579 lines
79 KiB
XML
579 lines
79 KiB
XML
<document id="AC93ECDE01CABD69B723A0F31AD2548A" ID-DOI="10.5281/zenodo.184529" ID-GBIF-Dataset="122d8bda-1530-4cad-83fe-ee016161039a" ID-ISSN="1175-5326" ID-Zenodo-Dep="184529" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459986298214" checkinUser="plazi" docAuthor="Kok, Philippe J. R." docDate="2008" docId="B0797419FF97A96DFF113C16B248FD65" docLanguage="en" docName="zt01909p015.pdf" docOrigin="Zootaxa 1909" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Arthrosaura hoogmoedi" docType="treatment" docVersion="7" lastPageNumber="14" masterDocId="4C400C61FF93A960FF863A16B126FFDF" masterDocTitle="A new highland species of Arthrosaura Boulenger, 1885 (Squamata: Gymnophthalmidae) from Maringma tepui on the border of Guyana and Brazil" masterLastPageNumber="15" masterPageNumber="1" pageNumber="5" updateTime="1698232308145" updateUser="plazi">
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<mods:mods id="A11105F53D820CD245210F438B65667D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="21EF77140EA3011E9E8B711E4D30A36A">
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<mods:title id="C1D4329BBD1AA43E959CDD5E6141D732">A new highland species of Arthrosaura Boulenger, 1885 (Squamata: Gymnophthalmidae) from Maringma tepui on the border of Guyana and Brazil</mods:title>
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<mods:name id="14E1F83CE65845BEBE166EAA5E0C9F2F" type="personal">
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<mods:roleTerm id="D606A01CF88F80749E6C23C1BE0AA15F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C4E5FD50A94D6BFD66EBC565054F442D">Kok, Philippe J. R.</mods:namePart>
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<mods:typeOfResource id="0CC16815752174DED7D223A948027F71">text</mods:typeOfResource>
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<mods:titleInfo id="43582B1FF8DE7E198F63DA1EF4A20158">
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<mods:title id="1DC95FCCF086D1F977E6926DF0397FE9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F867FA8F31C4EC28FAE01FAC08CB4EB4">
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<mods:date id="378ADA211F54AFA72F55545A3760ECC3">2008</mods:date>
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<mods:detail id="32AA689D3FABD6CCB814DC10D3A343A0" type="volume">
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<mods:number id="D82B52F805A934FA1F32B9A7832E5F00">1909</mods:number>
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<mods:start id="5B9CF3B94CB0D4530AC422DA10D7C78E">1</mods:start>
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<mods:end id="29396D56664BD61924665C93479E356A">15</mods:end>
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<mods:classification id="855C03C819E33A7B164EEB7BACFEA85F">journal article</mods:classification>
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<mods:identifier id="7222F04CD04F29B97FE829B1B0CEAB4A" type="DOI">10.5281/zenodo.184529</mods:identifier>
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<mods:identifier id="30C5BC1B5EB3E43ABD58585E9A1ED9E1" type="GBIF-Dataset">122d8bda-1530-4cad-83fe-ee016161039a</mods:identifier>
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<mods:identifier id="615247DB6A6FE33A4DBE367B8AD449DA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="EE2B690F931375C12F0DF47A1D22CC8A" type="Zenodo-Dep">184529</mods:identifier>
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<treatment id="B0797419FF97A96DFF113C16B248FD65" ID-DOI="http://doi.org/10.5281/zenodo.6233283" ID-GBIF-Taxon="119379319" ID-Zenodo-Dep="6233283" LSID="urn:lsid:plazi:treatment:B0797419FF97A96DFF113C16B248FD65" httpUri="http://treatment.plazi.org/id/B0797419FF97A96DFF113C16B248FD65" lastPageId="13" lastPageNumber="14" pageId="4" pageNumber="5">
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<subSubSection id="70CA9684FF97A964FF113C16B049F9E2" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph id="386FC50FFF97A964FF113C16B352F9C5" blockId="4.[151,628,1536,1597]" box="[151,628,1536,1562]" pageId="4" pageNumber="5">
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<heading id="63277263FF97A964FF113C16B352F9C5" bold="true" box="[151,628,1536,1562]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
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<emphasis id="0AA4191DFF97A964FF113C16B352F9C5" bold="true" box="[151,628,1536,1562]" pageId="4" pageNumber="5">
|
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<taxonomicName id="FFD0BE8CFF97A964FF113C16B09FF9C5" box="[151,441,1536,1562]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="hoogmoedi">
|
||
<emphasis id="0AA4191DFF97A964FF113C16B09FF9C5" bold="true" box="[151,441,1536,1562]" italics="true" pageId="4" pageNumber="5">Arthrosaura hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
species novum
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF97A964FF113C35B049F9E2" blockId="4.[151,628,1536,1597]" box="[151,367,1571,1597]" pageId="4" pageNumber="5">
|
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<figureCitation id="A0EBD98AFF97A964FF113C35B020F9E2" box="[151,262,1571,1597]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="5.[151,255,1961,1985]" captionStartId-1="8.[151,255,1965,1989]" captionTargetBox-0="[151,1436,194,1936]" captionTargetBox-1="[275,1319,188,1924]" captionTargetId-0="figure@5.[151,1436,194,1937]" captionTargetId-1="figure@8.[243,1344,173,1937]" captionTargetPageId-0="5" captionTargetPageId-1="8" captionText-0="FIGURE 5. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. A: Dorsolateral view. B: Dorsal view." captionText-1="FIGURE 6. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." httpUri-0="https://zenodo.org/record/184534/files/figure.png" httpUri-1="https://zenodo.org/record/184535/files/figure.png" pageId="4" pageNumber="5">Figs. 5–6</figureCitation>
|
||
,
|
||
<figureCitation id="A0EBD98AFF97A964FE943C35B007F9E2" box="[274,289,1571,1597]" captionStart="FIGURE 7" captionStartId="9.[151,255,465,489]" captionTargetBox="[161,1424,186,443]" captionTargetId="figure@9.[161,1426,186,443]" captionTargetPageId="9" captionText="FIGURE 7. Lower eyelid condition in Arthrosaura hoogmoedi sp. nov. and two related Arthrosaura species. A: Semitransparent, unpigmented in A. kockii (IRSNB 15397). B: Semi-transparent, partly pigmented in A. guianensis (IRSNB 17343). C: Completely opaque, completely pigmented in A. hoogmoedi sp. nov. (IRSNB 2653, holotype)." httpUri="https://zenodo.org/record/184536/files/figure.png" pageId="4" pageNumber="5">7</figureCitation>
|
||
C, 8–9
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="70CA9684FF97A964FF113C65B2C0F902" pageId="4" pageNumber="5" type="materials_examined">
|
||
<paragraph id="386FC50FFF97A964FF113C65B2C0F902" blockId="4.[151,1437,1651,1997]" pageId="4" pageNumber="5">
|
||
<emphasis id="0AA4191DFF97A964FF113C65B036F952" bold="true" box="[151,272,1651,1677]" pageId="4" pageNumber="5">
|
||
<typeStatus id="E76B7BADFF97A964FF113C65B02DF952" box="[151,267,1651,1677]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="5EC15DCAFF97A964FE9E3C65B049F952" box="[280,367,1651,1677]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="4" pageNumber="5">IRSNB</collectionCode>
|
||
2653 (field number PK 2031), an adult female collected by Philippe J. R. Kok, Paul Benjamin and Claudius Perry,
|
||
<date id="4C6EE3CFFF97A964FE533C8DB39AF96A" box="[469,700,1691,1717]" pageId="4" pageNumber="5" value="2007-11-27">27 November 2007</date>
|
||
at 11h00, summit plateau of Mount Maringma, Cuyuni-Mazaruni District,
|
||
<collectingCountry id="40C7859FFF97A964FEC73CD5B0BAF902" box="[321,412,1731,1757]" name="Guyana" pageId="4" pageNumber="5">Guyana</collectingCountry>
|
||
(
|
||
<geoCoordinate id="5DE4A3C8FF97A964FE2B3CD5B360F902" box="[429,582,1731,1757]" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="15" value="5.2166667">5° 12’ 60” N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5DE4A3C8FF97A964FDD43CD5B3D9F902" box="[594,767,1731,1757]" direction="west" orientation="longitude" pageId="4" pageNumber="5" precision="15" value="-60.585">60° 35’ 06” W</geoCoordinate>
|
||
,
|
||
<quantity id="FF2868EAFF97A964FC8C3CD5B244F902" box="[778,866,1731,1757]" metricMagnitude="3" metricUnit="m" metricValue="2.1119999999999997" pageId="4" pageNumber="5" unit="m" value="2112.0">2112 m</quantity>
|
||
elevation).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="70CA9684FF97A964FF433CFDB015F812" pageId="4" pageNumber="5" type="etymology">
|
||
<paragraph id="386FC50FFF97A964FF433CFDB015F812" blockId="4.[151,1437,1651,1997]" pageId="4" pageNumber="5">
|
||
<emphasis id="0AA4191DFF97A964FF433CFDB074F8DA" bold="true" box="[197,338,1771,1797]" pageId="4" pageNumber="5">Etymology.</emphasis>
|
||
The specific epithet is a noun in the genitive case, honoring Dutch herpetologist Marinus Steven Hoogmoed. Marinus was curator of reptiles and amphibians at the National Museum of Natural History (Leiden,
