233 lines
31 KiB
XML
233 lines
31 KiB
XML
<document id="6DECEFFB22246DC4C9F777024159C35A" ID-DOI="10.11646/zootaxa.3694.4.2" ID-GBIF-Dataset="8ce29893-89ba-4b0e-8941-287814ff4a8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="220826" ID-ZooBank="03DDE60B-89C5-4B20-B14F-21C6CF69BF15" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460658596926" checkinUser="plazi" docAuthor="Yin, Zi-Wei, Coulon, Georges & Li, Li-Zhen" docDate="2013" docId="03A2DE12975A713B34D9FE4377FAF996" docLanguage="en" docName="zt03694p342.pdf" docOrigin="Zootaxa 3694 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tmesiphodimerus Coulon and Yin, new genus" docType="treatment" docVersion="7" lastPageNumber="339" masterDocId="FF9BA66A975B7138344EFFD27679FFE3" masterDocTitle="Description of two new species of the genus Tmesiphodimerus gen. n. (Coleoptera: Staphylinidae: Pselaphinae) from South Asia" masterLastPageNumber="342" masterPageNumber="336" pageNumber="337" updateTime="1698347210820" updateUser="plazi">
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<mods:mods id="8CE44D82E5083068797576936CFBCA8E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6DEA4A11E12ADB2C093B5908E29175FD">
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<mods:title id="74A4846DE8A14BA245421C837107E7BA">Description of two new species of the genus Tmesiphodimerus gen. n. (Coleoptera: Staphylinidae: Pselaphinae) from South Asia</mods:title>
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<mods:name id="B7A2E896F1AA73EA491AFDC144B51A37" type="personal">
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<mods:role id="A514F772A24551EC3F4A5071E5323083">
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<mods:roleTerm id="CB7996EC0EC444E917B70447A21563F5">Author</mods:roleTerm>
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<mods:namePart id="EC7099C649B70ADB4C5F7098332CD439">Yin, Zi-Wei</mods:namePart>
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<mods:name id="54200DB5F216C022A3F493C1832663FB" type="personal">
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<mods:roleTerm id="E332C46CC329D0DDBE2C086DBCC6311E">Author</mods:roleTerm>
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<mods:namePart id="F5FC868EE53AF8FC263DAFFB99DA27DB">Coulon, Georges</mods:namePart>
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</mods:name>
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<mods:name id="4BD5A2B27E85B8BCD34747ABBC5C9397" type="personal">
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<mods:role id="67BBA775D082E5BA2B003E60861B7F1B">
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<mods:roleTerm id="245AE659856592A6FAFE5FF462E05F60">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="050AD5DB53B30141659F5082C98F6FF1">Li, Li-Zhen</mods:namePart>
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<mods:typeOfResource id="37E7D318C710987C726F59C4EFC56AAB">text</mods:typeOfResource>
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<mods:title id="A079E499C892F0C78BABADB0F186D11C">Zootaxa</mods:title>
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<mods:part id="06DE5E7471CAC876DD379A4ECBAB56F7">
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<mods:date id="8F0CFE49FD93BE02AE5FE9E01FA4C290">2013</mods:date>
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<mods:detail id="F3D43B50EED1205AF4907DD366BB66B2" type="volume">
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<mods:number id="6BBD531265ED6C26F677C8E5D99A7C80">3694</mods:number>
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<mods:detail id="0258F11CE4C62EE69EA644FC555B94DA" type="issue">
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<mods:number id="3A07E967D60467D0FB0AC5BE203D8F73">4</mods:number>
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<mods:start id="D14C0378993EBA0A809F668309274A05">336</mods:start>
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<mods:end id="95BEA9F8B918F309B0D5D9574A4516CB">342</mods:end>
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<mods:classification id="0DF4F1DEE865174DF0D42D647D4B8AC3">journal article</mods:classification>
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<mods:identifier id="B2F681DA6CA6237074263CB1136E4B50" type="DOI">10.11646/zootaxa.3694.4.2</mods:identifier>
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<mods:identifier id="CE142574012FE2CB51D4081C06EFF78E" type="GBIF-Dataset">8ce29893-89ba-4b0e-8941-287814ff4a8e</mods:identifier>
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<mods:identifier id="55F34D4FD6D435FB0DC27027BA1BB1D2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A297B5706FEAE3133681C2D9500727A6" type="Zenodo-Dep">220826</mods:identifier>
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<treatment id="03A2DE12975A713B34D9FE4377FAF996" ID-DOI="http://doi.org/10.5281/zenodo.6154984" ID-GBIF-Taxon="119574231" ID-Zenodo-Dep="6154984" LSID="urn:lsid:plazi:treatment:03A2DE12975A713B34D9FE4377FAF996" httpUri="http://treatment.plazi.org/id/03A2DE12975A713B34D9FE4377FAF996" lastPageId="3" lastPageNumber="339" pageId="1" pageNumber="337">
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<subSubSection id="C3113C8F975A713934D9FE437264FDF1" pageId="1" pageNumber="337" type="nomenclature">
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<paragraph id="8BB46F04975A713934D9FE4374B5FE48" blockId="1.[151,716,401,461]" box="[151,716,401,427]" pageId="1" pageNumber="337">
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<heading id="D0FCD868975A713934D9FE4374B5FE48" bold="true" box="[151,716,401,427]" fontSize="11" level="1" pageId="1" pageNumber="337" reason="1">
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<emphasis id="B97FB316975A713934D9FE4374B5FE48" bold="true" box="[151,716,401,427]" pageId="1" pageNumber="337">
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<taxonomicName id="4C0B1487975A713934D9FE437447FE48" ID-CoL="8XQDR" authority="Coulon and Yin" authorityName="Coulon and Yin" box="[151,574,401,427]" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="337" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B97FB316975A713934D9FE437716FE48" bold="true" box="[151,367,401,427]" italics="true" pageId="1" pageNumber="337">Tmesiphodimerus</emphasis>
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Coulon and Yin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24C0E6D975A71393605FE4374B5FE48" box="[587,716,401,427]" pageId="1" pageNumber="337" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB46F04975A713934D9FE667772FE2F" blockId="1.[151,716,401,461]" box="[151,267,436,461]" pageId="1" pageNumber="337">
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(
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<figureCitation id="13307381975A713934D1FE66777BFE2F" box="[159,258,436,461]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1941,1963]" captionStartId-1="2.[151,250,1903,1925]" captionStartId-2="4.[151,250,1917,1939]" captionStartId-3="5.[151,250,1940,1962]" captionTargetBox-0="[162,1429,771,1910]" captionTargetBox-1="[196,1336,196,1875]" captionTargetBox-2="[157,1416,666,1877]" captionTargetBox-3="[160,1427,797,1925]" captionTargetId-0="figure@1.[151,1436,762,1917]" captionTargetId-1="figure@2.[189,1341,193,1878]" captionTargetId-2="figure@4.[151,1436,639,1885]" captionTargetId-3="figure@5.[151,1436,788,1929]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Dorsal habitus of Tmesiphodimerus sinensis (A) and T. malaysianus (B). Scales (mm): 0.5." captionText-1="FIGURE 2. Diagnostic features of Tmesiphodimerus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Subantennal sulcus. E. Pronotum, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Left elytron. I. Meso- and metaventrite. J. Abdomen, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): J – L = 0.5; A – C, E – I = 0.2; D = 0.1." captionText-2="FIGURE 3. Diagnostic features of Tmesiphodimerus sinensis. A. Antenna. B. Maxillary palpus. C. Sternite IX. D. Aedeagus, in dorsal view. E. Same, in lateral view. F. Same, in ventral view. G. Parameres, in lateral view. H. Same, in ventral view. Scales (mm): A, D – F = 0.2; B, G, H = 0.1; C = 0.05." captionText-3="FIGURE 4. Diagnostic features of Tmesiphodimerus malaysianus. A. Antenna. B Subantennal sulcus. C. Maxillary palpus. D. Sternite IX. E. Aedeagus, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Parameres, in lateral view. I. Same, in ventral view. Scales (mm): A = 0.2; B, C, E – G = 0.1 mm; D, H, I = 0.05 mm." httpUri-0="https://zenodo.org/record/220827/files/figure.png" httpUri-1="https://zenodo.org/record/220828/files/figure.png" httpUri-2="https://zenodo.org/record/220829/files/figure.png" httpUri-3="https://zenodo.org/record/220830/files/figure.png" pageId="1" pageNumber="337">Figs 1–4</figureCitation>
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)
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</paragraph>
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<paragraph id="8BB46F04975A713934D9FE2F7264FDF1" blockId="1.[151,1053,508,530]" box="[151,1053,508,530]" pageId="1" pageNumber="337">
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<typeStatus id="54B0D1A6975A713934D9FE2F76B0FDF1" box="[151,201,509,530]" pageId="1" pageNumber="337">Type</typeStatus>
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species:
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<taxonomicName id="4C0B1487975A71393568FE2E74A0FDF1" authority="Yin and Coulon" authorityName="Yin and Coulon" box="[294,729,508,530]" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="337" phylum="Arthropoda" rank="species" species="sinensis" status="sp. nov.">
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<emphasis id="B97FB316975A71393568FE2E7449FDF1" box="[294,560,508,530]" italics="true" pageId="1" pageNumber="337">Tmesiphodimerus sinensis</emphasis>
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Yin and Coulon
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24C0E6D975A713936AAFE2E7519FDF1" box="[740,864,508,530]" pageId="1" pageNumber="337" rank="species">
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<emphasis id="B97FB316975A713936AAFE2E7519FDF1" bold="true" box="[740,864,508,530]" pageId="1" pageNumber="337">new species</emphasis>
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</taxonomicNameLabel>
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(here designated).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3113C8F975A713934D9FDEC747EFD20" pageId="1" pageNumber="337" type="diagnosis">
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<paragraph id="8BB46F04975A713934D9FDEC747EFD20" blockId="1.[151,1437,574,707]" pageId="1" pageNumber="337">
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<emphasis id="B97FB316975A713934D9FDEC7768FDB4" bold="true" box="[151,273,574,599]" pageId="1" pageNumber="337">Diagnosis.</emphasis>
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Head with small vertexal foveae, lacking frontal fovea; lateral branches of U-shaped cephalic sulci interrupted by an ocular plate (here termed) starting from the inner margin of the eyes. Maxillary palpomeres II and III each bearing an apicolateral tuft of short setae. Pronotum with small median and lateral antebasal foveae. Abdomen with tergite V longest.
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</paragraph>
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</subSubSection>
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<caption id="DF743F8C975A713934D9F84772CBF848" box="[151,1202,1941,1963]" httpUri="https://zenodo.org/record/220827/files/figure.png" pageId="1" pageNumber="337" targetBox="[162,1429,771,1910]" targetPageId="1">
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<paragraph id="8BB46F04975A713934D9F84772CBF848" blockId="1.[151,1202,1941,1963]" box="[151,1202,1941,1963]" pageId="1" pageNumber="337">
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<emphasis id="B97FB316975A713934D9F847776AF849" bold="true" box="[151,275,1941,1963]" pageId="1" pageNumber="337">FIGURE 1.</emphasis>
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Dorsal habitus of
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<taxonomicName id="4C0B1487975A71393583F84774A1F848" box="[461,728,1941,1963]" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="337" phylum="Arthropoda" rank="species" species="sinensis">
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<emphasis id="B97FB316975A71393583F84774A1F848" box="[461,728,1941,1963]" italics="true" pageId="1" pageNumber="337">Tmesiphodimerus sinensis</emphasis>
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</taxonomicName>
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(
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<emphasis id="B97FB316975A713936A9F8477480F849" bold="true" box="[743,761,1941,1962]" pageId="1" pageNumber="337">A</emphasis>
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) and
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<taxonomicName id="4C0B1487975A7139377DF84475B3F848" box="[819,970,1941,1963]" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="337" phylum="Arthropoda" rank="species" species="malaysianus">
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<emphasis id="B97FB316975A7139377DF84475B3F848" box="[819,970,1941,1963]" italics="true" pageId="1" pageNumber="337">T. malaysianus</emphasis>
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</taxonomicName>
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(
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<emphasis id="B97FB316975A71393797F8477593F849" bold="true" box="[985,1002,1941,1962]" pageId="1" pageNumber="337">B</emphasis>
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). Scales (mm): 0.5.
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</paragraph>
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</caption>
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<caption id="DF743F8C9759713A34D9F8BD77B9F802" httpUri="https://zenodo.org/record/220828/files/figure.png" pageId="2" pageNumber="338" targetBox="[196,1336,196,1875]" targetPageId="2">
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<paragraph id="8BB46F049759713A34D9F8BD77B9F802" blockId="2.[151,1436,1903,2017]" pageId="2" pageNumber="338">
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<emphasis id="B97FB3169759713A34D9F8BD776DF867" bold="true" box="[151,276,1903,1925]" pageId="2" pageNumber="338">FIGURE 2.</emphasis>
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Diagnostic features of
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<taxonomicName id="4C0B14879759713A3649F8BD74C2F866" box="[519,699,1903,1925]" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B97FB3169759713A3649F8BD74C2F866" box="[519,699,1903,1925]" italics="true" pageId="2" pageNumber="338">Tmesiphodimerus</emphasis>
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</taxonomicName>
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.
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<emphasis id="B97FB3169759713A3687F8BD749BF867" bold="true" box="[713,738,1903,1924]" pageId="2" pageNumber="338">A.</emphasis>
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Head, in dorsal view.
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<emphasis id="B97FB3169759713A3782F8BD759DF867" bold="true" box="[972,996,1903,1924]" pageId="2" pageNumber="338">B.</emphasis>
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Same, in lateral view.
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<emphasis id="B97FB3169759713A309DF8BD7292F866" bold="true" box="[1235,1259,1903,1925]" pageId="2" pageNumber="338">C.</emphasis>
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Same, in ventral view.
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<emphasis id="B97FB3169759713A349DF85C7695F840" bold="true" box="[211,236,1934,1955]" pageId="2" pageNumber="338">D.</emphasis>
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Subantennal sulcus.
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<emphasis id="B97FB3169759713A358FF85C77A1F840" bold="true" box="[449,472,1934,1955]" pageId="2" pageNumber="338">E.</emphasis>
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Pronotum, in dorsal view.
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<emphasis id="B97FB3169759713A36A7F85C7483F840" bold="true" box="[745,762,1934,1955]" pageId="2" pageNumber="338">F.</emphasis>
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Same, in lateral view.
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<emphasis id="B97FB3169759713A37ADF85F758EF840" bold="true" box="[995,1015,1933,1955]" pageId="2" pageNumber="338">G.</emphasis>
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Same, in ventral view.
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<emphasis id="B97FB3169759713A30A9F85C7378F840" bold="true" box="[1255,1281,1934,1955]" pageId="2" pageNumber="338">H.</emphasis>
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Left elytron.
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<emphasis id="B97FB3169759713A31C2F85C73E2F840" bold="true" box="[1420,1435,1934,1955]" pageId="2" pageNumber="338">I.</emphasis>
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Meso- and metaventrite.
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<emphasis id="B97FB3169759713A35D9F87F77D3F821" bold="true" box="[407,426,1965,1986]" pageId="2" pageNumber="338">J.</emphasis>
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Abdomen, in dorsal view.
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<emphasis id="B97FB3169759713A368FF87F74A2F821" bold="true" box="[705,731,1965,1986]" pageId="2" pageNumber="338">K.</emphasis>
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Same, in lateral view.
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<emphasis id="B97FB3169759713A3789F87F75A7F821" bold="true" box="[967,990,1965,1986]" pageId="2" pageNumber="338">L.</emphasis>
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Same, in ventral view. Scales (mm): J–L = 0.5; A–C, E–I = 0.2; D = 0.1.
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</paragraph>
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</caption>
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<subSubSection id="C3113C8F9758713B3489FF4575E5F9EB" pageId="3" pageNumber="339" type="description">
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<paragraph id="8BB46F049758713B3489FF4574F5FE16" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="339">
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<emphasis id="B97FB3169758713B3489FF457720FF53" bold="true" box="[199,345,151,176]" pageId="3" pageNumber="339">Description.</emphasis>
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Head transverse (
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<figureCitation id="133073819758713B367AFF457404FF53" box="[564,637,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,250,1941,1963]" captionTargetBox="[162,1429,771,1910]" captionTargetId="figure@1.[151,1436,762,1917]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Tmesiphodimerus sinensis (A) and T. malaysianus (B). Scales (mm): 0.5." httpUri="https://zenodo.org/record/220827/files/figure.png" pageId="3" pageNumber="339">Figs 1</figureCitation>
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A, B), frontal rostrum short and wide, slightly prominent antennal tubercles separated by wide, shallow median sulcus; with nude, punctiform vertexal foveae, lacking frontal fovea (
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<figureCitation id="133073819758713B34D1FF0D769AFF1B" box="[159,227,223,248]" captionStart="FIGURE 2" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[196,1336,196,1875]" captionTargetId="figure@2.[189,1341,193,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tmesiphodimerus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Subantennal sulcus. E. Pronotum, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Left elytron. I. Meso- and metaventrite. J. Abdomen, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): J – L = 0.5; A – C, E – I = 0.2; D = 0.1." httpUri="https://zenodo.org/record/220828/files/figure.png" pageId="3" pageNumber="339">Fig. 2</figureCitation>
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A); lacking postantennal notches at lateral margins; lateral postantennal pits developed into wide setose Ushaped sulci ventral to antennal insertions (
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<figureCitation id="133073819758713B3639FED674C3FEFF" box="[631,698,260,285]" captionStart="FIGURE 2" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[196,1336,196,1875]" captionTargetId="figure@2.[189,1341,193,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tmesiphodimerus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Subantennal sulcus. E. Pronotum, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Left elytron. I. Meso- and metaventrite. J. Abdomen, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): J – L = 0.5; A – C, E – I = 0.2; D = 0.1." httpUri="https://zenodo.org/record/220828/files/figure.png" pageId="3" pageNumber="339">Fig. 2</figureCitation>
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B), sulci interrupted by broad, smooth ocular plates (
|
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<figureCitation id="133073819758713B314DFED67333FEFF" box="[1283,1354,260,285]" captionStart="FIGURE 2" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[196,1336,196,1875]" captionTargetId="figure@2.[189,1341,193,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tmesiphodimerus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Subantennal sulcus. E. Pronotum, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Left elytron. I. Meso- and metaventrite. J. Abdomen, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): J – L = 0.5; A – C, E – I = 0.2; D = 0.1." httpUri="https://zenodo.org/record/220828/files/figure.png" pageId="3" pageNumber="339">Figs 2</figureCitation>
|
||
D, 4B); lacking ocular-mandibular carinae; postgenae rounded laterally, with tufts of dense, thick setae; gula (
|
||
<figureCitation id="133073819758713B3174FEF573F9FEA3" box="[1338,1408,295,320]" captionStart="FIGURE 2" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[196,1336,196,1875]" captionTargetId="figure@2.[189,1341,193,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tmesiphodimerus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Subantennal sulcus. E. Pronotum, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Left elytron. I. Meso- and metaventrite. J. Abdomen, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): J – L = 0.5; A – C, E – I = 0.2; D = 0.1." httpUri="https://zenodo.org/record/220828/files/figure.png" pageId="3" pageNumber="339">Fig. 2</figureCitation>
|
||
C) slightly swollen, gular foveae in small transverse impression, lacking gular carina or sulcus. Antennae with 11 antennomeres, clubs formed by apical four antennomeres. Maxillary palpi (
|
||
<figureCitation id="133073819758713B306DFEBD7208FE6B" box="[1059,1137,367,392]" captionStart="FIGURE 3" captionStartId="4.[151,250,1917,1939]" captionTargetBox="[157,1416,666,1877]" captionTargetId="figure@4.[151,1436,639,1885]" captionTargetPageId="4" captionText="FIGURE 3. Diagnostic features of Tmesiphodimerus sinensis. A. Antenna. B. Maxillary palpus. C. Sternite IX. D. Aedeagus, in dorsal view. E. Same, in lateral view. F. Same, in ventral view. G. Parameres, in lateral view. H. Same, in ventral view. Scales (mm): A, D – F = 0.2; B, G, H = 0.1; C = 0.05." httpUri="https://zenodo.org/record/220829/files/figure.png" pageId="3" pageNumber="339">Figs 3</figureCitation>
|
||
B, 4C) moderately large, palpomeres I minute, II elongate, pedunculate in basal halves, slightly to strongly prolonged apicolaterally with apical tuft of short setae, lateral margins of palpomeres III strongly lobed laterally, with tuft of short setae; IV broadly rounded laterally; palpal cone fused.
|
||
</paragraph>
|
||
<paragraph id="8BB46F049758713B3489FDD275C5FDDE" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="339">
|
||
Pronotum (
|
||
<figureCitation id="133073819758713B350AFE2D77F0FDFB" box="[324,393,511,536]" captionStart="FIGURE 2" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[196,1336,196,1875]" captionTargetId="figure@2.[189,1341,193,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tmesiphodimerus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Subantennal sulcus. E. Pronotum, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Left elytron. I. Meso- and metaventrite. J. Abdomen, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): J – L = 0.5; A – C, E – I = 0.2; D = 0.1." httpUri="https://zenodo.org/record/220828/files/figure.png" pageId="3" pageNumber="339">Figs 2</figureCitation>
|
||
E–G) with small, nude lateral and median antebasal foveae; disc with broadly rounded hump. Prosternite lacking paranotal carinae; with deep lateral procoxal foveae.
|
||
</paragraph>
|
||
<paragraph id="8BB46F049758713B3489FD9574CFFD4B" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="339">
|
||
Each elytron (
|
||
<figureCitation id="133073819758713B3520FD9577CCFD83" box="[366,437,583,608]" captionStart="FIGURE 2" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[196,1336,196,1875]" captionTargetId="figure@2.[189,1341,193,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tmesiphodimerus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Subantennal sulcus. E. Pronotum, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Left elytron. I. Meso- and metaventrite. J. Abdomen, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): J – L = 0.5; A – C, E – I = 0.2; D = 0.1." httpUri="https://zenodo.org/record/220828/files/figure.png" pageId="3" pageNumber="339">Fig. 2</figureCitation>
|
||
H) with two large basal foveae and small subbasal fovea; shallow, broad discal stria extending posteriorly to nearly two-thirds elytral length; sutural stria complete; lacking subhumeral fovea and marginal stria; lateroapical cleft widely notched.
|
||
</paragraph>
|
||
<paragraph id="8BB46F049758713B3489FD667749FCDB" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="339">
|
||
Mesoventrite (
|
||
<figureCitation id="133073819758713B3525FD6677C9FD2F" box="[363,432,692,717]" captionStart="FIGURE 2" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[196,1336,196,1875]" captionTargetId="figure@2.[189,1341,193,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tmesiphodimerus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Subantennal sulcus. E. Pronotum, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Left elytron. I. Meso- and metaventrite. J. Abdomen, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): J – L = 0.5; A – C, E – I = 0.2; D = 0.1." httpUri="https://zenodo.org/record/220828/files/figure.png" pageId="3" pageNumber="339">
|
||
Fig.
|
||
<date id="FFB549C49758713B35EFFD6677C9FD2F" box="[417,432,692,716]" pageId="3" pageNumber="339">2</date>
|
||
</figureCitation>
|
||
I) with prepectal foveae widely separated, median mesoventral foveae merged into wide opening, widely forked and deeply inserted internally, lateral mesoventral foveae broad, reaching to lateral branches of median mesoventral foveae; lateral mesocoxal foveae present; metaventrite lacking foveae, apex with looped notch.
|
||
</paragraph>
|
||
<paragraph id="8BB46F049758713B3489FC977311FC63" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="339">Legs simple, lacking spines; profemora thickened, setose ventrally; protibiae thickened to middle, slightly arcuate at apical third; third tarsomeres longest, second tarsomeres two-thirds length of third ones; tarsal claws subequal.</paragraph>
|
||
<paragraph id="8BB46F049758713B3489FC5E77DBFB9E" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="339">
|
||
Abdomen (
|
||
<figureCitation id="133073819758713B350BFC5E77F5FC47" box="[325,396,908,933]" captionStart="FIGURE 2" captionStartId="2.[151,250,1903,1925]" captionTargetBox="[196,1336,196,1875]" captionTargetId="figure@2.[189,1341,193,1878]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Tmesiphodimerus. A. Head, in dorsal view. B. Same, in lateral view. C. Same, in ventral view. D. Subantennal sulcus. E. Pronotum, in dorsal view. F. Same, in lateral view. G. Same, in ventral view. H. Left elytron. I. Meso- and metaventrite. J. Abdomen, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales (mm): J – L = 0.5; A – C, E – I = 0.2; D = 0.1." httpUri="https://zenodo.org/record/220828/files/figure.png" pageId="3" pageNumber="339">Figs 2</figureCitation>
|
||
J–L) with tergite V (second visible tergite) longest; tergite IV with deep, setose basal sulcus forming large pockets at lateral endings, lacking mediobasal fovea, basolateral foveae in sockets, with median and discal carinae extending through tergal length; tergite V with basal sulcus shallower and thinner, basolateral foveae present, with short lateral carinae and complete median carina; tergites VI–VII with basolateral foveae. Sternite V (third visible sternite) longest, IV–V with deep, setose basal sulci, basolateral foveae developed as cuticular pockets; sternite IV with lateral carinae complete, V with carinae extending to two-thirds sternal length; IV–VII with basolateral foveae.
|
||
</paragraph>
|
||
<paragraph id="8BB46F049758713B3489FB5574E1FB26" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="339">Males with enlarged antennal clubs, modified or simple. Aedeagus with median lobe nearly symmetric; parameres symmetric; dorsal diaphragm oval.</paragraph>
|
||
<paragraph id="8BB46F049758713B3489FB1D77BEFA7E" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="339">
|
||
<emphasis id="B97FB3169758713B3489FB1D77C9FB0B" bold="true" box="[199,432,1231,1256]" pageId="3" pageNumber="339">Comparative notes.</emphasis>
|
||
<taxonomicName id="4C0B14879758713B35F9FB1D7407FB0B" box="[439,638,1231,1256]" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="336" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97FB3169758713B35F9FB1D7407FB0B" box="[439,638,1231,1256]" italics="true" pageId="3" pageNumber="339">Tmesiphodimerus</emphasis>
|
||
</taxonomicName>
|
||
belongs to a group of genera centered on
|
||
<taxonomicName id="4C0B14879758713B3018FB1D73E1FB0B" authority="LeConte, 1849" authorityName="LeConte" authorityYear="1849" box="[1110,1432,1231,1256]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FB3169758713B3018FB1D729EFB0B" box="[1110,1255,1231,1256]" italics="true" pageId="3" pageNumber="339">Tmesiphorus</emphasis>
|
||
LeConte, 1849
|
||
</taxonomicName>
|
||
, characterized by the maxillary palpomeres II and III each bearing a lateral projection with a small apical tuft of setae. In
|
||
<taxonomicName id="4C0B14879758713B3545FAC57483FAD3" authority="Tmesiphomimus Lobl, 1964" authorityName="Tmesiphomimus Lobl" authorityYear="1964" box="[267,762,1303,1328]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FB3169758713B3545FAC577E5FAD3" box="[267,412,1303,1328]" italics="true" pageId="3" pageNumber="339">Tmesiphorus</emphasis>
|
||
,
|
||
<emphasis id="B97FB3169758713B35FCFAC57411FAD3" box="[434,616,1303,1328]" italics="true" pageId="3" pageNumber="339">Tmesiphomimus</emphasis>
|
||
Löbl, 1964
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B14879758713B370FFAC5720FFAD3" authority="Raffray, 1908" authorityName="Raffray" authorityYear="1908" box="[833,1142,1303,1328]" class="Insecta" family="Staphylinidae" genus="Dacnotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FB3169758713B370FFAC575BDFAD3" box="[833,964,1303,1328]" italics="true" pageId="3" pageNumber="339">Dacnotillus</emphasis>
|
||
Raffray, 1908
|
||
</taxonomicName>
|
||
, these lateral penicillate appendages are long and thin while they are much shorter and conical in
|
||
<taxonomicName id="4C0B14879758713B3782FAEE72EAFAB6" box="[972,1171,1340,1365]" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="336" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97FB3169758713B3782FAEE72EAFAB6" box="[972,1171,1340,1365]" italics="true" pageId="3" pageNumber="339">Tmesiphodimerus</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C0B14879758713B30F1FAEE76ABFA9B" authority="Raffray, 1897" authorityName="Raffray" authorityYear="1897" class="Insecta" family="Staphylinidae" genus="Ctenotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FB3169758713B30F1FAEE7343FAB6" box="[1215,1338,1340,1365]" italics="true" pageId="3" pageNumber="339">Ctenotillus</emphasis>
|
||
Raffray, 1897
|
||
</taxonomicName>
|
||
, the pubescence is made of small, scale-like setae, the elytra bear longitudinal ridges and the first two visible tergites are equal in length.
|
||
</paragraph>
|
||
<paragraph id="8BB46F049758713B3489FA7575E5F9EB" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="339">
|
||
The most important feature that separates
|
||
<taxonomicName id="4C0B14879758713B36D3FA75751DFA23" box="[669,868,1447,1472]" class="Insecta" family="Staphylinidae" genus="Tmesiphodimerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="336" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B97FB3169758713B36D3FA75751DFA23" box="[669,868,1447,1472]" italics="true" pageId="3" pageNumber="339">Tmesiphodimerus</emphasis>
|
||
</taxonomicName>
|
||
from all other tmesiphorine genera is the presence of a smooth, shiny ocular plate extending from the inner margin of the eyes towards the antennal insertions, and therefore dividing the subantennal sulcus into two separate branches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3113C8F9758713B3489F9C677FAF996" pageId="3" pageNumber="339" type="etymology">
|
||
<paragraph id="8BB46F049758713B3489F9C677FAF996" blockId="3.[151,1436,151,1653]" pageId="3" pageNumber="339">
|
||
<emphasis id="B97FB3169758713B3489F9C67737F9CE" bold="true" box="[199,334,1556,1581]" pageId="3" pageNumber="339">Etymology.</emphasis>
|
||
The generic name is a combination of ‘Tmesiph’, a common stem used in the tribe
|
||
<taxonomicName id="4C0B14879758713B30B0F9C673E1F9CE" box="[1278,1432,1556,1581]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="339" phylum="Arthropoda" rank="tribe" tribe="Tmesiphorini">Tmesiphorini</taxonomicName>
|
||
, and the Greek word ‘dimer’, meaning ‘divided to two parts’, referring to the unique shape of the subantennal sulci. Gender is masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |