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<mods:title id="7F2D26564CAE78DBDE2B1CB2A0BF1BD4">Not everything is what it seems: the study of type specimens of Cryptogenius Westwood, 1842 species (Coleoptera: Hybosoridae: Anaidinae) reveals a new species and new synonyms</mods:title>
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<emphasis id="0AB2C53BFFF4737C2595FCC73AC7E762" bold="true" box="[448,763,866,893]" italics="true" pageId="9" pageNumber="184">Cryptogenius westwoodi</emphasis>
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(
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, 
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)
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<emphasis id="0AB2C53BFFF4737C24F5FC6A3995E7F7" box="[160,425,975,1000]" italics="true" pageId="9" pageNumber="184">Cryptogenius miersianus</emphasis>
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[error]: 
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(not 1842a): 171 (
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); 
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<bibRefCitation id="5C5764D8FFF4737C279FFC6A38F0E015" author="Lacordaire JT" pageId="9" pageNumber="184" refId="ref8850" refString="Lacordaire JT. 1855. Histoire naturelle des insectes. Genera des coleopteres, ou expose methodique et critique de tous les genres proposes jusqu ̍ ici dans cet ordre d ̍ insectes. Tome troisieme. Contenant les familles des Pectinicornes et Lamellicornes. Librarie Encyclopedique de Roret, Paris. doi: 10.5962 / bhl. title. 8864." type="book" year="1855">Lacordaire 1855: 153</bibRefCitation>
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; 
<treatmentCitation id="B9673F38FFF4737C248FFC54398DE015" author="Harold E" box="[218,433,1009,1034]" page="1091" pageId="9" pageNumber="184" year="1869">
<bibRefCitation id="5C5764D8FFF4737C248FFC54398DE015" author="Harold E" box="[218,433,1009,1034]" pageId="9" pageNumber="184" pagination="979 - 1346" refId="ref8358" refString="Harold E. 1869. Tomo IV. Scarabaeidae. In: Gemminger &amp; Harold, editor. Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Monachii: Sumptu E. H. Gummi; p. 979 - 1346. doi: 10.5962 / bhl. title. 9089." type="book chapter" year="1869">Harold 1869: 1091</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="B9673F38FFF4737C2595FC543B06E015" author="Preudhomme de Borre FPCA" box="[448,826,1009,1034]" page="78" pageId="9" pageNumber="184" year="1886">
<bibRefCitation id="5C5764D8FFF4737C2595FC543B06E015" author="Preudhomme de Borre FPCA" box="[448,826,1009,1034]" pageId="9" pageNumber="184" pagination="56 - 82" refId="ref9063" refString="Preudhomme de Borre FPCA. 1886. Catalogue des Trogides decrits jusque ̍ a ce jour precede d ̍ un synopsis de leurs genres et d ̍ une esquisse de leur distribution geographique. Ann Soc Ent Belg. 30: 56 - 82." type="journal article" year="1886">Preudhomme de Borre 1886: 78</bibRefCitation>
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; 
<treatmentCitation id="B9673F38FFF4737C271CFC543C37E015" author="Arrow GJ" box="[841,1035,1009,1034]" page="501" pageId="9" pageNumber="184" year="1909">
<bibRefCitation id="5C5764D8FFF4737C271CFC543C37E015" author="Arrow GJ" box="[841,1035,1009,1034]" pageId="9" pageNumber="184" pagination="479 - 507" refId="ref7718" refString="Arrow GJ. 1909. On the characters and relationship of the less-known groups of lamellicorn Coleoptera, with descriptions of new species of Hybosorinae, etc. Trans Ent Soc Lond. 4: 479 - 507. doi: 10.1111 / j. 1365 - 2311.1909. tb 02183. x." type="journal article" year="1909">Arrow 1909: 501</bibRefCitation>
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, 
<treatmentCitation id="B9673F38FFF4737C204FFC543CBCE015" author="Arrow GJ" box="[1050,1152,1009,1034]" page="63" pageId="9" pageNumber="184" year="1912">
<bibRefCitation id="5C5764D8FFF4737C204FFC543CBCE015" author="Arrow GJ" box="[1050,1152,1009,1034]" pageId="9" pageNumber="184" pagination="52 - 66" refId="ref7772" refString="Arrow GJ. 1912. Trogidae. Coleopterorum Catalogue. 19 (43): 52 - 66." type="journal article" year="1912">1912: 63</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="B9673F38FFF4737C24F5FBB1398DE032" author="Blackwelder RE" box="[160,433,1044,1069]" page="219" pageId="9" pageNumber="184" year="1944">
<bibRefCitation id="5C5764D8FFF4737C24F5FBB1398DE032" author="Blackwelder RE" box="[160,433,1044,1069]" pageId="9" pageNumber="184" pagination="218 - 219" refId="ref7888" refString="Blackwelder RE. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America, pt. 2. Scarabaeidae, Troginae. Bull US Natl Mus. 185: 218 - 219." type="journal article" year="1944">Blackwelder 1944: 219</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="B9673F38FFF4737C2594FBB13AB7E032" author="Scholtz CH" box="[449,651,1044,1069]" page="19" pageId="9" pageNumber="184" year="1982">
<bibRefCitation id="5C5764D8FFF4737C2594FBB13AB7E032" author="Scholtz CH" box="[449,651,1044,1069]" pageId="9" pageNumber="184" pagination="1 - 27" refId="ref9143" refString="Scholtz CH. 1982. Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Mem Dep Agric Fish Repub S Afr. 54: 1 - 27." type="journal article" year="1982">Scholtz 1982: 19</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="B9673F38FFF4737C26C9FBB13BC4E032" author="Scholtz CH &amp; Hotman D &amp; Nel A" box="[668,1016,1044,1069]" page="193" pageId="9" pageNumber="184" year="1987">
<bibRefCitation id="5C5764D8FFF4737C26C9FBB13BC4E032" author="Scholtz CH &amp; Hotman D &amp; Nel A" box="[668,1016,1044,1069]" pageId="9" pageNumber="184" pagination="193 - 199" refId="ref9262" refString="Scholtz CH, D ̍ Hotman D, Nel A. 1987. Systematic position of the South American genus Cryptogenius Westwood (Coleoptera: Scarabaeoidea: Trogidae and Hybosoridae). Coleopt Bull. 41 (2): 193 - 199." type="journal article" year="1987">Scholtz et al. 1987: 193–194</bibRefCitation>
</treatmentCitation>
, 
<treatmentCitation id="B9673F38FFF4737C205CFBB13CBDE032" author="Scholtz CH &amp; Hotman D &amp; Evans AV &amp; Nel A" box="[1033,1153,1044,1069]" page="215" pageId="9" pageNumber="184" year="1988">
<bibRefCitation id="5C5764D8FFF4737C205CFBB13CBDE032" author="Scholtz CH &amp; Hotman D &amp; Evans AV &amp; Nel A" box="[1033,1153,1044,1069]" pageId="9" pageNumber="184" pagination="207 - 240" refId="ref9217" refString="Scholtz CH, D ̍ Hotman D, Evans AV, Nel A. 1988. Phylogeny and systematics of the Ochodaeidae (Insecta: Coleoptera: Scarabaeoidea). J Ent Soc Sth Afr. 51 (2): 207 - 240." type="journal article" year="1988">1988: 215</bibRefCitation>
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; 
<treatmentCitation id="B9673F38FFF4737C24F5FB923969E04F" author="Ratcliffe BC" box="[160,341,1079,1104]" page="7" pageId="9" pageNumber="184" year="2002">
<bibRefCitation id="5C5764D8FFF4737C24F5FB923969E04F" author="Ratcliffe BC" box="[160,341,1079,1104]" pageId="9" pageNumber="184" pagination="1 - 48" refId="ref9108" refString="Ratcliffe BC. 2002. A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa. 32 (1): 1 - 48. doi: 10.11646 / zootaxa. 32.1.1." type="journal article" year="2002">Ratcliffe 2002: 7</bibRefCitation>
</treatmentCitation>
; Ocampo 2006: 156–157; 
<treatmentCitation id="B9673F38FFF4737C262CFB923BE1E04F" author="Ocampo FC &amp; Ballerio A" box="[633,989,1079,1104]" page="181" pageId="9" pageNumber="184" year="2006">
<bibRefCitation id="5C5764D8FFF4737C262CFB923BE1E04F" author="Ocampo FC &amp; Ballerio A" box="[633,989,1079,1104]" pageId="9" pageNumber="184" pagination="178 - 209" refId="ref8959" refString="Ocampo FC, Ballerio A. 2006. Phylogenetic analysis of the scarab family Hybosoridae and monographic revision of the of the NewWorld subfamily Anaidinae (Coleoptera: Scarabaeoidea). 4. Catalog of the subfamilies Anaidinae, Ceratocanthinae, Hybosorinae, Liparochrinae, and Pachyplectrinae (Hybosoridae). Bull Univ Nebr State Mus. 19: 178 - 209." type="journal article" year="2006">Ocampo and Ballerio 2006: 181</bibRefCitation>
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; 
<treatmentCitation id="B9673F38FFF4737C27BDFB92394DE06D" author="Hielkema AJ &amp; Hielkema MA" page="15" pageId="9" pageNumber="184" year="2019">
<bibRefCitation id="5C5764D8FFF4737C27BDFB92394DE06D" author="Hielkema AJ &amp; Hielkema MA" pageId="9" pageNumber="184" pagination="1 - 306" refId="ref8409" refString="Hielkema AJ, Hielkema MA. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi. 0732: 1 - 306." type="book chapter" year="2019">Hielkema and Hielkema 2019: 15</bibRefCitation>
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.
