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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1052.66803" ID-GBIF-Dataset="d4c7ba8d-67d1-4e7e-81f8-8bf582104a99" ID-PMC="PMC8346418" ID-Pensoft-Pub="1313-2970-1052-25" ID-Pensoft-UUID="43B22A07D2085C8D994CC6C537BD3902" ID-PubMed="34393550" ID-ZooBank="0EF3C375ED5F4C70A134DBE1DA6A5C37" ModsDocID="1313-2970-1052-25" checkinTime="1627725770628" checkinUser="pensoft" docAuthor="Zhang, Jianshuang, Yu, Hao &amp; Li, Shuqiang" docDate="2021" docId="FABBED9EC0375C75BB6A0DDC4F7273DD" docLanguage="en" docName="ZooKeys 1052: 25-41" docOrigin="ZooKeys 1052" docPubDate="2021-07-30" docSource="http://dx.doi.org/10.3897/zookeys.1052.66803" docTitle="Femorbiona phami Yu &amp; Li 2021, sp. nov." docType="treatment" docUuid="B685CC15-07F5-431F-8A9B-B9A69906A55D" docUuidSource="ZooBank" docVersion="4" id="43B22A07D2085C8D994CC6C537BD3902" lastPageNumber="25" masterDocId="43B22A07D2085C8D994CC6C537BD3902" masterDocTitle="Femorbiona gen. nov., a new genus of sac spiders (Araneae, Clubionidae) from Southeast Asia" masterLastPageNumber="41" masterPageNumber="25" pageNumber="25" updateTime="1668150765872" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Femorbiona gen. nov., a new genus of sac spiders (Araneae, Clubionidae) from Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Jianshuang</mods:namePart>
<mods:affiliation>School of Life Sciences, Guizhou Normal University, Guiyang, Guizhou, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yu, Hao</mods:namePart>
<mods:affiliation>School of Biological Sciences, Guizhou Education University, Guiyang, Guizhou, China</mods:affiliation>
<mods:nameIdentifier type="email">insect1986@126.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
<mods:nameIdentifier type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-07-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1052</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>25</mods:start>
<mods:end>41</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1052.66803</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1052.66803</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1052-25</mods:identifier>
<mods:identifier type="ZooBank">0EF3C375ED5F4C70A134DBE1DA6A5C37</mods:identifier>
<mods:identifier type="Pensoft-UUID">43B22A07D2085C8D994CC6C537BD3902</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="183840199" LSID="urn:lsid:zoobank.org:act:B685CC15-07F5-431F-8A9B-B9A69906A55D" httpUri="http://treatment.plazi.org/id/FABBED9EC0375C75BB6A0DDC4F7273DD" lastPageNumber="25" pageId="0" pageNumber="25">
<subSubSection pageId="0" pageNumber="25" type="nomenclature">
<paragraph pageId="0" pageNumber="25">
<taxonomicName LSID="http://zoobank.org/B685CC15-07F5-431F-8A9B-B9A69906A55D" authority="Yu &amp; Li" authorityName="Yu &amp; Li" authorityYear="2021" family="Clubionidae" genus="Femorbiona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Femorbiona phami" order="Araneae" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="phami" status="sp. nov.">Femorbiona phami Yu &amp; Li</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="25">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="description">
<paragraph pageId="0" pageNumber="25">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Femorbiona phami sp. nov., holotype male palp A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, retrolateral view. Abbreviations: E = embolus; EB = embolic base; FA = femoral apophysis; PPA = prolateral patellar apophysis; VTA = ventral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66803.figure3" httpUri="https://binary.pensoft.net/fig/571993" pageId="0" pageNumber="25">Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Femorbiona phami sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: BS = bursa; CO = copulatory opening; SP = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E-H)." figureDoi="10.3897/zookeys.1052.66803.figure4" httpUri="https://binary.pensoft.net/fig/571994" pageId="0" pageNumber="25">, 4</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Femorbiona spp., left male palp, ventral view A F. brachyptera, paratype B F. phami sp. nov., holotype C F. shenzhen sp. nov., holotype. Abbreviations: C = conductor; E = embolus; FA = femoral apophysis; EB = embolic base; PPA = prolateral patellar apophysis. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1052.66803.figure7" httpUri="https://binary.pensoft.net/fig/571997" pageId="0" pageNumber="25">, 7B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="type material">
<paragraph pageId="0" pageNumber="25">Type material.</paragraph>
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" italics="true" pageId="0" pageNumber="25">Holotype</emphasis>
♂ (IZCAS Ar 34724), VIETNAM: Hai Phong: Cat Ba National Park: disturbed forest,
<geoCoordinate degrees="20" direction="north" minutes="48.258" orientation="latitude" precision="1" value="20.8043">20°48.258'N</geoCoordinate>
,
<geoCoordinate degrees="107" direction="east" minutes="00.581" orientation="longitude" precision="1" value="107.00968">107°00.581'E</geoCoordinate>
, ca. 80 m, 15.VII.2008, D.S. Pham leg.
<emphasis bold="true" italics="true" pageId="0" pageNumber="25">Paratypes</emphasis>
: 1♀ (IZCAS Ar 34725), same locality and collector as holotype,
<geoCoordinate degrees="20" direction="north" minutes="48.249" orientation="latitude" precision="1" value="20.80415">20°48.249'N</geoCoordinate>
,
<geoCoordinate degrees="107" direction="east" minutes="00.016" orientation="longitude" precision="1" value="107.00027">107°00.016'E</geoCoordinate>
, ca. 80 m, 16.VII.2008.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="diagnosis">
<paragraph pageId="0" pageNumber="25">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="25">
The males of
<taxonomicName lsidName="F. phami" pageId="0" pageNumber="25" rank="species" species="phami">
<emphasis italics="true" pageId="0" pageNumber="25">F. phami</emphasis>
</taxonomicName>
sp. nov. are most similar to those of
<taxonomicName lsidName="F. brachyptera" pageId="0" pageNumber="25" rank="species" species="brachyptera">
<emphasis italics="true" pageId="0" pageNumber="25">F. brachyptera</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3167.1.5" author="Zhu, BR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="25" pagination="53 - 56" refId="B14" refString="Zhu, BR, Ren, HQ, Chen, J, 2012. A new species of the genus Clubiona Latreille (Araneae, Clubionidae) from Hainan Island, China. Zootaxa 3167: 53 - 56, DOI: https://doi.org/10.11646/zootaxa.3167.1.5" title="A new species of the genus Clubiona Latreille (Araneae, Clubionidae) from Hainan Island, China." url="https://doi.org/10.11646/zootaxa.3167.1.5" volume="3167" year="2012">Zhu et al. 2012</bibRefCitation>
: 53, figs 2-4, 8-10; Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Femorbiona brachyptera, paratype male palp A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, retrolateral view. Abbreviations: C = conductor; E = embolus; EB = embolic base; FA = femoral apophysis; PPA = prolateral patellar apophysis; VTA = ventral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66803.figure1" httpUri="https://binary.pensoft.net/fig/571991" pageId="0" pageNumber="25">1A-E</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Femorbiona spp., left male palp, ventral view A F. brachyptera, paratype B F. phami sp. nov., holotype C F. shenzhen sp. nov., holotype. Abbreviations: C = conductor; E = embolus; FA = femoral apophysis; EB = embolic base; PPA = prolateral patellar apophysis. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1052.66803.figure7" httpUri="https://binary.pensoft.net/fig/571997" pageId="0" pageNumber="25">7A</figureCitation>
) by the general shape of the palpal femoral apophysis, which is shaped like a short bird wing in retrolateral view (cf. Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Femorbiona brachyptera, paratype male palp A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, retrolateral view. Abbreviations: C = conductor; E = embolus; EB = embolic base; FA = femoral apophysis; PPA = prolateral patellar apophysis; VTA = ventral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66803.figure1" httpUri="https://binary.pensoft.net/fig/571991" pageId="0" pageNumber="25">1B</figureCitation>
and Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Femorbiona phami sp. nov., holotype male palp A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, retrolateral view. Abbreviations: E = embolus; EB = embolic base; FA = femoral apophysis; PPA = prolateral patellar apophysis; VTA = ventral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66803.figure3" httpUri="https://binary.pensoft.net/fig/571993" pageId="0" pageNumber="25">3B</figureCitation>
), but can be distinguished from
<taxonomicName lsidName="F. brachyptera" pageId="0" pageNumber="25" rank="species" species="brachyptera">
<emphasis italics="true" pageId="0" pageNumber="25">F. brachyptera</emphasis>
</taxonomicName>
by having: (1) the distal tip of the femoral apophysis slightly curved (vs. not curved) (cf. Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Femorbiona spp., left male palp, ventral view A F. brachyptera, paratype B F. phami sp. nov., holotype C F. shenzhen sp. nov., holotype. Abbreviations: C = conductor; E = embolus; FA = femoral apophysis; EB = embolic base; PPA = prolateral patellar apophysis. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1052.66803.figure7" httpUri="https://binary.pensoft.net/fig/571997" pageId="0" pageNumber="25">7B</figureCitation>
and Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Femorbiona spp., left male palp, ventral view A F. brachyptera, paratype B F. phami sp. nov., holotype C F. shenzhen sp. nov., holotype. Abbreviations: C = conductor; E = embolus; FA = femoral apophysis; EB = embolic base; PPA = prolateral patellar apophysis. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1052.66803.figure7" httpUri="https://binary.pensoft.net/fig/571997" pageId="0" pageNumber="25">7A</figureCitation>
); (2) in ventral view, the embolar apex relatively long, over 1/3 the length of the embolic base and pointed prolatero-distally (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Femorbiona phami sp. nov., holotype male palp A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, retrolateral view. Abbreviations: E = embolus; EB = embolic base; FA = femoral apophysis; PPA = prolateral patellar apophysis; VTA = ventral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66803.figure3" httpUri="https://binary.pensoft.net/fig/571993" pageId="0" pageNumber="25">3D</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Femorbiona spp., left male palp, ventral view A F. brachyptera, paratype B F. phami sp. nov., holotype C F. shenzhen sp. nov., holotype. Abbreviations: C = conductor; E = embolus; FA = femoral apophysis; EB = embolic base; PPA = prolateral patellar apophysis. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1052.66803.figure7" httpUri="https://binary.pensoft.net/fig/571997" pageId="0" pageNumber="25">7B</figureCitation>
) (vs. relatively short tip, &lt;1/6 the length of the embolic base, pointed ventro-distally, as in Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Femorbiona brachyptera, paratype male palp A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, retrolateral view. Abbreviations: C = conductor; E = embolus; EB = embolic base; FA = femoral apophysis; PPA = prolateral patellar apophysis; VTA = ventral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66803.figure1" httpUri="https://binary.pensoft.net/fig/571991" pageId="0" pageNumber="25">1D</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Femorbiona spp., left male palp, ventral view A F. brachyptera, paratype B F. phami sp. nov., holotype C F. shenzhen sp. nov., holotype. Abbreviations: C = conductor; E = embolus; FA = femoral apophysis; EB = embolic base; PPA = prolateral patellar apophysis. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1052.66803.figure7" httpUri="https://binary.pensoft.net/fig/571997" pageId="0" pageNumber="25">7A</figureCitation>
); (3) conductor absent (vs. present) (cf. Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Femorbiona phami sp. nov., holotype male palp A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, retrolateral view. Abbreviations: E = embolus; EB = embolic base; FA = femoral apophysis; PPA = prolateral patellar apophysis; VTA = ventral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66803.figure3" httpUri="https://binary.pensoft.net/fig/571993" pageId="0" pageNumber="25">3C-E</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Femorbiona spp., left male palp, ventral view A F. brachyptera, paratype B F. phami sp. nov., holotype C F. shenzhen sp. nov., holotype. Abbreviations: C = conductor; E = embolus; FA = femoral apophysis; EB = embolic base; PPA = prolateral patellar apophysis. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1052.66803.figure7" httpUri="https://binary.pensoft.net/fig/571997" pageId="0" pageNumber="25">7B</figureCitation>
and Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Femorbiona brachyptera, paratype male palp A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, retrolateral view. Abbreviations: C = conductor; E = embolus; EB = embolic base; FA = femoral apophysis; PPA = prolateral patellar apophysis; VTA = ventral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66803.figure1" httpUri="https://binary.pensoft.net/fig/571991" pageId="0" pageNumber="25">1C-E</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Femorbiona spp., left male palp, ventral view A F. brachyptera, paratype B F. phami sp. nov., holotype C F. shenzhen sp. nov., holotype. Abbreviations: C = conductor; E = embolus; FA = femoral apophysis; EB = embolic base; PPA = prolateral patellar apophysis. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1052.66803.figure7" httpUri="https://binary.pensoft.net/fig/571997" pageId="0" pageNumber="25">7A</figureCitation>
). The female of
<taxonomicName lsidName="F. phami" pageId="0" pageNumber="25" rank="species" species="phami">
<emphasis italics="true" pageId="0" pageNumber="25">F. phami</emphasis>
</taxonomicName>
sp. nov. can be easily distinguished from all other congeners by the slightly sclerotised and pocket-like copulatory openings and by the absence of an epigynal hood (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Femorbiona phami sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: BS = bursa; CO = copulatory opening; SP = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E-H)." figureDoi="10.3897/zookeys.1052.66803.figure4" httpUri="https://binary.pensoft.net/fig/571994" pageId="0" pageNumber="25">4A, B</figureCitation>
) (vs. 2 circular copulatory openings located at the lateral border of hood, as in Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Femorbiona brachyptera, female and male paratypes, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: BS = bursa; CO = copulatory opening; H = hood; SP = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E-H)." figureDoi="10.3897/zookeys.1052.66803.figure2" httpUri="https://binary.pensoft.net/fig/571992" pageId="0" pageNumber="25">2A, B</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Femorbiona shenzhen sp. nov., male holotype and female paratype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: BS = bursa; CO = copulatory opening; H = hood; SP = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E-H)." figureDoi="10.3897/zookeys.1052.66803.figure6" httpUri="https://binary.pensoft.net/fig/571996" pageId="0" pageNumber="25">6A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="description">
<paragraph pageId="0" pageNumber="25">Description.</paragraph>
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Male.</emphasis>
Holotype (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Femorbiona phami sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: BS = bursa; CO = copulatory opening; SP = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E-H)." figureDoi="10.3897/zookeys.1052.66803.figure4" httpUri="https://binary.pensoft.net/fig/571994" pageId="0" pageNumber="25">4E, F</figureCitation>
): Total length 3.51; carapace 1.66 long, 1.28 wide; opisthosoma 1.85 long, 1.01 wide. Carapace orange, pars cephalica darker in ocular area, without distinct pattern. Eyes: both AER and PER slightly recurved in dorsal view. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.07, ALE 0.13, PME 0.10, PLE 0.12, AME-AME 0.08, AME-ALE 0.06, PME-PME 0.20, PME-PLE 0.12, MOQL 0.22, MOQA 0.21, MOQP 0.38. Chelicerae slightly darker than ocular area, with 6 promarginal and 5 retromarginal teeth. Labium and endites coloured as chelicerae, longer than wide. Sternum yellowish white. Legs coloured as sternum, without distinct markings. Leg measurements: I 3.56 (0.99, 1.48, 0.70, 0.39), II 4.11 (1.18, 1.69, 0.80, 0.44), III 3.27 (0.95, 1.17, 0.82, 0.33), IV 4.74 (1.32, 1.62, 1.36, 0.44). Abdomen elongate-oval, dorsally yellowish white, dorsum with 2 pairs of inconspicuous muscle depressions; venter white with no distinct pattern.
</paragraph>
<paragraph pageId="0" pageNumber="25">
Palp (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Femorbiona phami sp. nov., holotype male palp A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, retrolateral view. Abbreviations: E = embolus; EB = embolic base; FA = femoral apophysis; PPA = prolateral patellar apophysis; VTA = ventral tibial apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66803.figure3" httpUri="https://binary.pensoft.net/fig/571993" pageId="0" pageNumber="25">3A-E</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Femorbiona spp., left male palp, ventral view A F. brachyptera, paratype B F. phami sp. nov., holotype C F. shenzhen sp. nov., holotype. Abbreviations: C = conductor; E = embolus; FA = femoral apophysis; EB = embolic base; PPA = prolateral patellar apophysis. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1052.66803.figure7" httpUri="https://binary.pensoft.net/fig/571997" pageId="0" pageNumber="25">7B</figureCitation>
): Femur with short retrolateral apophysis originating proximally; FA ca. 1/3-1/4 length of femur, thin distally, wide basally. Patella 2
<normalizedToken originalValue="×">x</normalizedToken>
longer and 1.3-1.5
<normalizedToken originalValue="×">x</normalizedToken>
wider than tibia, with round, short prolateral apophysis. Tibia cup shaped, relatively short, &lt;1/3 of cymbium length, with papilliform, partly membranous, ventro-retrolateral apophysis. Tegulum elongate, oval, and inflated, ca. 1.6
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; sperm duct indistinct in ventral view. Embolus somewhat helical, more or less
<normalizedToken originalValue="Ƨ-shaped">Ƨ-shaped</normalizedToken>
in ventral view, base an enlarged tubercle, inserted at approximately the 10-11
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
position on tegulum, gradually tapered toward tip; embolar apex needle-like, prolaterally pointed, over 1/3 of base length.
</paragraph>
<caption doi="10.3897/zookeys.1052.66803.figure3" httpUri="https://binary.pensoft.net/fig/571993" pageId="0" pageNumber="25" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 3.</emphasis>
<taxonomicName authorityName="Yu &amp; Li" authorityYear="2021" family="Clubionidae" genus="Femorbiona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Femorbiona phami" order="Araneae" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="phami">
<emphasis italics="true" pageId="0" pageNumber="25">Femorbiona phami</emphasis>
</taxonomicName>
sp. nov., holotype male palp
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
prolateral view
<emphasis bold="true" pageId="0" pageNumber="25">B</emphasis>
retrolateral view
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
bulb, prolateral view
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
bulb, ventral view
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
bulb, retrolateral view. Abbreviations: E = embolus; EB = embolic base; FA = femoral apophysis; PPA = prolateral patellar apophysis; VTA = ventral tibial apophysis. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Female.</emphasis>
Paratype (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Femorbiona phami sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: BS = bursa; CO = copulatory opening; SP = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E-H)." figureDoi="10.3897/zookeys.1052.66803.figure4" httpUri="https://binary.pensoft.net/fig/571994" pageId="0" pageNumber="25">4G, H</figureCitation>
): total length 3.99; carapace 1.84 long, 1.43 wide; opisthosoma 2.15 long, 1.07 wide. Eye sizes and interdistances: AME 0.09, ALE 0.13, PME 0.12, PLE 0.10, AME-AME 0.06, AME-ALE 0.06, PME-PME 0.12, PME-PLE 0.15, MOQL 0.27, MOQA 0.27, MOQP 0.48. Legs yellowish white, without distinct markings. Leg measurements: I 4.10 (1.18, 1.70, 0.80, 0.42), II 4.60 (1.33, 1.92, 0.93, 0.42), III 3.72 (1.11, 1.28, 0.93, 0.40), IV 5.31 (1.52, 1.86, 1.47, 0.46). Similar to males but slightly smaller and lighter.
</paragraph>
<caption doi="10.3897/zookeys.1052.66803.figure4" httpUri="https://binary.pensoft.net/fig/571994" pageId="0" pageNumber="25" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 4.</emphasis>
<taxonomicName authorityName="Yu &amp; Li" authorityYear="2021" family="Clubionidae" genus="Femorbiona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Femorbiona phami" order="Araneae" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="phami">
<emphasis italics="true" pageId="0" pageNumber="25">Femorbiona phami</emphasis>
</taxonomicName>
sp. nov., female paratype and male holotype, epigyne (
<emphasis bold="true" pageId="0" pageNumber="25">A-D</emphasis>
), male habitus (
<emphasis bold="true" pageId="0" pageNumber="25">E, F</emphasis>
) and female habitus (
<emphasis bold="true" pageId="0" pageNumber="25">G, H</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
intact, ventral view
<emphasis bold="true" pageId="0" pageNumber="25">B</emphasis>
cleared, ventral view
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
cleared, dorsal view
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
cleared, dorsal view
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="25">F</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="25">G</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="25">H</emphasis>
ventral view. Abbreviations: BS = bursa; CO = copulatory opening; SP = spermatheca. Scale bars: 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="25">A-D</emphasis>
); 1 mm (
<emphasis bold="true" pageId="0" pageNumber="25">E-H</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="25">
Epigyne (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Femorbiona phami sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: BS = bursa; CO = copulatory opening; SP = spermatheca. Scale bars: 0.1 mm (A-D); 1 mm (E-H)." figureDoi="10.3897/zookeys.1052.66803.figure4" httpUri="https://binary.pensoft.net/fig/571994" pageId="0" pageNumber="25">4A-D</figureCitation>
). Epigynal plate wider than long, anterior and lateral margins not rebordered; spermathecae and bursae indistinct, copulatory ducts easily visible through integument in ventral view. Copulatory openings large, pocket-like, located on chitinous structures at the postero-lateral portion of epigynal plate, separated by ca. 1 diameter. Copulatory ducts thick, covered by large spermathecae in dorsal view, directed anteriorly, then connected to spermathecae. Spermathecae long, tubular, sinuous. Bursae oblong, translucent, surface smooth, close together, ca. 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="distribution">
<paragraph pageId="0" pageNumber="25">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="25">Known only from the type locality, Hai Phong, Vietnam.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="etymology">
<paragraph pageId="0" pageNumber="25">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="25">The specific name is a patronym after Dinh Sac Pham (Hanoi, Vietnam), collector of the type series; noun (name) in apposition.</paragraph>
</subSubSection>
</treatment>
</document>