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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.907.36563" ID-GBIF-Dataset="69422977-aa4d-4fe3-a450-8ef54d6fa905" ID-PMC="PMC7002455" ID-Pensoft-Pub="1313-2970-907-1" ID-Pensoft-UUID="9920267E73CF5E00B644DED1F101D965" ID-PubMed="32063727" ID-ZooBank="DBF339E5EBBC461994388359C769473F" ModsDocID="1313-2970-907-1" checkinTime="1580398733216" checkinUser="pensoft" docAuthor="Leger, Theo, Kehlmaier, Christian, Vairappan, Charles S. &amp; Nuss, Matthias" docDate="2020" docId="006C0C0DB5CB54BBAC61F308D63360E5" docLanguage="en" docName="ZooKeys 907: 1-99" docOrigin="ZooKeys 907" docSource="http://dx.doi.org/10.3897/zookeys.907.36563" docTitle="Hoploscopa jubata Leger &amp; Nuss 2020, sp. nov." docType="treatment" docUuid="F81F0837-5324-4B81-AC72-B0519F850301" docUuidSource="ZooBank" docVersion="4" id="9920267E73CF5E00B644DED1F101D965" lastPageNumber="1" masterDocId="9920267E73CF5E00B644DED1F101D965" masterDocTitle="Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding" masterLastPageNumber="99" masterPageNumber="1" pageNumber="1" updateTime="1668168367651" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Leger, Theo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kehlmaier, Christian</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vairappan, Charles S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nuss, Matthias</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>907</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>99</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.907.36563</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.907.36563</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-907-1</mods:identifier>
<mods:identifier type="ZooBank">DBF339E5EBBC461994388359C769473F</mods:identifier>
<mods:identifier type="Pensoft-UUID">9920267E73CF5E00B644DED1F101D965</mods:identifier>
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<treatment ID-GBIF-Taxon="161926076" LSID="urn:lsid:zoobank.org:act:F81F0837-5324-4B81-AC72-B0519F850301" httpUri="http://treatment.plazi.org/id/006C0C0DB5CB54BBAC61F308D63360E5" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="006C0C0D-B5CB-54BB-AC61-F308D63360E5" authority="Léger &amp; Nuss" authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa jubata" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jubata" status="sp. nov.">
Hoploscopa jubata
<normalizedToken originalValue="Léger">Leger</normalizedToken>
&amp; Nuss
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Habitus, venation and forewing pattern in Hoploscopa. 1 Habitus of Hoploscopa matheae sp. nov., holotype, ♀, Malaysia, Sabah, Kundasang, Kinabalu Mt. Lodge veranda, 6 ° 0 ' 42.15 &quot; N, 116 ° 32 ' 3.63 &quot; E, 1570 m (T. Leger &amp; R. Mally) (genitalia on slide TL 315 ♀) 2 wing venation of Hoploscopa species: H. jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Prov., nr Bulolo, Mt Susu Nat. Res., Araucaria For., 975 m, 27 - 28 Aug 1983 (S. Miller) (wing preparation TL 706) 3 nomenclature used for description of forewing pattern of Hoploscopa spp. Abbreviations: bp (basal patch), bs (basal streak), cf (costal field), dds (distal discoidal stigma), mcp (median cubital patch), mds (median discoidal stigma), mdp (median dorsal patch), pds (proximal discoidal stigma), pma (postmedian area), pml (postmedian line), pmp (postmedian patch), sbf (subterminal field), sbl (subterminal line)." figureDoi="10.3897/zookeys.907.36563.figures1-3" httpUri="https://binary.pensoft.net/fig/375756" pageId="0" pageNumber="1">Figs 2</figureCitation>
<figureCitation captionStart="Figures 34–45" captionStartId="F4" captionText="Figures 34 - 45. Forewing of Hoploscopa species. 34 Hoploscopa ignitamaculae sp. nov., paratype, ♂, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 30 - 31. iii. 2000 (A. Kallies &amp; C. Zorn) (genitalia on slide TL 509 ♂) 35 Hoploscopa albomaculata sp. nov., paratype, ♀, Indonesia, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 25 - 26. iii. 2000, (A. Kallies &amp; C. Zorn) (genitalia on slide TL 540 ♀) 36 Hoploscopa sumatrensis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 6 - 26. i. 1996 (A. Kallies) (genitalia on slide TL 755 ♂) 37 Hoploscopa mallyi sp. nov., paratype, ♂, Malaysia, Sabah, Mt Kinabalu, Mesilau, 2000 m, 14 - 17. xi. 2006 (W. Mey &amp; K. Ebert) 38 Hoploscopa gracilis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 754 ♂) 39 Hoploscopa agtuuganonensis sp. nov., holotype, ♀, Philippines, Mindanao, Mt Agtuuganon, 1050 m, 28. v- 7. vi. 1996 (W. Mey) (genitalia on slide TL 616 ♀) 40 Hoploscopa boleta sp. nov., paratype, ♀, Papua New Guinea, Morobe, Mt Kaindi, 2360 m, 3. x. 1992 (V. O. Becker) (genitalia on slide TL 448 ♀) 41 Hoploscopa pseudometacrossa sp. nov., holotype, ♀, Papua New Guinea, W. Hhl, Prv, near Mt. Hagen, Kuk Ag, Res. Sta., 1600 m, 19 - 20. viii. 1983 (Scott E. &amp; Pamela Miller) (genitalia on slide TL 689 ♀) 42 Hoploscopa metacrossa Hampson, ♀, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller). 43 Hoploscopa jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 44 Hoploscopa jubata sp. nov., paratype, ♀, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 45 Hindwing scent organ in Hoploscopa metacrossa. Paratype, ♂, Papua New Guinea, Morobe Province, Wau, 1200 m, 8 - 14. xii. 1976 (G. F. Hevel &amp; R. E. Dietz) (genitalia on slide TL 443 ♂)." figureDoi="10.3897/zookeys.907.36563.figures34-45" httpUri="https://binary.pensoft.net/fig/375759" pageId="0" pageNumber="1">, 43</figureCitation>
<figureCitation captionStart="Figures 34–45" captionStartId="F4" captionText="Figures 34 - 45. Forewing of Hoploscopa species. 34 Hoploscopa ignitamaculae sp. nov., paratype, ♂, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 30 - 31. iii. 2000 (A. Kallies &amp; C. Zorn) (genitalia on slide TL 509 ♂) 35 Hoploscopa albomaculata sp. nov., paratype, ♀, Indonesia, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 25 - 26. iii. 2000, (A. Kallies &amp; C. Zorn) (genitalia on slide TL 540 ♀) 36 Hoploscopa sumatrensis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 6 - 26. i. 1996 (A. Kallies) (genitalia on slide TL 755 ♂) 37 Hoploscopa mallyi sp. nov., paratype, ♂, Malaysia, Sabah, Mt Kinabalu, Mesilau, 2000 m, 14 - 17. xi. 2006 (W. Mey &amp; K. Ebert) 38 Hoploscopa gracilis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 754 ♂) 39 Hoploscopa agtuuganonensis sp. nov., holotype, ♀, Philippines, Mindanao, Mt Agtuuganon, 1050 m, 28. v- 7. vi. 1996 (W. Mey) (genitalia on slide TL 616 ♀) 40 Hoploscopa boleta sp. nov., paratype, ♀, Papua New Guinea, Morobe, Mt Kaindi, 2360 m, 3. x. 1992 (V. O. Becker) (genitalia on slide TL 448 ♀) 41 Hoploscopa pseudometacrossa sp. nov., holotype, ♀, Papua New Guinea, W. Hhl, Prv, near Mt. Hagen, Kuk Ag, Res. Sta., 1600 m, 19 - 20. viii. 1983 (Scott E. &amp; Pamela Miller) (genitalia on slide TL 689 ♀) 42 Hoploscopa metacrossa Hampson, ♀, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller). 43 Hoploscopa jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 44 Hoploscopa jubata sp. nov., paratype, ♀, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 45 Hindwing scent organ in Hoploscopa metacrossa. Paratype, ♂, Papua New Guinea, Morobe Province, Wau, 1200 m, 8 - 14. xii. 1976 (G. F. Hevel &amp; R. E. Dietz) (genitalia on slide TL 443 ♂)." figureDoi="10.3897/zookeys.907.36563.figures34-45" httpUri="https://binary.pensoft.net/fig/375759" pageId="0" pageNumber="1">, 44</figureCitation>
<figureCitation captionStart="Figures 82–86" captionStartId="F11" captionText="Figures 82 - 86. Male genitalia and abdomen features of Hoploscopa. 82 Hoploscopa jubata sp. nov., paratype, TL 542 ♂ 83 Tympanal organs of Hoploscopa sumatrensis sp. nov., paratype, TL 538 ♂ 84 Male tympanal organs of Hoploscopa metacrossa, TL 472 ♂ 85 Sternite VIII of Hoploscopa sp. near isarogensis, TL 626 ♂ 86 Sternite VIII of Hoploscopa metacrossa, TL 443 ♂. Abbreviations: tb (tympanic bridge), td (tympanic drum), tdp (tympanic depression), tm (tympanum), tr (transverse ridge)." figureDoi="10.3897/zookeys.907.36563.figures82-86" httpUri="https://binary.pensoft.net/fig/375766" pageId="0" pageNumber="1">, 82</figureCitation>
<figureCitation captionStart="Figures 119–122" captionStartId="F20" captionText="Figures 119 - 122. Female genitalia features of Hoploscopa. 119 Hoploscopa boleta sp. nov., paratype, TL 448 ♀ 120 Hoploscopa pseudometacrossa sp. nov., paratype, TL 435 ♀ 121 Hoploscopa metacrossa, paratype, TL 446 ♀ 122 Hoploscopa jubata sp. nov., paratype, TL 445 ♀." figureDoi="10.3897/zookeys.907.36563.figures119-122" httpUri="https://binary.pensoft.net/fig/375775" pageId="0" pageNumber="1">, 122</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: ♂, with labels: &quot;Papua New Guinea | Morobe Prov[ince]., n[ea]r Bulolo | Mt Susu Nat[ional]. Res[erve]., 975m | 27-28 Aug.1983, S. Miller | UV Lite, Araucaria For[est].&quot;; &quot; DNA voucher | Lepidoptera | MTD2016 | [vertically written:] no. 3164&quot;; &quot;TL444 | ♂&quot;. Deposited in USNM.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
s
</emphasis>
: 4 ♂, 24 ♀. Papua New Guinea. 4 ♂ (1 with genitalia on slide TL447 ♂, 1 with genitalia on slide TL542 ♂, 1 with wing preparation TL706), 21 ♀ (1 with DNA voucher MTD LEP3165 &amp; genitalia on slide TL445 ♂, 2 with genitalia on slide TL476 ♀ &amp; TL541 ♀), same data as holotype. 1 ♀, Morobe Province, Wau, Wau Ecological Institute, 12-24.vii.1983; 1 ♀, same data except 1-10.viii.1983; 1 ♀, Morobe Province, Wau, 1200 m, 8-14.xii.1976, at black light, leg. G. F. Hevel &amp; R. E. Dietz (USNM).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">This species shows a strong sexual dimorphism. In males, median and postmedian markings are white, and postmedian area is broadly suffused with white. In females, basal patch, median discoidal stigma and postmedian patch are reddish orange, and subterminal area is orange. Male hindwing displays an androconial organ on the dorsum. In male genitalia, gnathos projection is slender, ca. 1/3 of the uncus length, and juxta displays two conspicuous lateral projections at base, and a deeply indented apex. In female genitalia, the four loops formed by the ductus bursae is unique to this species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="similar species">
<paragraph pageId="0" pageNumber="1">Similar species.</paragraph>
<paragraph pageId="0" pageNumber="1">No similar species known.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Antennae dorsally brown. Proboscis brown. Maxillary palpi brown, base and inner side pale yellow. Labial palpi brown, ventral base and inner side pale yellow.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
(Figs
<figureCitation captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Habitus, venation and forewing pattern in Hoploscopa. 1 Habitus of Hoploscopa matheae sp. nov., holotype, ♀, Malaysia, Sabah, Kundasang, Kinabalu Mt. Lodge veranda, 6 ° 0 ' 42.15 &quot; N, 116 ° 32 ' 3.63 &quot; E, 1570 m (T. Leger &amp; R. Mally) (genitalia on slide TL 315 ♀) 2 wing venation of Hoploscopa species: H. jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Prov., nr Bulolo, Mt Susu Nat. Res., Araucaria For., 975 m, 27 - 28 Aug 1983 (S. Miller) (wing preparation TL 706) 3 nomenclature used for description of forewing pattern of Hoploscopa spp. Abbreviations: bp (basal patch), bs (basal streak), cf (costal field), dds (distal discoidal stigma), mcp (median cubital patch), mds (median discoidal stigma), mdp (median dorsal patch), pds (proximal discoidal stigma), pma (postmedian area), pml (postmedian line), pmp (postmedian patch), sbf (subterminal field), sbl (subterminal line)." figureDoi="10.3897/zookeys.907.36563.figures1-3" httpUri="https://binary.pensoft.net/fig/375756" pageId="0" pageNumber="1">2</figureCitation>
,
<figureCitation captionStart="Figures 34–45" captionStartId="F4" captionText="Figures 34 - 45. Forewing of Hoploscopa species. 34 Hoploscopa ignitamaculae sp. nov., paratype, ♂, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 30 - 31. iii. 2000 (A. Kallies &amp; C. Zorn) (genitalia on slide TL 509 ♂) 35 Hoploscopa albomaculata sp. nov., paratype, ♀, Indonesia, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 25 - 26. iii. 2000, (A. Kallies &amp; C. Zorn) (genitalia on slide TL 540 ♀) 36 Hoploscopa sumatrensis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 6 - 26. i. 1996 (A. Kallies) (genitalia on slide TL 755 ♂) 37 Hoploscopa mallyi sp. nov., paratype, ♂, Malaysia, Sabah, Mt Kinabalu, Mesilau, 2000 m, 14 - 17. xi. 2006 (W. Mey &amp; K. Ebert) 38 Hoploscopa gracilis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 754 ♂) 39 Hoploscopa agtuuganonensis sp. nov., holotype, ♀, Philippines, Mindanao, Mt Agtuuganon, 1050 m, 28. v- 7. vi. 1996 (W. Mey) (genitalia on slide TL 616 ♀) 40 Hoploscopa boleta sp. nov., paratype, ♀, Papua New Guinea, Morobe, Mt Kaindi, 2360 m, 3. x. 1992 (V. O. Becker) (genitalia on slide TL 448 ♀) 41 Hoploscopa pseudometacrossa sp. nov., holotype, ♀, Papua New Guinea, W. Hhl, Prv, near Mt. Hagen, Kuk Ag, Res. Sta., 1600 m, 19 - 20. viii. 1983 (Scott E. &amp; Pamela Miller) (genitalia on slide TL 689 ♀) 42 Hoploscopa metacrossa Hampson, ♀, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller). 43 Hoploscopa jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 44 Hoploscopa jubata sp. nov., paratype, ♀, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 45 Hindwing scent organ in Hoploscopa metacrossa. Paratype, ♂, Papua New Guinea, Morobe Province, Wau, 1200 m, 8 - 14. xii. 1976 (G. F. Hevel &amp; R. E. Dietz) (genitalia on slide TL 443 ♂)." figureDoi="10.3897/zookeys.907.36563.figures34-45" httpUri="https://binary.pensoft.net/fig/375759" pageId="0" pageNumber="1">43</figureCitation>
,
<figureCitation captionStart="Figures 34–45" captionStartId="F4" captionText="Figures 34 - 45. Forewing of Hoploscopa species. 34 Hoploscopa ignitamaculae sp. nov., paratype, ♂, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 30 - 31. iii. 2000 (A. Kallies &amp; C. Zorn) (genitalia on slide TL 509 ♂) 35 Hoploscopa albomaculata sp. nov., paratype, ♀, Indonesia, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 25 - 26. iii. 2000, (A. Kallies &amp; C. Zorn) (genitalia on slide TL 540 ♀) 36 Hoploscopa sumatrensis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 6 - 26. i. 1996 (A. Kallies) (genitalia on slide TL 755 ♂) 37 Hoploscopa mallyi sp. nov., paratype, ♂, Malaysia, Sabah, Mt Kinabalu, Mesilau, 2000 m, 14 - 17. xi. 2006 (W. Mey &amp; K. Ebert) 38 Hoploscopa gracilis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 754 ♂) 39 Hoploscopa agtuuganonensis sp. nov., holotype, ♀, Philippines, Mindanao, Mt Agtuuganon, 1050 m, 28. v- 7. vi. 1996 (W. Mey) (genitalia on slide TL 616 ♀) 40 Hoploscopa boleta sp. nov., paratype, ♀, Papua New Guinea, Morobe, Mt Kaindi, 2360 m, 3. x. 1992 (V. O. Becker) (genitalia on slide TL 448 ♀) 41 Hoploscopa pseudometacrossa sp. nov., holotype, ♀, Papua New Guinea, W. Hhl, Prv, near Mt. Hagen, Kuk Ag, Res. Sta., 1600 m, 19 - 20. viii. 1983 (Scott E. &amp; Pamela Miller) (genitalia on slide TL 689 ♀) 42 Hoploscopa metacrossa Hampson, ♀, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller). 43 Hoploscopa jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 44 Hoploscopa jubata sp. nov., paratype, ♀, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 45 Hindwing scent organ in Hoploscopa metacrossa. Paratype, ♂, Papua New Guinea, Morobe Province, Wau, 1200 m, 8 - 14. xii. 1976 (G. F. Hevel &amp; R. E. Dietz) (genitalia on slide TL 443 ♂)." figureDoi="10.3897/zookeys.907.36563.figures34-45" httpUri="https://binary.pensoft.net/fig/375759" pageId="0" pageNumber="1">44</figureCitation>
). Collar brown. Forewing length: 9-10 mm (♂ &amp; ♀); forewing ♂ ground colour brown; basal narrow white streak, distally marked with yellow and orange; median discoidal stigma white, filled with few red and yellow scales, together with cubital and dorsal patch forming a white Y; postmedian patch brown speckled with red, more or less marked, distally with costal white streak; postmedian line marked on costal half; postmedian area with broad white suffusion; subterminal line conspicuously incurved inward at CuA2, white; fringe chequered brown and pale yellow; forewing ♀ (Fig.
<figureCitation captionStart="Figures 34–45" captionStartId="F4" captionText="Figures 34 - 45. Forewing of Hoploscopa species. 34 Hoploscopa ignitamaculae sp. nov., paratype, ♂, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 30 - 31. iii. 2000 (A. Kallies &amp; C. Zorn) (genitalia on slide TL 509 ♂) 35 Hoploscopa albomaculata sp. nov., paratype, ♀, Indonesia, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 25 - 26. iii. 2000, (A. Kallies &amp; C. Zorn) (genitalia on slide TL 540 ♀) 36 Hoploscopa sumatrensis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 6 - 26. i. 1996 (A. Kallies) (genitalia on slide TL 755 ♂) 37 Hoploscopa mallyi sp. nov., paratype, ♂, Malaysia, Sabah, Mt Kinabalu, Mesilau, 2000 m, 14 - 17. xi. 2006 (W. Mey &amp; K. Ebert) 38 Hoploscopa gracilis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 754 ♂) 39 Hoploscopa agtuuganonensis sp. nov., holotype, ♀, Philippines, Mindanao, Mt Agtuuganon, 1050 m, 28. v- 7. vi. 1996 (W. Mey) (genitalia on slide TL 616 ♀) 40 Hoploscopa boleta sp. nov., paratype, ♀, Papua New Guinea, Morobe, Mt Kaindi, 2360 m, 3. x. 1992 (V. O. Becker) (genitalia on slide TL 448 ♀) 41 Hoploscopa pseudometacrossa sp. nov., holotype, ♀, Papua New Guinea, W. Hhl, Prv, near Mt. Hagen, Kuk Ag, Res. Sta., 1600 m, 19 - 20. viii. 1983 (Scott E. &amp; Pamela Miller) (genitalia on slide TL 689 ♀) 42 Hoploscopa metacrossa Hampson, ♀, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller). 43 Hoploscopa jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 44 Hoploscopa jubata sp. nov., paratype, ♀, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 45 Hindwing scent organ in Hoploscopa metacrossa. Paratype, ♂, Papua New Guinea, Morobe Province, Wau, 1200 m, 8 - 14. xii. 1976 (G. F. Hevel &amp; R. E. Dietz) (genitalia on slide TL 443 ♂)." figureDoi="10.3897/zookeys.907.36563.figures34-45" httpUri="https://binary.pensoft.net/fig/375759" pageId="0" pageNumber="1">44</figureCitation>
) ground colour brown; yellow basal patch crossed by orange streak; median trapezoid white patch, filled to a various extend with yellow and red scales, together with cubital and dorsal white patches forming a Y; postmedian patch roughly oval, reddish orange, basally edged with yellow, distally with costal white streak; postmedian line marked on costal half; postmedian area faintly speckled with white between postmedian patch and subterminal line; subterminal line white, incurved inwards at CuA2; subterminal field marked with reddish orange and yellow; fringe brown with pale yellow dots. Hindwing pale yellow; in males, presence of an androconial organ on the dorsum of the hindwing. Forelegs bronze. Midlegs with femur brown; tibia brown, distally pale yellow; tarsi bronze. Hindlegs with femur brown; tibia pale yellow, speckled with brown; tarsi bronze.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
.
</emphasis>
Male tergite I, II partially sclerotised; tergite III sclerotised, with two shallow depressions on each side of middle, bearing patches of modified scales. Sternum A8 posterior margin straight.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 3) (Fig.
<figureCitation captionStart="Figures 82–86" captionStartId="F11" captionText="Figures 82 - 86. Male genitalia and abdomen features of Hoploscopa. 82 Hoploscopa jubata sp. nov., paratype, TL 542 ♂ 83 Tympanal organs of Hoploscopa sumatrensis sp. nov., paratype, TL 538 ♂ 84 Male tympanal organs of Hoploscopa metacrossa, TL 472 ♂ 85 Sternite VIII of Hoploscopa sp. near isarogensis, TL 626 ♂ 86 Sternite VIII of Hoploscopa metacrossa, TL 443 ♂. Abbreviations: tb (tympanic bridge), td (tympanic drum), tdp (tympanic depression), tm (tympanum), tr (transverse ridge)." figureDoi="10.3897/zookeys.907.36563.figures82-86" httpUri="https://binary.pensoft.net/fig/375766" pageId="0" pageNumber="1">82</figureCitation>
). Uncus long and slender, basally narrow, medially slightly widened, narrowed at apical 1/4, apex spatula-shaped, ventrally with eight small ridges. Gnathos projection slender, ca. 1/3 of the uncus length. Valva gently bending dorsad from distal 1/3, dorsal margin slightly convex, apex roughly truncate, forming a furrow ring. Juxta mushroom-shaped, with base roughly rounded, slightly concave on its middle, with two conspicuous lateral extensions pointing outward, conspicuously narrowed at basal 1/4, apex bilobed. Saccus broad, triangular, pointing dorsad.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 3) (Fig.
<figureCitation captionStart="Figures 119–122" captionStartId="F20" captionText="Figures 119 - 122. Female genitalia features of Hoploscopa. 119 Hoploscopa boleta sp. nov., paratype, TL 448 ♀ 120 Hoploscopa pseudometacrossa sp. nov., paratype, TL 435 ♀ 121 Hoploscopa metacrossa, paratype, TL 446 ♀ 122 Hoploscopa jubata sp. nov., paratype, TL 445 ♀." figureDoi="10.3897/zookeys.907.36563.figures119-122" httpUri="https://binary.pensoft.net/fig/375775" pageId="0" pageNumber="1">122</figureCitation>
). Anterior apophyses with dorsal bump at posterior 1/3. Antrum sclerotisation short. Ductus bursae very long, forming four conspicuous loops bent before corpus opening. Corpus bursae small, globular, reticulated, medially with light broad sclerotised patch, with weak rounded sclerotisation at corpus opening. Thorn gently curved, with small dents pointing toward thorn base, basally with small outwardly projected extension.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Known from Wau and from Mount Susu in the Morobe Province (Papua New Guinea), at altitudes between 950 m and 1,200 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="phylogenetic relationships">
<paragraph pageId="0" pageNumber="1">Phylogenetic relationships.</paragraph>
<paragraph pageId="0" pageNumber="1">
See
<taxonomicName lsidName="H. boleta" pageId="0" pageNumber="1" rank="species" species="boleta">
<emphasis italics="true" pageId="0" pageNumber="1">H. boleta</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="H. pseudometacrossa" pageId="0" pageNumber="1" rank="species" species="pseudometacrossa">
<emphasis italics="true" pageId="0" pageNumber="1">H. pseudometacrossa</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
From the Latin
<emphasis italics="true" pageId="0" pageNumber="1">jubatus</emphasis>
, having a mane, referring to the dense hair covering on the inner side of the valva in male genitalia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">This species is the only one known from the genus to exhibit a pronounced sexual dimorphism.</paragraph>
<caption doi="10.3897/zookeys.907.36563.figures91-94" httpUri="https://binary.pensoft.net/fig/375768" pageId="0" pageNumber="1" start="Figures 91–94" startId="F13">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 91-94.</emphasis>
Female genitalia features of
<taxonomicName authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="1">91</emphasis>
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa parvimacula" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="parvimacula">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa parvimacula</emphasis>
</taxonomicName>
sp. nov., paratype, TL551 ♀
<emphasis bold="true" pageId="0" pageNumber="1">92</emphasis>
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa kinabaluensis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa kinabaluensis</emphasis>
</taxonomicName>
sp. nov., TL674 ♀
<emphasis bold="true" pageId="0" pageNumber="1">93</emphasis>
<taxonomicName authorityName="Nuss" authorityYear="1998" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa luteomacula" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="luteomacula">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa luteomacula</emphasis>
</taxonomicName>
, paratype, GU prep. Nuss 743 ♀
<emphasis bold="true" pageId="0" pageNumber="1">94</emphasis>
<taxonomicName baseAuthorityName="Rothschild" baseAuthorityYear="1915" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa obliqua" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="obliqua">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa obliqua</emphasis>
</taxonomicName>
, paratype, TL656 ♀.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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