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<document ID-DOI="10.5733/afin.051.0212" ID-ISSN="2305-2562" ID-Zenodo-Dep="7913777" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1683579024145" checkinUser="felipe" docAuthor="Londt, Jason G. H." docDate="2010" docId="FA455017FFF0AD6EFE2974BA4E61FDA4" docLanguage="en" docName="AfrInvertebr.51.2.431-482.pdf" docOrigin="African Invertebrates 51 (2)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.051.0212" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Acnephalomyia eremia Londt 2010, sp. n." docType="treatment" docVersion="3" lastPageNumber="452" masterDocId="067C286FFFE4AD7BFFED75294D5FFFC2" masterDocTitle="A review of Afrotropical Acnephalum Macquart, 1838, including the reinstatement of Sporadothrix Hermann, 1907 and descriptions of two new genera (Diptera: Asilidae: Stenopogoninae)" masterLastPageNumber="431" masterPageNumber="431" pageNumber="451" updateTime="1683633860393" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Afrotropical Acnephalum Macquart, 1838, including the reinstatement of Sporadothrix Hermann, 1907 and descriptions of two new genera (Diptera: Asilidae: Stenopogoninae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Londt, Jason G. H.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>51</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>431</mods:start>
<mods:end>431</mods:end>
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<mods:location>
<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.051.0212</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.051.0212</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7913777</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7913861" ID-Zenodo-Dep="7913861" LSID="urn:lsid:plazi:treatment:FA455017FFF0AD6EFE2974BA4E61FDA4" httpUri="http://treatment.plazi.org/id/FA455017FFF0AD6EFE2974BA4E61FDA4" lastPageId="21" lastPageNumber="452" pageId="20" pageNumber="451">
<subSubSection box="[452,795,403,430]" pageId="20" pageNumber="451" type="nomenclature">
<paragraph blockId="20.[452,795,403,430]" box="[452,795,403,430]" pageId="20" pageNumber="451">
<heading box="[452,795,403,430]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="451" reason="2">
<taxonomicName ID-CoL="89TV4" authority="Londt, 2010" authorityName="Londt" authorityYear="2010" box="[452,728,403,429]" class="Insecta" family="Asilidae" genus="Acnephalomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="451" phylum="Arthropoda" rank="species" species="eremia" status="sp. n.">
<emphasis bold="true" box="[452,728,403,429]" pageId="20" pageNumber="451">Acnephalomyia eremia</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[735,795,404,430]" pageId="20" pageNumber="451" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="21" lastPageNumber="452" pageId="20" pageNumber="451" type="description">
<paragraph blockId="20.[538,709,446,472]" box="[538,709,446,472]" pageId="20" pageNumber="451">
<figureCitation box="[538,670,446,472]" captionStart="Figs 3543" captionStartId="45.[181,220,1450,1472]" captionTargetBox="[202,1123,315,1428]" captionTargetId="figure-53@45.[183,1143,312,1433]" captionTargetPageId="45" captionText="Figs 3543. Acnephalomyia male terminalia: (3537) A. dorsalis (Macquart, 1838) Knersvlakte, North of Vanrhynsdorp, lateral, dorsal and ventral; (3840) A. eremia sp. n. holotype, lateral, dorsal and ventral; (4143) A. iota sp. n. holotype, lateral, dorsal and ventral. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7913793" httpUri="https://zenodo.org/record/7913793/files/figure.png" pageId="20" pageNumber="451">Figs 3840</figureCitation>
,
<figureCitation box="[681,709,446,472]" captionStart="Fig" captionStartId="49.[181,214,861,883]" captionTargetBox="[320,1014,266,831]" captionTargetId="figure-45@49.[319,1016,255,840]" captionTargetPageId="49" captionText="Fig. 66. Distribution of Acnephalomyia species: A. dorsalis (Macquart, 1838) circles; A. eremia sp. n. triangle." figureDoi="http://doi.org/10.5281/zenodo.7913807" httpUri="https://zenodo.org/record/7913807/files/figure.png" pageId="20" pageNumber="451">66</figureCitation>
</paragraph>
<paragraph blockId="20.[144,1104,488,1672]" pageId="20" pageNumber="451">
Etymology: From Greek
<taxonomicName authorityName="Londt" authorityYear="2010" box="[424,501,488,513]" class="Insecta" family="Asilidae" genus="Acnephalomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="451" phylum="Arthropoda" rank="species" species="eremia">
<emphasis box="[424,501,488,513]" italics="true" pageId="20" pageNumber="451">eremia</emphasis>
</taxonomicName>
(desert, wilderness); noun in apposition. Refers to the arid conditions under which the species has been found.
</paragraph>
<paragraph blockId="20.[144,1104,488,1672]" pageId="20" pageNumber="451">
Description (Condition of
<typeStatus box="[448,546,561,587]" pageId="20" pageNumber="451">holotype</typeStatus>
fair; antennae broken off beyond pedicels; right prothoracic and left meso- and metathoracic legs missing.):
</paragraph>
<paragraph blockId="20.[144,1104,488,1672]" pageId="20" pageNumber="451">
<emphasis box="[144,205,634,660]" italics="true" pageId="20" pageNumber="451">Head</emphasis>
: Dark red-brown to black, white setose, mostly apruinose. Antenna (broken off beyond pedicel): Scape and pedicel brown-orange. Segmental ratios 1.0:0.8:?:?:? scape and pedicel subequal in length, pale whitish setose (most macrosetae missing, but single macroseta on dorsal aspect of right pedicel suggests that these would be as long or longer than postpedicel). Face dark red-brown to black, apruinose except for narrow strips adjacent to eye margins, mystax white, covering entire face, but weakly dorsally (some setae appear to have been rubbed off). Frons, vertex and postocular region dark red-brown to black, apruinose except for central part of postocular region, white setose; angle subtended by eye margins at level of frons/vertex
<emphasis box="[917,929,891,916]" italics="true" pageId="20" pageNumber="451">c</emphasis>
. 17°. Proboscis red-brown, white setose. Palpus dark red-brown, 2-segmented, white setose.
</paragraph>
<paragraph blockId="20.[144,1104,488,1672]" pageId="20" pageNumber="451">
<emphasis box="[144,226,964,990]" italics="true" pageId="20" pageNumber="451">Thorax</emphasis>
: Dark red-brown to black, apruinose, mostly white setose. Pronotum mostly white setose, but with group of yellowish macrosetae medially. Mesonotum mostly white setose (a few pale yellowish setae posteromedially). Lateral macrosetae moderately developed, white (4
<emphasis box="[449,485,1060,1086]" italics="true" pageId="20" pageNumber="451">npl</emphasis>
, 5 or 6
<emphasis box="[576,623,1060,1086]" italics="true" pageId="20" pageNumber="451">spal</emphasis>
, 5 or 6
<emphasis box="[713,749,1060,1086]" italics="true" pageId="20" pageNumber="451">pal</emphasis>
). Pleura largely asetose except for numerous, long, somewhat wavy, white katatergals, dorsally situated anepisternals and katepisternals. Scutellum shiny dark red-brown apruinose with poorly-developed transverse, subapical groove.About 26 moderately developed white apical macrosetae accompanied by slightly shorter, fine, white setae; disc sparsely white setose. Legs: Mostly dark red-brown, but
<emphasis box="[455,496,1220,1246]" italics="true" pageId="20" pageNumber="451">fem</emphasis>
orange-brown ventrally. Major setae erect, translucent whitish, minor setae recumbent, white. Ventral parts of
<emphasis box="[768,801,1252,1277]" italics="true" pageId="20" pageNumber="451">tar</emphasis>
black setose. Claws black, longish (but shorter than tarsomere 5); empodia slender, yellow, about half length of claws; pulvilli tiny (about one-third length of empodium). Haltere brown-yellow, base slightly darker. Wing: 6.2×
<quantity box="[455,549,1348,1374]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="20" pageNumber="451" unit="mm" value="2.4">2.4 mm</quantity>
. Veins brown, membrane lacking microtrichiae; entirely transparent, unstained. Vein R
<subScript attach="left" box="[578,586,1398,1412]" fontSize="6" pageId="20" pageNumber="451">4</subScript>
with well-developed basal stump-vein.
</paragraph>
<paragraph blockId="20.[144,1104,488,1672]" lastBlockId="21.[181,1142,204,614]" lastPageId="21" lastPageNumber="452" pageId="20" pageNumber="451">
<emphasis box="[144,250,1421,1447]" italics="true" pageId="20" pageNumber="451">Abdomen</emphasis>
: Terga broader than long, mostly dark red-brown to black, but somewhat orange-brown laterally. Terga apruinose, but entirely pitted by setal sockets. Terga with long, recumbent white setae laterally. White setae extend along distal margins of terga gradually becoming shorter medially. Large areas of terga appear asetose, but are covered with tiny reddish-brown setae. Sterna dark red-brown, apruinose, longish white setose. Terminalia largely withdrawn between T6 and S6, distal parts somewhat obscured by setae. Genitalia (
<figureCitation box="[475,611,1613,1639]" captionStart="Figs 3543" captionStartId="45.[181,220,1450,1472]" captionTargetBox="[202,1123,315,1428]" captionTargetId="figure-53@45.[183,1143,312,1433]" captionTargetPageId="45" captionText="Figs 3543. Acnephalomyia male terminalia: (3537) A. dorsalis (Macquart, 1838) Knersvlakte, North of Vanrhynsdorp, lateral, dorsal and ventral; (3840) A. eremia sp. n. holotype, lateral, dorsal and ventral; (4143) A. iota sp. n. holotype, lateral, dorsal and ventral. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7913793" httpUri="https://zenodo.org/record/7913793/files/figure.png" pageId="20" pageNumber="451">Figs 3840</figureCitation>
):
<emphasis box="[630,703,1613,1639]" italics="true" pageId="20" pageNumber="451">Epand</emphasis>
moderately well-developed, jutting out to almost same level as achieved by outer lobe of
<emphasis box="[748,815,1646,1671]" italics="true" pageId="20" pageNumber="451">goncx</emphasis>
, distally broadly rounded and shallowly incised.
<emphasis box="[441,494,204,229]" italics="true" pageId="21" pageNumber="452">Proc</emphasis>
somewhat swollen in appearance and jutting out slightly beyond level achieved by
<emphasis box="[469,537,236,262]" italics="true" pageId="21" pageNumber="452">epand</emphasis>
(lateral view). Exterior lobe of
<emphasis box="[886,953,236,261]" italics="true" pageId="21" pageNumber="452">goncx</emphasis>
broadly rounded proximally, tapering distally to a broadly rounded, somewhat truncate, distal end; interior lobe rather long, jutting out to about same level as achieved by
<emphasis box="[987,1041,300,326]" italics="true" pageId="21" pageNumber="452">hypd</emphasis>
, slightly downturned distally.
<emphasis box="[415,476,332,358]" italics="true" pageId="21" pageNumber="452">Hypd</emphasis>
somewhat truncate basally, tapering to long finger­like, distally projecting medial lobe.
</paragraph>
<paragraph blockId="21.[181,1142,204,614]" pageId="21" pageNumber="452">
<materialsCitation collectingDate="1650-01" collectionCode="NMSA" country="Namibia" county="Dune" latitude="-23.766666" location="Namib Desert" longLatPrecision="1254" longitude="15.783334" municipality="Aug" pageId="21" pageNumber="452" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[181,269,397,419]" pageId="21" pageNumber="452">Holotype</typeStatus>
:
<specimenCount box="[277,294,397,418]" pageId="21" pageNumber="452" type="male"></specimenCount>
<collectingCountry box="[300,403,397,418]" name="Namibia" pageId="21" pageNumber="452">NAMIBIA</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:FA455017FFF0AD6EFE2974BA4E61FDA4:7733B7DAFFF1AD6EFE7374A44F7FFE60" box="[414,544,397,419]" country="Namibia" county="Dune" latitude="-23.766666" longLatPrecision="1254" longitude="15.783334" municipality="Aug" name="Namib Desert" pageId="21" pageNumber="452">Namib Desert</location>
SWA /
<location LSID="urn:lsid:plazi:treatment:FA455017FFF0AD6EFE2974BA4E61FDA4:7733B7DAFFF1AD6EFD8874A44F9EFE61" box="[613,705,397,419]" country="Namibia" county="Dune" latitude="-23.766666" longLatPrecision="1254" longitude="15.783334" municipality="Aug" name="Praetorius" pageId="21" pageNumber="452">Praetorius</location>
8
<collectingMunicipality box="[726,765,397,418]" pageId="21" pageNumber="452">Aug</collectingMunicipality>
82, ~
<collectingCounty box="[833,886,397,418]" pageId="21" pageNumber="452">Dune</collectingCounty>
<quantity box="[891,943,396,418]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="452" unit="km" value="15.0">15km</quantity>
/
<locationDeviation box="[960,1141,397,419]" location="The Far East" pageId="21" pageNumber="452">NW of The Far East</locationDeviation>
[
<geoCoordinate box="[187,261,420,442]" degrees="23" direction="south" minutes="46" orientation="latitude" pageId="21" pageNumber="452" precision="925" value="-23.766666">23°46'S</geoCoordinate>
:
<geoCoordinate box="[265,340,420,442]" degrees="15" direction="east" minutes="47" orientation="longitude" pageId="21" pageNumber="452" precision="925" value="15.783334">15°47'E</geoCoordinate>
] /
<date box="[364,425,420,442]" pageId="21" pageNumber="452" value="1650-01">
<collectingDate box="[364,425,420,442]" pageId="21" pageNumber="452" value="1650-01">I.1650</collectingDate>
</date>
(
<collectionCode box="[440,511,421,442]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="21" pageNumber="452" type="Museum">NMSA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="21.[181,1142,204,614]" pageId="21" pageNumber="452">
Distribution and phenology (Table 3): Known only from the
<typeStatus box="[888,936,455,481]" pageId="21" pageNumber="452">type</typeStatus>
locality (
<figureCitation box="[1049,1133,454,480]" captionStart="Fig" captionStartId="49.[181,214,861,883]" captionTargetBox="[320,1014,266,831]" captionTargetId="figure-45@49.[319,1016,255,840]" captionTargetPageId="49" captionText="Fig. 66. Distribution of Acnephalomyia species: A. dorsalis (Macquart, 1838) circles; A. eremia sp. n. triangle." figureDoi="http://doi.org/10.5281/zenodo.7913807" httpUri="https://zenodo.org/record/7913807/files/figure.png" pageId="21" pageNumber="452">Fig. 66</figureCitation>
) where it was collected in August.
</paragraph>
<paragraph blockId="21.[181,1142,204,614]" pageId="21" pageNumber="452">
Similar species: Male genitalia superficially similar to
<taxonomicName authorityName="Londt" authorityYear="2010" baseAuthorityName="Macquart" baseAuthorityYear="1838" box="[785,875,524,550]" class="Insecta" family="Asilidae" genus="Acnephalomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="452" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis box="[785,875,524,550]" italics="true" pageId="21" pageNumber="452">dorsalis</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Londt" authorityYear="2010" box="[888,983,524,550]" class="Insecta" family="Asilidae" genus="Acnephalomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="452" phylum="Arthropoda" rank="species" species="leukoros">
<emphasis box="[888,983,524,550]" italics="true" pageId="21" pageNumber="452">leukoros</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Londt" authorityYear="2010" box="[997,1096,524,550]" class="Insecta" family="Asilidae" genus="Acnephalomyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="452" phylum="Arthropoda" rank="species" species="probolos">
<emphasis box="[997,1096,524,550]" italics="true" pageId="21" pageNumber="452">probolos</emphasis>
</taxonomicName>
and
<taxonomicName authorityName=": Loew" authorityYear="1858" box="[181,307,556,582]" class="Insecta" family="Asilidae" genus="Acnephalum" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="452" phylum="Arthropoda" rank="species" species="platygaster">
<emphasis box="[181,307,556,582]" italics="true" pageId="21" pageNumber="452">platygaster</emphasis>
</taxonomicName>
, but shape of gonostylus in lateral view, and especially the distal extremity of outer lobe, is unique as is the shape of the hypandrium.
</paragraph>
</subSubSection>
</treatment>
</document>