1037 lines
108 KiB
XML
1037 lines
108 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1024.63678" ID-GBIF-Dataset="c542e28a-4470-4cd2-a945-a24f723f5253" ID-PMC="PMC7987703" ID-Pensoft-Pub="1313-2970-1024-137" ID-Pensoft-UUID="758C273A0D615743BEFABA016C9FC9F9" ID-PubMed="33786005" ID-ZooBank="EF866ABA11B14C18B2696B8DD97EC83B" ModsDocID="1313-2970-1024-137" checkinTime="1615979475054" checkinUser="pensoft" docAuthor="Wesener, Thomas, Wongthamwanich, Nattarin & Moritz, Leif" docDate="2021" docId="2AB443F1DFF45118AE5260106B9D2694" docLanguage="en" docName="ZooKeys 1024: 137-156" docOrigin="ZooKeys 1024" docPubDate="2021-03-16" docSource="http://dx.doi.org/10.3897/zookeys.1024.63678" docTitle="Glomeridesmus siamensis sp. nov." docType="treatment" docVersion="5" id="758C273A0D615743BEFABA016C9FC9F9" lastPageNumber="137" masterDocId="758C273A0D615743BEFABA016C9FC9F9" masterDocTitle="Description of the first species of Glomeridesmida from Thailand (Diplopoda, Glomeridesmida, Glomeridesmidae)" masterLastPageNumber="156" masterPageNumber="137" pageNumber="137" updateTime="1668149901420" updateUser="ExternalLinkService">
|
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
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<mods:titleInfo>
|
||
<mods:title>Description of the first species of Glomeridesmida from Thailand (Diplopoda, Glomeridesmida, Glomeridesmidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Wesener, Thomas</mods:namePart>
|
||
<mods:affiliation>Zoological Research Museum Alexander Koenig, Leibniz Institute for Animal Biodiversity, Adenauerallee 160, 53113 Bonn, Germany</mods:affiliation>
|
||
<mods:nameIdentifier type="email">t.wesener@leibniz-zfmk.de</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Wongthamwanich, Nattarin</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8165-7391</mods:nameIdentifier>
|
||
<mods:affiliation>Department of Biology, Faculty of Science, Srinakharinwirot University, Bangkok 10110, Thailand</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Moritz, Leif</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6028-5189</mods:nameIdentifier>
|
||
<mods:affiliation>Zoological Research Museum Alexander Koenig, Leibniz Institute for Animal Biodiversity, Adenauerallee 160, 53113 Bonn, Germany & University of Bonn, Institute for Evolutionary Biology and Ecology, An der Immenburg 1, 53121 Bonn, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2021</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2021-03-16</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>1024</mods:number>
|
||
</mods:detail>
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||
<mods:extent unit="page">
|
||
<mods:start>137</mods:start>
|
||
<mods:end>156</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
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||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.1024.63678</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1024.63678</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-1024-137</mods:identifier>
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<mods:identifier type="ZooBank">EF866ABA11B14C18B2696B8DD97EC83B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">758C273A0D615743BEFABA016C9FC9F9</mods:identifier>
|
||
</mods:mods>
|
||
<subSection pageId="0" pageNumber="137" type="results">
|
||
<treatment ID-GBIF-Taxon="180585834" LSID="urn:lsid:zoobank.org:act:9FDB569D-C0AC-4133-B271-62554FD2D757" httpUri="http://treatment.plazi.org/id/2AB443F1DFF45118AE5260106B9D2694" lastPageNumber="137" pageId="0" pageNumber="137">
|
||
<subSubSection pageId="0" pageNumber="137" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<taxonomicName LSID="http://zoobank.org/9FDB569D-C0AC-4133-B271-62554FD2D757" authority="sp. nov." authorityName="sp. nov." class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Glomeridesmus siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">Glomeridesmus siamensis sp. nov.</taxonomicName>
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">Figures 1</figureCitation>
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Glomeridesmus siamensis sp. nov., head, SEM A, B, D male holotype (ZCSWU Myr D 000011) A antennae B apical antennomeres C juvenile paratype (ZFMK MYR 10304), head, lateral view D antennal base and Toemoesvary organ. Abbreviations: I - VII = antennomeres, ad = apical disc, cl = clypeus, co = collum, epc = epicranium, fr = frons, gn = gnathochilarium, il = incisura lateralis, lab = labrum, mb = mandibular base, sc = sensory cones, to = Toemoesvary organ. Scale bars: 100 µm (A, C, D), 50 µm (B)." figureDoi="10.3897/zookeys.1024.63678.figure2" httpUri="https://binary.pensoft.net/fig/520750" pageId="0" pageNumber="137">, 2</figureCitation>
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">, 3</figureCitation>
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Glomeridesmus siamensis sp. nov., female paratype (ZFMK MYR 10302), legs and ovipositor, SEM A leg 23, posterior view B leg 23, ventral view C leg 2 with ovipositor, ventral view D ovipositors, posterior view. Abbreviations: ap = apical part of ovipositor, bp = basal part of ovipositor, cp = coxal pouch, cx = coxa, fe = femur, L 2 = leg 2, Lt = pleurite, pof = postfemur, prf = prefemur, sh = sheet, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 50 µm (A, B), 100 µm (C, D)." figureDoi="10.3897/zookeys.1024.63678.figure4" httpUri="https://binary.pensoft.net/fig/520752" pageId="0" pageNumber="137">, 4</figureCitation>
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), male sexual characters A telopod, anterior view B telopod, posterior view C gonopore on leg 2, posterior view D inner horns, anterior view. Abbreviations: I - IV = telopoditomeres, ff = immobile finger, gp = gonopore, ih = inner horns, ma = membranous area, pr = projection. Scale bars: 50 µm (A, B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure5" httpUri="https://binary.pensoft.net/fig/520753" pageId="0" pageNumber="137">, 5</figureCitation>
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011) mouthparts A gnathochilarium, ventral view B gnathochilarium, palps and central pad, frontal view C endochilarium, dorsal view D mandibular gnathal lobe, mesal view. Abbreviations: cp = central pad, ec = endochilarium, et = external tooth, gu = gula, ip = inner palp, ia = intermediate area, it = inner tooth, LL = lamella lingualis, Lp = lateral palp, me = mentum, mp = molar plate, pL = pectinate lamellae, stp = stipes. Scale bars: 100 µm (A), 50 µm (B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure6" httpUri="https://binary.pensoft.net/fig/520754" pageId="0" pageNumber="137">, 6</figureCitation>
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Glomeridesmus siamensis sp. nov., body-rings A, B, D juvenile paratype (ZFMK MYR 10304) A habitus, lateral view, arrow indicates drawn out lateral tip of tergite B detail of tergite margin, latero-ventral view, arrow indicates striae on tergite C male holotype (ZCSWU Myr D 000011), endotergum, arrow indicates posterior margin of tergite D posterior body-rings, lateral view, arrows indicate teeth on posterior margin of tergite. Abbreviations: co = collum, Lt = pleurite, te = tergite, tel = telson. Scale bars: 500 µm (A), 100 µm (B, C), 50 µm (D)." figureDoi="10.3897/zookeys.1024.63678.figure7" httpUri="https://binary.pensoft.net/fig/520755" pageId="0" pageNumber="137">, 7</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="137" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="137">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<materialsCitation ID-GBIF-Occurrence="3063614310" collectingDate="2017-07-27" collectionCode="ZCSWU" collectorName="Myr D" country="Thailand" county="Town" latitude="8.123278" location="Wesener" longLatPrecision="1" longitude="98.92191" municipality="Tiger Cave" specimenCode="THAI11" specimenCount="1" stateProvince="Krabi" typeStatus="holotype">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">
|
||
<typeStatus>Holotype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
</emphasis>
|
||
1 M,
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>ZCSWU</collectionCode>
|
||
<collectorName>Myr D</collectorName>
|
||
000011 (
|
||
<specimenCode collectionCode="THAI">THAI11</specimenCode>
|
||
)
|
||
</emphasis>
|
||
,
|
||
<collectingCountry name="Thailand">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion country="Thailand" name="Krabi">Krabi Province</collectingRegion>
|
||
, N. of
|
||
<collectingRegion country="Thailand" name="Krabi">Krabi</collectingRegion>
|
||
<collectingCounty>Town</collectingCounty>
|
||
, western aspect of
|
||
<collectingMunicipality>Tiger Cave</collectingMunicipality>
|
||
temple (Wat Tham Suea), overgrown rocks next to rubber plantation,
|
||
<geoCoordinate degrees="08" direction="north" minutes="07" orientation="latitude" precision="1" seconds="23.8" value="8.123278">08°07'23.8"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="098" direction="east" minutes="55" orientation="longitude" precision="1" seconds="18.9" value="98.92191">098°55'18.9"E</geoCoordinate>
|
||
. leg.
|
||
<date value="2017-07-27">
|
||
<collectingDate value="2017-07-27">27.VII.2017</collectingDate>
|
||
</date>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:2AB443F1DFF45118AE5260106B9D2694:F2FEEC8C740F24FB60051B95797602D9" country="Thailand" county="Town" latitude="8.123278" longLatPrecision="1" longitude="98.92191" municipality="Tiger Cave" name="Wesener" stateProvince="Krabi">Wesener</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:2AB443F1DFF45118AE5260106B9D2694:5D2DEE6E61A1EC1378766A8D86727432" country="Thailand" county="Town" latitude="8.123278" longLatPrecision="1" longitude="98.92191" municipality="Tiger Cave" name="Wongthamwanich" stateProvince="Krabi">Wongthamwanich</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:2AB443F1DFF45118AE5260106B9D2694:18FB25CBF9FC6295BD194374CE4AE914" country="Thailand" county="Town" latitude="8.123278" longLatPrecision="1" longitude="98.92191" municipality="Tiger Cave" name="Nawanetiwong" stateProvince="Krabi">Nawanetiwong</location>
|
||
, Moritz.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<materialsCitation ID-GBIF-Occurrence="3063614305" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="ZFMK, MYR" collectorName="Myr D" country="Thailand" county="Town" latitude="8.123278" location="Wesener" longLatPrecision="1" longitude="98.92191" municipality="Tiger Cave" specimenCount="1" specimenCount-immature="1" stateProvince="Krabi" typeStatus="paratype">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">
|
||
<typeStatus>Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
</emphasis>
|
||
<specimenCount type="immature">1 immature</specimenCount>
|
||
M,
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MYR10301">
|
||
<collectionCode>MYR</collectionCode>
|
||
10301
|
||
</accessionNumber>
|
||
</emphasis>
|
||
, same data as holotype; 1 F,
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MYR10302">
|
||
<collectionCode>MYR</collectionCode>
|
||
10302
|
||
</accessionNumber>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3063614311" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="ZCSWU" collectorName="Myr D" country="Thailand" specimenCount="5" specimenCount-juvenile="5" stateProvince="Krabi" typeStatus="paratype">
|
||
<specimenCount type="juvenile">5 juveniles</specimenCount>
|
||
,
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>ZCSWU</collectionCode>
|
||
<collectorName>Myr D</collectorName>
|
||
000012-16
|
||
</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3063614317" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="ZFMK" collectorName="Myr D" country="Thailand" county="Town" latitude="8.123278" location="Wesener" longLatPrecision="1" longitude="98.92191" municipality="Tiger Cave" specimenCode="MYR10303-10306" specimenCount="4" specimenCount-juvenile="4" stateProvince="Krabi" typeStatus="paratype">
|
||
<specimenCount type="juvenile">4 juveniles</specimenCount>
|
||
,
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||
<specimenCode collectionCode="MYR">MYR10303-10306</specimenCode>
|
||
</emphasis>
|
||
, same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="137" type="etymology">
|
||
<paragraph pageId="0" pageNumber="137">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Siamensis, noun in apposition, after the
|
||
<typeStatus>type</typeStatus>
|
||
locality in
|
||
<collectingCountry name="Thailand">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="137" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="137">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Small (
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5-7 mm</quantity>
|
||
) dark grey glomeridesmid with white legs (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">1A, B</figureCitation>
|
||
). Antennae short (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">1A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Glomeridesmus siamensis sp. nov., head, SEM A, B, D male holotype (ZCSWU Myr D 000011) A antennae B apical antennomeres C juvenile paratype (ZFMK MYR 10304), head, lateral view D antennal base and Toemoesvary organ. Abbreviations: I - VII = antennomeres, ad = apical disc, cl = clypeus, co = collum, epc = epicranium, fr = frons, gn = gnathochilarium, il = incisura lateralis, lab = labrum, mb = mandibular base, sc = sensory cones, to = Toemoesvary organ. Scale bars: 100 µm (A, C, D), 50 µm (B)." figureDoi="10.3897/zookeys.1024.63678.figure2" httpUri="https://binary.pensoft.net/fig/520750" pageId="0" pageNumber="137">2A-C</figureCitation>
|
||
). Coxal pouches starting at leg 9, all walking legs with apical spines and paronychium (Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3A-F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Glomeridesmus siamensis sp. nov., female paratype (ZFMK MYR 10302), legs and ovipositor, SEM A leg 23, posterior view B leg 23, ventral view C leg 2 with ovipositor, ventral view D ovipositors, posterior view. Abbreviations: ap = apical part of ovipositor, bp = basal part of ovipositor, cp = coxal pouch, cx = coxa, fe = femur, L 2 = leg 2, Lt = pleurite, pof = postfemur, prf = prefemur, sh = sheet, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 50 µm (A, B), 100 µm (C, D)." figureDoi="10.3897/zookeys.1024.63678.figure4" httpUri="https://binary.pensoft.net/fig/520752" pageId="0" pageNumber="137">4A-C</figureCitation>
|
||
). Last pleurite posterior margin well-rounded in female. Male sensory leg, podomere 1 with a conspicuous excavation on its mesal margin, not known from other
|
||
<taxonomicName authorityName="Gervais" authorityYear="1844" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Glomeridesmus</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3G, H</figureCitation>
|
||
). Male telopods unique: Inner horns slender, widely separated, a character only shared with
|
||
<taxonomicName class="Diplopoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="G. indus" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="indus">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">G. indus</emphasis>
|
||
</taxonomicName>
|
||
Carl, 1942. Telopod, podomere 2 rectangular, with a unique, strongly elongated and slender process protruding between movable (podomere 4) and immovable finger (process of podomere 3) (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), male sexual characters A telopod, anterior view B telopod, posterior view C gonopore on leg 2, posterior view D inner horns, anterior view. Abbreviations: I - IV = telopoditomeres, ff = immobile finger, gp = gonopore, ih = inner horns, ma = membranous area, pr = projection. Scale bars: 50 µm (A, B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure5" httpUri="https://binary.pensoft.net/fig/520753" pageId="0" pageNumber="137">5A, B</figureCitation>
|
||
). Podomere 3 with a slender immovable projection and podomere 4 forming a long movable finger. Immovable finger long and visibly laterally in
|
||
<taxonomicName class="Diplopoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="G. siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">G. siamensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">1C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), male sexual characters A telopod, anterior view B telopod, posterior view C gonopore on leg 2, posterior view D inner horns, anterior view. Abbreviations: I - IV = telopoditomeres, ff = immobile finger, gp = gonopore, ih = inner horns, ma = membranous area, pr = projection. Scale bars: 50 µm (A, B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure5" httpUri="https://binary.pensoft.net/fig/520753" pageId="0" pageNumber="137">5A, B</figureCitation>
|
||
), in contrast to South American species such as
|
||
<taxonomicName class="Diplopoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="G. spelaeus" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">G. spelaeus</emphasis>
|
||
</taxonomicName>
|
||
Iniesta & Wesener, 2012 and
|
||
<taxonomicName class="Diplopoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="G. arcostriatus" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="arcostriatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">G. arcostriatus</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Mauriès">Mauries</normalizedToken>
|
||
, 2020, where it is hidden in lateral view. Lamellae linguales of gnathochilarium fused with one another, but with a transverse suture towards mentum, lamellae with two central setae at margin and two lateral setae more posteriorly in
|
||
<taxonomicName class="Diplopoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="G. siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">G. siamensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011) mouthparts A gnathochilarium, ventral view B gnathochilarium, palps and central pad, frontal view C endochilarium, dorsal view D mandibular gnathal lobe, mesal view. Abbreviations: cp = central pad, ec = endochilarium, et = external tooth, gu = gula, ip = inner palp, ia = intermediate area, it = inner tooth, LL = lamella lingualis, Lp = lateral palp, me = mentum, mp = molar plate, pL = pectinate lamellae, stp = stipes. Scale bars: 100 µm (A), 50 µm (B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure6" httpUri="https://binary.pensoft.net/fig/520754" pageId="0" pageNumber="137">6A</figureCitation>
|
||
), while all four setae are marginal in
|
||
<taxonomicName class="Diplopoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="G. spelaeus" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="spelaeus">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">G. spelaeus</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3550.1.2" author="Moretti Iniesta, LF" journalOrPublisher="Zootaxa" pageId="0" pageNumber="137" pagination="26 - 42" refId="B21" refString="Moretti Iniesta, LF, Ferreira, RL, Wesener, T, 2012. The first troglobitic Glomeridesmus from Brazil, and a template for a modern taxonomic description of Glomeridesmida (Diplopoda). Zootaxa 3550: 26 - 42, DOI: https://doi.org/10.11646/zootaxa.3550.1.2" title="The first troglobitic Glomeridesmus from Brazil, and a template for a modern taxonomic description of Glomeridesmida (Diplopoda)." url="https://doi.org/10.11646/zootaxa.3550.1.2" volume="3550" year="2012">Iniesta et al. 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137" start="Figure 1" startId="F1">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Figure 1.</emphasis>
|
||
<taxonomicName class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Glomeridesmus siamensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., photographs
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A-C</emphasis>
|
||
male
|
||
<typeStatus>holotype</typeStatus>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>ZCSWU</collectionCode>
|
||
Myr D000011
|
||
</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
habitus, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
|
||
habitus, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C</emphasis>
|
||
telopods, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">D, E</emphasis>
|
||
female
|
||
<typeStatus>paratype</typeStatus>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MYR10302">
|
||
<collectionCode>MYR</collectionCode>
|
||
10302
|
||
</accessionNumber>
|
||
</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="137">D</emphasis>
|
||
head and anterior body-rings, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">E</emphasis>
|
||
head and anterior body-rings, ventral view. Not to scale.