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</paragraph>
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<emphasis id="0AB2C53BFFF4737C24F5FB3B38DFE0A7" bold="true" box="[160,227,1182,1208]" italics="true" pageId="9" pageNumber="184">Types</emphasis>
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<emphasis id="0AB2C53BFFF4737C24F5FB623936E0C4" bold="true" box="[160,266,1218,1243]" pageId="9" pageNumber="184">
<typeStatus id="E77DA78BFFF4737C24F5FB6238C7E0C7" box="[160,251,1223,1240]" pageId="9" pageNumber="184" type="holotype">HOLOTYPE</typeStatus>
(
</emphasis>
Ơ, 
<emphasis id="0AB2C53BFFF4737C257EFB6739B8E0C4" bold="true" box="[299,388,1218,1243]" pageId="9" pageNumber="184">CEMT) (</emphasis>
<figureCitation id="A0FD05ACFFF4737C25D0FB673A3DE0C4" box="[389,513,1218,1243]" captionStart="Figure 1" captionStartId="1.[160,225,1240,1263]" captionTargetBox="[163,1152,143,1206]" captionTargetId="figure-139@1.[162,1154,142,1208]" captionTargetPageId="1" captionText="Figure 1. Holotype of Cryptogenius westwoodi sp. nov. (Ò CEMT) in (a) dorsal (white arrow showing the diagonal carinae in the pronotum), (b) ventral and (c) lateral views (white arrow indicates the third carina extending from the humerus to declivous area). Habitus of male genitalia in (d) dorsal and (e) ventral view. (f) labels. Scale bars: a–c = 1 mm; d, e = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737075" httpUri="https://zenodo.org/record/7737075/files/figure.png" pageId="9" pageNumber="184">Figure 1a–f</figureCitation>
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| Chuyayacu. 
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<elevation id="B3EBFE1AFFF4737C25C5FB4039E5E0E1" box="[400,473,1253,1278]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="9" pageNumber="184" unit="m" value="810.0">810 m</elevation>
</quantity>
.
</specimenCount>
<date id="4C783FE9FFF4737C25B1FB403A0EE0E1" box="[484,562,1253,1278]" pageId="9" pageNumber="184" value="1999-05">
<collectingDate id="5C3CC601FFF4737C25B1FB403A0EE0E1" box="[484,562,1253,1278]" pageId="9" pageNumber="184" value="1999-05">V-1999</collectingDate>
</date>
| Fumigación dosel. PAraujo̍. Second label [red with black frame; printed]: ‘ 
<taxonomicName id="FFC662AAFFF4737C25CDFAA23A8FE13F" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[408,691,1287,1312]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="westwoodi">Cryptogenius westwoodi</taxonomicName>
| des. Costa-Silva &amp; | Vaz-de-Mello, 2022
</materialsCitation>
| 
<materialsCitation id="88AE1374FFF4737C24F5FA8F3B89E15C" ID-GBIF-Occurrence="4055506302" box="[160,949,1317,1347]" country="Ecuador" location="Chuyayacu" municipality="Ecuador" pageId="9" pageNumber="184" specimenCount="1" stateProvince="Pastaza" typeStatus="holotype">
<typeStatus id="E77DA78BFFF4737C24F5FA8F3926E15C" box="[160,282,1322,1347]" pageId="9" pageNumber="184" type="holotype">HOLOTYPE</typeStatus>
Ò̍ (
<figureCitation id="A0FD05ACFFF4737C2505FA8F3989E15C" box="[336,437,1322,1347]" captionStart="Figure 1" captionStartId="1.[160,225,1240,1263]" captionTargetBox="[163,1152,143,1206]" captionTargetId="figure-139@1.[162,1154,142,1208]" captionTargetPageId="1" captionText="Figure 1. Holotype of Cryptogenius westwoodi sp. nov. (Ò CEMT) in (a) dorsal (white arrow showing the diagonal carinae in the pronotum), (b) ventral and (c) lateral views (white arrow indicates the third carina extending from the humerus to declivous area). Habitus of male genitalia in (d) dorsal and (e) ventral view. (f) labels. Scale bars: a–c = 1 mm; d, e = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737075" httpUri="https://zenodo.org/record/7737075/files/figure.png" pageId="9" pageNumber="184">Figure 1f</figureCitation>
). 
<emphasis id="0AB2C53BFFF4737C259CFA8F3A5EE15C" bold="true" box="[457,610,1322,1347]" pageId="9" pageNumber="184">Type locality</emphasis>
: 
<collectingMunicipality id="D81D8353FFF4737C263AFA8F3AF1E15C" box="[623,717,1322,1347]" pageId="9" pageNumber="184">Ecuador</collectingMunicipality>
: 
<collectingRegion id="FA02D7CBFFF4737C2682FA8F3B11E15C" box="[727,813,1322,1347]" country="Ecuador" name="Pastaza" pageId="9" pageNumber="184">Pastaza</collectingRegion>
: 
<location id="3D194FF2FFF4737C276DFA8F3B89E15C" LSID="urn:lsid:plazi:treatment:B06FA83FFFF4737B2595FCC73A87E582:3D194FF2FFF4737C276DFA8F3B89E15C" box="[824,949,1322,1347]" country="Ecuador" municipality="Ecuador" name="Chuyayacu" pageId="9" pageNumber="184" stateProvince="Pastaza">Chuyayacu</location>
</materialsCitation>
.
</paragraph>
<paragraph id="38791929FFF473792495FAF73B28E692" blockId="9.[159,1157,1182,1728]" lastBlockId="12.[159,1156,142,653]" lastPageId="12" lastPageNumber="187" pageId="9" pageNumber="184">
<emphasis id="0AB2C53BFFF4737C2495FAF739EDE179" bold="true" box="[192,465,1357,1382]" pageId="9" pageNumber="184">
<typeStatus id="E77DA78BFFF4737C2495FAF73923E17C" box="[192,287,1362,1379]" pageId="9" pageNumber="184" type="paratype">PARATYPES</typeStatus>
(
<specimenCount id="2EC0D2A0FFF4737C2579FAE839F0E179" box="[300,460,1357,1382]" pageId="9" pageNumber="184" type="generic">17 specimens</specimenCount>
)
</emphasis>
: First label [white, typeset]: ‘ 
<collectingCountry id="40D159B9FFF4737C2754FAE83B4EE179" box="[769,882,1357,1382]" name="Venezuela" pageId="9" pageNumber="184">Venezuela</collectingCountry>
| 
<collectingRegion id="FA02D7CBFFF4737C27D2FAE83BECE179" box="[903,976,1357,1382]" country="Venezuela" name="Bolivar" pageId="9" pageNumber="184">Bolivar</collectingRegion>
| St. Elena-Paui | 10.- 
<date id="4C783FE9FFF4737C2497FACA390EE197" box="[194,306,1391,1416]" pageId="9" pageNumber="184" value="1977-12-09">12.9.1977</date>
̍. Second label [white, typeset]: ‘leg.Lindemann̍. Third label [yellow with black frame; printed]: ‘ 
<taxonomicName id="FFC662AAFFF4737C251EFA373A65E1B4" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[331,601,1426,1451]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="westwoodi">Cryptogenius westwoodi</taxonomicName>
| des. Costa-Silva &amp; | Vaz-de-Mello, 2022 | 
<typeStatus id="E77DA78BFFF4737C2047FA373CBEE1B4" box="[1042,1154,1426,1451]" pageId="9" pageNumber="184" type="paratype">PARATYPE</typeStatus>
Ò̍ 
<emphasis id="0AB2C53BFFF4737C2496FA1038E6E1D1" bold="true" box="[195,218,1461,1486]" pageId="9" pageNumber="184">(1</emphasis>
Ơ 
<emphasis id="0AB2C53BFFF4737C24A7FA103907E1D1" bold="true" box="[242,315,1461,1486]" pageId="9" pageNumber="184">CEMT)</emphasis>
. First label [white with black border,printed]:‘ 
<collectingCountry id="40D159B9FFF4737C274EFA103BB2E1D1" box="[795,910,1461,1486]" name="Ecuador" pageId="9" pageNumber="184">ECUADOR</collectingCountry>
: 
<collectingRegion id="FA02D7CBFFF4737C27C0FA103BD4E1D1" box="[917,1000,1461,1486]" country="Ecuador" name="Pastaza" pageId="9" pageNumber="184">Pastaza</collectingRegion>
,Arajuno,| B.P. Oglán, 
<quantity id="FF3EB4CCFFF4737C24B9FA72390FE1EF" box="[236,307,1495,1520]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="9" pageNumber="184" unit="m" value="640.0">640 m</quantity>
, 
<date id="4C783FE9FFF4737C2568FA72398EE1EF" box="[317,434,1495,1520]" pageId="9" pageNumber="184" value="2009-06-25">25-VI-2009</date>
, | Plato Amarillo. WChamorro y | GGallo̍. Second label [yellow with black frame; printed]: ‘ 
<taxonomicName id="FFC662AAFFF4737C25C2FA5F3A96E20C" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[407,682,1530,1555]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="westwoodi">Cryptogenius westwoodi</taxonomicName>
| des. Costa-Silva &amp; | Vaz-de-Mello, 2022 | 
<typeStatus id="E77DA78BFFF4737C24CAF9B83933E229" box="[159,271,1565,1590]" pageId="9" pageNumber="184" type="paratype">PARATYPE</typeStatus>
ƪ̍ 
<emphasis id="0AB2C53BFFF4737C256EF9B8396DE229" bold="true" box="[315,337,1565,1590]" pageId="9" pageNumber="184">
(