|
||
<collectingCountry id="40C7859FFF97A964FEBD3D2DB324F88A" box="[315,514,1851,1877]" name="Netherlands" pageId="4" pageNumber="5">The Netherlands</collectingCountry>
|
||
) for 38 years (
|
||
<date id="4C6EE3CFFF97A964FD323D2DB21AF88A" box="[692,828,1851,1877]" pageId="4" pageNumber="5" value="1966" valueMax="2004">1966–2004</date>
|
||
) before retiring as research associate at the Museu Paraense Emílio Goeldi in Belém,
|
||
<collectingCountry id="40C7859FFF97A964FDB33D75B359F8A2" box="[565,639,1891,1917]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
|
||
, where he is still very active. His contribution to the knowledge of the
|
||
<collectingCountry id="40C7859FFF97A964FF423D9DB03FF87A" box="[196,281,1931,1957]" name="Guyana" pageId="4" pageNumber="5">Guiana</collectingCountry>
|
||
Shield herpetofauna is extensive, and most of his papers on the lizard fauna of the area are still authoritative.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="6CAF9587FF96A965FF113DBFB038F83C" httpUri="https://zenodo.org/record/184534/files/figure.png" pageId="5" pageNumber="6" targetBox="[151,1436,194,1936]" targetPageId="5">
|
||
<paragraph id="386FC50FFF96A965FF113DBFB038F83C" blockId="5.[151,1435,1961,2019]" pageId="5" pageNumber="6">
|
||
<emphasis id="0AA4191DFF96A965FF113DBFB03CF81E" bold="true" box="[151,282,1961,1985]" pageId="5" pageNumber="6">FIGURE 5.</emphasis>
|
||
<taxonomicName id="FFD0BE8CFF96A965FEA73DBCB304F81E" box="[289,546,1962,1985]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="hoogmoedi" status="sp. nov.">
|
||
<emphasis id="0AA4191DFF96A965FEA73DBCB304F81E" box="[289,546,1962,1985]" italics="true" pageId="5" pageNumber="6">Arthrosaura hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1197A466FF96A965FDAC3DBFB3A6F81E" box="[554,640,1961,1985]" pageId="5" pageNumber="6" rank="species">
|
||
<emphasis id="0AA4191DFF96A965FDAC3DBFB3A6F81E" bold="true" box="[554,640,1961,1985]" pageId="5" pageNumber="6">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female holotype (IRSNB 2653), 59.3 mm SVL. A: Dorsolateral view. B: Dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="70CA9684FF95A966FF433A81B387F9DD" pageId="6" pageNumber="7" type="diagnosis">
|
||
<paragraph id="386FC50FFF95A966FF433A81B294FDC5" blockId="6.[151,1437,151,2018]" pageId="6" pageNumber="7">
|
||
<emphasis id="0AA4191DFF95A966FF433A81B063FF6E" bold="true" box="[197,325,151,177]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
|
||
The new species is assigned to the genus
|
||
<taxonomicName id="FFD0BE8CFF95A966FCAB3A8EB29BFF6E" box="[813,957,152,177]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="0AA4191DFF95A966FCAB3A8EB29BFF6E" box="[813,957,152,177]" italics="true" pageId="6" pageNumber="7">Arthrosaura</emphasis>
|
||
</taxonomicName>
|
||
because of the following combination of characters (modified from
|
||
<bibRefCitation id="5C41B8FEFF95A966FE5E3AD6B382FF05" author="Avila-Pires" box="[472,676,192,218]" pageId="6" pageNumber="14" refString="Avila-Pires, T. C. S. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen, Leiden, 299, 1 - 706." type="journal article" year="1995">Ávila-Pires 1995</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="5C41B8FEFF95A966FD613AD6B56CFF05" author="MacCulloch" box="[743,1098,192,218]" pageId="6" pageNumber="15" refString="MacCulloch, R. D. & Lathrop, A. (2001) A new species of Arthrosaura (Sauria: Teiidae) from the highlands of Guyana. Caribbean Journal of Science, 37, 174 - 181." type="journal article" year="2001">MacCulloch & Lathrop 2001</bibRefCitation>
|
||
): (1) distinctive ear opening and moveable eyelids, (2) limbs pentadactyl with all digits clawed, (3) nasal scales separated by a single frontonasal, (4) prefrontals present (although absent in
|
||
<taxonomicName id="FFD0BE8CFF95A966FD693B06B1F0FE8D" authority="Donnelly, McDiarmid & Myers, 1992" authorityName="Donnelly, McDiarmid & Myers" authorityYear="1992" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="synaptolepis">
|
||
<emphasis id="0AA4191DFF95A966FD693B06B53FFEF6" box="[751,1049,272,297]" italics="true" pageId="6" pageNumber="7">Arthrosaura synaptolepis</emphasis>
|
||
<bibRefCitation id="5C41B8FEFF95A966FBA63B06B1F0FE8D" author="Donnelly" pageId="6" pageNumber="14" refString="Donnelly, M. A., McDiarmid, R. W. & Myers, C. W. (1992) A new lizard of the genus Arthrosaura (Teiidae) from southern Venezuela. Proceedings of the Biological Society of Washington, 105, 821 - 833." type="journal article" year="1992">Donnelly, McDiarmid & Myers, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
), (5) frontoparietals, parietals and interparietal present, (6) parietals and interparietal longer than wide, (7) interparietal and parietals forming a straight posterior margin, (8) occipitals present, (9) second pair of enlarged genials in contact with two infralabials, (10) enlarged median pairs of gulars, (11) dorsal scales hexagonal, keeled, in transverse rows only, (12) ventral scales quadrangular, smooth, in both transverse and longitudinal rows, (13) tongue dorsally covered by scalelike papillae (although papillae are reported as interrupting oblique plicae in
|
||
<taxonomicName id="FFD0BE8CFF95A966FEE73816B283FDC5" authority="Myers & Donnelly, 2008" authorityName="Myers & Donnelly" authorityYear="2008" box="[353,933,512,538]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="montigena">
|
||
<emphasis id="0AA4191DFF95A966FEE73816B352FDC6" box="[353,628,512,537]" italics="true" pageId="6" pageNumber="7">Arthrosaura montigena</emphasis>
|
||
<bibRefCitation id="5C41B8FEFF95A966FDFA3816B283FDC5" author="Myers" box="[636,933,512,538]" pageId="6" pageNumber="15" refString="Myers, C. W. & Donnelly, M. A. (2008) The summit herpetofauna of Auyantepui, Venezuela: report from the Robert G. Goelet American Museum - Terramar expedition. Bulletin of the American Museum of Natural History, 308, 1 - 147." type="journal article" year="2008">Myers & Donnelly, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF95A966FF43383EB0E1FCED" blockId="6.[151,1437,151,2018]" pageId="6" pageNumber="7">In addition to the generic characteristics, the new species is characterized by the following features: four supraoculars; prefrontals in contact with loreals; palpebral disc completely opaque; 32–33 smooth temporal scales; scales around midbody 37; 3–4 middorsal scales distinctly narrower than adjacent dorsal scales; dorsal scales strongly keeled, in 29 transverse rows; trunk length 1.4x length of forelimb; overlap of limbs equivalent to seven lateral scales when they are adpressed along the body; dark brown dorsum with a black middorsal stripe from nape to tail, and a tan ill-defined dorsolateral line from nape to midbody; flanks black; ventrally cream with dark mottling.</paragraph>
|
||
<paragraph id="386FC50FFF95A966FF433956B387F9DD" blockId="6.[151,1437,151,2018]" pageId="6" pageNumber="7">
|
||
The new species is immediately distinguished from
|
||
<taxonomicName id="FFD0BE8CFF95A966FCC03956B57BFC86" box="[838,1117,832,857]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="montigena">
|
||
<emphasis id="0AA4191DFF95A966FCC03956B57BFC86" box="[838,1117,832,857]" italics="true" pageId="6" pageNumber="7">Arthrosaura montigena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="FFD0BE8CFF95A966FBF63956B00AFC5D" authority="O'Shaughnessy, 1881" authorityName="O'Shaughnessy" authorityYear="1881" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="reticulata">
|
||
<emphasis id="0AA4191DFF95A966FBF63956B420FC86" box="[1136,1286,832,857]" italics="true" pageId="6" pageNumber="7">A. reticulata</emphasis>
|
||
(
|
||
<bibRefCitation id="5C41B8FEFF95A966FA9C3956B005FC5D" author="O'Shaughnessy" pageId="6" pageNumber="15" refString="O'Shaughnessy, A. W. E. (1881) An account of the collection of lizards made by Mr. Buckley in Ecuador, and now in the British Museum, with descriptions of the new species. Proceedings of the Zoological Society of London, 1881, 227 - 245." type="proceedings paper" year="1881">O’Shaughnessy, 1881</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="FFD0BE8CFF95A966FEBF397EB0CBFC5E" box="[313,493,872,897]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="synaptolepis">
|
||
<emphasis id="0AA4191DFF95A966FEBF397EB0CBFC5E" box="[313,493,872,897]" italics="true" pageId="6" pageNumber="7">A. synaptolepis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="FFD0BE8CFF95A966FE7A397EB275FC5D" authority="Burt & Burt, 1931" authorityName="Burt & Burt" authorityYear="1931" box="[508,851,872,898]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="tyleri">
|
||
<emphasis id="0AA4191DFF95A966FE7A397EB37AFC5E" box="[508,604,872,897]" italics="true" pageId="6" pageNumber="7">A. tyleri</emphasis>
|
||
(
|
||
