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<mods:title id="EB7E837E8D7D281E38E2427F9A9B2554">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="B0FD47252F6C2D81CBAB2CFA25F7C09E">Bistroem, Olof</mods:namePart>
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<mods:namePart id="0AD30760D4EFEEAD9C208FE3F329892B">Bergsten, Johannes</mods:namePart>
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<treatment id="5B4DE85957F9931CBE531507697C1DE8" ID-GBIF-Taxon="127862796" LSID="urn:lsid:plazi:treatment:5B4DE85957F9931CBE531507697C1DE8" httpUri="http://treatment.plazi.org/id/5B4DE85957F9931CBE531507697C1DE8" lastPageId="124" lastPageNumber="125" pageId="122" pageNumber="123">
<subSubSection id="4D0002FBE96CB772199951F841005CE7" pageId="122" pageNumber="123" type="multiple">
<paragraph id="6B3A3FE82AF5CDAB50FDDE2453128899" pageId="122" pageNumber="123">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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<subSubSection id="2BDB2867F266AE6E859C9F25A8187DBE" pageId="122" pageNumber="123" type="nomenclature">
<paragraph id="582FB6612FE280181E40C147E0970B54" pageId="122" pageNumber="123">
<taxonomicName id="D3751AD91441539C0EB5C6874EB420DB" ID-CoL="3RM8X" authority="Guignot, 1952" authorityName="Guignot" authorityYear="1952" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mediocris" order="Coleoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="mediocris">Laccophilus mediocris Guignot, 1952</taxonomicName>
Figs 89-90, 278-279, 435, 552
</paragraph>
</subSubSection>
<subSubSection id="94FC0040E8DE3ADDC46F7AB8D632593C" pageId="123" pageNumber="124" type="reference_group">
<paragraph id="DD52331F484073BDEEC2E643F7ACD2AB" pageId="123" pageNumber="124">
<taxonomicName id="0A74C694E87A6B801652FCBD13E749E7" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mediocris" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="mediocris">
<pageBreakToken id="74AE7B3C6783F2EEA474F87D66FDFD60" pageId="123" pageNumber="124" start="start">Laccophilus</pageBreakToken>
mediocris
</taxonomicName>
<bibRefCitation id="9769F34E6F1C6C6125C29701A47B83F4" author="Guignot, F" journalOrPublisher="Bulletin Institut royal des Sciences Naturelles de Belgique" pageId="257" pageNumber="258" pagination="1 - 7" title="Description de nouveaux dytiscides africains." volume="28" year="1952 d">Guignot 1952d</bibRefCitation>
: 4 (original description, faunistics);
<bibRefCitation id="AFB47A7ABE7D64F71441BFFD53BDC831" author="Guignot, F" journalOrPublisher="Bulletin de l'Institut Francais d'Afrique Noire" pageId="258" pageNumber="259" pagination="83 - 92" title="Coleopteres Dytiscidae et Gyrinidae du Soudan francais." volume="18" year="1956 a">Guignot 1956a</bibRefCitation>
: 88 (faunistics);
<bibRefCitation id="B133E75BD238CA94D0CB41C856F86798" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="36 - 55" title="Dytiscidae from Nyasaland and Southern Rhodesia. III. Laccophilinae." volume="21" year="1958 b">Omer-Cooper 1958b</bibRefCitation>
: 51 (discussion);
<bibRefCitation id="4CC666C19DC202DE826C4D54E6214F12" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
: 562, 567, 568 (description, faunistics);
<bibRefCitation id="B0A20C423C70F3313E3E4AD60EB08A8D" author="Bilardo, A" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="254" pageNumber="255" pagination="93 - 130" title="Recoltes de coleopteres aquatiques Haliplidae et Dytiscidae dans le Gabon et la Cote d'Ivoire." volume="56" year="1978">Bilardo and Pederzani 1978</bibRefCitation>
: 119 (faunistics, description);
<bibRefCitation id="340A1B785652FF5A6B465037E945CA88" author="Medler, JT" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="259" pageNumber="260" pagination="1 - 919" title="Insects of Nigeria - Check list and bibliography." volume="30" year="1980">Medler 1980</bibRefCitation>
: 155 (faunistics, catalogue);
<bibRefCitation id="8E8C58F1B5E35736106061C0C6A2A310" author="Pederzani, F" journalOrPublisher="REDIA" pageId="261" pageNumber="262" pagination="65 - 88" title="Dytiscidae (Coleoptera) collected in the People's Republic of Congo by Mr. Giovanni Onore in the years 1977 - 1980." volume="65" year="1982">Pederzani and Rocchi 1982</bibRefCitation>
: 72 (faunistics);
<bibRefCitation id="A7DDB7D4EB8F384EB94071F5D0FFC939" author="Nilsson, AN" journalOrPublisher="Journal African Zoology" pageId="260" pageNumber="261" pagination="489 - 514" title="The diving beetles (Coleoptera, Dytiscidae) of Guinea Bissau in West Africa." volume="109" year="1995">Nilsson et al. 1995</bibRefCitation>
: 505 (faunistics);
<bibRefCitation id="4D42F9371C0B0864DF8955AEDEA7D3FF" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 246 (catalogue, faunistics);
<bibRefCitation id="5993122D56826DF304F7215672F245A6" author="Pederzani, F" journalOrPublisher="Quaderno di Studi e Notizie di Storia Naturale della Romana" pageId="261" pageNumber="262" pagination="35 - 41" title="Laccophiluscomoensis sp. n. from Ivory Coast (InsectaColeopteraDytiscidae)." volume="16" year="2002">Pederzani and Reintjes 2002</bibRefCitation>
: 40 (faunistics); Reintjes 2000: 67 (faunistics);
<bibRefCitation id="1CCC6D870F97CBA5EB9CA46D2A8DE286" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="195 - 238" title="Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (6 eme partie). Parc National des Plateaux Bateke (missions 2005 et 2006)." volume="149" year="2008">Bilardo and Rocchi 2008</bibRefCitation>
: 211, 236 (faunistics, biology);
<bibRefCitation id="10355E489E39C3BF9932F82D101E9A52" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="255" pageNumber="256" pagination="131 - 155" title="Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (9 eme partie). Parc National des Plateaux Bateke (missions 2010 et 2012)." volume="154" year="2013">Bilardo and Rocchi 2013</bibRefCitation>
: 141 (faunistics);
<bibRefCitation id="242057CAD74AA87A12835676963288C9" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 214 (catalogue, faunistics).
</paragraph>
<paragraph id="3436C0A0251FFE56FBDDADDC3384D2EA" pageId="123" pageNumber="124">
<taxonomicName id="6F5E7520B7CF75608BA6A7F7DE9B658C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus meii" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="meii">Laccophilus meii</taxonomicName>
<bibRefCitation id="169EFC3D8615F8B444AFB158B57C0677" author="Rocchi, S" journalOrPublisher="Roma" pageId="262" pageNumber="263" pagination="11 - 33" title="Contributo alla conoscenza dei ditiscidi di Guinea con descrizione di duo nuove specie (Coleoptera, Dytiscidae)." volumeTitle="Fragmenta Entomologica" year="2000">Rocchi 2000</bibRefCitation>
: 24 (original description, faunistics, discussion);
<bibRefCitation id="143FDB750DAD79CFD9A3ADE3F93F9DC7" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 246 (catalogue, faunistics);
<bibRefCitation id="3C94F5A28BF9EAC975233E9915637F1F" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 214 (catalogue, faunistics). New synonym.
</paragraph>
</subSubSection>
<subSubSection id="65FB64FA7E9D2F083A180493033FB0E7" pageId="123" pageNumber="124" type="type localities">
<paragraph id="F97BE7183668CD18BD92E9AA8E5E0470" pageId="123" pageNumber="124">Type localities.</paragraph>
<paragraph id="CB45D134E240D387DDF42C585EBF942E" pageId="123" pageNumber="124">
<taxonomicName id="5BCCB10077E801C801397ADA475E906D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mediocris" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="mediocris">Laccophilus mediocris</taxonomicName>
: Ivory Coast: Toumodi.
</paragraph>
<paragraph id="7D593546A3BE8F875F789F5A14708E4D" pageId="123" pageNumber="124">
<taxonomicName id="8D298931E0E94015F51F1353ABC59E63" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus meii" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="meii">Laccophilus meii</taxonomicName>
: Republic of Guinea: Faranah, Sidakoro.