<specimenCount id="2EC0D2A0FFF4737C2510F9B8396DE229" box="[325,337,1565,1590]" pageId="9" pageNumber="184" type="female">1</specimenCount>
</emphasis>
♀ 
<emphasis id="0AB2C53BFFF4737C253FF9B8398FE229" bold="true" box="[362,435,1565,1590]" pageId="9" pageNumber="184">
<collectionCode id="5ED781ECFFF4737C253FF9B83991E229" box="[362,429,1565,1590]" pageId="9" pageNumber="184">CEMT</collectionCode>
)
</emphasis>
. First label [white with black border, printed]: ‘ 
<collectingCountry id="40D159B9FFF4737C27C8F9B83C2CE229" box="[925,1040,1565,1590]" name="Ecuador" pageId="9" pageNumber="184">ECUADOR</collectingCountry>
: 
<collectingRegion id="FA02D7CBFFF4737C204DF9B83C48E229" box="[1048,1140,1565,1590]" country="Ecuador" name="Orellana" pageId="9" pageNumber="184">Orellana</collectingRegion>
, | Yasuní, Río Rumiyacu, | Apaica 
<quantity id="FF3EB4CCFFF4737C25B8F99A3A77E247" box="[493,587,1599,1624]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="9" pageNumber="184" unit="m" value="2220.0">2, 220 m</quantity>
, 76° | 42ʹ0̍W, 0°48ʹ36̎S, 10-X- | 2000. P Araujo fum̎. Second label [white]: ‘ 
<taxonomicName id="FFC662AAFFF4737C25DDF9C73A27E264" authorityName="Linnaeus" authorityYear="1758" box="[392,539,1634,1659]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="order">COLEOPTERA</taxonomicName>
| Scarabaeidae-Hybosorinae | 
<taxonomicName id="FFC662AAFFF4737C270AF9C73BD3E264" authorityName="Westwood" authorityYear="1842" box="[863,1007,1634,1659]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="genus">Cryptogenius</taxonomicName>
| miersianus | West, 1846 | Det: William Chamorro̍. Third label [yellow with black frame; printed]: ‘ 
<taxonomicName id="FFC662AAFFF4737C24F7F9023981E2DF" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[162,445,1703,1728]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="westwoodi">Cryptogenius westwoodi</taxonomicName>
| des. Costa-Silva &amp; | Vaz-de-Mello, 2022 | 
<typeStatus id="E77DA78BFFF4737C27E5F9023C1FE2DF" box="[944,1059,1703,1728]" pageId="9" pageNumber="184" type="paratype">PARATYPE</typeStatus>
ƪ̍ 
<emphasis id="0AB2C53BFFF4737C200DF9023C53E2DF" bold="true" box="[1112,1135,1703,1728]" pageId="9" pageNumber="184">
(
<specimenCount id="2EC0D2A0FFF4737C2036F9023C53E2DF" box="[1123,1135,1703,1728]" pageId="9" pageNumber="184" type="female">1</specimenCount>
</emphasis>
♀ 
<emphasis id="0AB2C53BFFF6737E24F5FC8738D5E724" bold="true" box="[160,233,802,827]" pageId="11" pageNumber="186">
<collectionCode id="5ED781ECFFF6737E24F5FC8738DFE724" box="[160,227,802,827]" pageId="11" pageNumber="186">CEMT</collectionCode>
)
</emphasis>
.First label [white, printed]:‘ 
<collectingCountry id="40D159B9FFF6737E265EFC873A42E724" box="[523,638,802,827]" name="Ecuador" pageId="11" pageNumber="186">ECUADOR</collectingCountry>
: 
<collectingRegion id="FA02D7CBFFF6737E26D0FC873ADDE724" box="[645,737,802,827]" country="Ecuador" name="Orellana" pageId="11" pageNumber="186">Orellana</collectingRegion>
,| Yasuní,Río Rumiyacu,| Apaika, 
<quantity id="FF3EB4CCFFF6737E206EFC873C43E724" box="[1083,1151,802,827]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="186" unit="m" value="220.0">220 m</quantity>
, 76°25̍ | 48̍W, 
<geoCoordinate id="5DF27FEEFFF6737E256DFCE039AAE741" box="[312,406,836,862]" degrees="0" direction="south" minutes="41" orientation="latitude" pageId="11" pageNumber="186" precision="15" seconds="16" value="-0.68777776">0°41ʹ16S</geoCoordinate>
, 08-X- | 2000, P Araujo fumig̍. Second label [yellow with black frame; printed]: ‘ 
<taxonomicName id="FFC662AAFFF6737E2554FCC23A32E79F" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[257,526,871,896]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="westwoodi">Cryptogenius westwoodi</taxonomicName>
| des.Costa-Silva &amp; | Vaz-de-Mello,2022 | 
<typeStatus id="E77DA78BFFF6737E2795FCC23C0CE79F" box="[960,1072,871,896]" pageId="11" pageNumber="186" type="paratype">PARATYPE</typeStatus>
Ò̍ 
<emphasis id="0AB2C53BFFF6737E200CFCC23C53E79F" bold="true" box="[1113,1135,871,896]" pageId="11" pageNumber="186">(1</emphasis>
Ơ 
<emphasis id="0AB2C53BFFF6737E24CAFC2F38D5E7BC" bold="true" box="[159,233,906,931]" pageId="11" pageNumber="186">CEMT)</emphasis>
. First label [white with black border, printed]: ‘ 
<collectingCountry id="40D159B9FFF6737E2685FC2F3B37E7BC" box="[720,779,906,931]" name="Brazil" pageId="11" pageNumber="186">Brasil</collectingCountry>
(MA), Bom Jardim | REBIO-Res, Biol. Gurupi | Armad. Luminosa Base̍. Second label [white with black border, printed]: ‘ 
<date id="4C783FE9FFF6737E204FFC0838EAE7F7" pageId="11" pageNumber="186" value="2010-06-05" valueMax="2010-06-15" valueMin="2010-06-05">05–15. vi. 2010</date>
, J. C. Silva, J. A. Silva, A. A. Santos, T. T. A. Silva̍. Third label [yellow with black frame; printed]: ‘ 
<taxonomicName id="FFC662AAFFF6737E2554FC573A32E014" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[257,526,1010,1035]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="westwoodi">Cryptogenius westwoodi</taxonomicName>
| des.Costa-Silva &amp; | Vaz-de-Mello,2022 | 
<typeStatus id="E77DA78BFFF6737E2795FC573C0CE014" box="[960,1072,1010,1035]" pageId="11" pageNumber="186" type="paratype">PARATYPE</typeStatus>
ƪ̍ 
<emphasis id="0AB2C53BFFF6737E200CFC573C53E014" bold="true" box="[1113,1135,1010,1035]" pageId="11" pageNumber="186">
(
<specimenCount id="2EC0D2A0FFF6737E2036FC573C53E014" box="[1123,1135,1010,1035]" pageId="11" pageNumber="186" type="female">1</specimenCount>
</emphasis>
♀ 
<emphasis id="0AB2C53BFFF6737E24CAFBB038D5E031" bold="true" box="[159,233,1045,1070]" pageId="11" pageNumber="186">
<collectionCode id="5ED781ECFFF6737E24CAFBB038DEE031" box="[159,226,1045,1070]" pageId="11" pageNumber="186">CEMT</collectionCode>
)
</emphasis>
. First label [white with black border, printed]: ‘ 
<collectingCountry id="40D159B9FFF6737E2688FBB03B11E031" box="[733,813,1045,1070]" name="Brazil" pageId="11" pageNumber="186">BRASIL</collectingCountry>
: 
<collectingRegion id="FA02D7CBFFF6737E2763FBB03BFDE031" box="[822,961,1045,1070]" country="Brazil" name="Mato Grosso" pageId="11" pageNumber="186">Mato Grosso</collectingRegion>
, | Cotriguaçu, Faz. São | Nicolau, PPBio 2, | 09°49ʹ12̍S, 58°15ʹ57̎W, | decayed log, 
<date id="4C783FE9FFF6737E2719FB923B88E04F" box="[844,948,1079,1104]" pageId="11" pageNumber="186" value="2018-10-27">27.x.2018</date>
, | Costa-Silva et al.̍ Second label [yellow with black frame; printed]: ‘ 
<taxonomicName id="FFC662AAFFF6737E26CAFBFF3B90E06C" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[671,940,1114,1139]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="westwoodi">Cryptogenius westwoodi</taxonomicName>
| des. Costa-Silva &amp; | Vaz-de-Mello, 2022 | 
<typeStatus id="E77DA78BFFF6737E25DDFBD839C4E089" box="[392,504,1149,1174]" pageId="11" pageNumber="186" type="paratype">PARATYPE</typeStatus>
ƪ̍ 
<emphasis id="0AB2C53BFFF6737E267FFBD83A7CE089" bold="true" box="[554,576,1149,1174]" pageId="11" pageNumber="186">
(
<specimenCount id="2EC0D2A0FFF6737E2661FBD83A7CE089" box="[564,576,1149,1174]" pageId="11" pageNumber="186" type="female">1</specimenCount>
</emphasis>
♀ 
<emphasis id="0AB2C53BFFF6737E2608FBD83A9AE089" bold="true" box="[605,678,1149,1174]" pageId="11" pageNumber="186">
<collectionCode id="5ED781ECFFF6737E2608FBD83A9CE089" box="[605,672,1149,1174]" pageId="11" pageNumber="186">CEMT</collectionCode>
)
</emphasis>
. First label [white, aged with black border, printed]: 
<collectingCountry id="40D159B9FFF6737E2554FB3A3966E0A7" box="[257,346,1183,1208]" name="Brazil" pageId="11" pageNumber="186">Brasilien</collectingCountry>
| 
<collectingRegion id="FA02D7CBFFF6737E253AFB3A39C4E0A7" box="[367,504,1183,1208]" country="Brazil" name="Mato Grosso" pageId="11" pageNumber="186">Mato Grosso</collectingRegion>
| Gleba Arinos – 
<quantity id="FF3EB4CCFFF6737E26E4FB3A3AC9E0A7" box="[689,757,1183,1208]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="186" unit="m" value="350.0">350 m</quantity>
| 
<date id="4C783FE9FFF6737E275FFB3A3B4EE0A7" box="[778,882,1183,1208]" pageId="11" pageNumber="186" value="1967-10-31">31.X.1967</date>
| leg. Viehmann̍. Second label [yellow with black frame; printed]: ‘ 
<taxonomicName id="FFC662AAFFF6737E261CFB673B6BE0C4" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[585,855,1218,1243]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="westwoodi">Cryptogenius westwoodi</taxonomicName>
| des. Costa-Silva &amp; | Vaz-deMello, 2022 | 
<typeStatus id="E77DA78BFFF6737E2564FB40399DE0E1" box="[305,417,1253,1278]" pageId="11" pageNumber="186" type="paratype">PARATYPE</typeStatus>
ƪ̍ 
<emphasis id="0AB2C53BFFF6737E259AFB4039DAE0E1" bold="true" box="[463,486,1253,1278]" pageId="11" pageNumber="186">
(
<specimenCount id="2EC0D2A0FFF6737E258FFB4039DAE0E1" box="[474,486,1253,1278]" pageId="11" pageNumber="186" type="female">1</specimenCount>
</emphasis>
♀ 
<emphasis id="0AB2C53BFFF6737E2655FB403A76E0E1" bold="true" box="[512,586,1253,1278]" pageId="11" pageNumber="186">
<collectionCode id="5ED781ECFFF6737E2655FB403A7FE0E1" box="[512,579,1253,1278]" pageId="11" pageNumber="186">CEMT</collectionCode>
)
</emphasis>
. First label [white, aged, handwritten]: ‘NG*̍. Second label [white, aged, handwritten]: ‘ 
<taxonomicName id="FFC662AAFFF6737E2651FAA23AABE13F" authorityName="Westwood" authorityYear="1842" box="[516,663,1287,1312]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="genus">Cryptogenius</taxonomicName>
| miersianus | Westw-̍. Third label [white with red border]:‘TYPE | WESTWOOD | Trans.Ent.Soc.| 4. P.171. | T.12.1.1 | Coll.Hope.Oxon̍. Fourth label [white, unknown handwriting]: ‘ 
<typeStatus id="E77DA78BFFF6737E2662FAE83A9FE179" box="[567,675,1357,1382]" pageId="11" pageNumber="186" type="lectotype">lectotype</typeStatus>
̍. Fifth label [white with black border]: ‘TYPE COL:353 ½ | 
<taxonomicName id="FFC662AAFFF6737E2571FACA3988E197" authorityName="Westwood" authorityYear="1842" box="[292,436,1391,1416]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="genus">Cryptogenius</taxonomicName>
| miersianus | Westw.| HOPE DEPT.OXFORD̍.Sixth label [yellow with black frame]: ‘ 
<taxonomicName id="FFC662AAFFF6737E257AFA373A01E1B4" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[303,573,1426,1451]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="westwoodi">Cryptogenius westwoodi</taxonomicName>
| des. Costa-Silva &amp; | Vaz-de-Mello,2022 | 
<typeStatus id="E77DA78BFFF6737E27A7FA373C5AE1B4" box="[1010,1126,1426,1451]" pageId="11" pageNumber="186" type="paratype">PARATYPE</typeStatus>
̍ 
<emphasis id="0AB2C53BFFF6737E2039FA37392CE1D1" bold="true" pageId="11" pageNumber="186">
(1 
<collectionCode id="5ED781ECFFF6737E24CAFA1038C2E1D1" box="[159,254,1461,1486]" pageId="11" pageNumber="186">OUMNH</collectionCode>
–
</emphasis>
<figureCitation id="A0FD05ACFFF6737E2543FA1039ADE1D1" box="[278,401,1461,1486]" captionStart="Figure 3" captionStartId="6.[160,225,784,807]" captionTargetBox="[169,1134,143,749]" captionTargetId="figure-170@6.[162,1154,142,752]" captionTargetPageId="6" captionText="Figure 3. Paratype of Cryptogenius westwoodi sp. nov. (1 OUMNH) in (a) dorsal and (b) lateral views. (c) labels. Scale bars: 1 mm. Photographs (a, b): Keita Matsumoto, BMNH." figureDoi="http://doi.org/10.5281/zenodo.7737081" httpUri="https://zenodo.org/record/7737081/files/figure.png" pageId="11" pageNumber="186">Figure 3a–c</figureCitation>
). First label [white, aged, typeset]: ‘Serra de Baturité | (
<collectingRegion id="FA02D7CBFFF6737E279EFA103C37E1D1" box="[971,1035,1461,1486]" country="Brazil" name="Ceara" pageId="11" pageNumber="186">Céara</collectingRegion>
) | Gounelle 1.1895̍.Second label [white, aged,typeset]:‘MUSÉUM PARIS | 1895 | COLL.E GOUNELLE̍.Third label [white, aged, handwritten]:‘ 
<taxonomicName id="FFC662AAFFF6737E25ABFA5F3AADE20C" authorityName="Westwood" authorityYear="1842" box="[510,657,1530,1555]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="genus">Cryptogenius</taxonomicName>
| miersianus | Westw̍.Fourth label [yellow with black frame, Vinícius Costa-Silva̍ s handwriting]: ‘ 
<taxonomicName id="FFC662AAFFF6737E26C9F9B83B13E229" authorityName="Westwood" authorityYear="1842" box="[668,815,1565,1590]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="genus">Cryptogenius</taxonomicName>
| westwoodi | Costa-Silva &amp; Vazde-Mello | 2022 | 
<typeStatus id="E77DA78BFFF6737E2502F99A39F7E247" box="[343,459,1599,1624]" pageId="11" pageNumber="186" type="paratype">PARATYPE</typeStatus>
̍ 
<emphasis id="0AB2C53BFFF6737E2587F99A39D4E247" bold="true" box="[466,488,1599,1624]" pageId="11" pageNumber="186">
(
<specimenCount id="2EC0D2A0FFF6737E2589F99A39D4E247" box="[476,488,1599,1624]" pageId="11" pageNumber="186" type="female">1</specimenCount>
</emphasis>
♀ 
<emphasis id="0AB2C53BFFF6737E2654F99A3A69E247" bold="true" box="[513,597,1599,1624]" pageId="11" pageNumber="186">
<collectionCode id="5ED781ECFFF6737E2654F99A3A72E247" box="[513,590,1599,1624]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="186">MNHN</collectionCode>
)
</emphasis>
. First label [white, printed]: ‘ 
<collectingCountry id="40D159B9FFF6737E2728F99A3BE1E247" box="[893,989,1599,1624]" name="French Guiana" pageId="11" pageNumber="186">GUYANE</collectingCountry>
| Approuague | Saut Mapaou | Battage̍. Second label [white, printed]: ‘ 
<date id="4C783FE9FFF6737E268AF9C73B67E264" box="[735,859,1634,1659]" pageId="11" pageNumber="186" value="2016-08-17">17.VIII.2016</date>
| J. Touroult leg̍. Third label [yellow with black frame, Vinícius Costa-Silva̍s handwriting]: ‘ 
<taxonomicName id="FFC662AAFFF6737E2718F9203BDDE281" authorityName="Westwood" authorityYear="1842" box="[845,993,1669,1694]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="genus">Cryptogenius</taxonomicName>
| westwoodi | Costa-Silva &amp; Vaz-de-Mello | 2022 | 
<typeStatus id="E77DA78BFFF6737E2675F9023AA8E2DF" box="[544,660,1703,1728]" pageId="11" pageNumber="186" type="paratype">PARATYPE</typeStatus>
̍ 
<emphasis id="0AB2C53BFFF6737E26C8F9023AC2E2DF" bold="true" box="[669,766,1703,1728]" pageId="11" pageNumber="186">
(1 
<collectionCode id="5ED781ECFFF6737E26EEF9023AF1E2DF" box="[699,717,1703,1728]" pageId="11" pageNumber="186">C</collectionCode>
]BH)
</emphasis>
. First label [white with black frame, printed]: ‘ 
<collectingCountry id="40D159B9FFF173792556FF2B395FE4B8" box="[259,355,142,167]" name="French Guiana" pageId="12" pageNumber="187">GUYANE</collectingCountry>
F., Saül, Grand | Boeuf Mort, 
<date id="4C783FE9FFF1737926C1FF2B3AD8E4B8" box="[660,740,142,167]" pageId="12" pageNumber="187" value="2007-11">xi-2007</date>
, | vitre [flight interceptation trap], P.-H. Dalens leg̍. Second label [yellow with black frame, Vinícius Costa-Silva̍s handwriting]: ‘ 
<taxonomicName id="FFC662AAFFF1737924F7FF713909E4F2" authorityName="Westwood" authorityYear="1842" box="[162,309,212,237]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="genus">Cryptogenius</taxonomicName>
| westwoodi | Costa-Silva &amp; Vaz-de-Mello | 2022 | 
<typeStatus id="E77DA78BFFF173792733FF713BE6E4F2" box="[870,986,212,237]" pageId="12" pageNumber="187" type="paratype">PARATYPE</typeStatus>
̍ 
<emphasis id="0AB2C53BFFF1737927B1FF713C7BE4F2" bold="true" box="[996,1095,212,237]" pageId="12" pageNumber="187">
(1 
<collectionCode id="5ED781ECFFF173792051FF713C2AE4F2" box="[1028,1046,212,237]" pageId="12" pageNumber="187">C</collectionCode>
]BH)
</emphasis>
. First label [white, printed]: ‘ 
<collectingCountry id="40D159B9FFF1737925DCFF5339D8E510" box="[393,484,246,271]" name="Bolivia" pageId="12" pageNumber="187">BOLIVIA</collectingCountry>
: 
<collectingRegion id="FA02D7CBFFF1737925B9FF533A62E510" box="[492,606,246,271]" country="Bolivia" name="Santa Cruz" pageId="12" pageNumber="187">Santa Cruz</collectingRegion>
| Amboro National Park | Los Volcanes, c. 
<quantity id="FF3EB4CCFFF173792041FF533C59E510" box="[1044,1125,246,271]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="187" unit="m" value="1000.0">1000 m</quantity>
| S 18° 06̍: W 63° 36̍| 
<date id="4C783FE9FFF173792534FEBC3A26E52D" box="[353,538,281,306]" pageId="12" pageNumber="187" value="2004-11-20" valueMax="2004-12-12" valueMin="2004-11-20">20/xi-12/xii/2004</date>
̍.Second label [white,printed]: ‘MV Light Sheet | Barcley.M. V.L. | &amp; Mendel, H. | BMNH(E) 2004–280̍. Third label [yellow with black frame, Vinícius CostaSilva̍s handwriting]: ‘ 
<taxonomicName id="FFC662AAFFF1737925DEFEFB3A1EE568" authorityName="Westwood" authorityYear="1842" box="[395,546,350,375]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="genus">Cryptogenius</taxonomicName>
/westwoodi/des. Costa-Silva &amp; Vaz-de-Mello/ 
<typeStatus id="E77DA78BFFF17379205DFEFB3CBEE568" box="[1032,1154,350,375]" pageId="12" pageNumber="187" type="paratype">PARATYPE</typeStatus>
2022̍ 
<emphasis id="0AB2C53BFFF1737924B7FE2438C4E585" bold="true" box="[226,248,385,410]" pageId="12" pageNumber="187">(1</emphasis>
Ơ 
<emphasis id="0AB2C53BFFF173792541FE243964E585" bold="true" box="[276,344,385,410]" pageId="12" pageNumber="187">
and 
<specimenCount id="2EC0D2A0FFF17379251CFE243964E585" box="[329,344,385,410]" pageId="12" pageNumber="187" type="female">3</specimenCount>
</emphasis>
♀ 
<emphasis id="0AB2C53BFFF173792526FE2439F9E585" bold="true" box="[371,453,385,410]" pageId="12" pageNumber="187">BMNH)</emphasis>
. First label [white, handwritten]: ‘Fr. 
<collectingCountry id="40D159B9FFF173792700FE243B96E585" box="[853,938,385,410]" name="French Guiana" pageId="12" pageNumber="187">Guyane</collectingCountry>
: Saint-Lourent | du-Maroni, Mont Itoupé, | 03°01ʹN, 53°05ʹW, | 
<date id="4C783FE9FFF17379263EFE013A91E5A2" box="[619,685,420,445]" pageId="12" pageNumber="187" value="2016-01">I.2016</date>
, SEAG leg̍. Second label [yellow with black frame, Vinícius Costa-Silva̍ s handwriting]: ‘ 
<taxonomicName id="FFC662AAFFF17379263EFE633AC2E5C0" authorityName="Westwood" authorityYear="1842" box="[619,766,454,479]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="genus">Cryptogenius</taxonomicName>
ƪ | westwoodi | des. Costa-Silva &amp; Vaz-de-Mello | 
<typeStatus id="E77DA78BFFF173792514FE4C398DE61D" box="[321,433,489,514]" pageId="12" pageNumber="187" type="paratype">PARATYPE</typeStatus>
2022̍ 
<emphasis id="0AB2C53BFFF1737925AFFE4C3A2CE61D" bold="true" box="[506,528,489,514]" pageId="12" pageNumber="187">
(
<specimenCount id="2EC0D2A0FFF173792651FE4C3A2CE61D" box="[516,528,489,514]" pageId="12" pageNumber="187" type="female">1</specimenCount>
</emphasis>
♀ 
<emphasis id="0AB2C53BFFF17379267EFE4C3A49E61D" bold="true" box="[555,629,489,514]" pageId="12" pageNumber="187">
<collectionCode id="5ED781ECFFF17379267EFE4C3A52E61D" box="[555,622,489,514]" pageId="12" pageNumber="187">CEMT</collectionCode>
)
</emphasis>
. First label [white, printed]: ‘ 
<collectingCountry id="40D159B9FFF1737927F3FE4C3BDFE61D" box="[934,995,489,514]" name="Brazil" pageId="12" pageNumber="187">Brasil</collectingCountry>
, RO, Itapuã do Oeste / Flona Jamari, Trilha Pedra / Grande, 09°11̍39.4̎S – / 63°04̍55.3̎W, 
<date id="4C783FE9FFF1737927E8FDA93C14E63A" box="[957,1064,524,549]" pageId="12" pageNumber="187" value="2014-10-06">06.x.2014</date>
̍. Second label [white, printed]:‘Facheamento,J.A. Rafael / F.F. Xavier F, R.M. Vieira &amp; / R.H. Aquino̍. Third label [yellow with black frame, Vinícius Costa-Silva̍s handwriting]: ‘ 
<taxonomicName id="FFC662AAFFF173792708FDF43BCCE675" authorityName="Westwood" authorityYear="1842" box="[861,1008,593,618]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="genus">Cryptogenius</taxonomicName>
| westwoodi | des. Costa-Silva &amp; Vaz-de-Mello | 
<typeStatus id="E77DA78BFFF173792655FDD13A4CE692" box="[512,624,628,653]" pageId="12" pageNumber="187" type="paratype">PARATYPE</typeStatus>
2023̍ (1 
<collectionCode id="5ED781ECFFF173792687FDD13B37E692" box="[722,779,628,653]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="12" pageNumber="187">INPA</collectionCode>
).
</paragraph>
<caption id="6CB949A1FFF7737F24F5FA103B51E25E" ID-DOI="http://doi.org/10.5281/zenodo.7737083" ID-Zenodo-Dep="7737083" httpUri="https://zenodo.org/record/7737083/files/figure.png" pageId="10" pageNumber="185" startId="10.[160,225,1461,1484]" targetBox="[163,1152,143,1428]" targetPageId="10">
<paragraph id="38791929FFF7737F24F5FA103B51E25E" blockId="10.[160,1156,1461,1601]" pageId="10" pageNumber="185">
<emphasis id="0AB2C53BFFF7737F24F5FA1038C5E1D3" bold="true" box="[160,249,1461,1484]" pageId="10" pageNumber="185">Figure 4.</emphasis>
Distribution map of known records of 
<taxonomicName id="FFC662AAFFF7737F2622FA103B7EE1D3" box="[631,834,1461,1484]" pageId="10" pageNumber="185">
<emphasis id="0AB2C53BFFF7737F2622FA103ACFE1D3" box="[631,755,1461,1484]" italics="true" pageId="10" pageNumber="185">Cryptogenius</emphasis>
species
</taxonomicName>
: 
<taxonomicName id="FFC662AAFFF7737F271EFA103BF5E1D3" authorityName="Westwood" authorityYear="1842" box="[843,969,1461,1484]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="miersianus">
<emphasis id="0AB2C53BFFF7737F271EFA103BF5E1D3" box="[843,969,1461,1484]" italics="true" pageId="10" pageNumber="185">C. miersianus</emphasis>
</taxonomicName>
(red circles) and 
<taxonomicName id="FFC662AAFFF7737F2024FA10393AE1F6" authority="Costa-Silva &amp; Vaz-de-Mello, 2023" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
<emphasis id="0AB2C53BFFF7737F2024FA10393AE1F6" italics="true" pageId="10" pageNumber="185">C. westwoodi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11817840FFF7737F2559FA773964E1F6" box="[268,344,1490,1513]" pageId="10" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
(blue circles). Red and blue stars represent records from the literature that possibly refer, according to the known geographical distribution, to 
<taxonomicName id="FFC662AAFFF7737F26A0FA553B48E218" authorityName="Westwood" authorityYear="1842" box="[757,884,1520,1543]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="miersianus">
<emphasis id="0AB2C53BFFF7737F26A0FA553B48E218" box="[757,884,1520,1543]" italics="true" pageId="10" pageNumber="185">C. miersianus</emphasis>
</taxonomicName>
and 
<taxonomicName id="FFC662AAFFF7737F27FCFA553C15E218" authority="Costa-Silva &amp; Vaz-de-Mello, 2023" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[937,1065,1520,1543]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
<emphasis id="0AB2C53BFFF7737F27FCFA553C15E218" box="[937,1065,1520,1543]" italics="true" pageId="10" pageNumber="185">C. westwoodi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11817840FFF7737F2064FA553CBDE218" box="[1073,1153,1520,1543]" pageId="10" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
, respectively, found in 
<bibRefCitation id="5C5764D8FFF7737F2529F9A83A27E23B" author="Ide S &amp; Costa C &amp; Vanin SA" box="[380,539,1549,1572]" pageId="10" pageNumber="185" pagination="757 - 765" refId="ref8758" refString="Ide S, Costa C, Vanin SA. 1990. Redescription of Cryptogenius fryi Arrow, 1909 with notes on sexual dimorphism and biological data (Coleoptera, Hybosoridae). Rev bras ent. 34 (4): 757 - 765." type="journal article" year="1990">Ide et al. (1990)</bibRefCitation>
, 
<bibRefCitation id="5C5764D8FFF7737F2670F9A83A85E23B" author="Ratcliffe BC" box="[549,697,1549,1572]" pageId="10" pageNumber="185" pagination="1 - 48" refId="ref9108" refString="Ratcliffe BC. 2002. A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa. 32 (1): 1 - 48. doi: 10.11646 / zootaxa. 32.1.1." type="journal article" year="2002">Ratcliffe (2002)</bibRefCitation>
and Ocampo (2006). These records illustrated with stars represent specimens not personally examined by the authors.
</paragraph>
</caption>
<caption id="6CB949A1FFF6737E24CAFD323AB4E6D4" ID-DOI="http://doi.org/10.5281/zenodo.7737087" ID-Zenodo-Dep="7737087" httpUri="https://zenodo.org/record/7737087/files/figure.png" pageId="11" pageNumber="186" startId="11.[159,224,663,686]" targetBox="[163,1153,143,630]" targetPageId="11">
<paragraph id="38791929FFF6737E24CAFD323AB4E6D4" blockId="11.[159,1155,660,715]" pageId="11" pageNumber="186">
<emphasis id="0AB2C53BFFF6737E24CAFD3238C7E6B1" bold="true" box="[159,251,663,686]" pageId="11" pageNumber="186">Figure 5.</emphasis>
Holotype of 
<taxonomicName id="FFC662AAFFF6737E25D5FD323AC9E6B1" authority="Krikken, 1975" authorityName="Krikken" authorityYear="1975" box="[384,757,663,686]" class="Insecta" family="Hybosoridae" genus="Cremastochilodius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="0AB2C53BFFF6737E25D5FD323A54E6B1" box="[384,616,663,686]" italics="true" pageId="11" pageNumber="186">Cremastochilodius tristis</emphasis>
<bibRefCitation id="5C5764D8FFF6737E263AFD323AC9E6B1" author="Krikken J" box="[623,757,663,686]" pageId="11" pageNumber="186" pagination="190 - 192" refId="ref8803" refString="Krikken J. 1975. Cremastochilodius tristis a new Scarabaeoid genus and species from Brazil. Entomol Ber. 35: 190 - 192." type="journal article" year="1975">Krikken, 1975</bibRefCitation>
</taxonomicName>
(ƪ RMNH) in (a) dorsal and (b) ventral view. (c) labels. Photographs: Oscar Vorst, RMNH.
</paragraph>
</caption>
<paragraph id="38791929FFF1737924F5FD1D38E4E6CD" blockId="12.[160,1156,696,792]" box="[160,216,696,722]" pageId="12" pageNumber="187">
<emphasis id="0AB2C53BFFF1737924F5FD1D38E4E6CD" bold="true" box="[160,216,696,722]" italics="true" pageId="12" pageNumber="187">Note</emphasis>
</paragraph>
<paragraph id="38791929FFF1737924F5FD793B04E707" blockId="12.[160,1156,696,792]" pageId="12" pageNumber="187">
The populations previously known as 
<taxonomicName id="FFC662AAFFF173792606FD793AD9E6EA" authorityName="Westwood" authorityYear="1842" box="[595,741,732,757]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="miersianus">
<emphasis id="0AB2C53BFFF173792606FD793AD9E6EA" box="[595,741,732,757]" italics="true" pageId="12" pageNumber="187">C. miersianus</emphasis>
</taxonomicName>
(sensu 
<bibRefCitation id="5C5764D8FFF17379276AFD793C2DE6EA" author="Scholtz CH &amp; Hotman D &amp; Nel A" box="[831,1041,732,757]" pageId="12" pageNumber="187" pagination="193 - 199" refId="ref9262" refString="Scholtz CH, D ̍ Hotman D, Nel A. 1987. Systematic position of the South American genus Cryptogenius Westwood (Coleoptera: Scarabaeoidea: Trogidae and Hybosoridae). Coleopt Bull. 41 (2): 193 - 199." type="journal article" year="1987">Scholtz et al. 1987</bibRefCitation>
; 
<bibRefCitation id="5C5764D8FFF17379204AFD7938E5E707" author="Ide S &amp; Costa C &amp; Vanin SA" pageId="12" pageNumber="187" pagination="757 - 765" refId="ref8758" refString="Ide S, Costa C, Vanin SA. 1990. Redescription of Cryptogenius fryi Arrow, 1909 with notes on sexual dimorphism and biological data (Coleoptera, Hybosoridae). Rev bras ent. 34 (4): 757 - 765." type="journal article" year="1990">Ide et al. 1990</bibRefCitation>
; Ocampo 2006) are now named 
<taxonomicName id="FFC662AAFFF17379261CFD5A3AE5E707" authority="Costa-Silva &amp; Vaz-de-Mello, 2023" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[585,729,767,792]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
<emphasis id="0AB2C53BFFF17379261CFD5A3AE5E707" box="[585,729,767,792]" italics="true" pageId="12" pageNumber="187">C. westwoodi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11817840FFF1737926B5FD5A3B04E707" box="[736,824,767,792]" pageId="12" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="70DC4AA2FFF1737924F5FCE63C1DE197" pageId="12" pageNumber="187" type="diagnosis">
<paragraph id="38791929FFF1737924F5FCE63928E742" blockId="12.[160,1156,835,1242]" box="[160,276,835,861]" pageId="12" pageNumber="187">
<emphasis id="0AB2C53BFFF1737924F5FCE63928E742" bold="true" box="[160,276,835,861]" italics="true" pageId="12" pageNumber="187">Diagnosis</emphasis>
</paragraph>
<paragraph id="38791929FFF1737924F5FCC23B05E0C5" blockId="12.[160,1156,835,1242]" pageId="12" pageNumber="187">
Size of 
<typeStatus id="E77DA78BFFF1737924A5FCC2396BE79F" box="[240,343,871,896]" pageId="12" pageNumber="187" type="holotype">holotype</typeStatus>
: Length: 
<quantity id="FF3EB4CCFFF1737925E8FCC23A2FE79F" box="[445,531,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="12" pageNumber="187" unit="mm" value="6.4">6.4 mm</quantity>
; width: 
<quantity id="FF3EB4CCFFF17379263FFCC23A92E79F" box="[618,686,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="187" unit="mm" value="3.0">3 mm</quantity>
. (
<figureCitation id="A0FD05ACFFF1737926EAFCC23B7CE79F" box="[703,832,871,896]" captionStart="Figure 1" captionStartId="1.[160,225,1240,1263]" captionTargetBox="[163,1152,143,1206]" captionTargetId="figure-139@1.[162,1154,142,1208]" captionTargetPageId="1" captionText="Figure 1. Holotype of Cryptogenius westwoodi sp. nov. (Ò CEMT) in (a) dorsal (white arrow showing the diagonal carinae in the pronotum), (b) ventral and (c) lateral views (white arrow indicates the third carina extending from the humerus to declivous area). Habitus of male genitalia in (d) dorsal and (e) ventral view. (f) labels. Scale bars: a–c = 1 mm; d, e = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737075" httpUri="https://zenodo.org/record/7737075/files/figure.png" pageId="12" pageNumber="187">Figure 1a–c</figureCitation>
). 
<taxonomicName id="FFC662AAFFF173792700FCC23C61E79F" authority="Costa-Silva &amp; Vaz-de-Mello, 2023" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[853,1117,871,896]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
<emphasis id="0AB2C53BFFF173792700FCC23C61E79F" box="[853,1117,871,896]" italics="true" pageId="12" pageNumber="187">Cryptogenius westwoodi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11817840FFF173792031FCC238EDE7BD" pageId="12" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
can be recognised as follows: clypeus deflexed, forming right angles with frons; pronotal disc flat and with surface densely areolate; lateral margins crenulate; diagonal carinae extended from the centre of disc into an acute projection near the posterior angles (
<figureCitation id="A0FD05ACFFF1737924ADFC54395DE015" box="[248,353,1009,1034]" captionStart="Figure 1" captionStartId="1.[160,225,1240,1263]" captionTargetBox="[163,1152,143,1206]" captionTargetId="figure-139@1.[162,1154,142,1208]" captionTargetPageId="1" captionText="Figure 1. Holotype of Cryptogenius westwoodi sp. nov. (Ò CEMT) in (a) dorsal (white arrow showing the diagonal carinae in the pronotum), (b) ventral and (c) lateral views (white arrow indicates the third carina extending from the humerus to declivous area). Habitus of male genitalia in (d) dorsal and (e) ventral view. (f) labels. Scale bars: a–c = 1 mm; d, e = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737075" httpUri="https://zenodo.org/record/7737075/files/figure.png" pageId="12" pageNumber="187">Figure 1a</figureCitation>
, white arrows). Elytra with surface flat and densely areolate; with three acute carina, the third extending from the humerus to declivous area (
<figureCitation id="A0FD05ACFFF173792799FBB13C0AE032" box="[972,1078,1044,1069]" captionStart="Figure 1" captionStartId="1.[160,225,1240,1263]" captionTargetBox="[163,1152,143,1206]" captionTargetId="figure-139@1.[162,1154,142,1208]" captionTargetPageId="1" captionText="Figure 1. Holotype of Cryptogenius westwoodi sp. nov. (Ò CEMT) in (a) dorsal (white arrow showing the diagonal carinae in the pronotum), (b) ventral and (c) lateral views (white arrow indicates the third carina extending from the humerus to declivous area). Habitus of male genitalia in (d) dorsal and (e) ventral view. (f) labels. Scale bars: a–c = 1 mm; d, e = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737075" httpUri="https://zenodo.org/record/7737075/files/figure.png" pageId="12" pageNumber="187">Figure 1c</figureCitation>
, white arrow). Outer margins of meso- and metatibiae serrate, with nine or more acute teeth pointed laterally (
<figureCitation id="A0FD05ACFFF173792524FBFC39C8E06D" box="[369,500,1113,1138]" captionStart="Figure 1" captionStartId="1.[160,225,1240,1263]" captionTargetBox="[163,1152,143,1206]" captionTargetId="figure-139@1.[162,1154,142,1208]" captionTargetPageId="1" captionText="Figure 1. Holotype of Cryptogenius westwoodi sp. nov. (Ò CEMT) in (a) dorsal (white arrow showing the diagonal carinae in the pronotum), (b) ventral and (c) lateral views (white arrow indicates the third carina extending from the humerus to declivous area). Habitus of male genitalia in (d) dorsal and (e) ventral view. (f) labels. Scale bars: a–c = 1 mm; d, e = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7737075" httpUri="https://zenodo.org/record/7737075/files/figure.png" pageId="12" pageNumber="187">Figure 1a,b</figureCitation>
). Male genitalia distinctive: parameres symmetrical and partially sclerotised; lateral margin not constricted near the apex; apex rounded and covered by sparce micropunctation. Further morphological details can be found in 
<bibRefCitation id="5C5764D8FFF1737924F5FB64395AE0C5" author="Westwood JO" box="[160,358,1217,1242]" pageId="12" pageNumber="187" pagination="161 - 180" refId="ref9682" refString="Westwood JO. 1846. On the Lamellicorn Beetles which possess exerted Mandibles and Labrum, and 10 - jointed Antennae (Part III). Trans Ent Soc Lond. IV: 161 - 180." type="journal article" year="1846">Westwood (1846)</bibRefCitation>
, 
<bibRefCitation id="5C5764D8FFF173792527FB643A72E0C5" author="Scholtz CH &amp; Hotman D &amp; Nel A" box="[370,590,1217,1242]" pageId="12" pageNumber="187" pagination="193 - 199" refId="ref9262" refString="Scholtz CH, D ̍ Hotman D, Nel A. 1987. Systematic position of the South American genus Cryptogenius Westwood (Coleoptera: Scarabaeoidea: Trogidae and Hybosoridae). Coleopt Bull. 41 (2): 193 - 199." type="journal article" year="1987">Scholtz et al. (1987)</bibRefCitation>
and Ocampo (2006).
</paragraph>
<paragraph id="38791929FFF1737924F5FAA339B8E13F" blockId="12.[160,1155,1286,1416]" box="[160,388,1286,1312]" pageId="12" pageNumber="187">
<emphasis id="0AB2C53BFFF1737924F5FAA339B8E13F" bold="true" box="[160,388,1286,1312]" italics="true" pageId="12" pageNumber="187">Sexual dimorphism</emphasis>
</paragraph>
<paragraph id="38791929FFF1737924F5FA8F3C1DE197" blockId="12.[160,1155,1286,1416]" pageId="12" pageNumber="187">
Males of 
<taxonomicName id="FFC662AAFFF173792556FA8F3A36E15C" authority="Costa-Silva &amp; Vaz-de-Mello, 2023" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[259,522,1322,1347]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
<emphasis id="0AB2C53BFFF173792556FA8F3A36E15C" box="[259,522,1322,1347]" italics="true" pageId="12" pageNumber="187">Cryptogenius westwoodi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11817840FFF173792645FA8F3A5AE15C" box="[528,614,1322,1347]" pageId="12" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
have the fourth ventrite with the same length as (or slightly shorter than) the third (
<figureCitation id="A0FD05ACFFF173792660FAE93A9CE17A" box="[565,672,1356,1381]" captionStart="Figure 6" captionStartId="13.[160,225,1243,1266]" captionTargetBox="[356,958,150,1224]" captionTargetId="figure-154@13.[355,960,142,1226]" captionTargetPageId="13" captionText="Figure 6. Extracted abdomen of two paratypes of Cryptogenius westwoodi sp. nov. showing the sexual dimorphism between (a) male, with V3 and V4 of equal length; and (b) female, with V4 longer than V3. Abbreviations: V1–4, first to fourth ventrites. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7737089" httpUri="https://zenodo.org/record/7737089/files/figure.png" pageId="12" pageNumber="187">Figure 6a</figureCitation>
), while the female has the fourth ventrite longer than the third (
<figureCitation id="A0FD05ACFFF1737925C9FACA3A3BE197" box="[412,519,1391,1416]" captionStart="Figure 6" captionStartId="13.[160,225,1243,1266]" captionTargetBox="[356,958,150,1224]" captionTargetId="figure-154@13.[355,960,142,1226]" captionTargetPageId="13" captionText="Figure 6. Extracted abdomen of two paratypes of Cryptogenius westwoodi sp. nov. showing the sexual dimorphism between (a) male, with V3 and V4 of equal length; and (b) female, with V4 longer than V3. Abbreviations: V1–4, first to fourth ventrites. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7737089" httpUri="https://zenodo.org/record/7737089/files/figure.png" pageId="12" pageNumber="187">Figure 6b</figureCitation>
). For additional characters, see 
<bibRefCitation id="5C5764D8FFF173792739FACA3C20E197" author="Ide S &amp; Costa C &amp; Vanin SA" box="[876,1052,1391,1416]" pageId="12" pageNumber="187" pagination="757 - 765" refId="ref8758" refString="Ide S, Costa C, Vanin SA. 1990. Redescription of Cryptogenius fryi Arrow, 1909 with notes on sexual dimorphism and biological data (Coleoptera, Hybosoridae). Rev bras ent. 34 (4): 757 - 765." type="journal article" year="1990">Ide et al. (1990)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="70DC4AA2FFF1737B24F5FA113B53E4F2" lastPageId="14" lastPageNumber="189" pageId="12" pageNumber="187" type="distribution">
<paragraph id="38791929FFF1737924F5FA113910E1D1" blockId="12.[160,1156,1460,1728]" box="[160,300,1460,1486]" pageId="12" pageNumber="187">
<emphasis id="0AB2C53BFFF1737924F5FA113910E1D1" bold="true" box="[160,300,1460,1486]" italics="true" pageId="12" pageNumber="187">Distribution</emphasis>
</paragraph>
<paragraph id="38791929FFF1737924F5FA7239D2E2DF" blockId="12.[160,1156,1460,1728]" pageId="12" pageNumber="187">
Certainly present throughout the Amazon, with recorded presence in 
<collectingCountry id="40D159B9FFF1737927B1FA723C18E1EF" box="[996,1060,1495,1520]" name="Brazil" pageId="12" pageNumber="187">Brazil</collectingCountry>
, 
<collectingCountry id="40D159B9FFF173792066FA723CBCE1EF" box="[1075,1152,1495,1520]" name="Bolivia" pageId="12" pageNumber="187">Bolivia</collectingCountry>
, 
<collectingCountry id="40D159B9FFF1737924F5FA5F38C2E20C" box="[160,254,1530,1555]" name="Ecuador" pageId="12" pageNumber="187">Ecuador</collectingCountry>
, 
<collectingCountry id="40D159B9FFF17379255EFA5F398DE20C" box="[267,433,1530,1555]" name="French Guiana" pageId="12" pageNumber="187">French Guiana</collectingCountry>
and 
<collectingCountry id="40D159B9FFF1737925A5FA5F3A5FE20C" box="[496,611,1530,1555]" name="Venezuela" pageId="12" pageNumber="187">Venezuela</collectingCountry>
(
<figureCitation id="A0FD05ACFFF173792620FA5F3AE8E20C" box="[629,724,1530,1555]" captionStart="Figure 4" captionStartId="10.[160,225,1461,1484]" captionTargetBox="[163,1152,143,1428]" captionTargetId="figure-3@10.[162,1154,142,1429]" captionTargetPageId="10" captionText="Figure 4. Distribution map of known records of Cryptogenius species: C. miersianus (red circles) and C. westwoodi sp. nov. (blue circles). Red and blue stars represent records from the literature that possibly refer, according to the known geographical distribution, to C. miersianus and C. westwoodi sp. nov., respectively, found in Ide et al. (1990), Ratcliffe (2002) and Ocampo (2006). These records illustrated with stars represent specimens not personally examined by the authors." figureDoi="http://doi.org/10.5281/zenodo.7737083" httpUri="https://zenodo.org/record/7737083/files/figure.png" pageId="12" pageNumber="187">Figure 4</figureCitation>
). 
<bibRefCitation id="5C5764D8FFF1737926BEFA5F3BA9E20C" author="Ratcliffe BC" box="[747,917,1530,1555]" pageId="12" pageNumber="187" pagination="1 - 48" refId="ref9108" refString="Ratcliffe BC. 2002. A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa. 32 (1): 1 - 48. doi: 10.11646 / zootaxa. 32.1.1." type="journal article" year="2002">Ratcliffe (2002)</bibRefCitation>
and Ocampo (2006) also mentioned records under the name of 
<taxonomicName id="FFC662AAFFF1737926DAF9B83B23E229" authorityName="Westwood" authorityYear="1842" box="[655,799,1565,1590]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="miersianus">
<emphasis id="0AB2C53BFFF1737926DAF9B83B23E229" box="[655,799,1565,1590]" italics="true" pageId="12" pageNumber="187">C. miersianus</emphasis>
</taxonomicName>
(now 
<taxonomicName id="FFC662AAFFF17379273DF9B83BC4E229" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[872,1016,1565,1590]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="westwoodi">
<emphasis id="0AB2C53BFFF17379273DF9B83BC4E229" box="[872,1016,1565,1590]" italics="true" pageId="12" pageNumber="187">C. westwoodi</emphasis>
</taxonomicName>
) from 
<collectingCountry id="40D159B9FFF173792010F9B838F1E247" name="Costa Rica" pageId="12" pageNumber="187">Costa Rica</collectingCountry>
and 
<collectingCountry id="40D159B9FFF173792552F99A3958E247" box="[263,356,1599,1624]" name="Panama" pageId="12" pageNumber="187">Panama</collectingCountry>
, without further information. Also, Ocampo (2006) mentioned a record from 
<collectingCountry id="40D159B9FFF17379248EF9C73975E264" box="[219,329,1634,1659]" name="Colombia" pageId="12" pageNumber="187">Colombia</collectingCountry>
, but no information about the specific locality or point on the map is given for this country. During this study, we were unable to find specimens from these countries and thus confirm the records.
</paragraph>
<caption id="6CB949A1FFF0737824F5FB7E3B33E132" ID-DOI="http://doi.org/10.5281/zenodo.7737089" ID-Zenodo-Dep="7737089" httpUri="https://zenodo.org/record/7737089/files/figure.png" pageId="13" pageNumber="188" startId="13.[160,225,1243,1266]" targetBox="[356,958,150,1224]" targetPageId="13">
<paragraph id="38791929FFF0737824F5FB7E3B33E132" blockId="13.[160,1157,1243,1325]" pageId="13" pageNumber="188">
<emphasis id="0AB2C53BFFF0737824F5FB7E38C5E0ED" bold="true" box="[160,249,1243,1266]" pageId="13" pageNumber="188">Figure 6.</emphasis>
Extracted abdomen of two paratypes of 
<taxonomicName id="FFC662AAFFF0737826DDFB7E3B4CE0ED" authority="Costa-Silva &amp; Vaz-de-Mello, 2023" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[648,880,1243,1266]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="188" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
<emphasis id="0AB2C53BFFF0737826DDFB7E3B4CE0ED" box="[648,880,1243,1266]" italics="true" pageId="13" pageNumber="188">Cryptogenius westwoodi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11817840FFF073782723FB7E3BFEE0ED" box="[886,962,1243,1266]" pageId="13" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
showing the sexual dimorphism between (a) male, with V3 and V4 of equal length; and (b) female, with V4 longer than V3. Abbreviations: V1–4, first to fourth ventrites. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="38791929FFF073782495FACC3BE7E1D8" blockId="13.[160,1157,1385,1722]" pageId="13" pageNumber="188">
Additionally, 
<bibRefCitation id="5C5764D8FFF073782507FACC39C7E19D" author="Ide S &amp; Costa C &amp; Vanin SA" box="[338,507,1385,1410]" pageId="13" pageNumber="188" pagination="757 - 765" refId="ref8758" refString="Ide S, Costa C, Vanin SA. 1990. Redescription of Cryptogenius fryi Arrow, 1909 with notes on sexual dimorphism and biological data (Coleoptera, Hybosoridae). Rev bras ent. 34 (4): 757 - 765." type="journal article" year="1990">Ide et al. (1990)</bibRefCitation>
mentioned a record of 
<taxonomicName id="FFC662AAFFF073782755FACC3B89E19D" authorityName="Arrow" authorityYear="1909" box="[768,949,1385,1410]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="188" phylum="Arthropoda" rank="species" species="fryi">
<emphasis id="0AB2C53BFFF073782755FACC3B89E19D" box="[768,949,1385,1410]" italics="true" pageId="13" pageNumber="188">Cryptogenius fryi</emphasis>
</taxonomicName>
from 
<collectingCountry id="40D159B9FFF0737827A3FACC3C15E19D" box="[1014,1065,1385,1410]" name="Peru" pageId="13" pageNumber="188">Peru</collectingCountry>
that we believe to be a 
<taxonomicName id="FFC662AAFFF07378251AFA2E39E3E1BB" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[335,479,1419,1444]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="188" phylum="Arthropoda" rank="species" species="westwoodi">
<emphasis id="0AB2C53BFFF07378251AFA2E39E3E1BB" box="[335,479,1419,1444]" italics="true" pageId="13" pageNumber="188">C. westwoodi</emphasis>
</taxonomicName>
(
<figureCitation id="A0FD05ACFFF0737825BBFA2E3A77E1BB" box="[494,587,1419,1444]" captionStart="Figure 4" captionStartId="10.[160,225,1461,1484]" captionTargetBox="[163,1152,143,1428]" captionTargetId="figure-3@10.[162,1154,142,1429]" captionTargetPageId="10" captionText="Figure 4. Distribution map of known records of Cryptogenius species: C. miersianus (red circles) and C. westwoodi sp. nov. (blue circles). Red and blue stars represent records from the literature that possibly refer, according to the known geographical distribution, to C. miersianus and C. westwoodi sp. nov., respectively, found in Ide et al. (1990), Ratcliffe (2002) and Ocampo (2006). These records illustrated with stars represent specimens not personally examined by the authors." figureDoi="http://doi.org/10.5281/zenodo.7737083" httpUri="https://zenodo.org/record/7737083/files/figure.png" pageId="13" pageNumber="188">Figure 4</figureCitation>
). Since we did not examine this specimen personally, we consider 
<collectingCountry id="40D159B9FFF073782537FA0B39A9E1D8" box="[354,405,1454,1479]" name="Peru" pageId="13" pageNumber="188">Peru</collectingCountry>
a potential record for this newly described species.
</paragraph>
<paragraph id="38791929FFF0737B2495FA743B53E4F2" blockId="13.[160,1157,1385,1722]" lastBlockId="14.[160,1155,142,237]" lastPageId="14" lastPageNumber="189" pageId="13" pageNumber="188">
The specimen found in Serra do Baturité (
<collectingRegion id="FA02D7CBFFF073782694FA743B3BE1F5" box="[705,775,1489,1514]" country="Brazil" name="Ceara" pageId="13" pageNumber="188">Ceará</collectingRegion>
, 
<collectingCountry id="40D159B9FFF073782743FA743B6AE1F5" box="[790,854,1489,1514]" name="Brazil" pageId="13" pageNumber="188">Brazil</collectingCountry>
) is the only record of 
<taxonomicName id="FFC662AAFFF07378203AFA74392BE213" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="188" phylum="Arthropoda" rank="species" species="westwoodi">
<emphasis id="0AB2C53BFFF07378203AFA74392BE213" italics="true" pageId="13" pageNumber="188">C. westwoodi</emphasis>
</taxonomicName>
outside of the Amazon. The region where this specimen was collected, named ‘Brejo de Altitude̍, is a refuge for forest species that are typically found in areas with higher humidity (i.e. Atlantic Forest and Amazon) (
<bibRefCitation id="5C5764D8FFF073782744F99C3BF7E24D" author="Porto KC &amp; Cabral JJP &amp; Tabarelli M" box="[785,971,1593,1618]" pageId="13" pageNumber="188" refId="ref9022" refString="Porto KC, Cabral JJP, Tabarelli M. 2004. Brejos de altitude em Pernambuco e Paraiba: historia natural, ecologia e conservacao. Brasilia: Ministerio do Meio Ambiente (Serie Biodiversidade); p. 324." type="journal volume" year="2004">Porto et al. 2004</bibRefCitation>
). This find, even unusual for the Amazonian fauna, is not a particularity of 
<taxonomicName id="FFC662AAFFF073782769F9FE3C17E26B" box="[828,1067,1627,1652]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="188" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="0AB2C53BFFF073782769F9FE3BF6E26B" box="[828,970,1627,1652]" italics="true" pageId="13" pageNumber="188">Cryptogenius</emphasis>
species.
</taxonomicName>
Several groups (e.g. dung beetles) were also reported for these isolated forests (
<bibRefCitation id="5C5764D8FFF0737827EEF9DB3C42E288" author="Araujo JF &amp; Silva FAB &amp; Moura RC" box="[955,1150,1662,1687]" pageId="13" pageNumber="188" pagination="1289 - 1303" refId="ref7666" refString="Araujo JF, Silva FAB, Moura RC. 2020. New records of relictual populations of dung beetle species (Coleoptera, Scarabaeidae) in the atlantic forest of the Brazilian Northeast. Check List. 16 (5): 1289 - 1303. doi: 10.15560 / 16.5.1289." type="journal article" year="2020">Araújo et al. 2020</bibRefCitation>
; 
<bibRefCitation id="5C5764D8FFF0737824F5F904394CE2A5" author="Cupello M &amp; Ribeiro-Costa CS &amp; Vaz-de-Mello FZ" box="[160,368,1697,1722]" pageId="13" pageNumber="188" pagination="1 - 85" refId="ref8171" refString="Cupello M, Ribeiro-Costa CS, Vaz-de-Mello FZ. 2020. Systematics of the enigmatic South American Streblopus Van Lansberge, 1874 dung beetles and their transatlantic origin: a case study on the role of dispersal events in the biogeographical history of the Scarabaeinae (Coleoptera: Scarabaeidae). Eur J Taxon. 603: 1 - 85." type="journal article" year="2020">Cupello et al. 2020</bibRefCitation>
) as an effect of a temporary connection between the Amazon and the Atlantic Forest in the last glacial cycle (
<bibRefCitation id="5C5764D8FFF3737B2618FF2B3B7DE4B8" author="De Oliveira PE &amp; Barreto AMF &amp; Suguiu K" box="[589,833,142,167]" pageId="14" pageNumber="189" pagination="319 - 337" refId="ref8232" refString="De Oliveira PE, Barreto AMF, Suguiu K. 1999. Late Pleistocene / Holocene climatic and vegetational history of the Brazilian caatinga: the fossil dunes of the middle Sa t o Francisco River. Palaeogeogr Palaeoclimatol Palaeoecol. 152: 319 - 337. doi: 10.1016 / S 0031 - 0182 (99) 00061 - 9." type="journal article" year="1999">De Oliveira et al. 1999</bibRefCitation>
). With the end of interglacial periods and the consequent retraction of these forests, some relict forests and populations of fauna and flora were isolated (
<bibRefCitation id="5C5764D8FFF3737B2607FF713B5EE4F2" author="Sobral-Souza T &amp; Lima-Ribeiro M &amp; Solferini VN" box="[594,866,212,237]" pageId="14" pageNumber="189" pagination="643 - 655" refId="ref9484" refString="Sobral-Souza T, Lima-Ribeiro M, Solferini VN. 2015. Biogeography of Neotropical Rainforests: past connections between Amazon and Atlantic Forest detected by ecological niche modeling. Evol Ecol. 29 (5): 643 - 655. doi: 10.1007 / s 10682 - 015 - 9780 - 9." type="journal article" year="2015">Sobral-Souza et al. 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="70DC4AA2FFF3737B24F5FEBE3A87E582" pageId="14" pageNumber="189" type="etymology">
<paragraph id="38791929FFF3737B24F5FEBE3921E52A" blockId="14.[160,1156,283,413]" box="[160,285,283,309]" pageId="14" pageNumber="189">
<emphasis id="0AB2C53BFFF3737B24F5FEBE3921E52A" bold="true" box="[160,285,283,309]" italics="true" pageId="14" pageNumber="189">Etymology</emphasis>
</paragraph>
<paragraph id="38791929FFF3737B24F5FE9A3A87E582" blockId="14.[160,1156,283,413]" pageId="14" pageNumber="189">
The specific name 
<taxonomicName id="FFC662AAFFF3737B2526FE9A39DBE547" authorityName="Costa-Silva &amp; Vaz-de-Mello" authorityYear="2023" box="[371,487,319,344]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="species" species="westwoodi">
<emphasis id="0AB2C53BFFF3737B2526FE9A39DBE547" box="[371,487,319,344]" italics="true" pageId="14" pageNumber="189">westwoodi</emphasis>
</taxonomicName>
, a masculine noun in the genitive case, is an eponym after the English entomologist John Obadiah Westwood (1805–1893), who described the genus 
<taxonomicName id="FFC662AAFFF3737B24B9FE213946E582" authorityName="Westwood" authorityYear="1842" box="[236,378,388,413]" class="Insecta" family="Hybosoridae" genus="Cryptogenius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="189" phylum="Arthropoda" rank="genus">
<emphasis id="0AB2C53BFFF3737B24B9FE213946E582" box="[236,378,388,413]" italics="true" pageId="14" pageNumber="189">Cryptogenius</emphasis>
</taxonomicName>
and countless other insects.
</paragraph>
</subSubSection>
</treatment>
</document>