<bibRefCitation id="5C41B8FEFF95A966FDED397EB26CFC5D" author="Burt" box="[619,842,872,898]" pageId="6" pageNumber="14" refString="Burt, C. E. & Burt M. D. (1931) South American lizards in the collection of the American Museum of Natural History. Bulletin of the American Museum of Natural History, 61, 227 - 395." type="journal article" year="1931">Burt & Burt, 1931</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="FFD0BE8CFF95A966FC14397EB54BFC5D" authority="Lidth" authorityName="Lidth" box="[914,1133,872,898]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="versteegii">
|
||
<emphasis id="0AA4191DFF95A966FC14397EB503FC5E" box="[914,1061,872,897]" italics="true" pageId="6" pageNumber="7">A. versteegii</emphasis>
|
||
Lidth
|
||
</taxonomicName>
|
||
de
|
||
<bibRefCitation id="5C41B8FEFF95A966FB1F397EB400FC5D" author="Jeude" box="[1177,1318,872,898]" pageId="6" pageNumber="15" refString="Lidth de Jeude, T. W. van (1904) Reptiles and Batrachians from Surinam. Notes of the Lyden Museum, 25, 83 - 94." type="journal article" year="1904">
|
||
Jeude,
|
||
<quantity id="FF2868EAFF95A966FB6D397EB400FC5D" box="[1259,1318,872,898]" metricMagnitude="1" metricUnit="m" metricValue="4.83616" pageId="6" pageNumber="7" unit="in" value="1904.0">1904</quantity>
|
||
</bibRefCitation>
|
||
in having four supraoculars (vs. three) and a black middorsal stripe from nape to tail (absent in the above-mentioned species). The new species agrees with
|
||
<taxonomicName id="FFD0BE8CFF95A966FDF339AEB585FC0D" authority="MacCulloch and Lathrop, 2001" authorityName="MacCulloch and Lathrop" authorityYear="2001" box="[629,1187,952,978]" class="Liliopsida" family="Rapateaceae" genus="Stegolepis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="guianensis">
|
||
<emphasis id="0AA4191DFF95A966FDF339AEB23EFC0E" box="[629,792,952,977]" italics="true" pageId="6" pageNumber="7">A. guianensis</emphasis>
|
||
<bibRefCitation id="5C41B8FEFF95A966FCA239AEB585FC0D" author="MacCulloch" box="[804,1187,952,978]" pageId="6" pageNumber="15" refString="MacCulloch, R. D. & Lathrop, A. (2001) A new species of Arthrosaura (Sauria: Teiidae) from the highlands of Guyana. Caribbean Journal of Science, 37, 174 - 181." type="journal article" year="2001">MacCulloch and Lathrop, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="FFD0BE8CFF95A966FB3239AEB439FC0E" box="[1204,1311,952,977]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="kockii">
|
||
<emphasis id="0AA4191DFF95A966FB3239AEB439FC0E" box="[1204,1311,952,977]" italics="true" pageId="6" pageNumber="7">A. kockii</emphasis>
|
||
</taxonomicName>
|
||
(Lidth de
|
||
<bibRefCitation id="5C41B8FEFF95A966FF1139F6B001FC25" author="Jeude" box="[151,295,992,1018]" pageId="6" pageNumber="15" refString="Lidth de Jeude, T. W. van (1904) Reptiles and Batrachians from Surinam. Notes of the Lyden Museum, 25, 83 - 94." type="journal article" year="1904">Jeude, 1904</bibRefCitation>
|
||
), and
|
||
<taxonomicName id="FFD0BE8CFF95A966FEEB39F6B260FC25" authority="Gorzula and Senaris, 1998" authorityName="Gorzula and Senaris" authorityYear="1998" box="[365,838,992,1018]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="testigensis">
|
||
<emphasis id="0AA4191DFF95A966FEEB39F6B321FC26" box="[365,519,992,1017]" italics="true" pageId="6" pageNumber="7">A. testigensis</emphasis>
|
||
Gorzula and Señaris,
|
||
<quantity id="FF2868EAFF95A966FC8D39F6B260FC25" box="[779,838,992,1018]" metricMagnitude="1" metricUnit="m" metricValue="5.07492" pageId="6" pageNumber="7" unit="in" value="1998.0">1998</quantity>
|
||
</taxonomicName>
|
||
in having four supraoculars.
|
||
<taxonomicName id="FFD0BE8CFF95A966FB1A39F6B190FBFE" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="guianensis">
|
||
<emphasis id="0AA4191DFF95A966FB1A39F6B190FBFE" italics="true" pageId="6" pageNumber="7">Arthrosaura guianensis</emphasis>
|
||
</taxonomicName>
|
||
mainly differs (characters of
|
||
<taxonomicName id="FFD0BE8CFF95A966FD9F3E1EB39BFBFE" box="[537,701,1032,1057]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="hoogmoedi">
|
||
<emphasis id="0AA4191DFF95A966FD9F3E1EB39BFBFE" box="[537,701,1032,1057]" italics="true" pageId="6" pageNumber="7">A. hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
in parentheses) in having keeled parietals and interparietal (a feeble keel on interparietal only), unpigmented tympanum (pigmented), lower eyelid with semitransparent disc (disc opaque), 45 scales around midbody (37), 40–42 keeled temporals (32–33, smooth), and absence of a black middorsal stripe (present).
|
||
<taxonomicName id="FFD0BE8CFF95A966FDAB3E96B22CFB46" box="[557,778,1152,1177]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="kockii">
|
||
<emphasis id="0AA4191DFF95A966FDAB3E96B22CFB46" box="[557,778,1152,1177]" italics="true" pageId="6" pageNumber="7">Arthrosaura kockii</emphasis>
|
||
</taxonomicName>
|
||
mainly differs (characters of
|
||
<taxonomicName id="FFD0BE8CFF95A966FBE33E96B42EFB46" box="[1125,1288,1152,1177]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="hoogmoedi">
|
||
<emphasis id="0AA4191DFF95A966FBE33E96B42EFB46" box="[1125,1288,1152,1177]" italics="true" pageId="6" pageNumber="7">A. hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
in parentheses) in having broadly keeled temporals (smooth), unpigmented tympanum (pigmented), lower eyelid with semitransparent disc (disc opaque), a wide light vertebral stripe (black middorsal stripe), and in having an immaculate venter (dark mottled).
|
||
<taxonomicName id="FFD0BE8CFF95A966FDBB3EEEB274FACE" box="[573,850,1272,1297]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="testigensis">
|
||
<emphasis id="0AA4191DFF95A966FDBB3EEEB274FACE" box="[573,850,1272,1297]" italics="true" pageId="6" pageNumber="7">Arthrosaura testigensis</emphasis>
|
||
</taxonomicName>
|
||
shares many characteristics with
|
||
<taxonomicName id="FFD0BE8CFF95A966FB683EEEB4B2FACE" box="[1262,1428,1272,1297]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="hoogmoedi">
|
||
<emphasis id="0AA4191DFF95A966FB683EEEB4B2FACE" box="[1262,1428,1272,1297]" italics="true" pageId="6" pageNumber="7">A. hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
, from which it mainly differs (characters of
|
||
<taxonomicName id="FFD0BE8CFF95A966FD233F36B26DFAE6" box="[677,843,1312,1337]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="hoogmoedi">
|
||
<emphasis id="0AA4191DFF95A966FD233F36B26DFAE6" box="[677,843,1312,1337]" italics="true" pageId="6" pageNumber="7">A. hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
in parentheses) in having less than 20 temporals (more than 30), lower eyelid with semitransparent disc (disc opaque), prefrontals separated from loreals by frontonasal-first supraocular contact (prefrontals in contact with loreals), weakly keeled dorsal scales (strongly keeled), middorsal scales not noticeably narrower than adjacent dorsal scales (3–4 middorsal scales distinctly narrower), and in having four narrow yellowish longitudinal stripes on dorsum (one middorsal black stripe and a tan ill-defined dorsolateral line).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="70CA9684FF95A96AFF433C19B051F83D" lastPageId="10" lastPageNumber="11" pageId="6" pageNumber="7" type="description">
|
||
<paragraph id="386FC50FFF95A966FF433C19B5C5F8C5" blockId="6.[151,1437,151,2018]" pageId="6" pageNumber="7">
|
||
<emphasis id="0AA4191DFF95A966FF433C19B30AF9F6" bold="true" box="[197,556,1551,1577]" pageId="6" pageNumber="7">
|
||
Description of the
|
||
<typeStatus id="E76B7BADFF95A966FE353C19B301F9F6" box="[435,551,1551,1577]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult female
|
||
<quantity id="FF2868EAFF95A966FD5A3C06B261F9F5" box="[732,839,1552,1578]" metricMagnitude="-2" metricUnit="m" metricValue="5.93" pageId="6" pageNumber="7" unit="mm" value="59.3">59.3 mm</quantity>
|
||
SVL. Head length 21 % SVL, 1.37x head width, head width 1.48x head height. Head slightly wider than neck, temporal region swollen, neck approximately as wide as anterior body. Snout blunt, rising gently posteriad. Body oval in transverse section, depressed, wider than high. Tail complete (regenerated distally), 97.0 mm, 1.63x SVL, circular in cross-section, tapering toward the tip. Limbs well-developed, moderately long, forelimbs 31% SVL, hindlimbs 48% SVL. Limbs pentadactyl, all fingers and toes with terminal claw. Overlap of limbs equivalent to seven lateral scales when they are adpressed along the body. Trunk length
|
||
<quantity id="FF2868EAFF95A966FD493D16B21DF8C5" box="[719,827,1792,1818]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="6" pageNumber="7" unit="mm" value="25.7">25.7 mm</quantity>
|
||
, 43 % SVL, 1.4x forelimbs (
|
||
<figureCitation id="A0EBD98AFF95A966FB0A3D16B5F2F8C5" box="[1164,1236,1792,1818]" captionStart="FIGURE 5" captionStartId="5.[151,255,1961,1985]" captionTargetBox="[151,1436,194,1936]" captionTargetId="figure@5.[151,1436,194,1937]" captionTargetPageId="5" captionText="FIGURE 5. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. A: Dorsolateral view. B: Dorsal view." httpUri="https://zenodo.org/record/184534/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF95A966FF433D3EB23EF84D" blockId="6.[151,1437,151,2018]" pageId="6" pageNumber="7">Tongue lanceolate, dorsally covered by oblique rows of small scalelike papillae; tip bifid, smooth; three well defined pairs of subequal infralingual plicae on the anterior part of the tongue, after fork. Anterior teeth conical, posterior teeth laterally compressed, tricuspid.</paragraph>
|
||
<paragraph id="386FC50FFF95A967FF433DB6B5C3FD4D" blockId="6.[151,1437,151,2018]" lastBlockId="7.[151,1437,152,1978]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
Rostral rectangular, twice as wide as high, visible from above, in broad contact with frontonasal. Frontonasal single, heptagonal, laterally in contact with nasal and in broad contact with loreal, not in contact with first supraocular. Prefrontals roughly pentagonal, as broad as long, with a long medial suture, laterally in broad contact with first supraocular, touching the loreal and the anterior point of the second supraocular. Frontal hexagonal, 2.1x as long as wide, slightly wider anteriorly, laterally markedly concave; laterally in contact over the entire length of second supraocular, separated from first supraocular by prefrontal-second supraocular contact, in point contact with third supraocular. Frontoparietals roughly hexagonal, 1.5x as long as wide with relatively broad medial suture; frontoparietals in contact with frontal, distinctly separated from second supraocular by frontal-third supraocular contact, in broad contact with third and fourth supraoculars, one parietal and interparietal. Interparietal 1.6x as long as wide, roughly pentagonal with curved sides converging anteriad, distinctly longer and narrower than parietals. Parietals and interparietal form a relatively straight posterior margin; anteriorly each is angulate. Interparietal with a short median, feeble, longitudinal keel. A row of seven occipitals of various shapes, the central largest; all of them feebly keeled, except the first left one. Four supraoculars, first smallest, about half the size of the fourth; third distinctly larger, second largest. Six superciliaries, first much larger than the others, last two smallest, others subequal (
|
||
<figureCitation id="A0EBD98AFF94A967FB08386EB5F0FD4D" box="[1166,1238,632,658]" captionStart="FIGURE 6" captionStartId="8.[151,255,1965,1989]" captionTargetBox="[275,1319,188,1924]" captionTargetId="figure@8.[243,1344,173,1937]" captionTargetPageId="8" captionText="FIGURE 6. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." httpUri="https://zenodo.org/record/184535/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF94A967FF4338B6B0FCFB95" blockId="7.[151,1437,152,1978]" pageId="7" pageNumber="8">
|
||
Nasal entire, with a very short division running ventrad from naris to supralabial on the left side; naris just below centre; directed laterally. Loreal trapezoidal, anteriorly inclined, in contact with nasal, frenocular, first superciliary, first supraocular, and prefrontal and frontonasal, separated from supralabials by frenocular-nasal contact and an odd, very small, quadrangular, supernumerary scale located at the intersection of nasal, frenocular, and first and second supralabials, present on both sides. Three small preoculars on the right side, two on the left, between first superciliary and first subocular. Frenocular quadrangular, longer than wide, subequal to loreal, separated from the first supralabial by a small supernumerary scale. Four suboculars on the right side, five on the left. Two postoculars, the lower smallest. Seven supralabials on both sides, fifth highest, with suture between third and fourth below middle of the eye; suture between second and third supralabials in contact with first subocular, suture between first and second supralabials in contact with a small supernumerary scale on both sides (
|
||
<figureCitation id="A0EBD98AFF94A967FE053E26B0EDFB95" box="[387,459,1072,1098]" captionStart="FIGURE 6" captionStartId="8.[151,255,1965,1989]" captionTargetBox="[275,1319,188,1924]" captionTargetId="figure@8.[243,1344,173,1937]" captionTargetPageId="8" captionText="FIGURE 6. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." httpUri="https://zenodo.org/record/184535/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF94A967FF433E4EB45FFB1D" blockId="7.[151,1437,152,1978]" pageId="7" pageNumber="8">
|
||
Upper eyelid with 10 ciliaries. Ocular recess with median row of five small scales separating median ciliaries from superciliaries. Lower eyelid with 12 ciliaries on the right side, 11 on the left side. Lower eyelid with a palpebral disc of four large, vertical, completely opaque (pigmented), palpebral scales (
|
||
<figureCitation id="A0EBD98AFF94A967FB603EBEB41FFB1D" box="[1254,1337,1192,1218]" captionStart="FIGURE 6" captionStartId="8.[151,255,1965,1989]" captionTargetBox="[275,1319,188,1924]" captionTargetId="figure@8.[243,1344,173,1937]" captionTargetPageId="8" captionText="FIGURE 6. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." httpUri="https://zenodo.org/record/184535/files/figure.png" pageId="7" pageNumber="8">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="A0EBD98AFF94A967FAC03EBEB470FB1D" box="[1350,1366,1192,1218]" captionStart="FIGURE 7" captionStartId="9.[151,255,465,489]" captionTargetBox="[161,1424,186,443]" captionTargetId="figure@9.[161,1426,186,443]" captionTargetPageId="9" captionText="FIGURE 7. Lower eyelid condition in Arthrosaura hoogmoedi sp. nov. and two related Arthrosaura species. A: Semitransparent, unpigmented in A. kockii (IRSNB 15397). B: Semi-transparent, partly pigmented in A. guianensis (IRSNB 17343). C: Completely opaque, completely pigmented in A. hoogmoedi sp. nov. (IRSNB 2653, holotype)." httpUri="https://zenodo.org/record/184536/files/figure.png" pageId="7" pageNumber="8">7</figureCitation>
|
||
C).
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF94A967FF433EC6B0DBFA6D" blockId="7.[151,1437,152,1978]" pageId="7" pageNumber="8">
|
||
Temporal region with 32 irregularly shaped, juxtaposed, smooth scales on the right side, 33 on the left. Temporals arranged in six ill-defined oblique rows, much larger in the upper part [a distinct cluster of 16 (on the right) and 18 (on the left) smaller temporals in the lower part]. Ear opening moderately large, vertically oval, posteriorly bordered by many small, smooth, mostly granular scales; anteriorly bordered by four (on the right side) or six (on the left side) smooth, enlarged, juxtaposed scales. Auditory meatus moderately deep, tympanum pigmented (
|
||
<figureCitation id="A0EBD98AFF94A967FE213F8EB0C9FA6D" box="[423,495,1432,1458]" captionStart="FIGURE 6" captionStartId="8.[151,255,1965,1989]" captionTargetBox="[275,1319,188,1924]" captionTargetId="figure@8.[243,1344,173,1937]" captionTargetPageId="8" captionText="FIGURE 6. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." httpUri="https://zenodo.org/record/184535/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF94A967FF433FD6B0E0F92D" blockId="7.[151,1437,152,1978]" pageId="7" pageNumber="8">
|
||
Mental trapezoidal, its posterior edge slightly concave. Postmental large, roughly heptagonal, laterally in contact with first and second infralabials. Three pairs of large genials, first two pairs in contact with each other medially, in lateral contact with infralabials, medial suture of second pair twice as long as that of first pair; third pair in contact with infralabials, but separated medially by small scales. A pair of enlarged postgenials separated medially by scales of varying sizes and widely separated from infralabials. Six infralabials on the right side, five on the left side. Six rows of two transversely enlarged gulars. Collar row with nine scales, decreasing in size laterally, forming a scalloped margin. Gular fold distinct, concealing three rows of very small scales (
|
||
<figureCitation id="A0EBD98AFF94A967FEB53CCEB0A1F92D" box="[307,391,1752,1778]" captionStart="FIGURE 6" captionStartId="8.[151,255,1965,1989]" captionTargetBox="[275,1319,188,1924]" captionTargetId="figure@8.[243,1344,173,1937]" captionTargetPageId="8" captionText="FIGURE 6. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." httpUri="https://zenodo.org/record/184535/files/figure.png" pageId="7" pageNumber="8">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="A0EBD98AFF94A967FE153CCEB084F92D" box="[403,418,1752,1778]" captionStart="FIGURE 8" captionStartId="9.[151,255,1701,1725]" captionTargetBox="[163,1426,600,1683]" captionTargetId="figure@9.[163,1428,600,1684]" captionTargetPageId="9" captionText="FIGURE 8. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. A: Ventral view in life. B: Detail of the gular region in life." httpUri="https://zenodo.org/record/184537/files/figure.png" pageId="7" pageNumber="8">8</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF94A967FF433D16B42AF865" blockId="7.[151,1437,152,1978]" pageId="7" pageNumber="8">Head scales with numerous small pits, especially concentrated on rostral, frontonasal, supraoculars and lateral head scales, absent on ventral head scales and body scales. Scales on nape posterior to the row of occipitals longer than wide, keeled, mucronate, in transverse rows, grading posteriorly into dorsal body scales. Sides of neck with distinctly smaller granular scales, juxtaposed, not in rows, slightly decreasing in size posteriorly. Three (left side) or four rows (right side) of distinctly enlarged scales before arm insertion.</paragraph>
