treatments-xml/data/29/6C/61/296C61683D0036743DC46CA5232736C4.xml

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<mods:title id="83D9B886C65275961A621A028BEDA1DD">A revision of the new world stiletto fly genus Ataenogera Kröber (Diptera: Therevidae: Phycinae) with the description of two new species</mods:title>
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<mods:namePart id="ACD480308319D237490AA104D8AEC461">Hauser, Martin</mods:namePart>
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<mods:namePart id="796CB50B1CD00BEFF39FC334C8B7C6E3">Webb, Donald W.</mods:namePart>
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<subSubSection id="E9DF83F53D0036763DC46CA5216633C2" pageId="16" pageNumber="57" type="nomenclature">
<paragraph id="A17AD07E3D0036763DC46CA5220333A7" blockId="16.[151,517,391,452]" box="[151,517,391,417]" pageId="16" pageNumber="57">
<heading id="FA3267123D0036763DC46CA5220333A7" bold="true" box="[151,517,391,417]" fontSize="11" level="1" pageId="16" pageNumber="57" reason="1">
<taxonomicName id="66C5ABFD3D0036763DC46CA5220333A7" ID-CoL="J7VM" authority="Lyneborg" authorityName="Lyneborg" box="[151,517,391,417]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="93B10C6C3D0036763DC46CA5220333A7" bold="true" box="[151,517,391,417]" pageId="16" pageNumber="57">
<emphasis id="93B10C6C3D0036763DC46CA5218333A7" bold="true" box="[151,389,391,417]" italics="true" pageId="16" pageNumber="57">Ataenogera grandis</emphasis>
Lyneborg
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A17AD07E3D0036763DC46C88216633C2" blockId="16.[151,517,391,452]" box="[151,352,426,452]" pageId="16" pageNumber="57">
(
<figureCitation id="39FECCFB3D0036763DCC6C88212B33C2" box="[159,301,426,452]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="16" pageNumber="57">Figs. 30–38</figureCitation>
, 
<figureCitation id="39FECCFB3D0036763C696C88215F33C2" box="[314,345,426,452]" captionStart="FIGURE 56" captionStartId="23.[151,255,1614,1638]" captionTargetBox="[260,1323,172,1590]" captionTargetId="figure@23.[258,1329,167,1590]" captionTargetPageId="23" captionText="FIGURE 56. Distribution map; Ataenogera abdominalis (closed circle), A. grandis (open circle), A. irwini (closed square), A. minuta (asterisk)." httpUri="https://zenodo.org/record/177673/files/figure.png" pageId="16" pageNumber="57">56</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E9DF83F53D0036763DC46CD823773037" pageId="16" pageNumber="57" type="description">
<paragraph id="A17AD07E3D0036763DC46CD823773037" blockId="16.[151,1435,506,561]" pageId="16" pageNumber="57">
<taxonomicName id="66C5ABFD3D0036763DC46CD821523017" authority="Lyneborg" authorityName="Lyneborg" box="[151,340,506,529]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="93B10C6C3D0036763DC46CD820EF3017" box="[151,233,506,529]" italics="true" pageId="16" pageNumber="57">grandis</emphasis>
<bibRefCitation id="C554AD8F3D0036763DBD6CD821523017" author="Lyneborg" box="[238,340,506,529]" pageId="16" pageNumber="66" refString="Lyneborg, L. (2002) Two new species of Ataenogera Krober from South America (Dipt., Therevidae). Entomologist's Monthly Magazine, 138, 103 - 108." type="journal article" year="2002">Lyneborg</bibRefCitation>
</taxonomicName>
(2002:104, fem. desc., key; 2002:105, 
<figureCitation id="39FECCFB3D0036763FA76CD823573017" box="[756,849,506,529]" captionStart="FIGURES 1 – 9" captionStartId="5.[151,269,1669,1693]" captionTargetBox="[271,1340,349,1654]" captionTargetId="figure@5.[227,1360,307,1664]" captionTargetPageId="5" captionText="FIGURES 1 – 9. Ataenogera abdominalis. 1. male frons. 2. female frons. 3. epandrium, cerci, hypoproct, ventral view. 4. hypandrium, gonocoxite, gonostylus, ventral view. 5. aedeagus, gonocoxite, gonostylus, dorsal view. 6. aedeagus dorsal view. 7. aedeagus ventral view. 8. aedeagus, lateral view. 9. female reproductive organs. Scale = 0.1 mm. Abbreviations: C = cercus, CSpD = common spermathecal duct, DA = dorsal apodeme, EA = ejaculatory apodeme, Epa = epandrium, F = furca, Gx = gonocoxite, GxA = gonocoxal apodeme, Gs = gonostylus, Hpt = hypoproct, H = hypandrium, Dp = distiphallus, Sp = spermatheca, VA = ventral apodeme." httpUri="https://zenodo.org/record/177667/files/figure.png" pageId="16" pageNumber="57">Figs.1–2</figureCitation>
fem. ant., max. palp.) (
<taxonomicName id="66C5ABFD3D003676391A6CD824C33017" box="[1097,1221,506,529]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="93B10C6C3D003676391A6CD824C33017" box="[1097,1221,506,529]" italics="true" pageId="16" pageNumber="57">Ataenogera</emphasis>
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). 