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Abbreviations</emphasis>
|
||
: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to =
|
||
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
|
||
organ, tp = telopod, vu = vulva.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="137" type="description">
|
||
<paragraph pageId="0" pageNumber="137">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<materialsCitation ID-GBIF-Occurrence="3063614314" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="ZCSWU, ZFMK, MYR" collectorName="Based & Myr D" specimenCount="2" typeStatus="holotype">
|
||
<collectorName>Based</collectorName>
|
||
on male
|
||
<typeStatus>holotype</typeStatus>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>ZCSWU</collectionCode>
|
||
<collectorName>Myr D</collectorName>
|
||
000011
|
||
</emphasis>
|
||
) and female
|
||
<typeStatus>paratype</typeStatus>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MYR10302">
|
||
<collectionCode>MYR</collectionCode>
|
||
10302
|
||
</accessionNumber>
|
||
</emphasis>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Measurements</emphasis>
|
||
:
|
||
</emphasis>
|
||
Largest adult female (20+
|
||
<collectingCountry name="American Samoa">AS</collectingCountry>
|
||
tergites, 35+1 leg pairs): length: ~
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="6.0" unit="mm" value="6.0">6 mm</quantity>
|
||
; width (midbody): 1.0 mm. Male (19+
|
||
<collectingCountry name="American Samoa">AS</collectingCountry>
|
||
tergites, 33+1+
|
||
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||
leg pairs,
|
||
<typeStatus>holotype</typeStatus>
|
||
male): length: ~
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
|
||
; width (midbody):
|
||
<quantity metricMagnitude="-4" metricUnit="m" metricValue="9.0" unit="mm" value="0.9">0.9 mm</quantity>
|
||
; immature (?) male (19+
|
||
<collectingCountry name="American Samoa">AS</collectingCountry>
|
||
tergites, 32+1+
|
||
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
|
||
,): length: ~
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
|
||
, width:
|
||
<quantity metricMagnitude="-4" metricUnit="m" metricValue="8.0" unit="mm" value="0.8">0.8 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Colour in ethanol</emphasis>
|
||
:
|
||
</emphasis>
|
||
Tergites and dorsal side of head grayish-brown (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">1</figureCitation>
|
||
), legs, antennae and ventral side whitish (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">1A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Head</emphasis>
|
||
:
|
||
</emphasis>
|
||
General shape typical for the family (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">1D, E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Glomeridesmus siamensis sp. nov., head, SEM A, B, D male holotype (ZCSWU Myr D 000011) A antennae B apical antennomeres C juvenile paratype (ZFMK MYR 10304), head, lateral view D antennal base and Toemoesvary organ. Abbreviations: I - VII = antennomeres, ad = apical disc, cl = clypeus, co = collum, epc = epicranium, fr = frons, gn = gnathochilarium, il = incisura lateralis, lab = labrum, mb = mandibular base, sc = sensory cones, to = Toemoesvary organ. Scale bars: 100 µm (A, C, D), 50 µm (B)." figureDoi="10.3897/zookeys.1024.63678.figure2" httpUri="https://binary.pensoft.net/fig/520750" pageId="0" pageNumber="137">2C</figureCitation>
|
||
). Epicranium glabrous; frons, clypeus and labrum with several isolated setae (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Glomeridesmus siamensis sp. nov., head, SEM A, B, D male holotype (ZCSWU Myr D 000011) A antennae B apical antennomeres C juvenile paratype (ZFMK MYR 10304), head, lateral view D antennal base and Toemoesvary organ. Abbreviations: I - VII = antennomeres, ad = apical disc, cl = clypeus, co = collum, epc = epicranium, fr = frons, gn = gnathochilarium, il = incisura lateralis, lab = labrum, mb = mandibular base, sc = sensory cones, to = Toemoesvary organ. Scale bars: 100 µm (A, C, D), 50 µm (B)." figureDoi="10.3897/zookeys.1024.63678.figure2" httpUri="https://binary.pensoft.net/fig/520750" pageId="0" pageNumber="137">2C</figureCitation>
|
||
). Antennae widely separated by a distance slightly longer than twice width of first antennomere (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">1A, E</figureCitation>
|
||
). Antennae and organ of
|
||
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
|
||
both surrounded by individual cuticular rims, both on an elevated plateau (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Glomeridesmus siamensis sp. nov., head, SEM A, B, D male holotype (ZCSWU Myr D 000011) A antennae B apical antennomeres C juvenile paratype (ZFMK MYR 10304), head, lateral view D antennal base and Toemoesvary organ. Abbreviations: I - VII = antennomeres, ad = apical disc, cl = clypeus, co = collum, epc = epicranium, fr = frons, gn = gnathochilarium, il = incisura lateralis, lab = labrum, mb = mandibular base, sc = sensory cones, to = Toemoesvary organ. Scale bars: 100 µm (A, C, D), 50 µm (B)." figureDoi="10.3897/zookeys.1024.63678.figure2" httpUri="https://binary.pensoft.net/fig/520750" pageId="0" pageNumber="137">2D</figureCitation>
|
||
). Genae (area below the antennae) almost non-existent, with incisura lateralis. Undivided basal joint of mandible large and visible in dorsal view.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1024.63678.figure2" httpUri="https://binary.pensoft.net/fig/520750" pageId="0" pageNumber="137" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Figure 2.</emphasis>
|
||
<taxonomicName class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Glomeridesmus siamensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., head, SEM
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A, B, D</emphasis>
|
||
male
|
||
<typeStatus>holotype</typeStatus>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>ZCSWU</collectionCode>
|
||
Myr D000011
|
||
</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
antennae
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
|
||
apical antennomeres
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C</emphasis>
|
||
juvenile
|
||
<typeStatus>paratype</typeStatus>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MYR10304">
|
||
<collectionCode>MYR</collectionCode>
|
||
10304
|
||
</accessionNumber>
|
||
</emphasis>
|
||
), head, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">D</emphasis>
|
||
antennal base and
|
||
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
|
||
organ.