</paragraph>
</subSubSection>
<subSubSection id="73BABE0AC8C64A79C2CA9A0CD3FBF382" pageId="123" pageNumber="124" type="type material">
<paragraph id="60222A632DF57358AD416D1FDDD47626" pageId="123" pageNumber="124">Type material studied</paragraph>
<paragraph id="1C8B2BEDD492924FF0D46484E53033ED" pageId="123" pageNumber="124">
(8 exs.)
<taxonomicName id="8E5067FBDA81D2F1E3AC3749A2E6D4D2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mediocris" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="mediocris">Laccophilus mediocris</taxonomicName>
. Holotype: male: &quot;Cote
<normalizedToken id="E59483E0F54E03B7F21890584C9FAE00" originalValue="d’Ivoire">d'Ivoire</normalizedToken>
Toumodi /male symbol / Wehnckei Shp det. Gschwendt. / Type /
<taxonomicName id="B1A676C4C7EC722F683543EC28ABCFFF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mediocris" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="mediocris">Laccophilus mediocris</taxonomicName>
Guign. Type, male&quot; (MNHN). - Paratypes: &quot;Cote
<normalizedToken id="CB59CFBA99162674D9BDED4F280E82F6" originalValue="d’Ivoire">d'Ivoire</normalizedToken>
Toumodi / Museum Paris 12-1930-IV-1931 Ch. Alluaud &amp; P.A. Chappuis / female symbol / Paratype&quot; (3 exs. MNHN; habitus in Fig. 435); Mali: &quot;Soudan Francais Bamako / Museum Paris 12-1930-IV-1931 Ch. Alluaud &amp; P.A. Chappuis / male symbol / Paratype&quot; (1 ex. MNHN).
</paragraph>
<paragraph id="B4F742E8FDC293C55D3293D8430BE8D6" pageId="123" pageNumber="124">
<taxonomicName id="99B5EF170B7F8EAEEF9BC4BB10D9A984" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus meii" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="meii">Laccophilus meii</taxonomicName>
: Holotype: male:
<normalizedToken id="752729E0FAFF97FC657CDDD128B33860" originalValue="“République">&quot;Republique</normalizedToken>
de
<normalizedToken id="F2914D7A3BAFEBD3C6F7820951975BE2" originalValue="Guinée">Guinee</normalizedToken>
, PNHN 10
<geoCoordinate id="49E86BC0EA82C3971EB894533D6A483D" direction="north" orientation="latitude" precision="15" value="10.252222">10o15'08&quot;N</geoCoordinate>
,
<geoCoordinate id="3BD5541C3ED6728826A35F40F2E149E3" direction="west" orientation="longitude" precision="15" value="-10.472221">10o28'20&quot;W</geoCoordinate>
Faranah, Sidakoro, mare, 12.1. 1996 leg. M. Mei&quot; (in acqua con fondo fangoso e molto detrito) / Holotypus /
<taxonomicName id="CE372E3DEBF6A1A6BDD07EDA02153797" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus meii" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="meii">Laccophilus meii</taxonomicName>
sp. n. det. S. Rocchi 1997&quot; (MZUL). - Paratypes, female with same data as holotype but labelled
<normalizedToken id="97D6DB0F3D575D05821FFD4C2DCC5166" originalValue="“Paratypus”">&quot;Paratypus&quot;</normalizedToken>
(1 ex. MZUL); male:
<normalizedToken id="3FCD90DB9A50034A418820AA158D4305" originalValue="“République">&quot;Republique</normalizedToken>
de
<normalizedToken id="55952F749755D23C527C170CE70CC5A2" originalValue="Guinée">Guinee</normalizedToken>
, PNHN9,
<geoCoordinate id="F34E07FCD02FE3ED8276D78D5F6B59F7" direction="north" orientation="latitude" precision="15" value="10.463889">10o27'50&quot;N</geoCoordinate>
,
<geoCoordinate id="C3CE6721A031807BF0DB7FA3E028CA1B" direction="west" orientation="longitude" precision="15" value="-10.440556">10o26'26&quot;W</geoCoordinate>
, Faranah, Somorya F. Koffin 18.1. 1996 leg. M. Mei / Paratypus /
<taxonomicName id="99A396A51900603B0BFBA6F423AC52C3" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus meii" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="meii">Laccophilus meii</taxonomicName>
sp. n. det. S. Rocchi 1997&quot; (1 ex. CSR).
</paragraph>
</subSubSection>
<subSubSection id="6FEA97C0D78C96233A86BF0DC30EF92F" pageId="123" pageNumber="124" type="materials_examined">
<paragraph id="AEA55D17F099D81FD4E1FFAFD31EFA4A" pageId="123" pageNumber="124">Additional material studied</paragraph>
<paragraph id="24B4C9DEE50DFD017CB6F365E103BF41" pageId="123" pageNumber="124">
(22 exs.). Ivory-Coast:
<normalizedToken id="5C36C8183CA1604CBA2F009DC690979B" originalValue="“Comoé">&quot;Comoe</normalizedToken>
NP,
<geoCoordinate id="205CD660B27AC2AC6801B3E49EF7BA52" direction="north" orientation="latitude" precision="5555" value="8.5">N8,5°</geoCoordinate>
-
<geoCoordinate id="FF7E717AF17E8BCC7D2D013CCB94E45E" direction="west" orientation="longitude" precision="5555" value="-3.5">W3,5°</geoCoordinate>
Reintjes / 20.2. 1999&quot; (1 ex. NMW); same data but &quot;5.1. 1999&quot; (3 exs. NMW); &quot;Touba, a la
<normalizedToken id="3AA4D2FAB22DDA9D47C48D2FE78C73DA" originalValue="lumière">lumiere</normalizedToken>
4. 2002 Moretto /
<taxonomicName id="6ADB8FDDBD96225A09703A1BCBE8F504" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mediocris" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="mediocris">Laccophilus mediocris</taxonomicName>
det. Rocchi 2002&quot; (1 ex. CSR). - Ghana: &quot;Ashanti Reg. Kwadaso, agric. st.,
<geoCoordinate id="51B3A3A33B2E16FC1D6A84E8A3154FA7" direction="north" orientation="latitude" precision="555" value="6.42">6.42N-</geoCoordinate>
<geoCoordinate id="57893BC9327EEF2604400A6E82ACF281" direction="west" orientation="longitude" precision="555" value="-1.39">1.39W</geoCoordinate>
/ 26.2. 1969 light trap
<normalizedToken id="5B87887E2594C3ABC0E7EBB515E3A30B" originalValue="Endrödy-Younga”">Endroedy-Younga&quot;</normalizedToken>
(1 ex. TMSA, 1 ex. MZH); &quot;N Reg., Damongo game res.
<geoCoordinate id="E4455185B2B68CE79D8CDCCF82CF85F3" direction="north" orientation="latitude" precision="555" value="9.04">9.04N-</geoCoordinate>
<geoCoordinate id="DA208FC201A0816B3F607FE757A310F0" direction="west" orientation="longitude" precision="555" value="-1.48">1.48W</geoCoordinate>
/ at light 12.8. 1971
<normalizedToken id="93B84D48FFE5E0DD0A5C245D75ABB320" originalValue="Endrödy-Younga”">Endroedy-Younga&quot;</normalizedToken>
(1 ex. TMSA). - Nigeria: &quot;Kontagora 3.IV. 1963 JOC.&quot; (3 exs. AMGS); &quot;Stream, road Kaduna-Kontagora 3.IV. 1963 JOC.&quot; (1 ex. AMGS); &quot;River, Enugo rd about 47 mi. from Makurdi 24.4. 1963 JOC.&quot; (3 exs. AMGS); &quot;Stream Enugo-Makurdi 24.IV. 1963 JOC.&quot; (1 ex. AMGS); &quot;Stream 86 mi. from Makurdi on Jos road 25.IV. 1963&quot; (5 exs. MNHN). - Congo: &quot;Ewo 5. 1979 Onore /
<taxonomicName id="D4A4EA63024ED4CDC5837B24D3FD0C6A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mediocris" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="mediocris">Laccophilus mediocris</taxonomicName>
Guignot det. Pederzani&quot; (1 ex. CSR).
</paragraph>
</subSubSection>
<subSubSection id="CF1A6AB1580B4859441EA0BE45D84BF6" pageId="123" pageNumber="124" type="comments on synonymy">
<paragraph id="A54109BF850DF6EFE41BE46B7C9EA645" pageId="123" pageNumber="124">Comments on synonymy.</paragraph>
<paragraph id="03606B0E9721F5A91FCD05FFB7A86268" pageId="123" pageNumber="124">
Male holotypes of
<taxonomicName id="1BAB4B5D3CA4E087E62948CBA3B3A182" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mediocris" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="mediocris">Laccophilus mediocris</taxonomicName>
and
<taxonomicName id="49664F15937F17C9C00B4574EB4673BA" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus meii" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="meii">Laccophilus meii</taxonomicName>
have been studied and compared. Both external and male genital features seem to be identical. Accordingly the two species are regarded conspecific, and
<taxonomicName id="44FE88542FB685E1CDB9430963528991" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mediocris" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="mediocris">Laccophilus mediocris</taxonomicName>
, which is the older name, is the valid name of the species.
<taxonomicName id="9E297D7DBAE3E3CEBB6E79B11DB9269A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mediocris" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="mediocris">Laccophilus mediocris</taxonomicName>
seems to have been overlooked when
<taxonomicName id="4F74568B51B29E8B8B5B871EB1C33CB6" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus meii" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="meii">Laccophilus meii</taxonomicName>
was introduced as a new species because no reference to it is given to it in the original description (
<bibRefCitation id="57877B551518CF9F474FB7694D986A15" author="Rocchi, S" journalOrPublisher="Roma" pageId="262" pageNumber="263" pagination="11 - 33" title="Contributo alla conoscenza dei ditiscidi di Guinea con descrizione di duo nuove specie (Coleoptera, Dytiscidae)." volumeTitle="Fragmenta Entomologica" year="2000">Rocchi 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C98FC7E01D9CC8D9AFF46F224738C224" pageId="123" pageNumber="124" type="diagnosis">
<paragraph id="4BA3074DC3FDA6ECDDDD675B1DB091CD" pageId="123" pageNumber="124">Diagnosis.</paragraph>
<paragraph id="B2E049692F47B2852D4D43F27DF4BA89" pageId="123" pageNumber="124">
<taxonomicName id="D58116FCBB5FF246B5A265924B630A09" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus mediocris" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="mediocris">Laccophilus mediocris</taxonomicName>
is especially characterized by its peculiar, twisted penis, which is different from all other African
<taxonomicName id="A564DA0B20E436B3B2774A1F1FBD47DC" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
species. Additionally, apical portion of penis comparatively delicate, narrow and moderately twisted; extreme apex finely hooked.
</paragraph>
</subSubSection>
<subSubSection id="801B843BBE8E47B894B5A5918988EF76" pageId="123" pageNumber="124" type="description">
<paragraph id="1B6040B594515E73692D039038371E18" pageId="123" pageNumber="124">Description.</paragraph>
<paragraph id="FAA4BC354596E5FF31D5B05465E3ED91" pageId="123" pageNumber="124">Body length 3.6-4.1 mm, width 2.0-2.2 mm. Elytra with vague colour pattern; irrorations of elytra somewhat weakly developed and in part vague. In frontal half some vague, pale areas may be discerned (Fig. 435).</paragraph>
<paragraph id="DA10F346E2909AFB4DDF86FF7F0E9659" pageId="123" pageNumber="124">Head: Pale ferrugineous. Submat, finely microsculptured; reticulation in part indistinctly double. Large meshes indistinct; when discernible they are only slightly coarser than fine meshes; may include 3-6 fine meshes. At eyes with very fine, scattered punctures. Area of punctures extends a short distance towards centre of head.</paragraph>
<paragraph id="395C936EC5967B5B7801270DBF88BEAC" pageId="123" pageNumber="124">Pronotum: Pale ferrugineous; distinct colour pattern lacking. Finely microsculptured and reticulation indistinctly double. Anteriorly and laterally with very fine punctures.</paragraph>
<paragraph id="7A83C2B99385DFE4F65C6A12EC97ABDC" pageId="123" pageNumber="124">Elytra: Pale ferrugineous, with extensive but rather indistinct, ferrugineous irrorations (Fig. 435). Slightly mat, densely microsculptured. Difference between fine and coarse meshes very small. Rows of punctures hardly visible, almost absent. Discal row of punctures consists of fine, somewhat irregularly located punctures. Dorsolateral and lateral row of punctures only with a few, scattered punctures discernible.</paragraph>
<paragraph id="5E1310EE5EFD2696C66D05B03882AF1D" pageId="123" pageNumber="124">Ventral aspect: Pale ferrugineous; no distinct colour pattern discernible. Almost impunctate. Rather shiny, although very finely, in part indistinctly microsculptured. Semitransverse furrows of metacoxal plates shallow and quite indistinct. Abdomen basally with curved striae. Prosternal process slender and pointed. Apical ventrite with asymmetric (located on one side) knob (Fig. 89).</paragraph>
<paragraph id="6A09677E4B6B130FF21BCEB84C6292B8" pageId="123" pageNumber="124">Legs: Pro- and mesotarsus slightly enlarged, provided with protruding suckers.</paragraph>
<paragraph id="08480D1F4CEB49DF7AD2528AB54D0CC1" pageId="123" pageNumber="124">Male genitalia: Penis twisted, in lateral aspect with apical portion of penis comparatively delicate, narrow and moderately undulate; extreme apex finely hooked (Figs 274-275).</paragraph>
<paragraph id="699C1229E1FA8BD287B4BBA031D87C3A" pageId="123" pageNumber="124">Female: Apical ventrite almost symmetric lacking lateral knob (Fig. 90). Pro- and mesotarsus narrower than in male.</paragraph>
</subSubSection>
<subSubSection id="7A5EBA93E404962FC3E436509CE875A5" pageId="124" pageNumber="125" type="distribution">
<paragraph id="759ABABCCBBDCCE719D94AC372D8E417" pageId="124" pageNumber="125">
<pageBreakToken id="E3DC5BD0065213B938D1E1212DF5C3AF" pageId="124" pageNumber="125" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph id="1280DCF438717B519150E4EBF74DBA90" pageId="124" pageNumber="125">
Mali, Guinea, Ivory Coast, Ghana, Nigeria, Congo (Fig. 552). Non-verified country-records are Zaire (
<bibRefCitation id="50A3A6198C1CA74EB5B9BC9468D21546" author="Guignot, F" journalOrPublisher="Bulletin de l'Institut Francais d'Afrique Noire" pageId="258" pageNumber="259" pagination="83 - 92" title="Coleopteres Dytiscidae et Gyrinidae du Soudan francais." volume="18" year="1956 a">Guignot 1956a</bibRefCitation>
), Guinea-Bissau (
<bibRefCitation id="7EA2218E665C124A4BF10A415DCDEBA0" author="Nilsson, AN" journalOrPublisher="Journal African Zoology" pageId="260" pageNumber="261" pagination="489 - 514" title="The diving beetles (Coleoptera, Dytiscidae) of Guinea Bissau in West Africa." volume="109" year="1995">Nilsson et al. 1995</bibRefCitation>
) and Gabon (
<bibRefCitation id="BD2B513921BD94CAFD266AC50EB07302" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="195 - 238" title="Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (6 eme partie). Parc National des Plateaux Bateke (missions 2005 et 2006)." volume="149" year="2008">Bilardo and Rocchi 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="48E87493AAA7D42C109A27D2F52DC323" pageId="124" pageNumber="125" type="collecting circumstances">
<paragraph id="CFB9544C1A827E7F9E8B125935B0924D" pageId="124" pageNumber="125">Collecting circumstances.</paragraph>
<paragraph id="CC8408A6AEB9785B882DFA1F8B20AE01" pageId="124" pageNumber="125">
Rather insufficiently documented. When describing
<taxonomicName id="B69C4F89B883619EA1B8778CF1FD3518" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus meii" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="meii">Laccophilus meii</taxonomicName>
,
<bibRefCitation id="936CAE20241DCCD71680665E0E7128A9" author="Rocchi, S" journalOrPublisher="Roma" pageId="262" pageNumber="263" pagination="11 - 33" title="Contributo alla conoscenza dei ditiscidi di Guinea con descrizione di duo nuove specie (Coleoptera, Dytiscidae)." volumeTitle="Fragmenta Entomologica" year="2000">Rocchi (2000)</bibRefCitation>
, however, added some data on the sampling localities: the species was collected in water with a muddy bottom and much debris and moreover, in a pool along river bed of Koffin River.
</paragraph>
</subSubSection>
</treatment>
</document>
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