<typeStatus id="7E7E6EDC3D00367639886CD825163017" box="[1243,1296,506,529]" pageId="16" pageNumber="57">Type</typeStatus>
locality 
<collectingCountry id="D9D290EE3D003676383F6CD820E23037" name="Brazil" pageId="16" pageNumber="57">Brazil</collectingCountry>
, Itatiaya. 
<typeStatus id="7E7E6EDC3D0036763C1F6F3821A93037" box="[332,431,538,561]" pageId="16" pageNumber="57" type="holotype">Holotype</typeStatus>
female in USNM by original designation.
</paragraph>
</subSubSection>
<subSubSection id="E9DF83F53D0036763DC46F40213331BA" pageId="16" pageNumber="57" type="diagnosis">
<paragraph id="A17AD07E3D0036763DC46F40213331BA" blockId="16.[151,1437,610,1999]" pageId="16" pageNumber="57">
<emphasis id="93B10C6C3D0036763DC46F402111307A" bold="true" box="[151,279,610,636]" pageId="16" pageNumber="57">Diagnosis.</emphasis>
<taxonomicName id="66C5ABFD3D0036763C4D6F402201307D" box="[286,519,610,635]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="93B10C6C3D0036763C4D6F402201307D" box="[286,519,610,635]" italics="true" pageId="16" pageNumber="57">Ataenogera grandis</emphasis>
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shares with 
<emphasis id="93B10C6C3D0036763FC96F4023C0307D" box="[666,966,610,635]" italics="true" pageId="16" pageNumber="57">
<taxonomicName id="66C5ABFD3D0036763FC96F40234A307D" box="[666,844,610,635]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="argentifrons">A. argentifrons</taxonomicName>
, 
<taxonomicName id="66C5ABFD3D0036763E046F4023C7307D" box="[855,961,610,635]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="irwini">A. irwini</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="66C5ABFD3D0036763EAD6F402476307D" box="[1022,1136,610,635]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="93B10C6C3D0036763EAD6F402476307D" box="[1022,1136,610,635]" italics="true" pageId="16" pageNumber="57">A. minuta</emphasis>
</taxonomicName>
the absence of lanceolate setae on the male and female abdomens. It differs from 
<taxonomicName id="66C5ABFD3D0036763E756FA823D030A5" box="[806,982,650,675]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="argentifrons">
<emphasis id="93B10C6C3D0036763E756FA823D030A5" box="[806,982,650,675]" italics="true" pageId="16" pageNumber="57">A. argentifrons</emphasis>
</taxonomicName>
in having the frons (
<figureCitation id="39FECCFB3D00367639996FA8255E30A2" box="[1226,1368,650,676]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="16" pageNumber="57">Figs. 30–31</figureCitation>
) convergent dorsally, dark reddish brown, glossy, pubescence gray ventral to ocellar tubercle and white, dense dorsal and ventral to antennal bases with setae dorsolateral to the antennal bases; the coxal macrosetae dark reddish brown; the male gonocoxite lacking a pair of large falcate projections posterolaterally and having the anterior margin of the female furca (
<figureCitation id="39FECCFB3D0036763F196E0822A63142" box="[586,672,810,836]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="16" pageNumber="57">Fig. 38</figureCitation>
) open. It differs from 
<taxonomicName id="66C5ABFD3D0036763EFF6E0824123145" box="[940,1044,810,835]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="93B10C6C3D0036763EFF6E0824123145" box="[940,1044,810,835]" italics="true" pageId="16" pageNumber="57">A. irwini</emphasis>
</taxonomicName>
in having the anepisternum covered with gray pubescence and with setae over the entire anepisternum. It differs from 
<taxonomicName id="66C5ABFD3D00367639E96E702529316D" box="[1210,1327,850,875]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="93B10C6C3D00367639E96E702529316D" box="[1210,1327,850,875]" italics="true" pageId="16" pageNumber="57">A. minuta</emphasis>
</taxonomicName>
in being much larger; possessing two pairs of dorsocentral macrosetae; and the male distiphallus (
<figureCitation id="39FECCFB3D00367639FF6E5825073192" box="[1196,1281,890,916]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="16" pageNumber="57">Fig. 37</figureCitation>
) with a large dorsal carina.