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Abbreviations</emphasis>
|
||
: I - VII = antennomeres, ad = apical disc, cl = clypeus, co = collum, epc = epicranium, fr = frons, gn = gnathochilarium, il = incisura lateralis, lab = labrum, mb = mandibular base, sc = sensory cones, to =
|
||
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
|
||
organ. Scale bars: 100
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A, C, D</emphasis>
|
||
), 50
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Antennae consisting of seven joints, each covered with numerous setae. Size of joints 1 = 2 = 3 = 4 = 5> 6 (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Glomeridesmus siamensis sp. nov., head, SEM A, B, D male holotype (ZCSWU Myr D 000011) A antennae B apical antennomeres C juvenile paratype (ZFMK MYR 10304), head, lateral view D antennal base and Toemoesvary organ. Abbreviations: I - VII = antennomeres, ad = apical disc, cl = clypeus, co = collum, epc = epicranium, fr = frons, gn = gnathochilarium, il = incisura lateralis, lab = labrum, mb = mandibular base, sc = sensory cones, to = Toemoesvary organ. Scale bars: 100 µm (A, C, D), 50 µm (B)." figureDoi="10.3897/zookeys.1024.63678.figure2" httpUri="https://binary.pensoft.net/fig/520750" pageId="0" pageNumber="137">2A</figureCitation>
|
||
). Apical disc carrying four long apical cones; sensilla basiconica apparently absent (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Glomeridesmus siamensis sp. nov., head, SEM A, B, D male holotype (ZCSWU Myr D 000011) A antennae B apical antennomeres C juvenile paratype (ZFMK MYR 10304), head, lateral view D antennal base and Toemoesvary organ. Abbreviations: I - VII = antennomeres, ad = apical disc, cl = clypeus, co = collum, epc = epicranium, fr = frons, gn = gnathochilarium, il = incisura lateralis, lab = labrum, mb = mandibular base, sc = sensory cones, to = Toemoesvary organ. Scale bars: 100 µm (A, C, D), 50 µm (B)." figureDoi="10.3897/zookeys.1024.63678.figure2" httpUri="https://binary.pensoft.net/fig/520750" pageId="0" pageNumber="137">2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
|
||
organs as large as antennal base, of well-rounded oval shape, interior covered by a sclerotized plate; surrounded by cuticular rim and several very short setae (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Glomeridesmus siamensis sp. nov., head, SEM A, B, D male holotype (ZCSWU Myr D 000011) A antennae B apical antennomeres C juvenile paratype (ZFMK MYR 10304), head, lateral view D antennal base and Toemoesvary organ. Abbreviations: I - VII = antennomeres, ad = apical disc, cl = clypeus, co = collum, epc = epicranium, fr = frons, gn = gnathochilarium, il = incisura lateralis, lab = labrum, mb = mandibular base, sc = sensory cones, to = Toemoesvary organ. Scale bars: 100 µm (A, C, D), 50 µm (B)." figureDoi="10.3897/zookeys.1024.63678.figure2" httpUri="https://binary.pensoft.net/fig/520750" pageId="0" pageNumber="137">2C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Gnathochilarium with very broad gula (hypostoma). Cardines very small, separated from basal part of gnathochilarium. Proximal fourth of mentum towards gula distinctly elevated, transverse suture visible between mentum and lamellae linguales. Lamellae linguales fused to one another, not separated by a longitudinal suture, apically with four long setae, central pair at apical margin, lateral two setae located more posteriorly. Whole surface of gnathochilarium covered with few, isolated setae. Stipites laterally without sclerotized ledge. Stipites apically with 2 extra-long setae (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011) mouthparts A gnathochilarium, ventral view B gnathochilarium, palps and central pad, frontal view C endochilarium, dorsal view D mandibular gnathal lobe, mesal view. Abbreviations: cp = central pad, ec = endochilarium, et = external tooth, gu = gula, ip = inner palp, ia = intermediate area, it = inner tooth, LL = lamella lingualis, Lp = lateral palp, me = mentum, mp = molar plate, pL = pectinate lamellae, stp = stipes. Scale bars: 100 µm (A), 50 µm (B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure6" httpUri="https://binary.pensoft.net/fig/520754" pageId="0" pageNumber="137">6A</figureCitation>
|
||
). Stipites lateral palpi slender, carrying 4 slender sensory cones; inner palpi very wide, carrying 12-15 long, tube-shaped sensory cones interspersed between long cuticular fibres (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011) mouthparts A gnathochilarium, ventral view B gnathochilarium, palps and central pad, frontal view C endochilarium, dorsal view D mandibular gnathal lobe, mesal view. Abbreviations: cp = central pad, ec = endochilarium, et = external tooth, gu = gula, ip = inner palp, ia = intermediate area, it = inner tooth, LL = lamella lingualis, Lp = lateral palp, me = mentum, mp = molar plate, pL = pectinate lamellae, stp = stipes. Scale bars: 100 µm (A), 50 µm (B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure6" httpUri="https://binary.pensoft.net/fig/520754" pageId="0" pageNumber="137">6B</figureCitation>
|
||
). Central pads (modified central palpi of lamellae linguales) large, mesally pointing towards and touching one another, in ventral view covered with numerous long, tube-shaped sensory cones and long cuticular fibers (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011) mouthparts A gnathochilarium, ventral view B gnathochilarium, palps and central pad, frontal view C endochilarium, dorsal view D mandibular gnathal lobe, mesal view. Abbreviations: cp = central pad, ec = endochilarium, et = external tooth, gu = gula, ip = inner palp, ia = intermediate area, it = inner tooth, LL = lamella lingualis, Lp = lateral palp, me = mentum, mp = molar plate, pL = pectinate lamellae, stp = stipes. Scale bars: 100 µm (A), 50 µm (B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure6" httpUri="https://binary.pensoft.net/fig/520754" pageId="0" pageNumber="137">6B</figureCitation>
|
||
). Basally of central pads on endochilarium with two parallel rows of 9 sensory cones along median furrow (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011) mouthparts A gnathochilarium, ventral view B gnathochilarium, palps and central pad, frontal view C endochilarium, dorsal view D mandibular gnathal lobe, mesal view. Abbreviations: cp = central pad, ec = endochilarium, et = external tooth, gu = gula, ip = inner palp, ia = intermediate area, it = inner tooth, LL = lamella lingualis, Lp = lateral palp, me = mentum, mp = molar plate, pL = pectinate lamellae, stp = stipes. Scale bars: 100 µm (A), 50 µm (B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure6" httpUri="https://binary.pensoft.net/fig/520754" pageId="0" pageNumber="137">6C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Mandible</emphasis>
|
||
:
|
||
</emphasis>
|
||
Basal joint massive, undivided (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">1D, E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Glomeridesmus siamensis sp. nov., head, SEM A, B, D male holotype (ZCSWU Myr D 000011) A antennae B apical antennomeres C juvenile paratype (ZFMK MYR 10304), head, lateral view D antennal base and Toemoesvary organ. Abbreviations: I - VII = antennomeres, ad = apical disc, cl = clypeus, co = collum, epc = epicranium, fr = frons, gn = gnathochilarium, il = incisura lateralis, lab = labrum, mb = mandibular base, sc = sensory cones, to = Toemoesvary organ. Scale bars: 100 µm (A, C, D), 50 µm (B)." figureDoi="10.3897/zookeys.1024.63678.figure2" httpUri="https://binary.pensoft.net/fig/520750" pageId="0" pageNumber="137">2C</figureCitation>
|
||
). Apical joint (mandibular gnathal lobe) damaged during preparation, but with long slender external tooth and a long condylus. Inner tooth 4-combed. Seven rows of pectinate lamellae, large intermediate area covered by scale-like spines. Additional intermediate area located at a lower level next to molar plate with more elongated spine-plates. Molar plate plane, without indentations or grooves, margins entirely surrounded by thick membranous fringes (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011) mouthparts A gnathochilarium, ventral view B gnathochilarium, palps and central pad, frontal view C endochilarium, dorsal view D mandibular gnathal lobe, mesal view. Abbreviations: cp = central pad, ec = endochilarium, et = external tooth, gu = gula, ip = inner palp, ia = intermediate area, it = inner tooth, LL = lamella lingualis, Lp = lateral palp, me = mentum, mp = molar plate, pL = pectinate lamellae, stp = stipes. Scale bars: 100 µm (A), 50 µm (B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure6" httpUri="https://binary.pensoft.net/fig/520754" pageId="0" pageNumber="137">6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Trunk</emphasis>
|
||
:
|
||
</emphasis>
|
||
