A96D24534E004B30AA2F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D4CD342ADB9FEB4426D69A075865E571">
<mods:title id="1E6110EE64CB5D603A06AD6964106E24">Bombardier beetles of genus Brachinus Weber, 1801 (Carabidae: Brachininae Brachinini) from India</mods:title>
</mods:titleInfo>
<mods:name id="83652A15B8B28B073F2A4F9C75C063A7" type="personal">
<mods:role id="A2453590B869A1E71884750087AB85E5">
<mods:roleTerm id="9B67C7223BFE0ABF84138CDB20157DFA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E9B32729FA11ED17942BE92230610783">Akhil, S. V.</mods:namePart>
<mods:affiliation id="7A57526432E0B8ABA8E580881657DD43">Entomology Research Lab, P. G. &amp; Research Department of Zoology, St. Joseph’s College (Autonomous), Devagiri, Calicut, Kerala, 673008, India</mods:affiliation>
</mods:name>
<mods:name id="DC51B85BFA14C16C72952786E3692227" type="personal">
<mods:role id="1997AE2775A6F53A92D18150FCFF5DEB">
<mods:roleTerm id="6D05B3B90B85B4B5E6B855E2C0190AE4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="915C99EC42F1E2CC6272AE9E11CA20A8">Divya, M.</mods:namePart>
<mods:affiliation id="D6F54FFA11AE50E8AB2B1725AD5FFDA7">Entomology Research Lab, P. G. &amp; Research Department of Zoology, St. Joseph’s College (Autonomous), Devagiri, Calicut, Kerala, 673008, India &amp; divyamanikandan 123 @ gmail. com; https: // orcid. org / 0000 - 0003 - 4594 - 5991</mods:affiliation>
</mods:name>
<mods:name id="86FC9F1E687CC7D549FAA46C90308FE3" type="personal">
<mods:role id="5C58637A157A0FD66775F0281A71A356">
<mods:roleTerm id="0388E1937E63846A7CE90B1C5B89CAC6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A423AFF647D0F2CB4D7184E787AF6E05">Sabu, K. Thomas</mods:namePart>
<mods:affiliation id="6FC44CCFB9DBBC28C6620C51C3EC26E6">Entomology Research Lab, P. G. &amp; Research Department of Zoology, St. Joseph’s College (Autonomous), Devagiri, Calicut, Kerala, 673008, India &amp; sabukthomas @ gmail. com; https: // orcid. org / 0000 - 0002 - 8701 - 0484</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1386DF9695E8770C1BCCD7414CB6BC5B">text</mods:typeOfResource>
<mods:relatedItem id="B26AD70A1C5D958D930AF5D17ABEB996" type="host">
<mods:titleInfo id="C03A0C30348FE835F290F03750AFD512">
<mods:title id="ADC7D2FA04422FA77B8493690BF619E0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D8A5E9D9C75F5187A6C69CB6A9CC0BA4">
<mods:date id="99A8896263DC2F2D9C7908D07119CC1A">2020</mods:date>
<mods:detail id="9ED8EDB408ED8F0F822D26DB58F89F12" type="pubDate">
<mods:number id="F69F936523F80401493B349992DF082A">2020-07-20</mods:number>
</mods:detail>
<mods:detail id="53590A0518EB3FB59C0134C496BF23D7" type="volume">
<mods:number id="1798AE9A76DFC80DE5F7E51ECE9BCB63">4816</mods:number>
</mods:detail>
<mods:detail id="ACC0D88DD53C993237DE35E0666A89FD" type="issue">
<mods:number id="7EC403EE20BDD888F67E6876D44A670C">4</mods:number>
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<mods:extent id="B9334C21E2FF5EE3D7AD07C6644A1051" unit="page">
<mods:start id="8AD91A37FA8D8AA1AE9E406A725DFC8E">576</mods:start>
<mods:end id="46A842914C8B7C88EDABBB8E043BB5C3">600</mods:end>
</mods:extent>
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<mods:classification id="31A4EDA1346C9C8B911603732437983B">journal article</mods:classification>
<mods:identifier id="2F098FCAD96105DF5B84A4AFB55DF9E5" type="DOI">10.11646/zootaxa.4816.4.7</mods:identifier>
<mods:identifier id="C4F0E4CF1C6443EFEA17B3B9785BDBC8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="251D5B3D85119C2C2B7F1A5D9FFDFD8D" type="Zenodo-Dep">3954685</mods:identifier>
<mods:identifier id="FEC09CC76EADFABF325D1515A352695E" type="ZooBank">5154FD08-0AED-4826-A275-37AA9FBF4874</mods:identifier>
</mods:mods>
<treatment id="6B0687C5A660FFF1ADD3FEAEE3779CB3" LSID="urn:lsid:plazi:treatment:6B0687C5A660FFF1ADD3FEAEE3779CB3" httpUri="http://treatment.plazi.org/id/6B0687C5A660FFF1ADD3FEAEE3779CB3" lastPageNumber="583" pageId="7" pageNumber="583">
<subSubSection id="ABB56558A660FFF1ADD3FEAEE5009948" box="[151,868,260,287]" pageId="7" pageNumber="583" type="nomenclature">
<paragraph id="E31036D3A660FFF1ADD3FEAEE5009948" blockId="7.[151,868,260,323]" box="[151,868,260,287]" pageId="7" pageNumber="583">
<heading id="B85881BFA660FFF1ADD3FEAEE5009948" bold="true" box="[151,868,260,287]" fontSize="11" level="1" pageId="7" pageNumber="583" reason="1">
<taxonomicName id="24AF4D50A660FFF1ADD3FEAEE5009948" ID-CoL="MT27" authority="Chaudoir, 1876" authorityName="Chaudoir" authorityYear="1876" box="[151,868,260,287]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="583" phylum="Arthropoda" rank="species" species="hexagrammus" subGenus="Metabrachinus">
<emphasis id="D1DBEAC1A660FFF1ADD3FEAEE5009948" bold="true" box="[151,868,260,287]" pageId="7" pageNumber="583">
<emphasis id="D1DBEAC1A660FFF1ADD3FEAEE4FA9949" bold="true" box="[151,670,260,287]" italics="true" pageId="7" pageNumber="583">Brachinus (Metabrachinus) hexagrammus</emphasis>
<bibRefCitation id="873E4B22A660FFF1AFE1FEAEE5009948" author="Chaudoir, M. de" box="[677,868,260,286]" pageId="7" pageNumber="583" pagination="11 - 104" refId="ref18332" refString="Chaudoir, M. de (1876) Monographie des brachynides. Annales de la Societe Entomologique de Belgique, 19, 11 - 104." type="journal article" year="1876">Chaudoir, 1876</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ABB56558A660FFF1ADD3FE82E6819915" box="[151,229,296,323]" pageId="7" pageNumber="583" type="description">
<paragraph id="E31036D3A660FFF1ADD3FE82E6819915" blockId="7.[151,868,260,323]" box="[151,229,296,323]" pageId="7" pageNumber="583">
<figureCitation id="7B942A56A660FFF1ADD3FE82E6819915" box="[151,229,296,323]" captionStart="FIGURE 1" captionStartId="14.[151,250,1885,1910]" captionTargetBox="[241,1348,192,1860]" captionTargetId="figure@14.[230,1356,181,1860]" captionTargetPageId="14" captionText="FIGURE 1. Dorsal habitus of Indian Brachinus species: a) B. dryas Andrewes, 1936 (holotype, BMNH); b) B. inops Andrewes, 1932 (holotype, BMNH); c) B. hexagrammus Chaudoir, 1876 (syntype, MNHN); d) B. sexmaculatus Dejean, 1825 (syntype, MNHN); e) B. tetracolon Chuadoir, 1876 (syntype, MNHN); f) B. tetragrammus Chaudoir, 1876 (syntype, MNHN); g) B. pictus (Hope, 1833) (ZSIM); h) B. cinctellus Chaudoir, 1876 (syntype, MNHN); i) B. eucosmus Andrewes, 1937 (holotype, BMNH)" figureDoi="http://doi.org/10.5281/zenodo.3954687" httpUri="https://zenodo.org/record/3954687/files/figure.png" pageId="7" pageNumber="583">Fig. 1c</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="ABB56558A660FFF1ADD3FEC7E2EA99D3" box="[151,1166,364,389]" pageId="7" pageNumber="583" type="reference_group">
<paragraph id="E31036D3A660FFF1ADD3FEC7E2EA99D3" blockId="7.[151,1166,364,389]" box="[151,1166,364,389]" pageId="7" pageNumber="583">
<treatmentCitationGroup id="C3BF11FDA660FFF1ADD3FEC7E2EA99D3" box="[151,1166,364,389]" pageId="7" pageNumber="583">
<treatmentCitation id="620E10C2A660FFF1ADD3FEC7E40499D3" author="Chaudoir, M. de" box="[151,608,364,389]" page="55" pageId="7" pageNumber="583" year="1876">
<taxonomicName id="24AF4D50A660FFF1ADD3FEC7E40499D3" ID-CoL="MT27" authority="Chaudoir, 1876: 55" authorityName="Chaudoir" authorityPageNumber="55" authorityYear="1876" box="[151,608,364,389]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="583" phylum="Arthropoda" rank="species" species="hexagrammus">
<emphasis id="D1DBEAC1A660FFF1ADD3FEC7E7F099D3" box="[151,404,365,389]" italics="true" pageId="7" pageNumber="583">Brachinus hexagrammus</emphasis>
<bibRefCitation id="873E4B22A660FFF1ACDEFEC6E40499D3" author="Chaudoir, M. de" box="[410,608,364,389]" pageId="7" pageNumber="583" pagination="11 - 104" refId="ref18332" refString="Chaudoir, M. de (1876) Monographie des brachynides. Annales de la Societe Entomologique de Belgique, 19, 11 - 104." type="journal article" year="1876">Chaudoir, 1876: 55</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="620E10C2A660FFF1AF2DFEC6E52099D3" author="Andrewes, H. E." box="[617,836,364,389]" page="49" pageId="7" pageNumber="583" year="1930">
<bibRefCitation id="873E4B22A660FFF1AF2DFEC6E52099D3" author="Andrewes, H. E." box="[617,836,364,389]" pageId="7" pageNumber="583" refId="ref17586" refString="Andrewes, H. E. (1930 a) Catalogue of Indian Insects. Part 18. Carabidae. Government of India, Central Publication Branch, Calcutta, 389 pp." type="book" year="1930">Andrewes, 1930a: 49</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="620E10C2A660FFF1AE0BFEC6E27999D3" author="Hrdlicka, J." box="[847,1053,364,389]" page="215" pageId="7" pageNumber="583" year="2003">
<bibRefCitation id="873E4B22A660FFF1AE0BFEC6E27999D3" author="Hrdlicka, J." box="[847,1053,364,389]" pageId="7" pageNumber="583" pagination="212 - 218" refId="ref18818" refString="Hrdlicka, J. (2003) Brachininae. In: Lobl I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata- Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 212 - 218." type="book chapter" year="2003">Hrdlička, 2003: 215</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="620E10C2A660FFF1A963FEC7E2EA99D3" author="Hrdlicka, J." box="[1063,1166,364,389]" page="476" pageId="7" pageNumber="583" year="2017">
<bibRefCitation id="873E4B22A660FFF1A963FEC7E2EA99D3" author="Hrdlicka, J." box="[1063,1166,364,389]" pageId="7" pageNumber="583" pagination="471 - 480" refId="ref18865" refString="Hrdlicka, J. (2017) Brachininae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata- Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden, pp. 471 - 480." type="book chapter" year="2017">2017: 476</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="ABB56558A660FFF1ADD3FE19E5319A0B" pageId="7" pageNumber="583" type="materials_examined">
<paragraph id="E31036D3A660FFF1ADD3FE19E4F79998" blockId="7.[151,1437,434,1253]" box="[151,659,434,462]" pageId="7" pageNumber="583">
<materialsCitation id="53C73C8EA660FFF1ADD3FE19E4F79998" box="[151,659,434,462]" collectorName="Dr. Bacon." country="India" location="India" pageId="7" pageNumber="583" specimenCount="1" typeStatus="holotype">
<emphasis id="D1DBEAC1A660FFF1ADD3FE19E752999B" bold="true" box="[151,310,434,461]" pageId="7" pageNumber="583">
<typeStatus id="3C148871A660FFF1ADD3FE19E6B6999B" box="[151,210,435,461]" pageId="7" pageNumber="583">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="9BB87643A660FFF1AC78FE19E71E999B" box="[316,378,435,461]" name="India" pageId="7" pageNumber="583">India</collectingCountry>
: Bengal, coll.
<collectorName id="4E5A5305A660FFF1AF5EFE1EE4F79998" box="[538,659,436,462]" pageId="7" pageNumber="583">Dr. Bacon.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="E31036D3A660FFF1AD83FE7DE79499A4" blockId="7.[151,1437,434,1253]" box="[199,496,470,498]" pageId="7" pageNumber="583">
<materialsCitation id="53C73C8EA660FFF1AD83FE7DE78899A4" box="[199,492,470,498]" collectionCode="MNHN" pageId="7" pageNumber="583" specimenCount="1" typeStatus="holotype">
<emphasis id="D1DBEAC1A660FFF1AD83FE7DE7E899A7" bold="true" box="[199,396,470,497]" pageId="7" pageNumber="583">
<typeStatus id="3C148871A660FFF1AD83FE7DE76699A7" box="[199,258,471,497]" pageId="7" pageNumber="583">Type</typeStatus>
depository.
</emphasis>
<collectionCode id="85BEAE16A660FFF1ACD7FE72E78899A4" box="[403,492,472,498]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="583">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="E31036D3A660FFF1AD83FE50E5319A0B" blockId="7.[151,1437,434,1253]" pageId="7" pageNumber="583">
<emphasis id="D1DBEAC1A660FFF1AD83FE50E7DB9A43" bold="true" box="[199,447,506,533]" pageId="7" pageNumber="583">Specimens examined</emphasis>
(n = 7).
<materialsCitation id="53C73C8EA660FFF1AF64FE51E5359A0B" collectingDate="2015-09-08" collectingDateMax="2017-11-17" collectingDateMin="2015-09-08" collectorName="Ind. Or. Bor. &amp; Dr. Bacon &amp; S. V. Akhil &amp; Thangavel" country="India" county="Ettimadai" determinerName="Chaudoir" location="Anand" municipality="Hand Picking" pageId="7" pageNumber="583" specimenCount="11" specimenCount-female="3" specimenCount-male="2" stateProvince="Tamil Nadu" typeStatus="syntype">
<typeStatus id="3C148871A660FFF1AF64FE51E4189A43" box="[544,636,507,533]" pageId="7" pageNumber="583" type="syntype">Syntype</typeStatus>
labelled “
<collectorName id="4E5A5305A660FFF1AFA8FE51E51B9A43" box="[748,895,507,533]" pageId="7" pageNumber="583">Ind. Or. Bor.</collectorName>
’,coll.
<collectorName id="4E5A5305A660FFF1AE86FE56E2529A40" box="[962,1078,508,534]" pageId="7" pageNumber="583">Dr. Bacon</collectorName>
, det.
<determinerName id="16CCDA17A660FFF1A936FE51E2859A43" box="[1138,1249,507,533]" pageId="7" pageNumber="583">Chaudoir</determinerName>
”;
<specimenCount id="F5A9FD5AA660FFF1A9B1FE50E31A9A43" box="[1269,1406,506,533]" pageId="7" pageNumber="583" type="generic">5 specimens</specimenCount>
(
<specimenCount id="F5A9FD5AA660FFF1A8C9FE51E6BF9A6F" pageId="7" pageNumber="583" type="male">2 males</specimenCount>
,
<specimenCount id="F5A9FD5AA660FFF1ADA2FDB5E7309A6C" box="[230,340,543,570]" pageId="7" pageNumber="583" type="female">3 females</specimenCount>
) labelled “
<collectingCountry id="9BB87643A660FFF1AC88FDB5E4689A6F" box="[460,524,543,569]" name="India" pageId="7" pageNumber="583">India</collectingCountry>
:
<collectingRegion id="216BF831A660FFF1AF52FDB5E4FB9A6F" box="[534,671,543,569]" country="India" name="Tamil Nadu" pageId="7" pageNumber="583">Tamil Nadu</collectingRegion>
:
<collectingCounty id="0A714E5FA660FFF1AFEDFDB5E57D9A6F" box="[681,793,543,569]" pageId="7" pageNumber="583">Ettimadai</collectingCounty>
,
<collectingMunicipality id="0374ACA9A660FFF1AE67FDB5E5DB9A6F" box="[803,959,543,569]" pageId="7" pageNumber="583">Hand Picking</collectingMunicipality>
,
<date id="97111013A660FFF1AE8EFDB4E2289A6F" box="[970,1100,542,569]" pageId="7" pageNumber="583" value="2015-09-08">
<collectingDate id="8755E9FBA660FFF1AE8EFDB4E2289A6F" box="[970,1100,542,569]" pageId="7" pageNumber="583" value="2015-09-08">08.IX.2015</collectingDate>
</date>
, coll.
<collectorName id="4E5A5305A660FFF1A9CBFDB5E36D9A6F" box="[1167,1289,543,569]" pageId="7" pageNumber="583">S.V. Akhil</collectorName>
”;
<specimenCount id="F5A9FD5AA660FFF1A85AFDB5E3F89A6F" box="[1310,1436,543,569]" pageId="7" pageNumber="583" type="generic">1 specimen</specimenCount>
labelled “
<collectingCountry id="9BB87643A660FFF1AC45FDE9E7259A0B" box="[257,321,579,605]" name="India" pageId="7" pageNumber="583">India</collectingCountry>
:
<collectingRegion id="216BF831A660FFF1AC08FDE9E7C19A0B" box="[332,421,579,605]" country="India" name="Gujarat" pageId="7" pageNumber="583">Gujarat</collectingRegion>
:
<location id="E6706008A660FFF1ACE9FDE9E79F9A0B" LSID="urn:lsid:plazi:treatment:6B0687C5A660FFF1ADD3FEAEE3779CB3:E6706008A660FFF1ACE9FDE9E79F9A0B" box="[429,507,579,605]" country="India" county="Ettimadai" municipality="Hand Picking" name="Anand" pageId="7" pageNumber="583" stateProvince="Tamil Nadu">Anand</location>
,
<date id="97111013A660FFF1AF41FDE9E4E39A0B" box="[517,647,579,605]" pageId="7" pageNumber="583" value="2017-11-17">
<collectingDate id="8755E9FBA660FFF1AF41FDE9E4E39A0B" box="[517,647,579,605]" pageId="7" pageNumber="583" value="2017-11-17">17.XI.2017</collectingDate>
</date>
, coll.
<collectorName id="4E5A5305A660FFF1AF8EFDE9E5229A0B" box="[714,838,579,605]" pageId="7" pageNumber="583">Thangavel</collectorName>
”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ABB56558A660FFF1AD83FDCCE7239CCB" pageId="7" pageNumber="583" type="description">
<paragraph id="E31036D3A660FFF1AD83FDCCE4729A9F" blockId="7.[151,1437,434,1253]" pageId="7" pageNumber="583">
<emphasis id="D1DBEAC1A660FFF1AD83FDCCE73D9AD7" bold="true" box="[199,345,614,641]" pageId="7" pageNumber="583">Description.</emphasis>
Length
<quantity id="24579B36A660FFF1ACFCFDCCE4279AD4" box="[440,579,614,642]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="7" pageNumber="583" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.5–9.5 mm</quantity>
. Head, pronotum and antennal segments 2–11 reddish brown; antennal segment one, legs and six spots on elytra yellowish red; pronotum with brownish black lateral margins; scutellum reddish brown; eyes and elytra black.
</paragraph>
<paragraph id="E31036D3A660FFF1AD83FD79E2679B63" blockId="7.[151,1437,434,1253]" pageId="7" pageNumber="583">Head short, stout, rough, closely punctated, sparsely pubescent; neck highly wrinkled; eyes protruding; last segment of palpi narrowed and sub-truncate at apex; antennae short and stout, not reaching the middle of the elytra; length of antennal segment 3 equals the combined length of segments 1 and 2.</paragraph>
<paragraph id="E31036D3A660FFF1AD83FCEAE7F19B90" blockId="7.[151,1437,434,1253]" pageId="7" pageNumber="583">Pronotum broad, wider than long; disc laterally with anterior two-thirds convex, posterior third slightly divergent; hind angles pointed, sharp, but not protruding laterally; disc with surface highly punctated, wrinkled, pubescent with distantly placed setae; median groove shallow but visible in the mid region of disc, erased near to base and apex of pronotum.</paragraph>
<paragraph id="E31036D3A660FFF1AD83FC65E7239CCB" blockId="7.[151,1437,434,1253]" pageId="7" pageNumber="583">Elytra broad, longer than wide, widest just before apex; scutellum pointed; humerus straight, corners rounded; apex straight, without re-entrant angle; striations broad, weak, shallow, rounded, clear towards base but faint towards apex, intervals shallow, narrow; surface highly punctated, densely pubescent; three yellowish red spots on each elytron, larger square shaped spot just below the humerus extending between the fourth and eighth striations, smaller, rounded spot just below the middle of elytra, oval shaped spot along the apical edge of elytra, not covering the apical edge.</paragraph>
</subSubSection>
<subSubSection id="ABB56558A660FFF1AD83FB0CE3779CB3" pageId="7" pageNumber="583" type="distribution">
<paragraph id="E31036D3A660FFF1AD83FB0CE3779CB3" blockId="7.[151,1437,434,1253]" pageId="7" pageNumber="583">
<emphasis id="D1DBEAC1A660FFF1AD83FB0CE7059C97" bold="true" box="[199,353,1190,1217]" pageId="7" pageNumber="583">Distribution.</emphasis>
<collectingCountry id="9BB87643A660FFF1AC23FB0DE7DF9C97" box="[359,443,1191,1217]" name="India" pageId="7" pageNumber="583">INDIA</collectingCountry>
: Bengal;
<collectingRegion id="216BF831A660FFF1AF66FB0DE41F9C97" box="[546,635,1191,1217]" country="India" name="Gujarat" pageId="7" pageNumber="583">Gujarat</collectingRegion>
: Anand;
<collectingRegion id="216BF831A660FFF1AF9FFB0DE5079C97" box="[731,867,1191,1217]" country="India" name="Tamil Nadu" pageId="7" pageNumber="583">Tamil Nadu</collectingRegion>
: Ettimadai;
<collectingRegion id="216BF831A660FFF1AEA3FB0DE2DA9C97" box="[999,1214,1191,1217]" country="India" name="Arunachal Pradesh" pageId="7" pageNumber="583">Arunachal Pradesh</collectingRegion>
;
<collectingRegion id="216BF831A660FFF1A98DFB0DE3FC9C97" box="[1225,1432,1191,1217]" country="India" name="Himachal Pradesh" pageId="7" pageNumber="583">Himachal Pradesh</collectingRegion>
; Kashmir;
<collectingRegion id="216BF831A660FFF1AC42FB61E73E9CB3" box="[262,346,1227,1253]" country="India" name="Sikkim" pageId="7" pageNumber="583">Sikkim</collectingRegion>
, Uttarpradesh.
<collectingCountry id="9BB87643A660FFF1AF40FB61E4129CB3" box="[516,630,1227,1253]" name="Bhutan" pageId="7" pageNumber="583">BHUTAN</collectingCountry>
(
<bibRefCitation id="873E4B22A660FFF1AFC1FB61E54F9CB3" author="Hrdlicka, J." box="[645,811,1227,1253]" pageId="7" pageNumber="583" pagination="212 - 218" refId="ref18818" refString="Hrdlicka, J. (2003) Brachininae. In: Lobl I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata- Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 212 - 218." type="book chapter" year="2003">Hrdlička 2003</bibRefCitation>
,
<bibRefCitation id="873E4B22A660FFF1AE72FB61E5179CB3" author="Hrdlicka, J." box="[822,883,1227,1253]" pageId="7" pageNumber="583" pagination="471 - 480" refId="ref18865" refString="Hrdlicka, J. (2017) Brachininae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata- Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden, pp. 471 - 480." type="book chapter" year="2017">2017</bibRefCitation>
).
<collectingCountry id="9BB87643A660FFF1AEC1FB61E26F9CB3" box="[901,1035,1227,1253]" name="Pakistan" pageId="7" pageNumber="583">PAKISTAN</collectingCountry>
(
<bibRefCitation id="873E4B22A660FFF1A95DFB61E2DB9CB3" author="Hrdlicka, J." box="[1049,1215,1227,1253]" pageId="7" pageNumber="583" pagination="212 - 218" refId="ref18818" refString="Hrdlicka, J. (2003) Brachininae. In: Lobl I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata- Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 212 - 218." type="book chapter" year="2003">Hrdlička 2003</bibRefCitation>
,
<bibRefCitation id="873E4B22A660FFF1A98EFB61E3639CB3" author="Hrdlicka, J." box="[1226,1287,1227,1253]" pageId="7" pageNumber="583" pagination="471 - 480" refId="ref18865" refString="Hrdlicka, J. (2017) Brachininae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata- Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden, pp. 471 - 480." type="book chapter" year="2017">2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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