</paragraph>
</subSubSection>
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<emphasis id="93B10C6C3D0036763D956EE821BC31E2" bold="true" box="[198,442,970,996]" pageId="16" pageNumber="57">Description of male</emphasis>
</paragraph>
<paragraph id="A17AD07E3D0036763D956ED0217D360A" blockId="16.[151,1437,610,1999]" box="[198,379,1010,1036]" pageId="16" pageNumber="57">Variation (n=2)</paragraph>
<paragraph id="A17AD07E3D0036763D95693822203632" blockId="16.[151,1437,610,1999]" box="[198,550,1050,1076]" pageId="16" pageNumber="57">
Body length 8.2–8.4, 
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.
</paragraph>
<paragraph id="A17AD07E3D0036763D95696024ED37C2" blockId="16.[151,1437,610,1999]" pageId="16" pageNumber="57">
<emphasis id="93B10C6C3D0036763D9569602116365A" bold="true" box="[198,272,1090,1116]" pageId="16" pageNumber="57">Head.</emphasis>
Head length 0.74–0.76, 
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. Eyes dichoptic (
<figureCitation id="39FECCFB3D0036763E39696023C6365A" box="[874,960,1090,1116]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="16" pageNumber="57">Fig. 30</figureCitation>
) separated by distance equal to width of ocellar tubercle; ommatidia of equal size. Frons (
<figureCitation id="39FECCFB3D0036763FB8694823423682" box="[747,836,1130,1156]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="16" pageNumber="57">Fig. 30</figureCitation>
) convergent dorsally, dark reddish brown, glossy, pubescence gray ventral to ocellar tubercle and white, dense dorsal and ventral to antennal bases; setae white, short dorsolateral to antennal base; area lateral to antennal bases lacking black pubescent band or spot. Antenna dark brown, pubescence light gray on scape and pedicel, antenna/head length 1.7; scape cylindrical, length 0.24–0.30, 
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, width 
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, length/width 2.0–2.5, 2.25, scape/pedicel length 2.4–3.0, 2.7, scape/flagellum width 0.86, setae dark reddish brown, short, macrosetae dark reddish brown; pedicel length 
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, width 0.12–0.14, 
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, length/width 0.71; flagellum length 
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, width 
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, length/ width 6.6, flagellum/scape length 3.8. Maxillary palpus dark reddish brown; setae white. Occipital setae white, absent dorsomedially but extending across dorsal margin of occiput, macrosetae absent.
</paragraph>
<paragraph id="A17AD07E3D0036763D9568F022E83501" blockId="16.[151,1437,610,1999]" pageId="16" pageNumber="57">
<emphasis id="93B10C6C3D0036763D9568F0212C37EA" bold="true" box="[198,298,1490,1516]" pageId="16" pageNumber="57">Thorax.</emphasis>
Macrosetae 2 np, 1 sa, 1 pa, 2 dc, 1 sc. Mesonotum black, glossy; vittae indistinct; setae white, short, erect and suberect. Pleurae dark reddish brown, pubescence white, anepimeron glossy; setae white over entire anepisternum and laterotergite. Scutellum dark reddish brown, glossy; setae absent. 
<emphasis id="93B10C6C3D00367639916B012502343A" box="[1218,1284,1571,1596]" italics="true" pageId="16" pageNumber="57">Wing.</emphasis>
Length 5.3– 5.5, 
<quantity id="663D7D9B3D0036763D9D6B6821363462" box="[206,304,1610,1636]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="16" pageNumber="57" unit="mm" value="5.4">5.4 mm</quantity>
, width 1.7–2.0, 
<quantity id="663D7D9B3D0036763CA96B68226F3462" box="[506,617,1610,1636]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="16" pageNumber="57" unit="mm" value="1.85">1.85 mm</quantity>
, length/width 2.7–3.2, 2.95. Membrane hyaline; veins dark brown; pterostigma dark brown. Cell m3 closed, petiolate. Halter dark brown. 
<emphasis id="93B10C6C3D0036763E9C6B512408348A" box="[975,1038,1651,1676]" italics="true" pageId="16" pageNumber="57">Legs.</emphasis>
Coxa dark reddish brown, pubescence white, dense; setae white; apical macrosetae dark reddish brown, not extending around anteroventral margin, anteromedial macroseta absent on forecoxa, posterolateral macroseta absent of hindcoxa. Femora, tibiae, and tarsi dark reddish brown, midfemora paler.
</paragraph>
<paragraph id="A17AD07E3D0036773D956A3724B2332C" blockId="16.[151,1437,610,1999]" lastBlockId="17.[151,1436,152,298]" lastPageId="17" lastPageNumber="58" pageId="16" pageNumber="57">
<emphasis id="93B10C6C3D0036763D956A37214E3529" bold="true" box="[198,328,1813,1839]" pageId="16" pageNumber="57">Abdomen.</emphasis>
Dark reddish brown, glossy, posterior margin white on tergites 1–6; dorsal setae dark brown, short with white elongate setae dorsally on tergites 1–2 and across posterior margins of tergites 1–4; tergite 7 and sternites 5–8 lacking lanceolate setae. 
<taxonomicName id="66C5ABFD3D0036763FCA6A44231F3579" box="[665,793,1894,1919]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="16" pageNumber="57" phylum="Tracheophyta" rank="genus">
<emphasis id="93B10C6C3D0036763FCA6A44231F3579" box="[665,793,1894,1919]" italics="true" pageId="16" pageNumber="57">Terminalia</emphasis>
</taxonomicName>
(MEI 147884). Epandrium (
<figureCitation id="39FECCFB3D00367639206A4724CB3579" box="[1139,1229,1893,1919]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="16" pageNumber="57">Fig. 32</figureCitation>
), anterior margin truncate with median emargination; setae dark reddish brown. Hypoproct ventral view (
<figureCitation id="39FECCFB3D00367639C16AAF24E135A1" box="[1170,1255,1933,1959]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="16" pageNumber="57">Fig. 32</figureCitation>
) attached laterally to sides of epandrium; setae yellow, scattered. Hypandrium (
<figureCitation id="39FECCFB3D0036763EDC6A9723E235C9" box="[911,996,1973,1999]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="16" pageNumber="57">Fig. 33</figureCitation>
) glabrous. Gonocoxite (
<figureCitation id="39FECCFB3D00367638526A97255035C9" box="[1281,1366,1973,1999]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="16" pageNumber="57">Fig. 33</figureCitation>
) rectangular, sides broadly rounded, posterolateral corner short, not extended posteriorly as outer gonocoxal process, setae brown. Gonostylus dorsal view (
<figureCitation id="39FECCFB3D0136773F836DE2232F32DC" box="[720,809,192,218]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="17" pageNumber="58">Fig. 34</figureCitation>
) broad with two falcate projections. Aedeagus with dorsal apodeme (
<figureCitation id="39FECCFB3D0136773C336DCA21BE3304" box="[352,440,232,258]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="17" pageNumber="58">Fig. 35</figureCitation>
), oval; ventral apodeme (
<figureCitation id="39FECCFB3D0136773FB06DCA233C3304" box="[739,826,232,258]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="17" pageNumber="58">Fig. 36</figureCitation>
) broad, rounded anteriorly; distiphallus dorsal view (
<figureCitation id="39FECCFB3D0136773DCC6C3220F2332C" box="[159,244,272,298]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="17" pageNumber="58">Fig. 35</figureCitation>
) with narrow dorsal carina, lateral view (
<figureCitation id="39FECCFB3D0136773F8A6C322329332C" box="[729,815,272,298]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="17" pageNumber="58">Fig. 37</figureCitation>
) sinuate with large dorsal carina.
</paragraph>
<caption id="F5BA80F63D0136773DC46B82248634FB" httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="17" pageNumber="58" targetBox="[307,1264,355,1661]" targetPageId="17">
<paragraph id="A17AD07E3D0136773DC46B82248634FB" blockId="17.[151,1437,1696,1789]" pageId="17" pageNumber="58">
<emphasis id="93B10C6C3D0136773DC46B82215834BE" bold="true" box="[151,350,1696,1720]" pageId="17" pageNumber="58">FIGURES 30–38.</emphasis>
<emphasis id="93B10C6C3D0136773C366B83224434BE" box="[357,578,1697,1720]" italics="true" pageId="17" pageNumber="58">
<taxonomicName id="66C5ABFD3D0136773C366B83223834BE" box="[357,574,1697,1720]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="grandis">Ataenogera grandis</taxonomicName>
.
</emphasis>
30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm.
</paragraph>
</caption>
<paragraph id="A17AD07E3D0136773D956A7A21E83574" blockId="17.[198,633,1880,2027]" box="[198,494,1880,1906]" pageId="17" pageNumber="58">
<emphasis id="93B10C6C3D0136773D956A7A21E83574" bold="true" box="[198,494,1880,1906]" pageId="17" pageNumber="58">Redescription of female</emphasis>
</paragraph>
<paragraph id="A17AD07E3D0136773D956AA3218435C5" blockId="17.[198,633,1880,2027]" pageId="17" pageNumber="58">Similar to male except for following. Variation (n=3).</paragraph>
<paragraph id="A17AD07E3D0136773D956AF3224B35ED" blockId="17.[198,633,1880,2027]" box="[198,589,2001,2027]" pageId="17" pageNumber="58">Body Length 9.6–10.5, 10.0 mm.</paragraph>
<paragraph id="A17AD07E3D0236743D956DB521FE33A4" blockId="18.[151,1437,151,1218]" pageId="18" pageNumber="59">
<emphasis id="93B10C6C3D0236743D956DB5211632B7" bold="true" box="[198,272,151,177]" pageId="18" pageNumber="59">Head.</emphasis>
Length 0.96–1.02, 1.00 mm. Eyes (
<figureCitation id="39FECCFB3D0236743F986DBA232532B4" box="[715,803,152,178]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="18" pageNumber="59">Fig. 31</figureCitation>
) separated by distance greater than width of ocellar tubercle. Frons (
<figureCitation id="39FECCFB3D0236743C056DE221AC32DC" box="[342,426,192,218]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="18" pageNumber="59">Fig. 31</figureCitation>
) glossy, pubescence whitish gray, dense dorsal and ventral to base of antenna and with an area of gray pubescence ventrolateral to ocellar tubercle; setae sparse. Antenna/head length 1.7–1.9, 1.8; scape length 0.24–0.28, 
<quantity id="663D7D9B3D0236743CE56C322224332C" box="[438,546,272,298]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="18" pageNumber="59" unit="mm" value="0.26">0.26 mm</quantity>
, width 0.16–0.19, 
<quantity id="663D7D9B3D0236743FAE6C32236C332C" box="[765,874,272,298]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="18" pageNumber="59" unit="mm" value="0.18">0.18 mm</quantity>
, length/width 1.4–1.6, 1.5, scape/pedicel length 2.4–2.8, 2.6, scape/flagellum width 1.0–1.2, 1.1; pedicel length 
<quantity id="663D7D9B3D0236743EDA6C1A23F03354" box="[905,1014,312,338]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="59" unit="mm" value="0.1">0.10 mm</quantity>
, width 0.16–0.18, 
<quantity id="663D7D9B3D02367439816C1A25383354" box="[1234,1342,312,338]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="59" unit="mm" value="0.17">0.17 mm</quantity>
, length/ width 0.56–0.63, 0.60; flagellum length 1.38–1.60, 1.49, width 
<quantity id="663D7D9B3D0236743EDE6C4223FC337C" box="[909,1018,352,378]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="59" unit="mm" value="0.16">0.16 mm</quantity>
, length/width 8.6–10.0, 9.3, flagellum/scape length 5.8–6.2, 6.0.