Collum (tergite 1) wider than head, shape similar to following tergites (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">1A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Tergites 2-19 (20) very thin, difficult to distinguish from one another. Width from tergite 1 to 8 gradually increasing, then decreasing to last tergite (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">1A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Glomeridesmus siamensis sp. nov., body-rings A, B, D juvenile paratype (ZFMK MYR 10304) A habitus, lateral view, arrow indicates drawn out lateral tip of tergite B detail of tergite margin, latero-ventral view, arrow indicates striae on tergite C male holotype (ZCSWU Myr D 000011), endotergum, arrow indicates posterior margin of tergite D posterior body-rings, lateral view, arrows indicate teeth on posterior margin of tergite. Abbreviations: co = collum, Lt = pleurite, te = tergite, tel = telson. Scale bars: 500 µm (A), 100 µm (B, C), 50 µm (D)." figureDoi="10.3897/zookeys.1024.63678.figure7" httpUri="https://binary.pensoft.net/fig/520755" pageId="0" pageNumber="137">7A</figureCitation>
|
||
). Each tergite bearing 6-8 longitudinal striae. Distances between striae increasing towards posterior margin. Surface of tergites with sparse isolated setae (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Glomeridesmus siamensis sp. nov., body-rings A, B, D juvenile paratype (ZFMK MYR 10304) A habitus, lateral view, arrow indicates drawn out lateral tip of tergite B detail of tergite margin, latero-ventral view, arrow indicates striae on tergite C male holotype (ZCSWU Myr D 000011), endotergum, arrow indicates posterior margin of tergite D posterior body-rings, lateral view, arrows indicate teeth on posterior margin of tergite. Abbreviations: co = collum, Lt = pleurite, te = tergite, tel = telson. Scale bars: 500 µm (A), 100 µm (B, C), 50 µm (D)." figureDoi="10.3897/zookeys.1024.63678.figure7" httpUri="https://binary.pensoft.net/fig/520755" pageId="0" pageNumber="137">7A, B</figureCitation>
|
||
). Tergite 11 onwards posterior-lateral edge pronounced into a sharp-edged tip, becoming a longer and spine-like process at tergite 17. Limbus (posterior margin of tergite) smooth, without any structures. Endotergum (underside of posterior margin of tergite) smooth area without any specific structures (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Glomeridesmus siamensis sp. nov., body-rings A, B, D juvenile paratype (ZFMK MYR 10304) A habitus, lateral view, arrow indicates drawn out lateral tip of tergite B detail of tergite margin, latero-ventral view, arrow indicates striae on tergite C male holotype (ZCSWU Myr D 000011), endotergum, arrow indicates posterior margin of tergite D posterior body-rings, lateral view, arrows indicate teeth on posterior margin of tergite. Abbreviations: co = collum, Lt = pleurite, te = tergite, tel = telson. Scale bars: 500 µm (A), 100 µm (B, C), 50 µm (D)." figureDoi="10.3897/zookeys.1024.63678.figure7" httpUri="https://binary.pensoft.net/fig/520755" pageId="0" pageNumber="137">7C</figureCitation>
|
||
). Posterior margin of tergite 19 and 20 with several short teeth (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Glomeridesmus siamensis sp. nov., body-rings A, B, D juvenile paratype (ZFMK MYR 10304) A habitus, lateral view, arrow indicates drawn out lateral tip of tergite B detail of tergite margin, latero-ventral view, arrow indicates striae on tergite C male holotype (ZCSWU Myr D 000011), endotergum, arrow indicates posterior margin of tergite D posterior body-rings, lateral view, arrows indicate teeth on posterior margin of tergite. Abbreviations: co = collum, Lt = pleurite, te = tergite, tel = telson. Scale bars: 500 µm (A), 100 µm (B, C), 50 µm (D)." figureDoi="10.3897/zookeys.1024.63678.figure7" httpUri="https://binary.pensoft.net/fig/520755" pageId="0" pageNumber="137">7D</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Pleurites of rectangular shape, posterior margin overlapping first third of subsequent pleurite (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Glomeridesmus siamensis sp. nov., body-rings A, B, D juvenile paratype (ZFMK MYR 10304) A habitus, lateral view, arrow indicates drawn out lateral tip of tergite B detail of tergite margin, latero-ventral view, arrow indicates striae on tergite C male holotype (ZCSWU Myr D 000011), endotergum, arrow indicates posterior margin of tergite D posterior body-rings, lateral view, arrows indicate teeth on posterior margin of tergite. Abbreviations: co = collum, Lt = pleurite, te = tergite, tel = telson. Scale bars: 500 µm (A), 100 µm (B, C), 50 µm (D)." figureDoi="10.3897/zookeys.1024.63678.figure7" httpUri="https://binary.pensoft.net/fig/520755" pageId="0" pageNumber="137">7B</figureCitation>
|
||
), wider than long. Pleurites similar to one another, but last pleurite on posterior margin with a well-rounded central indentation, providing space for the movements of the last leg.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Legs</emphasis>
|
||
:
|
||
</emphasis>
|
||
In all except first coxae fused with stigmatic plates (typical for
|
||
<taxonomicName authorityName="Gervais" authorityYear="1844" class="Diplopoda" kingdom="Animalia" lsidName="" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="order">Glomeridesmida</taxonomicName>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.1016/j.asd.2013.11.003" author="Wesener, T" journalOrPublisher="Arthropod Structure and Development" pageId="0" pageNumber="137" pagination="87 - 95" refId="B43" refString="Wesener, T, Sierwald, P, Waegele, J-W, 2014. Sternites and spiracles - The unclear homology of ventral sclerites in the basal millipede order Glomeridesmida (Myriapoda, Diplopoda). Arthropod Structure and Development 43: 87 - 95, DOI: https://doi.org/10.1016/j.asd.2013.11.003" title="Sternites and spiracles - The unclear homology of ventral sclerites in the basal millipede order Glomeridesmida (Myriapoda, Diplopoda)." url="https://doi.org/10.1016/j.asd.2013.11.003" volume="43" year="2014">Wesener et al. 2014</bibRefCitation>
|
||
) (Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3A-H</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Glomeridesmus siamensis sp. nov., female paratype (ZFMK MYR 10302), legs and ovipositor, SEM A leg 23, posterior view B leg 23, ventral view C leg 2 with ovipositor, ventral view D ovipositors, posterior view. Abbreviations: ap = apical part of ovipositor, bp = basal part of ovipositor, cp = coxal pouch, cx = coxa, fe = femur, L 2 = leg 2, Lt = pleurite, pof = postfemur, prf = prefemur, sh = sheet, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 50 µm (A, B), 100 µm (C, D)." figureDoi="10.3897/zookeys.1024.63678.figure4" httpUri="https://binary.pensoft.net/fig/520752" pageId="0" pageNumber="137">4A-C</figureCitation>
|
||
). Starting at coxo-stigmatic plate 9 (male
|
||
<typeStatus>holotype</typeStatus>
|
||
) or 11 (female
|
||
<typeStatus>paratype</typeStatus>
|
||
), every second coxa up to pair 21 (male) or 25 (female) apically with an eversible coxal pouch posteriorly of prefemur (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Glomeridesmus siamensis sp. nov., female paratype (ZFMK MYR 10302), legs and ovipositor, SEM A leg 23, posterior view B leg 23, ventral view C leg 2 with ovipositor, ventral view D ovipositors, posterior view. Abbreviations: ap = apical part of ovipositor, bp = basal part of ovipositor, cp = coxal pouch, cx = coxa, fe = femur, L 2 = leg 2, Lt = pleurite, pof = postfemur, prf = prefemur, sh = sheet, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 50 µm (A, B), 100 µm (C, D)." figureDoi="10.3897/zookeys.1024.63678.figure4" httpUri="https://binary.pensoft.net/fig/520752" pageId="0" pageNumber="137">4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Figure 3.</emphasis>
|
||
<taxonomicName class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Glomeridesmus siamensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male
|
||
<typeStatus>holotype</typeStatus>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>ZCSWU</collectionCode>
|
||
Myr D000011
|
||
</emphasis>
|
||
), legs, SEM
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
leg 1, posterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
|
||
leg 1, tarsus, arrow indicated parasitic fungi
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C</emphasis>
|
||
leg 2 with gonopore, posterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">D</emphasis>
|
||
leg 2, tarsus
|
||
<emphasis bold="true" pageId="0" pageNumber="137">E</emphasis>
|
||
leg 34, posterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">F</emphasis>
|
||
leg 34, tarsus
|
||
<emphasis bold="true" pageId="0" pageNumber="137">G</emphasis>
|
||
sensory leg (leg 35), posterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">H</emphasis>
|
||
sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I.
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Abbreviations</emphasis>
|
||
: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A, C, E, G</emphasis>
|
||
), 20
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
|
||
), 10
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">D, F</emphasis>
|
||
), 50
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">H</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Leg-pair 1 with a well-rounded elliptical sternite visible in oral view. Femur 1.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than wide, tarsus slender and elongated,>7 times longer than wide (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3A</figureCitation>
|
||
). Apical half of tarsus without any spines or setae, except 1 or 2 apical spines, claw and paronychium (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Leg 2 (male) femur 1.7, tarsus>10
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than wide (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3C</figureCitation>
|
||
). Tarsus in apical half without any spines or setae except for single apical spine, claw and paronychium (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Legs 3-32 similar to first legs, e.g. midbody leg femur 1.8, tarsus>7 times longer than wide. Tarsus with 1 or 2 apical spines, claw and paronychium (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Penultimate leg with coxosternite narrow, with stigma opening and sternal part located below (instead of lateral to) of coxal part. Femur slightly longer than prefemur, 1.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than wide, tarsus shorter,>7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than wide, apically with claw, apical spine and paronychium (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3E, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Ultimate leg pair (sensory leg) modified, consisting of a free large sternite, fused coxa-stigmatic plates plus 3 podomeres (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3G</figureCitation>
|
||
). First podomere with a conspicuous mesal bend. Last two podomeres extending postero-laterally (confused with telopod by pre-1941 authors). Podomere 3 apically with a large, long spine, longer than the podomere (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3H</figureCitation>
|
||
). Both legs widely separated from one another, connected by large sternite. Coxa-stigmatic plates large, almost as long as all 3 podomeres combined (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Anal shield</emphasis>
|
||
:
|
||
</emphasis>
|
||
glabrous, with a well-rounded edge. Subanal plate located behind last pair of legs, large and hyaline.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Sexual characters</emphasis>
|
||
: Female
|
||
</emphasis>
|
||
: second coxae on posterior side with prominent ovipositors protruding back to leg pair 5. Coxa protruding mesally as a short lobe. Ovipositors basally supported by an undivided plate (sternite?) (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Glomeridesmus siamensis sp. nov., female paratype (ZFMK MYR 10302), legs and ovipositor, SEM A leg 23, posterior view B leg 23, ventral view C leg 2 with ovipositor, ventral view D ovipositors, posterior view. Abbreviations: ap = apical part of ovipositor, bp = basal part of ovipositor, cp = coxal pouch, cx = coxa, fe = femur, L 2 = leg 2, Lt = pleurite, pof = postfemur, prf = prefemur, sh = sheet, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 50 µm (A, B), 100 µm (C, D)." figureDoi="10.3897/zookeys.1024.63678.figure4" httpUri="https://binary.pensoft.net/fig/520752" pageId="0" pageNumber="137">4C</figureCitation>
|
||
). Ovipositor with a basal part consisting of 21 or 22 segments of (eversible?) half-rings, each carrying isolated setae in a regular distance to each other. Half rings anteriorly and posteriorly interrupted by two different, much slender tubes, also consisting of half-rings, running along the whole length of the ovipositor. Apical part of ovipositor with opening surrounded by four larger plates, each arising out of a tube of rings. All four plates covered by longer setae (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Glomeridesmus siamensis sp. nov., female paratype (ZFMK MYR 10302), legs and ovipositor, SEM A leg 23, posterior view B leg 23, ventral view C leg 2 with ovipositor, ventral view D ovipositors, posterior view. Abbreviations: ap = apical part of ovipositor, bp = basal part of ovipositor, cp = coxal pouch, cx = coxa, fe = femur, L 2 = leg 2, Lt = pleurite, pof = postfemur, prf = prefemur, sh = sheet, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 50 µm (A, B), 100 µm (C, D)." figureDoi="10.3897/zookeys.1024.63678.figure4" httpUri="https://binary.pensoft.net/fig/520752" pageId="0" pageNumber="137">4D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1024.63678.figure4" httpUri="https://binary.pensoft.net/fig/520752" pageId="0" pageNumber="137" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Figure 4.</emphasis>
|
||
<taxonomicName class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Glomeridesmus siamensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., female
|
||
<typeStatus>paratype</typeStatus>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MYR10302">
|
||
<collectionCode>MYR</collectionCode>
|
||
10302
|
||
</accessionNumber>
|
||
</emphasis>
|
||
), legs and ovipositor, SEM
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
leg 23, posterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
|
||
leg 23, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C</emphasis>
|
||
leg 2 with ovipositor, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">D</emphasis>
|
||
ovipositors, posterior view.
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Abbreviations</emphasis>
|
||
: ap = apical part of ovipositor, bp = basal part of ovipositor, cp = coxal pouch, cx = coxa, fe = femur, L2 = leg 2, Lt = pleurite, pof = postfemur, prf = prefemur, sh = sheet, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 50
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A, B</emphasis>
|
||
), 100
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C, D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Male</emphasis>
|
||
: second coxa with gonopore located mesally (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3C</figureCitation>
|
||
). Gonopore flanked by two sclerotized, plates resembling an open bivalve shell. Apical part of both plates with 2 or 3 longer setae (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), male sexual characters A telopod, anterior view B telopod, posterior view C gonopore on leg 2, posterior view D inner horns, anterior view. Abbreviations: I - IV = telopoditomeres, ff = immobile finger, gp = gonopore, ih = inner horns, ma = membranous area, pr = projection. Scale bars: 50 µm (A, B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure5" httpUri="https://binary.pensoft.net/fig/520753" pageId="0" pageNumber="137">5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Male telopod</emphasis>
|
||
consisting of syncoxite with inner horns and 4 podomeres (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Glomeridesmus siamensis sp. nov., photographs A-C male holotype (ZCSWU Myr D 000011) A habitus, dorsal view B habitus, ventral view C telopods, ventral view D, E female paratype (ZFMK MYR 10302) D head and anterior body-rings, lateral view E head and anterior body-rings, ventral view. Not to scale. Abbreviations: at = antennae, co = collum, gn = gnathochilarium, hd = head, ih = inner horns, mb = mandibular base, sL = sensory leg, to = Toemoesvary organ, tp = telopod, vu = vulva." figureDoi="10.3897/zookeys.1024.63678.figure1" httpUri="https://binary.pensoft.net/fig/520749" pageId="0" pageNumber="137">1C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), male sexual characters A telopod, anterior view B telopod, posterior view C gonopore on leg 2, posterior view D inner horns, anterior view. Abbreviations: I - IV = telopoditomeres, ff = immobile finger, gp = gonopore, ih = inner horns, ma = membranous area, pr = projection. Scale bars: 50 µm (A, B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure5" httpUri="https://binary.pensoft.net/fig/520753" pageId="0" pageNumber="137">5A, D</figureCitation>
|
||
). Syncoxite covering basal podomeres laterally and in anterior view, medially rising into a smooth glabrous process. Process apically with two long inner horns; horns basally fused, completely separating more apically, diverging and running parallel to one another. Each horn apically slightly widening and spoon-shaped. Whole surface of horns sparsely covered by isolated, minute setae (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), male sexual characters A telopod, anterior view B telopod, posterior view C gonopore on leg 2, posterior view D inner horns, anterior view. Abbreviations: I - IV = telopoditomeres, ff = immobile finger, gp = gonopore, ih = inner horns, ma = membranous area, pr = projection. Scale bars: 50 µm (A, B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure5" httpUri="https://binary.pensoft.net/fig/520753" pageId="0" pageNumber="137">5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1024.63678.figure5" httpUri="https://binary.pensoft.net/fig/520753" pageId="0" pageNumber="137" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Figure 5.</emphasis>
|
||
<taxonomicName class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Glomeridesmus siamensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male
|
||
<typeStatus>holotype</typeStatus>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>ZCSWU</collectionCode>
|
||
Myr D000011
|
||
</emphasis>
|
||
), male sexual characters
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
telopod, anterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
|
||
telopod, posterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C</emphasis>
|
||
gonopore on leg 2, posterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">D</emphasis>
|
||
inner horns, anterior view.
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Abbreviations</emphasis>
|
||
: I - IV = telopoditomeres, ff = immobile finger, gp = gonopore, ih = inner horns, ma = membranous area, pr = projection. Scale bars: 50
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A, B, D</emphasis>
|
||
), 20
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="137">Telopod podomere 1 largest and most massive.</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Podomere 2 wide, rectangular, apically with a large membranous area and mesally with a long and slender process. Membranous area well rounded, consisting of several inverse membranous folds. Slender process strongly elongated, as long as podomere 2, protruding between movable and immovable finger, inner surface excavated (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), male sexual characters A telopod, anterior view B telopod, posterior view C gonopore on leg 2, posterior view D inner horns, anterior view. Abbreviations: I - IV = telopoditomeres, ff = immobile finger, gp = gonopore, ih = inner horns, ma = membranous area, pr = projection. Scale bars: 50 µm (A, B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure5" httpUri="https://binary.pensoft.net/fig/520753" pageId="0" pageNumber="137">5A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1024.63678.figure6" httpUri="https://binary.pensoft.net/fig/520754" pageId="0" pageNumber="137" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Figure 6.</emphasis>
|
||
<taxonomicName class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Glomeridesmus siamensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male
|
||
<typeStatus>holotype</typeStatus>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>ZCSWU</collectionCode>
|
||
Myr D000011
|
||
</emphasis>
|
||
) mouthparts
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
gnathochilarium, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
|
||
gnathochilarium, palps and central pad, frontal view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C</emphasis>
|
||
endochilarium, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">D</emphasis>
|
||
mandibular gnathal lobe, mesal view.
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Abbreviations</emphasis>
|
||
: cp = central pad, ec = endochilarium, et = external tooth, gu = gula, ip = inner palp, ia = intermediate area, it = inner tooth, LL = lamella lingualis, Lp = lateral palp, me = mentum, mp = molar plate, pL = pectinate lamellae, stp = stipes. Scale bars: 100
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
), 50
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B, D</emphasis>
|
||
), 20
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Podomere 3 long and slender, only 1/3 of width of podomere 2, inserting laterally on podomere 2. Posterior-lateral aspect of podomere
|
||
<quantity metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
|
||
apical part with slender, finger-shaped and well-rounded process called immobile finger, which makes up ca. half the length of podomere 3, inner surface of immobile finger excavated. Length of podomere 3, without immobile finger, 2 times longer than wide, with few isolated setae at margins in apical half (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), male sexual characters A telopod, anterior view B telopod, posterior view C gonopore on leg 2, posterior view D inner horns, anterior view. Abbreviations: I - IV = telopoditomeres, ff = immobile finger, gp = gonopore, ih = inner horns, ma = membranous area, pr = projection. Scale bars: 50 µm (A, B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure5" httpUri="https://binary.pensoft.net/fig/520753" pageId="0" pageNumber="137">5A, B</figureCitation>
|
||
). Immobile finger protruding up to 3/4 of length of podomere 4, called mobile finger (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), male sexual characters A telopod, anterior view B telopod, posterior view C gonopore on leg 2, posterior view D inner horns, anterior view. Abbreviations: I - IV = telopoditomeres, ff = immobile finger, gp = gonopore, ih = inner horns, ma = membranous area, pr = projection. Scale bars: 50 µm (A, B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure5" httpUri="https://binary.pensoft.net/fig/520753" pageId="0" pageNumber="137">5A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Mobile finger (podomere 4), articulated to podomere 3 at base of immovable finger (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), male sexual characters A telopod, anterior view B telopod, posterior view C gonopore on leg 2, posterior view D inner horns, anterior view. Abbreviations: I - IV = telopoditomeres, ff = immobile finger, gp = gonopore, ih = inner horns, ma = membranous area, pr = projection. Scale bars: 50 µm (A, B, D), 20 µm (C)." figureDoi="10.3897/zookeys.1024.63678.figure5" httpUri="https://binary.pensoft.net/fig/520753" pageId="0" pageNumber="137">5A, B</figureCitation>
|
||
), slender and glabrous, 5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than wide, posteriorly with an excavated area; at apex with a single long spine reaching 2/3 of length of immobile finger
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="137" type="immature male">
|
||
<paragraph pageId="0" pageNumber="137">Immature male.</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">The immature male has the same number of tergites, but one walking leg pair less than the mature male. The immature male is almost 1/3 shorter than the mature male. Coxal pouches starting at leg pair 9 as in mature male. Last pleurite with small well-rounded indentation at posterior margin. Telopod only slightly different from mature male: process of podomere 2 shorter, podomere 3 and 4 slenderer, immobile finger almost protruding up to apical end of movable finger.</paragraph>
|
||
<caption doi="10.3897/zookeys.1024.63678.figure7" httpUri="https://binary.pensoft.net/fig/520755" pageId="0" pageNumber="137" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Figure 7.</emphasis>
|
||
<taxonomicName class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Glomeridesmus siamensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., body-rings
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A, B, D</emphasis>
|
||
juvenile
|
||
<typeStatus>paratype</typeStatus>
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
(
|
||
<collectionCode country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MYR10304">
|
||
<collectionCode>MYR</collectionCode>
|
||
10304
|
||
</accessionNumber>
|
||
</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
habitus, lateral view, arrow indicates drawn out lateral tip of tergite
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
|
||
detail of tergite margin, latero-ventral view, arrow indicates striae on tergite
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C</emphasis>
|
||
male
|
||
<typeStatus>holotype</typeStatus>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>ZCSWU</collectionCode>
|
||
Myr D000011
|
||
</emphasis>
|
||
), endotergum, arrow indicates posterior margin of tergite
|
||
<emphasis bold="true" pageId="0" pageNumber="137">D</emphasis>
|
||
posterior body-rings, lateral view, arrows indicate teeth on posterior margin of tergite.
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Abbreviations</emphasis>
|
||
: co = collum, Lt = pleurite, te = tergite, tel = telson. Scale bars: 500
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
), 100
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B, C</emphasis>
|
||
), 50
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="137">D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="137" type="ecology">
|
||
<paragraph pageId="0" pageNumber="137">Ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
All specimens were collected by hand in the morning during rainy season in
|
||
<collectingRegion country="Thailand" name="Krabi">Krabi province</collectingRegion>
|
||
, the second reported area for
|
||
<taxonomicName authorityName="Gervais" authorityYear="1844" class="Diplopoda" kingdom="Animalia" lsidName="" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="order">Glomeridesmida</taxonomicName>
|
||
in
|
||
<collectingCountry name="Thailand">Thailand</collectingCountry>
|
||
(the first in
|
||
<bibRefCitation author="Shelley, R" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="137" pagination="1 - 11" refId="B36" refString="Shelley, R, 2011. The milliped order Glomeridesmida (Diplopoda: Pentazonia: Limacomorpha) in Oceania, the East Indies, and southeastern Asia; first records from Palau, the Philippines, Vanuatu, New Britain, the Island of New Guinea, Cambodia, Thailand, and Borneo and Sulawesi, Indonesia. Insecta Mundi 0196: 1 - 11" title="The milliped order Glomeridesmida (Diplopoda: Pentazonia: Limacomorpha) in Oceania, the East Indies, and southeastern Asia; first records from Palau, the Philippines, Vanuatu, New Britain, the Island of New Guinea, Cambodia, Thailand, and Borneo and Sulawesi, Indonesia." volume="0196" year="2011">Shelley 2011</bibRefCitation>
|
||
). They were found in a karst landscape covered with evergreen forest (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitat and type locality of Glomeridesmus siamensis sp. nov. A karstic landscape around Krabi Town, view from Tiger Cave temple (Wat Tham Suea) B, C slope underneath Tiger Cave temple at which Glomeridesmus siamensis sp. nov. was found, arrow indicates the overgrown rock from which the specimens were collected." figureDoi="10.3897/zookeys.1024.63678.figure8" httpUri="https://binary.pensoft.net/fig/520756" pageId="0" pageNumber="137">8A, B</figureCitation>
|
||
) in soils at a depth of
|
||
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10-20 mm</quantity>
|
||
near horizontal roots on a partially overgrown rock. Despite numerous attempts, no
|
||
<taxonomicName authorityName="Gervais" authorityYear="1844" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Glomeridesmus</emphasis>
|
||
</taxonomicName>
|
||
was discovered outside the less than 30
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
<quantity metricMagnitude="-1" metricUnit="m" metricValue="3.0" unit="cm" value="30.0">30 centimeter</quantity>
|
||
of habitat, showing that their distribution might be dependent on some unknown microhabitat requirements. The habitat was a flat area on a western aspect (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitat and type locality of Glomeridesmus siamensis sp. nov. A karstic landscape around Krabi Town, view from Tiger Cave temple (Wat Tham Suea) B, C slope underneath Tiger Cave temple at which Glomeridesmus siamensis sp. nov. was found, arrow indicates the overgrown rock from which the specimens were collected." figureDoi="10.3897/zookeys.1024.63678.figure8" httpUri="https://binary.pensoft.net/fig/520756" pageId="0" pageNumber="137">8C</figureCitation>
|
||
). Average air temperature and relative humidity during observation was approximately 29 °C and 85%, respectively. The specimens were fast moving (at least as fast as
|
||
<taxonomicName authorityName="Gervais" authorityYear="1844" class="Diplopoda" kingdom="Animalia" lsidName="" order="Chordeumatida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="order">Chordeumatida</taxonomicName>
|
||
), capable of bending their flexible body 180° to quickly change directions. Furthermore, the male
|
||
<typeStatus>holotype</typeStatus>
|
||
carried an
|
||
<normalizedToken originalValue="‘amphoromorph’">'amphoromorph'</normalizedToken>
|
||
fungus on one of its legs (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Glomeridesmus siamensis sp. nov., male holotype (ZCSWU Myr D 000011), legs, SEM A leg 1, posterior view B leg 1, tarsus, arrow indicated parasitic fungi C leg 2 with gonopore, posterior view D leg 2, tarsus E leg 34, posterior view F leg 34, tarsus G sensory leg (leg 35), posterior view H sensory leg (leg 35), detail of podomeres, arrow indicated dent in podomere I. Abbreviations: I-III = podomeres, as = apical spine, cl = claw, cx = coxa, fe = femur, gp = gonopore, pn = paronychium, pof = postfemur, prf = prefemur, sp = spine, st = sternite, ta = tarsus, ti = tibia, v = stigmatic plate. Scale bars: 100 µm (A, C, E, G), 20 µm (B), 10 µm (D, F), 50 µm (H)." figureDoi="10.3897/zookeys.1024.63678.figure3" httpUri="https://binary.pensoft.net/fig/520751" pageId="0" pageNumber="137">3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1024.63678.figure8" httpUri="https://binary.pensoft.net/fig/520756" pageId="0" pageNumber="137" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Figure 8.</emphasis>
|
||
Habitat and
|
||
<typeStatus>type</typeStatus>
|
||
locality of
|
||
<taxonomicName class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Glomeridesmus siamensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
karstic landscape around
|
||
<collectingRegion country="Thailand" name="Krabi">Krabi</collectingRegion>
|
||
Town, view from Tiger Cave temple (Wat Tham Suea)
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B, C</emphasis>
|
||
slope underneath Tiger Cave temple at which
|
||
<taxonomicName class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus siamensis" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Glomeridesmus siamensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. was found, arrow indicates the overgrown rock from which the specimens were collected.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="137" type="additional locality data of unidentified glomeridesmus from thailand">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Additional locality data of unidentified
|
||
<taxonomicName authorityName="Gervais" authorityYear="1844" class="Diplopoda" family="Glomeridesmidae" genus="Glomeridesmus" kingdom="Animalia" lsidName="Glomeridesmus" order="Glomeridesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">Glomeridesmus</taxonomicName>
|
||
from
|
||
<collectingCountry name="Thailand">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<materialsCitation ID-GBIF-Occurrence="3063614308" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NHMD" specimenCount="2">
|
||
<specimenCount type="generic">2 imm.</specimenCount>
|
||
F;
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>NHMD</collectionCode>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3063614316" collectingDate="1981-09-30" collectionCode="NHMD" collectorName="Doi Suthep N. P. & Konthathan & Montha Than" elevation="650" location="Chiang Mai Province" specimenCount="1" stateProvince="Chiang Mai">
|
||
<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
|
||
,
|
||
<collectorName>Doi Suthep N. P.</collectorName>
|
||
;
|
||
<collectorName>Konthathan</collectorName>
|
||
(
|
||
<collectorName>Montha Than</collectorName>
|
||
waterfall?);
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">600-700 m</elevation>
|
||
</quantity>
|
||
; leg.
|
||
<collectionCode>NHMD</collectionCode>
|
||
,
|
||
<date value="1981-09-30">
|
||
<collectingDate value="1981-09-30">30.ix.1981</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation ID-GBIF-Occurrence="3063614309" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NHMD" specimenCount="1" stateProvince="Chiang Mai">
|
||
<specimenCount type="generic">1 imm.</specimenCount>
|
||
F.;
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>NHMD</collectionCode>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3063614302" collectingDate="1981-09-26" collectionCode="NHMD" collectorName="Doi Suthep N. P. & Montha Than" elevation="650" location="Chiang Mai Province" specimenCount="1" stateProvince="Chiang Mai">
|
||
<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai Province</collectingRegion>
|
||
,
|
||
<collectorName>Doi Suthep N. P.</collectorName>
|
||
;
|
||
<collectorName>Montha Than</collectorName>
|
||
;
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">600-700 m</elevation>
|
||
</quantity>
|
||
; leg.
|
||
<collectionCode>NHMD</collectionCode>
|
||
,
|
||
<date value="1981-09-26">
|
||
<collectingDate value="1981-09-26">26.ix.1981</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation ID-GBIF-Occurrence="3063614304" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NHMD" specimenCount="1" stateProvince="Chiang Mai">
|
||
<specimenCount type="generic">1 juv.</specimenCount>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>NHMD</collectionCode>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3063614315" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Birgit Degerbol & Doi Suthep" elevation="1150" location="Lok." municipality="Doi Suthep" specimenCount="1" stateProvince="Chiang Mai">
|
||
<collectingMunicipality>Doi Suthep</collectingMunicipality>
|
||
,
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">1150 m</elevation>
|
||
</quantity>
|
||
;
|
||
<location LSID="urn:lsid:plazi:treatment:2AB443F1DFF45118AE5260106B9D2694:9F38150D4E0FB774CEB3E2ED921EB8EA" municipality="Doi Suthep" name="Lok." stateProvince="Chiang Mai">Lok.</location>
|
||
3a; 29/9.1958;
|
||
<collectorName>
|
||
Birgit
|
||
<normalizedToken originalValue="Degerbøl">Degerbol</normalizedToken>
|
||
</collectorName>
|
||
leg. Zool. Mus. Kbh. 1/7 59.
|
||
<collectorName>Doi Suthep</collectorName>
|
||
,
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.055" metricValueMax="12.0" metricValueMin="0.11" unit="m" value="605.5" valueMax="1200.0" valueMin="11.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.055" metricValueMax="12.0" metricValueMin="0.11" unit="m" value="605.5" valueMax="1200.0" valueMin="11.0">11-1200 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation ID-GBIF-Occurrence="3063614312" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NHMD" specimenCount="1" stateProvince="Chiang Mai">
|
||
<specimenCount type="generic">1 juv.</specimenCount>
|
||
(16 pairs of legs);
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>NHMD</collectionCode>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3063614301" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NHMD" collectorName="No. Kasens & Birgit Degerbol & Zool. Mus. Kbh." location="Doi Suthep" specimenCount="1" stateProvince="Chiang Mai">
|
||
<location LSID="urn:lsid:plazi:treatment:2AB443F1DFF45118AE5260106B9D2694:9090505B73E16A9C11CF39512CDD6CBC" name="Doi Suthep" stateProvince="Chiang Mai">Doi Suthep</location>
|
||
,
|
||
<collectorName>
|
||
<emphasis bold="true" pageId="0" pageNumber="137">No.</emphasis>
|
||
Kasens
|
||
</collectorName>
|
||
DSal; 14/9.1959,
|
||
<collectorName>
|
||
Birgit
|
||
<normalizedToken originalValue="Degerbøl">Degerbol</normalizedToken>
|
||
</collectorName>
|
||
leg.,
|
||
<collectorName>Zool. Mus. Kbh.</collectorName>
|
||
1/7 59. 1 mature F (with nemertini parasite);
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>NHMD</collectionCode>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3063614307" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Birgit Degerbol & Zool. Mus. Kbh." location="No." municipality="Doi Suthep" specimenCount="1" stateProvince="Chiang Mai">
|
||
<collectingMunicipality>Doi Suthep</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:2AB443F1DFF45118AE5260106B9D2694:2F9C1489A20E7ED64EFBD2FEFCBE131E" municipality="Doi Suthep" name="No." stateProvince="Chiang Mai">No.</location>
|
||
<normalizedToken originalValue="sigteprøve">sigteprove</normalizedToken>
|
||
(= sifting sample) 3a?; 2/8.1958,
|
||
<collectorName>
|
||
Birgit
|
||
<normalizedToken originalValue="Degerbøl">Degerbol</normalizedToken>
|
||
</collectorName>
|
||
leg.,
|
||
<collectorName>Zool. Mus. Kbh.</collectorName>
|
||
1/7 59
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation ID-GBIF-Occurrence="3063614306" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NHMD" specimenCount="1" stateProvince="Chiang Mai">
|
||
<specimenCount type="generic">1 juv.</specimenCount>
|
||
(8 pairs of legs);
|
||
<emphasis bold="true" pageId="0" pageNumber="137">
|
||
<collectionCode>NHMD</collectionCode>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3063614313" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="No. Lok. & Birgit Degerbol & Zool. Mus. Kbh." elevation="1150" location="Doi Suthep" specimenCount="1" stateProvince="Chiang Mai">
|
||
<location LSID="urn:lsid:plazi:treatment:2AB443F1DFF45118AE5260106B9D2694:23543BA1388F6960C19394E76560386D" name="Doi Suthep" stateProvince="Chiang Mai">Doi Suthep</location>
|
||
,
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">1150 m</elevation>
|
||
</quantity>
|
||
;
|
||
<collectorName>
|
||
<emphasis bold="true" pageId="0" pageNumber="137">No.</emphasis>
|
||
Lok.
|
||
</collectorName>
|
||
3a; 29/9.1958;
|
||
<collectorName>
|
||
Birgit
|
||
<normalizedToken originalValue="Degerbøl">Degerbol</normalizedToken>
|
||
</collectorName>
|
||
leg.,
|
||
<collectorName>Zool. Mus. Kbh.</collectorName>
|
||
1/7 59
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |