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<mods:title id="890E74C6C33B288CB00B60D73594E5B0">A taxonomic review of the genus Ashibusa Matsumura, 1931 (Lepidoptera: Cosmopterigidae) from China</mods:title>
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<mods:namePart id="62C718C6C13F1688D1692F632C1751BF">Li, Houhun</mods:namePart>
<mods:affiliation id="B913D5DDD53A740185F520B3F92E45F0">College of Life Sciences, Nankai University, Tianjin 300071, China &amp; College of Life and Geographic Sciences, Kashi University, Kashi 844000, China; Xinjiang Key Laboratory of Biological Resources and Ecology of Pamirs Plateau, Kashi 844000, China</mods:affiliation>
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<subSubSection id="3FE765289D1EFFF8FF37F8BEFE598027" box="[151,493,1884,1911]" pageId="0" pageNumber="265" type="nomenclature">
<paragraph id="774236A39D1EFFF8FF37F8BEFE598027" blockId="0.[151,493,1884,1911]" box="[151,493,1884,1911]" pageId="0" pageNumber="265">
<heading id="2C0A81CF9D1EFFF8FF37F8BEFE598027" bold="true" box="[151,493,1884,1911]" fontSize="11" level="1" pageId="0" pageNumber="265" reason="1">
<taxonomicName id="B0FD4D209D1EFFF8FF37F8BEFE598027" ID-CoL="9233F" authority="Matsumura, 1931" authorityName="Matsumura" authorityYear="1931" box="[151,493,1884,1911]" class="Insecta" family="Cosmopterigidae" genus="Ashibusa" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="4589EAB19D1EFFF8FF37F8BEFE598027" bold="true" box="[151,493,1884,1911]" pageId="0" pageNumber="265">
<emphasis id="4589EAB19D1EFFF8FF37F8BEFEB38026" bold="true" box="[151,263,1884,1911]" italics="true" pageId="0" pageNumber="265">Ashibusa</emphasis>
Matsumura, 1931
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="3FE765289D1EFFF8FF37F843FE4080E8" box="[151,500,1953,1977]" pageId="0" pageNumber="265" type="reference_group">
<paragraph id="774236A39D1EFFF8FF37F843FE4080E8" blockId="0.[151,670,1953,2008]" box="[151,500,1953,1977]" pageId="0" pageNumber="265">
<taxonomicName id="B0FD4D209D1EFFF8FF37F843FE0480E8" ID-CoL="9233F" authority="Matsumura, 1931: 1087" authorityName="Matsumura" authorityPageNumber="1087" authorityYear="1931" box="[151,432,1953,1977]" class="Insecta" family="Cosmopterigidae" genus="Ashibusa" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="4589EAB19D1EFFF8FF37F843FF4080E8" box="[151,244,1953,1977]" italics="true" pageId="0" pageNumber="265">Ashibusa</emphasis>
Matsumura, 1931
</taxonomicName>
: 1087.
</paragraph>
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<subSubSection id="3FE765289D1EFFF9FF37F822FB2E8006" lastPageId="1" lastPageNumber="266" pageId="0" pageNumber="265" type="materials_examined">
<paragraph id="774236A39D1EFFF9FF37F822FE2E810A" blockId="0.[151,670,1953,2008]" lastBlockId="1.[151,1437,1420,2023]" lastPageId="1" lastPageNumber="266" pageId="0" pageNumber="265">
<materialsCitation id="C7953CFE9D1EFFF9FF37F822FE23810A" collectionCode="R" lastPageId="1" lastPageNumber="266" pageId="0" pageNumber="265" specimenCode="R4, R5" specimenCount="1" typeStatus="holotype">
<typeStatus id="A84688019D1EFFF8FF37F822FF7D8089" box="[151,201,1984,2008]" pageId="0" pageNumber="265">Type</typeStatus>
species:
<taxonomicName id="B0FD4D209D1EFFF9FE86F822FF4282F7" authority="Matsumura, 1931 Generic" authorityName="Matsumura" authorityYear="1931" class="Insecta" family="Cosmopterigidae" genus="Ashibusa" kingdom="Animalia" lastPageId="1" lastPageNumber="266" order="Lepidoptera" pageId="0" pageNumber="265" phylum="Arthropoda" rank="species" species="jezoensis">
<emphasis id="4589EAB19D1EFFF8FE86F822FE518086" box="[294,485,1984,2008]" italics="true" pageId="0" pageNumber="265">Ashibusa jezoensis</emphasis>
Matsumura, 1931
<emphasis id="4589EAB19D1FFFF9FF37FA6EFF4282F7" bold="true" box="[151,246,1420,1446]" pageId="1" pageNumber="266">Generic</emphasis>
</taxonomicName>
<emphasis id="4589EAB19D1FFFF9FF5FFA6FFE3082F6" bold="true" box="[255,388,1421,1447]" pageId="1" pageNumber="266">characters.</emphasis>
Head with scales appressed, frons protruding forward (
<figureCitation id="EFC62A269D1FFFF9FC5FFA6FFBD282F6" box="[1023,1126,1421,1447]" captionStart="FIGURES 13" captionStartId="1.[151,264,1289,1313]" captionTargetBox="[151,1436,181,1260]" captionTargetId="figure-17@1.[151,1436,181,1265]" captionTargetPageId="1" captionText="FIGURES 13. Head of Ashibusa sinensis. 1. From front view; 2. From lateral view; 3. Wing venation of A. sinensis, slide No. ZD21430. Scales 12 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10224517" httpUri="https://zenodo.org/record/10224517/files/figure.png" pageId="1" pageNumber="266">Figs 12</figureCitation>
). Antennae longer than 3/4 length of forewing, scape with pectens in basal half. Labial palpi slender, third palpomere slightly longer than second palpomere. Forewing ground color yellow to brown in general, distally with a short white streak from costal margin to termen;
<specimenCode id="275B9ED89D1FFFF9FEC8FA1BFE368148" box="[360,386,1529,1561]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="266">
R
<subScript id="EB7934E69D1FFFF9FEDAF9E9FE368148" attach="left" box="[378,386,1547,1561]" fontSize="6" pageId="1" pageNumber="266">4</subScript>
</specimenCode>
,
<specimenCode id="275B9ED89D1FFFF9FE31FA1BFE1F8148" box="[401,427,1529,1561]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="266">
R
<subScript id="EB7934E69D1FFFF9FE03F9E9FE1F8148" attach="left" box="[419,427,1547,1561]" fontSize="6" pageId="1" pageNumber="266">5</subScript>
</specimenCode>
and M
<subScript id="EB7934E69D1FFFF9FE5CF9E9FDB08148" attach="left" box="[508,516,1547,1561]" fontSize="6" pageId="1" pageNumber="266">1</subScript>
stalked, CuA
<subScript id="EB7934E69D1FFFF9FD3BF9E9FD178148" attach="left" box="[667,675,1547,1561]" fontSize="6" pageId="1" pageNumber="266">2</subScript>
prolonged into a strong thorn near tornus (
<figureCitation id="EFC62A269D1FFFF9FB27FA1BFB788142" box="[1159,1228,1529,1555]" captionStart="FIGURES 13" captionStartId="1.[151,264,1289,1313]" captionTargetBox="[151,1436,181,1260]" captionTargetId="figure-17@1.[151,1436,181,1265]" captionTargetPageId="1" captionText="FIGURES 13. Head of Ashibusa sinensis. 1. From front view; 2. From lateral view; 3. Wing venation of A. sinensis, slide No. ZD21430. Scales 12 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10224517" httpUri="https://zenodo.org/record/10224517/files/figure.png" pageId="1" pageNumber="266">Fig. 3</figureCitation>
). Metathorax with a pair of bundles of setae from its anterolateral corner reaching third abdominal segment in male. Tibiae of hindlegs with whorls of bristles
</materialsCitation>
.
</paragraph>
<caption id="2382662B9D1FFFF9FF37FAEBFE568214" ID-DOI="http://doi.org/10.5281/zenodo.10224517" ID-Zenodo-Dep="10224517" httpUri="https://zenodo.org/record/10224517/files/figure.png" pageId="1" pageNumber="266" startId="1.[151,264,1289,1313]" targetBox="[151,1436,181,1260]" targetPageId="1" targetType="figure">
<paragraph id="774236A39D1FFFF9FF37FAEBFE568214" blockId="1.[151,1437,1289,1349]" pageId="1" pageNumber="266">
<emphasis id="4589EAB19D1FFFF9FF37FAEBFE8E8270" bold="true" box="[151,314,1289,1313]" pageId="1" pageNumber="266">FIGURES 13.</emphasis>
Head of
<emphasis id="4589EAB19D1FFFF9FE37FAEBFDE58270" box="[407,593,1289,1313]" italics="true" pageId="1" pageNumber="266">
<taxonomicName id="B0FD4D209D1FFFF9FE37FAEBFDF88270" box="[407,588,1289,1313]" class="Insecta" family="Cosmopterigidae" genus="Ashibusa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="266" phylum="Arthropoda" rank="species" species="sinensis">Ashibusa sinensis</taxonomicName>
.
</emphasis>
1. From front view; 2. From lateral view; 3. Wing venation of
<taxonomicName id="B0FD4D209D1FFFF9FB69FAEBFA818270" box="[1225,1333,1289,1313]" class="Insecta" family="Cosmopterigidae" genus="Ashibusa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="266" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="4589EAB19D1FFFF9FB69FAEBFA818270" box="[1225,1333,1289,1313]" italics="true" pageId="1" pageNumber="266">A. sinensis</emphasis>
</taxonomicName>
, slide No. ZD21430. Scales 12 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="774236A39D1FFFF9FF67F987FD1981BA" blockId="1.[151,1437,1420,2023]" pageId="1" pageNumber="266">Male abdomen and genitalia. Pleural lobes of eighth segment subelliptical in general, with a trifurcate sclerite at base. Left brachium of socius shorter than right brachium, right brachium often with a dorsal process. Left valvella long and slender, right valvella reduced or very short. Valvae narrow basally, widened distally, with dense setae on ventral surface. Phallus stout, without cornutus.</paragraph>
<paragraph id="774236A39D1FFFF9FF67F917FB2E8006" blockId="1.[151,1437,1420,2023]" pageId="1" pageNumber="266">Female genitalia. Papillae anales membranous, obtuse apically. Apophyses posteriores longer than apophyses anteriores. Ductus bursae extremely slender. Corpus bursae longer than wide; ductus seminalis arising from posterior end of corpus bursae; signum absent, or being one or two rows of small denticles if present.</paragraph>
</subSubSection>
<subSubSection id="3FE765289D1FFFF9FF67F882FF6680B6" pageId="1" pageNumber="266" type="diagnosis">
<paragraph id="774236A39D1FFFF9FF67F882FF6680B6" blockId="1.[151,1437,1420,2023]" pageId="1" pageNumber="266">
<emphasis id="4589EAB19D1FFFF9FF67F882FEF6802B" bold="true" box="[199,322,1888,1914]" pageId="1" pageNumber="266">Diagnosis.</emphasis>
Species of
<taxonomicName id="B0FD4D209D1FFFF9FE69F883FD84802A" authorityName="Matsumura, 1931:" authorityYear="1087" box="[457,560,1889,1915]" class="Insecta" family="Cosmopterigidae" genus="Ashibusa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis id="4589EAB19D1FFFF9FE69F883FD84802A" box="[457,560,1889,1915]" italics="true" pageId="1" pageNumber="266">Ashibusa</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="B0FD4D209D1FFFF9FC9DF883FC47802A" authorityName="Meyrick" authorityYear="1913" box="[829,1011,1889,1915]" class="Insecta" family="Stathmopodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="266" phylum="Arthropoda" rank="subFamily" subFamily="Stathmopodinae">Stathmopodinae</taxonomicName>
in
<taxonomicName id="B0FD4D209D1FFFF9FBBBF883FB65802A" box="[1051,1233,1889,1915]" class="Insecta" family="Stathmopodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="266" phylum="Arthropoda" rank="family">Stathmopodidae</taxonomicName>
in the hind tibiae with whorls of bristles and the adults erecting hindlegs at rest. Species of
<taxonomicName id="B0FD4D209D1FFFF9FC1FF867FB9280CE" authorityName="Matsumura, 1931:" authorityYear="1087" box="[959,1062,1925,1951]" class="Insecta" family="Cosmopterigidae" genus="Ashibusa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis id="4589EAB19D1FFFF9FC1FF867FB9280CE" box="[959,1062,1925,1951]" italics="true" pageId="1" pageNumber="266">Ashibusa</emphasis>
</taxonomicName>
can be distinguished by the shape of the head, the labial palpi and the forewings, and the strongly asymmetrical male genitalia (
<bibRefCitation id="136C4B529D1FFFF9FB64F84BFF7180B6" author="Zhang, Z. W. &amp; Li, H. H." pageId="1" pageNumber="266" pagination="335 - 343" refId="ref11131" refString="Lepidoptera. Pt. 2. Dal'nauka, Vladivostok, pp. 257 - 286. Zhang, Z. W. &amp; Li, H. H. (2009) Taxonomic study of the genus Ashibusa Matsumura (Lepidoptera, Cosmopterigidae), with description of six new species in China. Deutsche Entomologische Zeitschrift, 56 (2), 335 - 343. https: // doi. org / 10.1002 / mmnd. 200900031" type="journal article" year="2009">Zhang &amp; Li, 2009: 335</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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