</paragraph>
<paragraph id="A17AD07E3D0236743D956C8D234C3044" blockId="18.[151,1437,151,1218]" pageId="18" pageNumber="59">
<emphasis id="93B10C6C3D0236743D956C8D212C33CF" bold="true" box="[198,298,431,457]" pageId="18" pageNumber="59">Thorax.</emphasis>
Macrosetae 2 np, 1 sa, 1 pa, 2 dc, 1 sc. Mesonotum with light covering of gray pubescence; setae black, short. Anepisternum dark reddish brown, glossy with medial dorsoventral band of white pubescence. 
<emphasis id="93B10C6C3D0236743DC46F2220DF301F" box="[151,217,512,537]" italics="true" pageId="18" pageNumber="59">Wing.</emphasis>
Length 
<quantity id="663D7D9B3D0236743C136F222198301C" box="[320,414,512,538]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="18" pageNumber="59" unit="mm" value="8.8">8.8 mm</quantity>
, width 2.3–2.6, 
<quantity id="663D7D9B3D0236743F316F2222C7301C" box="[610,705,512,538]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="59" unit="mm" value="2.5">2.5 mm</quantity>
, length/width 3.4–3.8, 3.6. Membrane pale brown, becoming darker apically. 
<emphasis id="93B10C6C3D0236743C016F0A21973047" box="[338,401,552,577]" italics="true" pageId="18" pageNumber="59">Legs.</emphasis>
Femora dark reddish brown to black.
</paragraph>
<paragraph id="A17AD07E3D0236743D956F6D22EA30E4" blockId="18.[151,1437,151,1218]" pageId="18" pageNumber="59">
<emphasis id="93B10C6C3D0236743D956F6D214E306F" bold="true" box="[198,328,591,617]" pageId="18" pageNumber="59">Abdomen.</emphasis>
Dark reddish brown, glossy, tergites 1–3 with posterior margin whitish yellow; dorsal setae dark brown, short, white elongate laterally on tergite 1 and across posterior margins of tergites 1–3; sternites 5–7 lacking dark brown lanceolate setae. 
<taxonomicName id="66C5ABFD3D0236743FD06F82230430BF" box="[643,770,672,697]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="18" pageNumber="59" phylum="Tracheophyta" rank="genus">
<emphasis id="93B10C6C3D0236743FD06F82230430BF" box="[643,770,672,697]" italics="true" pageId="18" pageNumber="59">Terminalia</emphasis>
</taxonomicName>
(MEI 155341). Furca (
<figureCitation id="39FECCFB3D02367439446F82246B30BC" box="[1047,1133,672,698]" captionStart="FIGURES 30 – 38" captionStartId="17.[151,269,1696,1720]" captionTargetBox="[307,1264,355,1661]" captionTargetId="figure@17.[307,1280,354,1672]" captionTargetPageId="17" captionText="FIGURES 30 – 38. Ataenogera grandis. 30. male frons. 31. female frons. 32. epandrium, cerci, hypoproct, ventral view. 33. hypandrium, gonocoxite, gonostylus, ventral view. 34. gonostylus, dorsal view. 35. aedeagus, dorsal view. 36. aedeagus, ventral view. 37. aedeagus, lateral view. 38. female reproductive organs. Scale = 0.1 mm." httpUri="https://zenodo.org/record/177670/files/figure.png" pageId="18" pageNumber="59">Fig. 38</figureCitation>
) broadly triangular, anterior margin open; length 
<quantity id="663D7D9B3D0236743CE86FEA222330E4" box="[443,549,712,738]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="59" unit="mm" value="0.5">0.50 mm</quantity>
, width 
<quantity id="663D7D9B3D0236743F2F6FEA22E130E4" box="[636,743,712,738]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="18" pageNumber="59" unit="mm" value="0.44">0.44 mm</quantity>
.
</paragraph>
<paragraph id="A17AD07E3D0236743D956FCD2191310F" blockId="18.[151,1437,151,1218]" box="[198,407,751,777]" pageId="18" pageNumber="59">
<emphasis id="93B10C6C3D0236743D956FCD2191310F" bold="true" box="[198,407,751,777]" pageId="18" pageNumber="59">Immature stages</emphasis>
</paragraph>
<paragraph id="A17AD07E3D0236743D956E3A21443134" blockId="18.[151,1437,151,1218]" box="[198,322,792,818]" pageId="18" pageNumber="59">Unknown.</paragraph>
</subSubSection>
<subSubSection id="E9DF83F53D0236743D956E1D22FB31AC" pageId="18" pageNumber="59" type="distribution">
<paragraph id="A17AD07E3D0236743D956E1D23C8315C" blockId="18.[151,1437,151,1218]" box="[198,974,831,858]" pageId="18" pageNumber="59">
<emphasis id="93B10C6C3D0236743D956E1D2161315F" bold="true" box="[198,359,831,857]" pageId="18" pageNumber="59">Distribution.</emphasis>
<taxonomicName id="66C5ABFD3D0236743C3D6E622251315F" box="[366,599,832,857]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="93B10C6C3D0236743C3D6E622251315F" box="[366,599,832,857]" italics="true" pageId="18" pageNumber="59">Ataenogera grandis</emphasis>
</taxonomicName>
is known from 
<collectingCountry id="D9D290EE3D0236743E426E62235F315C" box="[785,857,832,858]" name="Brazil" pageId="18" pageNumber="59">Brazil</collectingCountry>
(
<figureCitation id="39FECCFB3D0236743E3A6E6223C6315C" box="[873,960,832,858]" captionStart="FIGURE 56" captionStartId="23.[151,255,1614,1638]" captionTargetBox="[260,1323,172,1590]" captionTargetId="figure@23.[258,1329,167,1590]" captionTargetPageId="23" captionText="FIGURE 56. Distribution map; Ataenogera abdominalis (closed circle), A. grandis (open circle), A. irwini (closed square), A. minuta (asterisk)." httpUri="https://zenodo.org/record/177673/files/figure.png" pageId="18" pageNumber="59">Fig. 56</figureCitation>
).
</paragraph>
<paragraph id="A17AD07E3D0236743D956E4522FB31AC" blockId="18.[151,1437,151,1218]" pageId="18" pageNumber="59">
<emphasis id="93B10C6C3D0236743D956E4521F03187" bold="true" box="[198,502,871,897]" pageId="18" pageNumber="59">Habitats and phenology.</emphasis>
<taxonomicName id="66C5ABFD3D0236743CAE6E4A22E03187" box="[509,742,872,897]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="93B10C6C3D0236743CAE6E4A22E03187" box="[509,742,872,897]" italics="true" pageId="18" pageNumber="59">Ataenogera grandis</emphasis>
</taxonomicName>
has been handnetted or collected in a Malaise trap in January, February, July, August, October, and December.
</paragraph>
</subSubSection>
<subSubSection id="E9DF83F53D0236743D956E95232736C4" pageId="18" pageNumber="59" type="description">
<paragraph id="A17AD07E3D0236743D956E95232736C4" blockId="18.[151,1437,151,1218]" pageId="18" pageNumber="59">
<emphasis id="93B10C6C3D0236743D956E9521D531D7" bold="true" box="[198,467,951,977]" pageId="18" pageNumber="59">Specimens examined.</emphasis>
<emphasis id="93B10C6C3D0236743C8F6E9A229731D7" box="[476,657,952,977]" italics="true" pageId="18" pageNumber="59">
<typeStatus id="7E7E6EDC3D0236743C8F6E9A221531D7" box="[476,531,952,977]" pageId="18" pageNumber="59">Type</typeStatus>
specimen.
</emphasis>
The 
<typeStatus id="7E7E6EDC3D0236743F826E9A233E31D4" box="[721,824,952,978]" pageId="18" pageNumber="59" type="holotype">holotype</typeStatus>
female of 
<taxonomicName id="66C5ABFD3D0236743EEF6E9A252731D4" authority="Lyneborg" authorityName="Lyneborg" box="[956,1313,952,978]" class="Insecta" family="Therevidae" genus="Ataenogera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="93B10C6C3D0236743EEF6E9A24A131D7" box="[956,1191,952,977]" italics="true" pageId="18" pageNumber="59">Ataenogera grandis</emphasis>
Lyneborg
</taxonomicName>
is labeled &quot;Itatiaya– 
<quantity id="663D7D9B3D0236743C416EC2215131FC" box="[274,343,992,1018]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="59" unit="m" value="700.0">700m</quantity>
, E. Rio- 
<collectingCountry id="D9D290EE3D0236743C906EC2220831FC" box="[451,526,992,1018]" name="Brazil" pageId="18" pageNumber="59">Brazil</collectingCountry>
, 
<date id="D57BF6BE3D0236743F486EC2227831FC" box="[539,638,992,1018]" pageId="18" pageNumber="59" value="1934-01-07">7.i.1934</date>
, J. F. Zikán&quot; (MEI 147914) and is in the USNM. 
<emphasis id="93B10C6C3D023674399C6EC2259A31FF" box="[1231,1436,992,1017]" italics="true" pageId="18" pageNumber="59">Other specimens.</emphasis>
<emphasis id="93B10C6C3D0236743DC4692521223627" bold="true" box="[151,292,1031,1057]" pageId="18" pageNumber="59">AMAZONAS</emphasis>
. Parque Nacional Jau [-1.895, -61.586], 
<date id="D57BF6BE3D0236743E57692A23FD3624" box="[772,1019,1032,1058]" pageId="18" pageNumber="59" value="2001-07-29" valueMax="2001-08-08" valueMin="2001-07-29">29.VII–08.VIII.2001</date>
, Henriques, Vidal, 
<specimenCount id="B7C31BF73D023674398D692A24F9361D" box="[1246,1279,1032,1058]" pageId="18" pageNumber="59" type="generic">1♂</specimenCount>
MEI 147878 (INHS). 
<emphasis id="93B10C6C3D0236743C57690D21E4364F" bold="true" box="[260,482,1071,1097]" pageId="18" pageNumber="59">RIO de JANEIRO.</emphasis>
Itatiaya [-22.5, -44.571], 
<date id="D57BF6BE3D0236743E7F69122399364C" box="[812,927,1072,1098]" pageId="18" pageNumber="59" value="1939-01-16">16.I.1939</date>
, J. F. Zikán, 1Ψ MEI 155341 (USNM); 6. 
<date id="D57BF6BE3D0236743DF9697A21223674" box="[170,292,1112,1138]" pageId="18" pageNumber="59" value="1940-02-01">II.1940, 1</date>
Ψ MEI 155345 (USNM); 24. 
<date id="D57BF6BE3D0236743FD4697A22F33674" box="[647,757,1112,1138]" pageId="18" pageNumber="59" value="1945-01-01">I.1945, 1</date>
Ψ MEI 155342 (USNM); 26. 
<date id="D57BF6BE3D0236743905697A24E33674" box="[1110,1253,1112,1138]" pageId="18" pageNumber="59" value="1948-12-01">XII.1948, 1</date>
Ψ MEI 155343 (USNM). 
<emphasis id="93B10C6C3D0236743C5C695D219A369F" bold="true" box="[271,412,1151,1177]" pageId="18" pageNumber="59">RONDONIA.</emphasis>
Guajara-Mirim, Rio Ouro Preto, Bananal [-10.973, -65.094], 
<date id="D57BF6BE3D023674392869A22526369C" box="[1147,1312,1152,1178]" pageId="18" pageNumber="59" value="1995-10-20" valueMax="1995-10-27" valueMin="1995-10-20">20-27.X.1995</date>
, Rafael, J. A., Henriques, A. L., 
<specimenCount id="B7C31BF73D0236743CC6698A21B336BD" box="[405,437,1192,1218]" pageId="18" pageNumber="59" type="generic">1♂</specimenCount>
MEI 147884 (INPA), Malaise.
</paragraph>
</subSubSection>
</treatment>
</document>