|
||
<caption id="6CAF9587FF9BA968FF113DBBB0D9F838" httpUri="https://zenodo.org/record/184535/files/figure.png" pageId="8" pageNumber="9" targetBox="[275,1319,188,1924]" targetPageId="8">
|
||
<paragraph id="386FC50FFF9BA968FF113DBBB0D9F838" blockId="8.[151,1436,1965,2023]" pageId="8" pageNumber="9">
|
||
<emphasis id="0AA4191DFF9BA968FF113DBBB03FF81A" bold="true" box="[151,281,1965,1989]" pageId="8" pageNumber="9">FIGURE 6.</emphasis>
|
||
<taxonomicName id="FFD0BE8CFF9BA968FE983DB8B338F81A" box="[286,542,1966,1989]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="hoogmoedi" status="sp. nov.">
|
||
<emphasis id="0AA4191DFF9BA968FE983DB8B338F81A" box="[286,542,1966,1989]" italics="true" pageId="8" pageNumber="9">Arthrosaura hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1197A466FF9BA968FDA23DBBB35EF81A" box="[548,632,1965,1989]" pageId="8" pageNumber="9" rank="species">
|
||
<emphasis id="0AA4191DFF9BA968FDA23DBBB35EF81A" bold="true" box="[548,632,1965,1989]" pageId="8" pageNumber="9">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female holotype (IRSNB 2653), 59.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6CAF9587FF9AA969FF113BC7B427FDF1" httpUri="https://zenodo.org/record/184536/files/figure.png" pageId="9" pageNumber="10" targetBox="[161,1424,186,443]" targetPageId="9">
|
||
<paragraph id="386FC50FFF9AA969FF113BC7B427FDF1" blockId="9.[151,1436,465,558]" pageId="9" pageNumber="10">
|
||
<emphasis id="0AA4191DFF9AA969FF113BC7B03CFE36" bold="true" box="[151,282,465,489]" pageId="9" pageNumber="10">FIGURE 7.</emphasis>
|
||
Lower eyelid condition in
|
||
<taxonomicName id="FFD0BE8CFF9AA969FDC63BC4B267FE36" box="[576,833,466,489]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hoogmoedi" status="sp. nov.">
|
||
<emphasis id="0AA4191DFF9AA969FDC63BC4B267FE36" box="[576,833,466,489]" italics="true" pageId="9" pageNumber="10">Arthrosaura hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1197A466FF9AA969FCCF3BC7B2B9FE36" box="[841,927,465,489]" pageId="9" pageNumber="10" rank="species">
|
||
<emphasis id="0AA4191DFF9AA969FCCF3BC7B2B9FE36" bold="true" box="[841,927,465,489]" pageId="9" pageNumber="10">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and two related
|
||
<taxonomicName id="FFD0BE8CFF9AA969FBD23BC4B5F1FE36" box="[1108,1239,466,489]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="0AA4191DFF9AA969FBD23BC4B5F1FE36" box="[1108,1239,466,489]" italics="true" pageId="9" pageNumber="10">Arthrosaura</emphasis>
|
||
</taxonomicName>
|
||
species. A: Semitransparent, unpigmented in
|
||
<taxonomicName id="FFD0BE8CFF9AA969FE4D3BE2B30CFDD4" box="[459,554,500,523]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="kockii">
|
||
<emphasis id="0AA4191DFF9AA969FE4D3BE2B30CFDD4" box="[459,554,500,523]" italics="true" pageId="9" pageNumber="10">A. kockii</emphasis>
|
||
</taxonomicName>
|
||
(IRSNB 15397). B: Semi-transparent, partly pigmented in
|
||
<taxonomicName id="FFD0BE8CFF9AA969FB2A3BE2B41BFDD4" box="[1196,1341,500,523]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="guianensis">
|
||
<emphasis id="0AA4191DFF9AA969FB2A3BE2B41BFDD4" box="[1196,1341,500,523]" italics="true" pageId="9" pageNumber="10">A. guianensis</emphasis>
|
||
</taxonomicName>
|
||
(IRSNB 17343). C: Completely opaque, completely pigmented in
|
||
<taxonomicName id="FFD0BE8CFF9AA969FD7A3801B2B6FDF1" box="[764,912,535,558]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hoogmoedi" status="sp. nov.">
|
||
<emphasis id="0AA4191DFF9AA969FD7A3801B2B6FDF1" box="[764,912,535,558]" italics="true" pageId="9" pageNumber="10">A. hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1197A466FF9AA969FC113800B2CAFDF1" box="[919,1004,534,558]" pageId="9" pageNumber="10" rank="species">
|
||
<emphasis id="0AA4191DFF9AA969FC113800B2CAFDF1" bold="true" box="[919,1004,534,558]" pageId="9" pageNumber="10">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(IRSNB 2653, holotype).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6CAF9587FF9AA969FF113CB3B0D7F900" httpUri="https://zenodo.org/record/184537/files/figure.png" pageId="9" pageNumber="10" targetBox="[163,1426,600,1683]" targetPageId="9">
|
||
<paragraph id="386FC50FFF9AA969FF113CB3B0D7F900" blockId="9.[151,1436,1701,1759]" pageId="9" pageNumber="10">
|
||
<emphasis id="0AA4191DFF9AA969FF113CB3B03FF962" bold="true" box="[151,281,1701,1725]" pageId="9" pageNumber="10">FIGURE 8.</emphasis>
|
||
<taxonomicName id="FFD0BE8CFF9AA969FE993CB0B306F962" box="[287,544,1702,1725]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hoogmoedi" status="sp. nov.">
|
||
<emphasis id="0AA4191DFF9AA969FE993CB0B306F962" box="[287,544,1702,1725]" italics="true" pageId="9" pageNumber="10">Arthrosaura hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0AA4191DFF9AA969FDA33CB3B35DF962" bold="true" box="[549,635,1701,1725]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="1197A466FF9AA969FDA33CB3B35DF962" box="[549,635,1701,1725]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype (IRSNB 2653), 59.3 mm SVL. A: Ventral view in life. B: Detail of the gular region in life.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="386FC50FFF9AA96AFF433D05B295FEF5" blockId="9.[151,1437,1811,2037]" lastBlockId="10.[151,1437,152,2018]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
Dorsals imbricate, appearing hexagonal because of imbrication, elongate, strongly keeled, mucronate, in transverse rows only, 29 rows between the interparietal and posterior edge of hindlimbs (including row of occipitals). Three to four middorsal scales, between anterior margin of forelimbs and anterior margin of hindlimbs, distinctly narrower than adjacent dorsals and very similar to laterals (
|
||
<figureCitation id="A0EBD98AFF9AA969FBA63D9DB541F87A" box="[1056,1127,1931,1957]" captionStart="FIGURE 9" captionStartId="11.[151,255,1881,1905]" captionTargetBox="[151,1436,194,1855]" captionTargetId="figure@11.[151,1436,194,1857]" captionTargetPageId="11" captionText="FIGURE 9. Arthrosaura hoogmoedi sp. nov., preserved female holotype (IRSNB 2653), 59.3 mm SVL. A: Detail of dorsal scales at midbody showing the 3 – 4 middorsal scales noticeably narrower than contiguous dorsal scales. B: Detail of perianal region." httpUri="https://zenodo.org/record/184538/files/figure.png" pageId="9" pageNumber="10">Fig. 9</figureCitation>
|
||
A), 37 scales around midbody. A zone of small granular scales extending from anteriorly of forelimb insertion to above arm and becoming wider in the axilla. Small granular scales at hindlimb insertion. Flank scales similar to, but narrower than, most dorsal scales (except three to four middorsal ones, see above); no abrupt demarcation between dorsals and laterals. Lateral scales becoming smaller toward groin. Below the laterals, two rows of ventrolateral scales (see
|
||
<bibRefCitation id="5C41B8FEFF99A96AFE9B3AFEB363FEDD" author="Myers" box="[285,581,232,258]" pageId="10" pageNumber="15" refString="Myers, C. W. & Donnelly, M. A. (2008) The summit herpetofauna of Auyantepui, Venezuela: report from the Robert G. Goelet American Museum - Terramar expedition. Bulletin of the American Museum of Natural History, 308, 1 - 147." type="journal article" year="2008">Myers & Donnelly 2008</bibRefCitation>
|
||
); some short, smaller scales intercalated between the transverse rows of laterals; no distinct demarcation between laterals and ventrolaterals.
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF99A96AFF433B2EB076FDC5" blockId="10.[151,1437,152,2018]" pageId="10" pageNumber="11">
|
||
Ventrals imbricate, smooth, longer than wide, with posterior margins slightly rounded, arranged in transverse and longitudinal rows (
|
||
<figureCitation id="A0EBD98AFF99A96AFE6D3B76B316FEA5" box="[491,560,352,378]" captionStart="FIGURE 8" captionStartId="9.[151,255,1701,1725]" captionTargetBox="[163,1426,600,1683]" captionTargetId="figure@9.[163,1428,600,1684]" captionTargetPageId="9" captionText="FIGURE 8. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. A: Ventral view in life. B: Detail of the gular region in life." httpUri="https://zenodo.org/record/184537/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
|
||
A). Eighteen transverse rows between collar and preanal plate, six longitudinal rows of ventrals at midbody (10 rows including the ventrolaterals). Preanal plate consisting of eight scales, three anterior scales, five posterior scales (four elongate large scales and a small distal medial triangular scale); lateral posterior preanal scales narrow, almost as long as adjacent scales. No preanal or femoral pores (
|
||
<figureCitation id="A0EBD98AFF99A96AFF603816B00BFDC5" box="[230,301,512,538]" captionStart="FIGURE 9" captionStartId="11.[151,255,1881,1905]" captionTargetBox="[151,1436,194,1855]" captionTargetId="figure@11.[151,1436,194,1857]" captionTargetPageId="11" captionText="FIGURE 9. Arthrosaura hoogmoedi sp. nov., preserved female holotype (IRSNB 2653), 59.3 mm SVL. A: Detail of dorsal scales at midbody showing the 3 – 4 middorsal scales noticeably narrower than contiguous dorsal scales. B: Detail of perianal region." httpUri="https://zenodo.org/record/184538/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF99A96AFF43383EB211FCD5" blockId="10.[151,1437,152,2018]" pageId="10" pageNumber="11">Scales on tail keeled, mucronate (except distally), imbricate, appearing hexagonal because of imbrication, similar to, but slightly shorter than, dorsal scales, in transverse rows only except two middorsal caudal scales that form an almost complete longitudinal row from the seventh dorsal caudal scale (from posterior margin of hindlimbs) to tip of tail. Regenerated portion of tail (distal 31.0 mm) with much shorter scales ventrally and dorsally. Pigmented subcaudal scales smooth, imbricate, with rounded ends, in both transverse and longitudinal rows, becoming keeled laterally, grading into laterals.</paragraph>
|
||
<paragraph id="386FC50FFF99A96AFF43390EB38FFBAD" blockId="10.[151,1437,152,2018]" pageId="10" pageNumber="11">Dorsal surfaces of upper and lower arms with smooth, variably polygonal scales, ventral surface of upper arms with small, juxtaposed, granular scales, ventral surface of lower arms with scales similar to those on dorsal surface, except for a band on inner side with small juxtaposed granular scales. Thighs with a row of enlarged, smooth, hexagonal scales on anterior surface, bordered ventrally by two rows of slightly smaller, smooth, polygonal scales; bordered dorsally by 2–3 rows of keeled scales, diminishing in size dorsad, grading into small, granular, juxtaposed scales, which cover posterodorsal and posterior surfaces of thighs. Shanks ventrally and posteriorly with large, smooth, polygonal, imbricate scales; scales on anterior surface smaller, keeled, imbricate, grading into small, granular, juxtaposed scales, which cover distal portion of anterior side (near the ankle) and dorsal portion of shanks.</paragraph>
|
||
<paragraph id="386FC50FFF99A96AFF433E96B562FAE5" blockId="10.[151,1437,152,2018]" pageId="10" pageNumber="11">Dorsal surface of hands, feet, and digits with imbricate, smooth scales, except on the posterodorsal surface of feet where 2–3 scales are keeled; palms and soles covered with small roundish or elongate scales, juxtaposed, not projecting anteriorly. Two enlarged thenar scales on inner margin of palm below pollex, two enlarged hypothenar scales on outer margin of palm. Subdigital lamellae protuberant, slightly divided basally on some digits, single on distal halves of most digits; lamellae rounded laterally.</paragraph>
|
||
<paragraph id="386FC50FFF99A96AFF433F5EB24BFA6D" blockId="10.[151,1437,152,2018]" pageId="10" pageNumber="11">
|
||
Subdigital lamellae as follows (Roman numbers indicate digits, Arabic numbers indicate paired or single lamellae on left/right; lower ungual sheath scale is omitted from the counts): hands
|
||
<date id="4C6EE3CFFF99A96AFBEE3F79B5ACFA55" box="[1128,1162,1391,1418]" pageId="10" pageNumber="11">
|
||
<emphasis id="0AA4191DFF99A96AFBEE3F79B555FA56" bold="true" box="[1128,1139,1391,1417]" pageId="10" pageNumber="11">I</emphasis>
|
||
4
|
||
</date>
|
||
/4,
|
||
<date id="4C6EE3CFFF99A96AFB293F79B5FDFA55" box="[1199,1243,1391,1418]" pageId="10" pageNumber="11">
|
||
<emphasis id="0AA4191DFF99A96AFB293F79B5E0FA56" bold="true" box="[1199,1222,1391,1417]" pageId="10" pageNumber="11">II</emphasis>
|
||
8
|
||
</date>
|
||
/8,
|
||
<date id="4C6EE3CFFF99A96AFA873F79B449FA55" box="[1281,1391,1391,1418]" pageId="10" pageNumber="11" value="1913-03-14">
|
||
<emphasis id="0AA4191DFF99A96AFA873F79B405FA56" bold="true" box="[1281,1315,1391,1417]" pageId="10" pageNumber="11">III</emphasis>
|
||
14/13
|
||
</date>
|
||
,
|
||
<date id="4C6EE3CFFF99A96AFAFD3F79B1FAFA6D" pageId="10" pageNumber="11" value="1914-04-15">
|
||
<emphasis id="0AA4191DFF99A96AFAFD3F79B4BAFA56" bold="true" box="[1403,1436,1391,1417]" pageId="10" pageNumber="11">IV</emphasis>
|
||
15/14
|
||
</date>
|
||
,
|
||
<emphasis id="0AA4191DFF99A96AFF6E3F81B1DBFA6E" bold="true" box="[232,253,1431,1457]" pageId="10" pageNumber="11">V</emphasis>
|
||
8/8; feet
|
||
<date id="4C6EE3CFFF99A96AFEEB3F81B0A8FA6D" box="[365,398,1431,1458]" pageId="10" pageNumber="11">
|
||
<emphasis id="0AA4191DFF99A96AFEEB3F81B05FFA6E" bold="true" box="[365,377,1431,1457]" pageId="10" pageNumber="11">I</emphasis>
|
||
6
|
||
</date>
|
||
/6,
|
||
<date id="4C6EE3CFFF99A96AFE323F81B321FA6D" box="[436,519,1431,1458]" pageId="10" pageNumber="11" value="1919-02-09">
|
||
<emphasis id="0AA4191DFF99A96AFE323F81B0EDFA6E" bold="true" box="[436,459,1431,1457]" pageId="10" pageNumber="11">II</emphasis>
|
||
9/19
|
||
</date>
|
||
,
|
||
<date id="4C6EE3CFFF99A96AFD923F81B3A5FA6D" box="[532,643,1431,1458]" pageId="10" pageNumber="11" value="1914-03-14">
|
||
<emphasis id="0AA4191DFF99A96AFD923F81B310FA6E" bold="true" box="[532,566,1431,1457]" pageId="10" pageNumber="11">III</emphasis>
|
||
14/14
|
||
</date>
|
||
,
|
||
<date id="4C6EE3CFFF99A96AFD093F81B3DBFA6D" box="[655,765,1431,1458]" pageId="10" pageNumber="11" value="1920-04-20">
|
||
<emphasis id="0AA4191DFF99A96AFD093F81B396FA6E" bold="true" box="[655,688,1431,1457]" pageId="10" pageNumber="11">IV</emphasis>
|
||
20/20
|
||
</date>
|
||
,
|
||
<date id="4C6EE3CFFF99A96AFC8F3F81B24BFA6D" box="[777,877,1431,1458]" pageId="10" pageNumber="11" value="1911-05-11">
|
||
<emphasis id="0AA4191DFF99A96AFC8F3F81B238FA6E" bold="true" box="[777,798,1431,1457]" pageId="10" pageNumber="11">V</emphasis>
|
||
11/11.
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF99A96AFF433FA9B376F8B5" blockId="10.[151,1437,152,2018]" pageId="10" pageNumber="11">
|
||
<emphasis id="0AA4191DFF99A96AFF433FA9B32EFA06" bold="true" box="[197,520,1471,1497]" pageId="10" pageNumber="11">
|
||
Color of
|
||
<typeStatus id="E76B7BADFF99A96AFEBC3FA9B08AFA06" box="[314,428,1471,1497]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
|
||
in life.
|
||
</emphasis>
|
||
Dorsum dark brown, with a black middorsal stripe (about two-three dorsal scales wide) extending from nape to tail, where it quickly becomes inconspicuous due to the dark coloration of the tail; subtle, irregular, tan dorsolateral line extending from nape to about midbody; flanks black, lower part of flanks peppered with reddish brown, and with some small pale brown spots ventrolaterally, forming an irregular ventrolateral line. Axilla region and underside of arms marbled with reddish brown and black. Head blackish brown with tan dorsolateral lines extending above eyes, from the anterior part of the loreals to the parietals. Tip of snout paler; diffuse paler blotches on lower part of lateral surface of head. Tympanum black, surrounded by some tan scales. Arms black, legs and tail blackish brown. Iris black (
|
||
<figureCitation id="A0EBD98AFF99A96AFBEB3CCEB592F92D" box="[1133,1204,1752,1778]" captionStart="FIGURE 5" captionStartId="5.[151,255,1961,1985]" captionTargetBox="[151,1436,194,1936]" captionTargetId="figure@5.[151,1436,194,1937]" captionTargetPageId="5" captionText="FIGURE 5. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. A: Dorsolateral view. B: Dorsal view." httpUri="https://zenodo.org/record/184534/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
|
||
). Tongue dark gray. Palms and soles black.
|
||
<collectingCountry id="40C7859FFF99A96AFE2D3D16B0C1F8C5" box="[427,487,1792,1818]" name="China" pageId="10" pageNumber="11">Chin</collectingCountry>
|
||
, throat, chest, belly, and ventral surface of upper legs cream with dark mottling; lower legs ventrally reddish brown with dark mottling; tail ventrally black, except proximally where it is dirty white peppered with black (
|
||
<figureCitation id="A0EBD98AFF99A96AFE623D46B30FF8B5" box="[484,553,1872,1898]" captionStart="FIGURE 8" captionStartId="9.[151,255,1701,1725]" captionTargetBox="[163,1426,600,1683]" captionTargetId="figure@9.[163,1428,600,1684]" captionTargetPageId="9" captionText="FIGURE 8. Arthrosaura hoogmoedi sp. nov., female holotype (IRSNB 2653), 59.3 mm SVL. A: Ventral view in life. B: Detail of the gular region in life." httpUri="https://zenodo.org/record/184537/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF99A96AFF433D61B051F83D" blockId="10.[151,1437,152,2018]" pageId="10" pageNumber="11">
|
||
<emphasis id="0AA4191DFF99A96AFF433D61B357F84E" bold="true" box="[197,625,1911,1937]" pageId="10" pageNumber="11">
|
||
Color of
|
||
<typeStatus id="E76B7BADFF99A96AFEB33D61B081F84E" box="[309,423,1911,1937]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
|
||
in preservative.
|
||
</emphasis>
|
||
After four months in 70 % ethanol, color pattern is similar to that in life. The dorsolateral line became white and is more conspicuous, the cream coloration of the ventral surface also became white.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="6CAF9587FF98A96BFF113D4FB07AF86A" httpUri="https://zenodo.org/record/184538/files/figure.png" pageId="11" pageNumber="12" targetBox="[151,1436,194,1855]" targetPageId="11">
|
||
<paragraph id="386FC50FFF98A96BFF113D4FB07AF86A" blockId="11.[151,1436,1881,1973]" pageId="11" pageNumber="12">
|
||
<emphasis id="0AA4191DFF98A96BFF113D4FB03DF8AE" bold="true" box="[151,283,1881,1905]" pageId="11" pageNumber="12">FIGURE 9.</emphasis>
|
||
<taxonomicName id="FFD0BE8CFF98A96BFEA23D4FB301F8AF" box="[292,551,1881,1904]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hoogmoedi" status="sp. nov.">
|
||
<emphasis id="0AA4191DFF98A96BFEA23D4FB301F8AF" box="[292,551,1881,1904]" italics="true" pageId="11" pageNumber="12">Arthrosaura hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1197A466FF98A96BFDA93D4FB3A1F8AE" box="[559,647,1881,1905]" pageId="11" pageNumber="12" rank="species">
|
||
<emphasis id="0AA4191DFF98A96BFDA93D4FB3A1F8AE" bold="true" box="[559,647,1881,1905]" pageId="11" pageNumber="12">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, preserved female holotype (IRSNB 2653), 59.3 mm SVL. A: Detail of dorsal scales at midbody showing the 3–4 middorsal scales noticeably narrower than contiguous dorsal scales. B: Detail of perianal region.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6CAF9587FF9FA96CFF113E23B3D6FBB0" httpUri="https://zenodo.org/record/184539/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1435,194,1053]" targetPageId="12">
|
||
<paragraph id="386FC50FFF9FA96CFF113E23B3D6FBB0" blockId="12.[151,1436,1077,1136]" pageId="12" pageNumber="13">
|
||
<emphasis id="0AA4191DFF9FA96CFF113E23B000FB92" bold="true" box="[151,294,1077,1101]" pageId="12" pageNumber="13">FIGURE 10.</emphasis>
|
||
A patch of
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFE223E20B3DBFB92" box="[420,765,1078,1101]" class="Gastropoda" family="Cancellariidae" genus="Brocchinia" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="tatei">
|
||
<emphasis id="0AA4191DFF9FA96CFE223E20B369FB92" box="[420,591,1078,1101]" italics="true" pageId="12" pageNumber="13">Brocchinia tatei</emphasis>
|
||
(Bromeliaceae)
|
||
</taxonomicName>
|
||
, habitat on the summit plateau of Maringma tepui in which the holotype of
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFE9E3E4FB33EFBAF" box="[280,536,1113,1136]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hoogmoedi" status="sp. nov.">
|
||
<emphasis id="0AA4191DFF9FA96CFE9E3E4FB33EFBAF" box="[280,536,1113,1136]" italics="true" pageId="12" pageNumber="13">Arthrosaura hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1197A466FF9FA96CFD983E4EB352FBAF" box="[542,628,1112,1136]" pageId="12" pageNumber="13" rank="species">
|
||
<emphasis id="0AA4191DFF9FA96CFD983E4EB352FBAF" bold="true" box="[542,628,1112,1136]" pageId="12" pageNumber="13">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was found.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="70CA9684FF9FA96CFF433EB5B5B3FA0A" pageId="12" pageNumber="13" type="distribution">
|
||
<paragraph id="386FC50FFF9FA96CFF433EB5B5B3FA0A" blockId="12.[151,1437,1187,2013]" pageId="12" pageNumber="13">
|
||
<emphasis id="0AA4191DFF9FA96CFF433EB5B0D8FB62" bold="true" box="[197,510,1187,1213]" pageId="12" pageNumber="13">Distribution and ecology.</emphasis>
|
||
The species is currently known only from the
|
||
<typeStatus id="E76B7BADFF9FA96CFB9D3EB5B568FB62" box="[1051,1102,1187,1213]" pageId="12" pageNumber="13">type</typeStatus>
|
||
locality, the summit of Maringma tepui in
|
||
<collectingCountry id="40C7859FFF9FA96CFECF3EDDB082FB3A" box="[329,420,1227,1253]" name="Guyana" pageId="12" pageNumber="13">Guyana</collectingCountry>
|
||
(
|
||
<figureCitation id="A0EBD98AFF9FA96CFE333EDDB0DCFB3A" box="[437,506,1227,1253]" captionStart="FIGURE 1" captionStartId="1.[151,255,1709,1733]" captionTargetBox="[151,1435,674,1685]" captionTargetId="figure@1.[151,1436,674,1685]" captionTargetPageId="1" captionText="FIGURE 1. Area map of Maringma tepui showing the type locality of Arthrosaura hoogmoedi sp. nov. (blue dot). Coordinates and boundary with Brazil are approximate and based on the available map (note that there is a substantial difference in the country boundary between this map and that shown in Google Earth). Maps elaborated after “ Roraima Sheet 40 SE ” published by the Survey Department of Guyana, 1972, and a radar image of South America by NASA / JPL / NIMA available at http: // photojournal. jpl. nasa. gov / catalog / PIA 03388." httpUri="https://zenodo.org/record/184530/files/figure.png" pageId="12" pageNumber="13">Fig. 1</figureCitation>
|
||
). The only specimen available was collected in late morning (11h00), during a sunny spell, in a large bromeliad (
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFDBE3EE5B3D1FAD3" box="[568,759,1267,1292]" class="Gastropoda" family="Cancellariidae" genus="Brocchinia" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="tatei">
|
||
<emphasis id="0AA4191DFF9FA96CFDBE3EE5B3D1FAD3" box="[568,759,1267,1292]" italics="true" pageId="12" pageNumber="13">Brocchinia tatei</emphasis>
|
||
</taxonomicName>
|
||
), about
|
||
<quantity id="FF2868EAFF9FA96CFCD83EE5B28FFAD2" box="[862,937,1267,1293]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="13" unit="cm" value="60.0">60 cm</quantity>
|
||
above the ground (
|
||
<figureCitation id="A0EBD98AFF9FA96CFB1E3EE5B5D4FAD2" box="[1176,1266,1267,1293]" captionStart="FIGURE 10. A" captionStartId="12.[151,255,1077,1101]" captionTargetBox="[151,1435,194,1053]" captionTargetId="figure@12.[151,1436,194,1054]" captionTargetPageId="12" captionText="FIGURE 10. A patch of Brocchinia tatei (Bromeliaceae), habitat on the summit plateau of Maringma tepui in which the holotype of Arthrosaura hoogmoedi sp. nov. was found." httpUri="https://zenodo.org/record/184539/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
|
||
). Possibly the specimen was hunting for some insects and spiders that became numerous once the sun chased away the mist generally present all over the summit. One other specimen was observed on the same day (in the afternoon) in another patch of bromeliads, but that specimen escaped. Interestingly, Gorzula and Señaris (1998) and
|
||
<bibRefCitation id="5C41B8FEFF9FA96CFF113F85B0E3FA72" author="Fuentes" box="[151,453,1427,1453]" pageId="12" pageNumber="15" refString="Fuentes, O. & Rivas, G. (2000) Geographic distribution. Euspondylus goeleti. Herpetological Review, 31, 181 - 182." type="journal article" year="2000">Fuentes and Rivas (2000)</bibRefCitation>
|
||
also reported collection of gymnophthalmids in
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFB823F85B5E7FA73" box="[1028,1217,1427,1452]" class="Gastropoda" family="Cancellariidae" genus="Brocchinia" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="tatei">
|
||
<emphasis id="0AA4191DFF9FA96CFB823F85B5E7FA73" box="[1028,1217,1427,1452]" italics="true" pageId="12" pageNumber="13">Brocchinia tatei</emphasis>
|
||
</taxonomicName>
|
||
. This plant probably plays an important role as a refuge and food source for highland gymnophthalmids.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="70CA9684FF9FA96DFF433FF5B248FD65" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph id="386FC50FFF9FA96CFF433FF5B07AF932" blockId="12.[151,1437,1187,2013]" pageId="12" pageNumber="13">
|
||
<emphasis id="0AA4191DFF9FA96CFF433FF5B077FA22" bold="true" box="[197,337,1507,1533]" pageId="12" pageNumber="13">Discussion.</emphasis>
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFEDC3FF5B350FA23" box="[346,630,1507,1532]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hoogmoedi">
|
||
<emphasis id="0AA4191DFF9FA96CFEDC3FF5B350FA23" box="[346,630,1507,1532]" italics="true" pageId="12" pageNumber="13">Arthrosaura hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
shares many morphological characteristics with
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFB3B3FF5B47CFA23" box="[1213,1370,1507,1532]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="testigensis">
|
||
<emphasis id="0AA4191DFF9FA96CFB3B3FF5B47CFA23" box="[1213,1370,1507,1532]" italics="true" pageId="12" pageNumber="13">A. testigensis</emphasis>
|
||
</taxonomicName>
|
||
from Los Testigos (Estado Bolívar,
|
||
<collectingCountry id="40C7859FFF9FA96CFE713C1DB354F9FA" box="[503,626,1547,1573]" name="Venezuela" pageId="12" pageNumber="13">Venezuela</collectingCountry>
|
||
) a group of tepuis east of Auyán-tepui and situated about
|
||
<quantity id="FF2868EAFF9FA96CFA923C1DB44BF9FA" box="[1300,1389,1547,1573]" metricMagnitude="5" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="km" value="180.0">180 km</quantity>
|
||
airline NWW from Maringma tepui. In having a black middorsal stripe, the new species resembles a putative new taxon (“
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFEA93C4DB099F9AB" box="[303,447,1627,1652]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="0AA4191DFF9FA96CFEA93C4DB099F9AB" box="[303,447,1627,1652]" italics="true" pageId="12" pageNumber="13">Arthrosaura</emphasis>
|
||
</taxonomicName>
|
||
species a”) reported by Gorzula and Señaris (1998: 124) from the Chimantá Massif (about
|
||
<quantity id="FF2868EAFF9FA96CFF6D3C95B061F942" box="[235,327,1667,1693]" metricMagnitude="5" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="km" value="180.0">180 km</quantity>
|
||
airline W from Maringma tepui), unfortunately the absence of diagnostic characters and the poor quality of the photograph provided by Gorzula and Señaris (1998, photograph #89) does not allow further comparison.
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF9FA96DFF433CEDB047FF05" blockId="12.[151,1437,1187,2013]" lastBlockId="13.[151,1437,152,698]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||
Without formally naming them,
|
||
<bibRefCitation id="5C41B8FEFF9FA96CFDC03CEDB2E3F8CA" author="MacCulloch" box="[582,965,1787,1813]" pageId="12" pageNumber="15" refString="MacCulloch, R. D. & Lathrop, A. (2001) A new species of Arthrosaura (Sauria: Teiidae) from the highlands of Guyana. Caribbean Journal of Science, 37, 174 - 181." type="journal article" year="2001">MacCulloch and Lathrop (2001)</bibRefCitation>
|
||
postulated the existence of two species groups of
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFE923D35B082F8E3" box="[276,420,1827,1852]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="0AA4191DFF9FA96CFE923D35B082F8E3" box="[276,420,1827,1852]" italics="true" pageId="12" pageNumber="13">Arthrosaura</emphasis>
|
||
</taxonomicName>
|
||
, one group containing the species having four supraoculars, a relatively short body with relatively long legs (which I propose to name the
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFC993D5DB243F8BB" box="[799,869,1867,1892]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="kockii">
|
||
<emphasis id="0AA4191DFF9FA96CFC993D5DB243F8BB" box="[799,869,1867,1892]" italics="true" pageId="12" pageNumber="13">kockii</emphasis>
|
||
</taxonomicName>
|
||
group), the other group containing species with three supraoculars, gracile body and relatively short legs (which I propose to name the
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFB4C3D65B41DF853" box="[1226,1339,1907,1932]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="reticulata">
|
||
<emphasis id="0AA4191DFF9FA96CFB4C3D65B41DF853" box="[1226,1339,1907,1932]" italics="true" pageId="12" pageNumber="13">reticulata</emphasis>
|
||
</taxonomicName>
|
||
group). Among the
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFEA23D8DB04CF86B" box="[292,362,1947,1972]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="kockii">
|
||
<emphasis id="0AA4191DFF9FA96CFEA23D8DB04CF86B" box="[292,362,1947,1972]" italics="true" pageId="12" pageNumber="13">kockii</emphasis>
|
||
</taxonomicName>
|
||
group, the presence of keeled temporals in
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFCFE3D8DB571F86B" box="[888,1111,1947,1972]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="kockii">
|
||
<emphasis id="0AA4191DFF9FA96CFCFE3D8DB571F86B" box="[888,1111,1947,1972]" italics="true" pageId="12" pageNumber="13">Arthrosaura kockii</emphasis>
|
||
</taxonomicName>
|
||
(a lowland species) and
|
||
<taxonomicName id="FFD0BE8CFF9FA96CFA053D8DB032F803" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="guianensis">
|
||
<emphasis id="0AA4191DFF9FA96CFA053D8DB032F803" italics="true" pageId="12" pageNumber="13">A. guianensis</emphasis>
|
||
</taxonomicName>
|
||
(an upland and highland species) might indicate a close relationship between these two taxa, while the smooth temporals could indicate a close relationship between
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFC1C3A8EB51BFF6E" box="[922,1085,152,177]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hoogmoedi">
|
||
<emphasis id="0AA4191DFF9EA96DFC1C3A8EB51BFF6E" box="[922,1085,152,177]" italics="true" pageId="13" pageNumber="14">A. hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFBF03A8EB437FF6E" box="[1142,1297,152,177]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="testigensis">
|
||
<emphasis id="0AA4191DFF9EA96DFBF03A8EB437FF6E" box="[1142,1297,152,177]" italics="true" pageId="13" pageNumber="14">A. testigensis</emphasis>
|
||
</taxonomicName>
|
||
, both being highland species.
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF9EA96DFF433AFEB2B8FE15" blockId="13.[151,1437,152,698]" pageId="13" pageNumber="14">
|
||
The completely opaque condition of the lower eyelid in
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFCE13AFEB5A5FEDE" box="[871,1155,232,257]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hoogmoedi">
|
||
<emphasis id="0AA4191DFF9EA96DFCE13AFEB5A5FEDE" box="[871,1155,232,257]" italics="true" pageId="13" pageNumber="14">Arthrosaura hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
is shared only with the recently described
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFEF53B06B329FEF6" box="[371,527,272,297]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="montigena">
|
||
<emphasis id="0AA4191DFF9EA96DFEF53B06B329FEF6" box="[371,527,272,297]" italics="true" pageId="13" pageNumber="14">A. montigena</emphasis>
|
||
</taxonomicName>
|
||
from Auyantepui (
|
||
<bibRefCitation id="5C41B8FEFF9EA96DFD683B06B529FEF5" author="Myers" box="[750,1039,272,298]" pageId="13" pageNumber="15" refString="Myers, C. W. & Donnelly, M. A. (2008) The summit herpetofauna of Auyantepui, Venezuela: report from the Robert G. Goelet American Museum - Terramar expedition. Bulletin of the American Museum of Natural History, 308, 1 - 147." type="journal article" year="2008">Myers & Donnelly 2008</bibRefCitation>
|
||
) and is a rare trait within the family
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFF383B2EB08DFE8D" box="[190,427,312,338]" class="Reptilia" family="Gymnophthalmidae" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="family">Gymnophthalmidae</taxonomicName>
|
||
. It is unknown whether this is an adaptive character to high UV radiation and intense sunshine sometimes occurring on high elevation tepuis, or whether it is a plesiomorphic character state (secondarily lost in other species).
|
||
<figureCitation id="A0EBD98AFF9EA96DFD873B9EB342FE7D" box="[513,612,392,418]" captionStart="FIGURE 7" captionStartId="9.[151,255,465,489]" captionTargetBox="[161,1424,186,443]" captionTargetId="figure@9.[161,1426,186,443]" captionTargetPageId="9" captionText="FIGURE 7. Lower eyelid condition in Arthrosaura hoogmoedi sp. nov. and two related Arthrosaura species. A: Semitransparent, unpigmented in A. kockii (IRSNB 15397). B: Semi-transparent, partly pigmented in A. guianensis (IRSNB 17343). C: Completely opaque, completely pigmented in A. hoogmoedi sp. nov. (IRSNB 2653, holotype)." httpUri="https://zenodo.org/record/184536/files/figure.png" pageId="13" pageNumber="14">Figure 7</figureCitation>
|
||
shows a comparison of lower eyelid condition between
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFB7F3B9EB4BAFE7E" box="[1273,1436,392,417]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hoogmoedi">
|
||
<emphasis id="0AA4191DFF9EA96DFB7F3B9EB4BAFE7E" box="[1273,1436,392,417]" italics="true" pageId="13" pageNumber="14">A. hoogmoedi</emphasis>
|
||
</taxonomicName>
|
||
and two related
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFED53BA6B0C5FE16" box="[339,483,432,457]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis id="0AA4191DFF9EA96DFED53BA6B0C5FE16" box="[339,483,432,457]" italics="true" pageId="13" pageNumber="14">Arthrosaura</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFDC93BA6B390FE16" box="[591,694,432,457]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="kockii">
|
||
<emphasis id="0AA4191DFF9EA96DFDC93BA6B390FE16" box="[591,694,432,457]" italics="true" pageId="13" pageNumber="14">A. kockii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFD693BA6B2ABFE16" box="[751,909,432,457]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="guianensis">
|
||
<emphasis id="0AA4191DFF9EA96DFD693BA6B2ABFE16" box="[751,909,432,457]" italics="true" pageId="13" pageNumber="14">A. guianensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF9EA96DFF433BCEB357FDB5" blockId="13.[151,1437,152,698]" pageId="13" pageNumber="14">
|
||
Recently
|
||
<bibRefCitation id="5C41B8FEFF9EA96DFEB23BCEB383FE2D" author="Myers" box="[308,677,472,498]" pageId="13" pageNumber="15" refString="Myers, C. W. & Donnelly, M. A. (2008) The summit herpetofauna of Auyantepui, Venezuela: report from the Robert G. Goelet American Museum - Terramar expedition. Bulletin of the American Museum of Natural History, 308, 1 - 147." type="journal article" year="2008">Myers and Donnelly (2008: 97)</bibRefCitation>
|
||
provided a key to the species of
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFBA73BCEB597FE2E" box="[1057,1201,472,497]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis id="0AA4191DFF9EA96DFBA73BCEB597FE2E" box="[1057,1201,472,497]" italics="true" pageId="13" pageNumber="14">Arthrosaura</emphasis>
|
||
</taxonomicName>
|
||
. Unfortunately they inverted some characters of
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFE623816B3A6FDC6" box="[484,640,512,537]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="testigensis">
|
||
<emphasis id="0AA4191DFF9EA96DFE623816B3A6FDC6" box="[484,640,512,537]" italics="true" pageId="13" pageNumber="14">A. testigensis</emphasis>
|
||
</taxonomicName>
|
||
with those of
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFCAD3816B2EDFDC6" box="[811,971,512,537]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="guianensis">
|
||
<emphasis id="0AA4191DFF9EA96DFCAD3816B2EDFDC6" box="[811,971,512,537]" italics="true" pageId="13" pageNumber="14">A. guianensis</emphasis>
|
||
</taxonomicName>
|
||
and their key should be used with caution: interparietal and parietals are keeled in
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFD18383EB21AFD9E" box="[670,828,552,577]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="guianensis">
|
||
<emphasis id="0AA4191DFF9EA96DFD18383EB21AFD9E" box="[670,828,552,577]" italics="true" pageId="13" pageNumber="14">A. guianensis</emphasis>
|
||
</taxonomicName>
|
||
, but not in
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFC46383EB57CFD9E" box="[960,1114,552,577]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="testigensis">
|
||
<emphasis id="0AA4191DFF9EA96DFC46383EB57CFD9E" box="[960,1114,552,577]" italics="true" pageId="13" pageNumber="14">A. testigensis</emphasis>
|
||
</taxonomicName>
|
||
, and temporals are small in
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFF113846B012FDB6" box="[151,308,592,617]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="guianensis">
|
||
<emphasis id="0AA4191DFF9EA96DFF113846B012FDB6" box="[151,308,592,617]" italics="true" pageId="13" pageNumber="14">A. guianensis</emphasis>
|
||
</taxonomicName>
|
||
, but large in
|
||
<taxonomicName id="FFD0BE8CFF9EA96DFE493846B34CFDB6" box="[463,618,592,617]" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="testigensis">
|
||
<emphasis id="0AA4191DFF9EA96DFE493846B34CFDB6" box="[463,618,592,617]" italics="true" pageId="13" pageNumber="14">A. testigensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="386FC50FFF9EA96DFF43386EB248FD65" blockId="13.[151,1437,152,698]" pageId="13" pageNumber="14">The only other lizard collected on the summit of Maringma tepui belongs to a possibly new gymnophthalmid genus, which will be treated in a separate publication.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |