treatments-xml/data/FD/74/87/FD7487B1FFBCFFF5FF205F8838E1FA56.xml

<document id="8A472286F7C56FE2A0ADD7104139C8EE" ID-CLB-Dataset="56235" ID-DOI="10.11646/zootaxa.5262.1.1" ID-GBIF-Dataset="e24a8032-296e-442d-80ae-e0bf8c7237f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7795161" ID-ZooBank="F05805FD-BC5B-4236-BF04-C6AA5C37F2D7" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680528957709" checkinUser="plazi" docAuthor="Hou, Yanmeng, Feng, Zegang &amp; Zhang, Feng" docDate="2023" docId="FD7487B1FFBCFFF5FF205F8838E1FA56" docLanguage="en" docName="zootaxa.5262.1.1.pdf" docOrigin="Zootaxa 5262 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5262.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Tyrannochthonius duo Hou &amp; Feng &amp; Zhang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="48" masterDocId="014DFFC9FF96FFDAFFB75F103A10FFB3" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="158" masterPageNumber="1" pageNumber="43" updateTime="1701997099389" updateUser="plazi">
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<mods:title id="61608BF4AC4E9C4C2CBEE4842DE43F50">Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="357668702A306BD3E03DD539FD6825BC">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="8F25EAEDD272A546520DD715CC8A33D2">Feng, Zegang</mods:namePart>
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<mods:namePart id="174B9A0C323798C899BB4D2C8E35F45E">Zhang, Feng</mods:namePart>
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<paragraph id="756236A7FFBCFFF0FF205F8838F6FF03" blockId="42.[151,742,152,214]" box="[151,742,152,179]" pageId="42" pageNumber="43">
<heading id="2E2A81CBFFBCFFF0FF205F8838F6FF03" bold="true" box="[151,742,152,179]" fontSize="11" level="2" pageId="42" pageNumber="43" reason="4">
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<taxonomicName id="B2DD4D24FFBCFFF0FF205F883BBCFF00" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[151,428,152,179]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="duo" status="sp. nov.">
<emphasis id="47A9EAB5FFBCFFF0FF205F883BBCFF00" bold="true" box="[151,428,152,179]" italics="true" pageId="42" pageNumber="43">Tyrannochthonius duo</emphasis>
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<taxonomicNameLabel id="5C9A57CEFFBCFFF0FE045F883801FF01" box="[435,529,152,178]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
‹Ξ⁂Ɵṁae›
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<paragraph id="756236A7FFBCFFF0FF205FAC3B04FF65" blockId="42.[151,742,152,214]" box="[151,276,188,214]" pageId="42" pageNumber="43">
<figureCitation id="EDE62A22FFBCFFF0FF205FAC3B04FF65" box="[151,276,188,214]" captionStart-0="FIGURE 32" captionStart-1="FIGURE 33" captionStart-2="FIGURE 34" captionStart-3="FIGURE 35" captionStart-4="FIGURE 36" captionStartId-0="42.[151,250,1762,1786]" captionStartId-1="43.[151,250,1295,1319]" captionStartId-2="44.[151,250,1943,1967]" captionStartId-3="45.[151,250,1943,1967]" captionStartId-4="46.[151,250,1943,1967]" captionTargetBox-0="[160,1428,706,1738]" captionTargetBox-1="[167,1422,424,1259]" captionTargetBox-2="[179,1408,186,1920]" captionTargetBox-3="[179,1396,181,1905]" captionTargetBox-4="[179,1408,181,1919]" captionTargetId-0="figure-196@42.[160,1428,706,1738]" captionTargetId-1="figure-141@43.[151,1436,417,1271]" captionTargetId-2="figure-20@44.[179,1408,181,1920]" captionTargetId-3="figure-18@45.[179,1408,181,1919]" captionTargetId-4="figure-20@46.[179,1408,181,1919]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionTargetPageId-2="44" captionTargetPageId-3="45" captionTargetPageId-4="46" captionText-0="FIGURE 32. Liangfeng Cave, type locality of Tyrannochthonius duo sp. nov., A. Entrance; B. Inside the cave entrance; C. Area where T. duo sp. nov. specimens were collected; D. Live male of T. duo sp. nov. in its natural environment." captionText-1="FIGURE 33. Tyrannochthonius duo sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." captionText-2="FIGURE 34. Tyrannochthonius duo sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." captionText-3="FIGURE 35. Tyrannochthonius duo sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." captionText-4="FIGURE 36. Tyrannochthonius duo sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7795234" figureDoi-1="http://doi.org/10.5281/zenodo.7795236" figureDoi-2="http://doi.org/10.5281/zenodo.7795238" figureDoi-3="http://doi.org/10.5281/zenodo.7795240" figureDoi-4="http://doi.org/10.5281/zenodo.7795242" httpUri-0="https://zenodo.org/record/7795234/files/figure.png" httpUri-1="https://zenodo.org/record/7795236/files/figure.png" httpUri-2="https://zenodo.org/record/7795238/files/figure.png" httpUri-3="https://zenodo.org/record/7795240/files/figure.png" httpUri-4="https://zenodo.org/record/7795242/files/figure.png" pageId="42" pageNumber="43">Figs 32–36</figureCitation>
</paragraph>
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<subSubSection id="3DC7652CFFBCFFF0FF205E143BAFFE1D" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="756236A7FFBCFFF0FF205E143BAFFE1D" blockId="42.[151,1437,260,683]" pageId="42" pageNumber="43">
<emphasis id="47A9EAB5FFBCFFF0FF205E143B58FEAD" bold="true" box="[151,328,260,286]" pageId="42" pageNumber="43">Type material.</emphasis>
<materialsCitation id="C5B53CFAFFBCFFF0FEE65E143B68FE39" ID-GBIF-Occurrence="4071438334" collectingDate="2022-07-04" collectionCode="MHBU-HBUARA" collectorName="Yanmeng Hou &amp; Nana Zhan &amp; Long Lin &amp; Jianzhou Sun" country="China" county="Jiangkou County" elevation="408" latitude="27.827179" location="Mengjiatun Village" longLatPrecision="1" longitude="108.86204" municipality="Nuxi Town" pageId="42" pageNumber="43" specimenCode="MHBU-HBUARA#2022-305, 01" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
<typeStatus id="AA668805FFBCFFF0FEE65E143BA9FEAD" box="[337,441,260,286]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
♂ (Ps.- 
<specimenCode id="257B9EDCFFBCFFF0FDA75E14397CFEAC" box="[528,876,260,287]" collectionCode="MHBU-HBUARA" pageId="42" pageNumber="43">MHBU-HBUARA#2022-305</specimenCode>
- 
<specimenCode id="257B9EDCFFBCFFF0FCC35E14399EFEAC" box="[884,910,260,287]" collectionCode="MHBU-HBUARA" pageId="42" pageNumber="43">01</specimenCode>
): 
<collectingCountry id="0DCA7637FFBCFFF0FC145E1439F9FEAD" box="[931,1001,260,286]" name="China" pageId="42" pageNumber="43">China</collectingCountry>
, 
<collectingRegion id="B719F845FFBCFFF0FC405E143ED7FEAD" box="[1015,1223,260,286]" country="China" name="Guizhou" pageId="42" pageNumber="43">Guizhou Province</collectingRegion>
, 
<collectingCounty id="9C034E2BFFBCFFF0FB615E143F89FEAD" box="[1238,1433,260,286]" pageId="42" pageNumber="43">Jiangkou County</collectingCounty>
, 
<collectingMunicipality id="9506ACDDFFBCFFF0FF205E383B09FEF0" box="[151,281,296,323]" pageId="42" pageNumber="43">Nuxi Town</collectingMunicipality>
, 
<location id="7002607CFFBCFFF0FE925E383812FEF1" LSID="urn:lsid:plazi:treatment:FD7487B1FFBCFFF5FF205F8838E1FA56:7002607CFFBCFFF0FE925E383812FEF1" box="[293,514,296,322]" country="China" county="Jiangkou County" latitude="27.827179" longLatPrecision="1" longitude="108.86204" municipality="Nuxi Town" name="Mengjiatun Village" pageId="42" pageNumber="43" stateProvince="Guizhou">Mengjiatun Village</location>
, 
<location id="7002607CFFBCFFF0FDBA5E3838D5FEF1" LSID="urn:lsid:plazi:treatment:FD7487B1FFBCFFF5FF205F8838E1FA56:7002607CFFBCFFF0FDBA5E3838D5FEF1" box="[525,709,296,322]" country="China" county="Jiangkou County" latitude="27.827179" longLatPrecision="1" longitude="108.86204" municipality="Nuxi Town" name="Liangfeng Cave" pageId="42" pageNumber="43" stateProvince="Guizhou">Liangfeng Cave</location>
, under stones and detritus in the deep zone (Temperature: 13°C, Humidity: 80%) [
<geoCoordinate id="10E95060FFBCFFF0FEEC5E5E3818FEDA" box="[347,520,334,361]" degrees="27" direction="north" minutes="49" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="37.84" value="27.827179">27°49′37.84″N</geoCoordinate>
, 
<geoCoordinate id="10E95060FFBCFFF0FDA35E5E38DCFEDA" box="[532,716,334,361]" degrees="108" direction="east" minutes="51" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="43.34" value="108.86204">108°51′43.34″E</geoCoordinate>
], 
<elevation id="FEF0D194FFBCFFF0FD695E5C3971FED5" box="[734,865,332,359]" metricMagnitude="2" metricUnit="m" metricValue="4.08" pageId="42" pageNumber="43" unit="m" value="408.0">
<quantity id="B2259B42FFBCFFF0FD695E5C3935FED4" box="[734,805,332,359]" metricMagnitude="2" metricUnit="m" metricValue="4.08" pageId="42" pageNumber="43" unit="m" value="408.0">408 m</quantity>
a.s.l.
</elevation>
, 
<date id="01631067FFBCFFF0FCDA5E5C39E2FED5" box="[877,1010,332,358]" pageId="42" pageNumber="43" value="2022-07-04">
<collectingDate id="1127E98FFFBCFFF0FCDA5E5C39E2FED5" box="[877,1010,332,358]" pageId="42" pageNumber="43" value="2022-07-04">4 July 2022</collectingDate>
</date>
, 
<collectorName id="D8285371FFBCFFF0FC4B5E5D3E8FFED4" box="[1020,1183,333,359]" pageId="42" pageNumber="43">Yanmeng Hou</collectorName>
, 
<collectorName id="D8285371FFBCFFF0FB1D5E5D3F38FED5" box="[1194,1320,332,359]" pageId="42" pageNumber="43">Nana Zhan</collectorName>
, 
<collectorName id="D8285371FFBCFFF0FA845E5D3F8CFED5" box="[1331,1436,332,359]" pageId="42" pageNumber="43">Long Lin</collectorName>
&amp; 
<collectorName id="D8285371FFBCFFF0FF045E603B59FE39" box="[179,329,368,394]" pageId="42" pageNumber="43">Jianzhou Sun</collectorName>
leg.
</materialsCitation>
<materialsCitation id="C5B53CFAFFBCFFF0FEC95E613BABFE1D" ID-GBIF-Occurrence="4071438306" collectingDate="2022-07-04" collectionCode="MSWU-HBUARA" collectorName="Yanmeng Hou &amp; Nana Zhan &amp; Long Lin &amp; Jianzhou Sun" country="China" county="Jiangkou County" elevation="408" latitude="27.827179" location="Mengjiatun Village" longLatPrecision="1" longitude="108.86204" municipality="Nuxi Town" pageId="42" pageNumber="43" specimenCode="MSWU-HBUARA#2022-305, 02" specimenCount="2" specimenCount-female="2" stateProvince="Guizhou" typeStatus="paratype">
<typeStatus id="AA668805FFBCFFF0FEC95E613BFFFE38" box="[382,495,369,395]" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="63DBFD2EFFBCFFF0FE4E5E603832FE38" box="[505,546,368,395]" pageId="42" pageNumber="43" type="female">2 ♀</specimenCount>
(Ps.- 
<specimenCode id="257B9EDCFFBCFFF0FDE25E6039BEFE38" box="[597,942,368,395]" collectionCode="MSWU-HBUARA" pageId="42" pageNumber="43">MSWU-HBUARA#2022-305</specimenCode>
- 
<specimenCode id="257B9EDCFFBCFFF0FC015E6039C2FE38" box="[950,978,368,395]" collectionCode="MSWU-HBUARA" pageId="42" pageNumber="43">02</specimenCode>
&amp; HBUARA#2022-305-03), all with the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFBCFFF0FF705EA8387CFE44" pageId="42" pageNumber="43" type="etymology">
<paragraph id="756236A7FFBCFFF0FF705EA8387CFE44" blockId="42.[151,1437,260,683]" pageId="42" pageNumber="43">
<emphasis id="47A9EAB5FFBCFFF0FF705EA83B5EFE61" bold="true" box="[199,334,440,466]" pageId="42" pageNumber="43">Etymology.</emphasis>
The specific name is derived from the Latin word “ 
<taxonomicName id="B2DD4D24FFBCFFF0FC2B5EA839D6FE61" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[924,966,440,466]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="duo">
<emphasis id="47A9EAB5FFBCFFF0FC2B5EA839D6FE61" box="[924,966,440,466]" italics="true" pageId="42" pageNumber="43">duo</emphasis>
</taxonomicName>
”, meaning dual, which refers to the presence of two well-developed anterior eyes.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFBCFFF0FF705D103F0CFD19" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="756236A7FFBCFFF0FF705D103F0CFD19" blockId="42.[151,1437,260,683]" pageId="42" pageNumber="43">
<emphasis id="47A9EAB5FFBCFFF0FF705D103B95FDAD" bold="true" box="[199,389,512,542]" pageId="42" pageNumber="43">
Diagnosis (J 
<specimenCount id="63DBFD2EFFBCFFF0FEE95D123B64FDAD" box="[350,372,514,542]" pageId="42" pageNumber="43" type="female">♀</specimenCount>
).
</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace with two anterior corneate eyes, posterior eyes reduced into eyespots; anterior margin of carapace thin, finely denticulated, epistome small, pointed, triangular; posterior margin of carapace with 2 setae; tergites I–II each with 4 setae. Pedipalps slender, femur 5.85 (♂), 5.79–6.08 (
<specimenCount id="63DBFD2EFFBCFFF0FD815D7C3858FD35" box="[566,584,620,646]" pageId="42" pageNumber="43" type="female">♀</specimenCount>
) times longer than broad; chela 6.94 (♂), 5.85–6.56 (
<specimenCount id="63DBFD2EFFBCFFF0FB065D7C3ED3FD35" box="[1201,1219,620,646]" pageId="42" pageNumber="43" type="female">♀</specimenCount>
) times longer than broad; both chelal fingers with intercalary teeth; chemosensory setae present on dorsum of chelal hand.
</paragraph>
</subSubSection>
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<paragraph id="756236A7FFBCFFF0FF2059F23EC8F8AD" blockId="42.[151,1436,1762,1822]" pageId="42" pageNumber="43">
<emphasis id="47A9EAB5FFBCFFF0FF2059F23B0FF949" bold="true" box="[151,287,1762,1786]" pageId="42" pageNumber="43">FIGURE 32.</emphasis>
Liangfeng Cave, type locality of 
<taxonomicName id="B2DD4D24FFBCFFF0FDD859F23941F949" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[623,849,1762,1786]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="duo" status="sp. nov.">
<emphasis id="47A9EAB5FFBCFFF0FDD859F23941F949" box="[623,849,1762,1786]" italics="true" pageId="42" pageNumber="43">Tyrannochthonius duo</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFBCFFF0FCE159F239B6F949" bold="true" box="[854,934,1762,1786]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="5C9A57CEFFBCFFF0FCE159F239B6F949" box="[854,934,1762,1786]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Entrance; B. Inside the cave entrance; C. Area where 
<taxonomicName id="B2DD4D24FFBCFFF0FF6C58173B0AF8AD" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[219,282,1798,1822]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="duo" status="sp. nov.">
<emphasis id="47A9EAB5FFBCFFF0FF6C58173B0AF8AD" box="[219,282,1798,1822]" italics="true" pageId="42" pageNumber="43">T. duo</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFBCFFF0FE9658163B61F8AD" bold="true" box="[289,369,1798,1822]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="5C9A57CEFFBCFFF0FE9658163B61F8AD" box="[289,369,1798,1822]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected; D. Live male of 
<taxonomicName id="B2DD4D24FFBCFFF0FC985817397EF8AD" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[815,878,1798,1822]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="duo" status="sp. nov.">
<emphasis id="47A9EAB5FFBCFFF0FC985817397EF8AD" box="[815,878,1798,1822]" italics="true" pageId="42" pageNumber="43">T. duo</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFBCFFF0FCC3581639D5F8AD" bold="true" box="[884,965,1798,1822]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="5C9A57CEFFBCFFF0FCC3581639D5F8AD" box="[884,965,1798,1822]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in its natural environment.
</paragraph>
</caption>
<subSubSection id="3DC7652CFFBCFFF5FF70585C3EE4FAE6" lastPageId="47" lastPageNumber="48" pageId="42" pageNumber="43" type="description">
<paragraph id="756236A7FFBCFFF0FF70585C3999F8D5" blockId="42.[151,1436,1868,2039]" box="[199,905,1868,1895]" pageId="42" pageNumber="43">
<emphasis id="47A9EAB5FFBCFFF0FF70585C3B49F8D5" bold="true" box="[199,345,1868,1894]" pageId="42" pageNumber="43">Description.</emphasis>
Male (
<typeStatus id="AA668805FFBCFFF0FE1F585C381DF8D5" box="[424,525,1868,1894]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
) (
<figureCitation id="EDE62A22FFBCFFF0FD93585C389FF8D5" box="[548,655,1868,1894]" captionStart="FIGURE 32" captionStartId="42.[151,250,1762,1786]" captionTargetBox="[160,1428,706,1738]" captionTargetId="figure-196@42.[160,1428,706,1738]" captionTargetPageId="42" captionText="FIGURE 32. Liangfeng Cave, type locality of Tyrannochthonius duo sp. nov., A. Entrance; B. Inside the cave entrance; C. Area where T. duo sp. nov. specimens were collected; D. Live male of T. duo sp. nov. in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.7795234" httpUri="https://zenodo.org/record/7795234/files/figure.png" pageId="42" pageNumber="43">Figs 32D</figureCitation>
, 
<figureCitation id="EDE62A22FFBCFFF0FD2C585C38DDF8D5" box="[667,717,1868,1894]" captionStart="FIGURE 33" captionStartId="43.[151,250,1295,1319]" captionTargetBox="[167,1422,424,1259]" captionTargetId="figure-141@43.[151,1436,417,1271]" captionTargetPageId="43" captionText="FIGURE 33. Tyrannochthonius duo sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795236" httpUri="https://zenodo.org/record/7795236/files/figure.png" pageId="42" pageNumber="43">33A</figureCitation>
, 
<figureCitation id="EDE62A22FFBCFFF0FD6E585C3937F8D5" box="[729,807,1868,1894]" captionStart="FIGURE 34" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[179,1408,186,1920]" captionTargetId="figure-20@44.[179,1408,181,1920]" captionTargetPageId="44" captionText="FIGURE 34. Tyrannochthonius duo sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795238" httpUri="https://zenodo.org/record/7795238/files/figure.png" pageId="42" pageNumber="43">34A–F</figureCitation>
, 
<figureCitation id="EDE62A22FFBCFFF0FC85585C3940F8D4" box="[818,848,1868,1895]" captionStart="FIGURE 35" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[179,1396,181,1905]" captionTargetId="figure-18@45.[179,1408,181,1919]" captionTargetPageId="45" captionText="FIGURE 35. Tyrannochthonius duo sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795240" httpUri="https://zenodo.org/record/7795240/files/figure.png" pageId="42" pageNumber="43">35</figureCitation>
, 
<figureCitation id="EDE62A22FFBCFFF0FCEB585C396CF8D5" box="[860,892,1868,1894]" captionStart="FIGURE 36" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@46.[179,1408,181,1919]" captionTargetPageId="46" captionText="FIGURE 36. Tyrannochthonius duo sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795242" httpUri="https://zenodo.org/record/7795242/files/figure.png" pageId="42" pageNumber="43">36</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFBCFFF0FF7058603EFFF839" blockId="42.[151,1436,1868,2039]" box="[199,1263,1904,1931]" pageId="42" pageNumber="43">Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</paragraph>
<paragraph id="756236A7FFBCFFF1FF7058843B3DFE3A" blockId="42.[151,1436,1868,2039]" lastBlockId="43.[151,1437,151,393]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
Cephalothorax (
<figureCitation id="EDE62A22FFBCFFF0FECB58843BF8F81D" box="[380,488,1940,1966]" captionStart="FIGURE 34" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[179,1408,186,1920]" captionTargetId="figure-20@44.[179,1408,181,1920]" captionTargetPageId="44" captionText="FIGURE 34. Tyrannochthonius duo sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795238" httpUri="https://zenodo.org/record/7795238/files/figure.png" pageId="42" pageNumber="43">Figs 34C</figureCitation>
, 
<figureCitation id="EDE62A22FFBCFFF0FE425884383AF81C" box="[501,554,1940,1967]" captionStart="FIGURE 35" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[179,1396,181,1905]" captionTargetId="figure-18@45.[179,1408,181,1919]" captionTargetPageId="45" captionText="FIGURE 35. Tyrannochthonius duo sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795240" httpUri="https://zenodo.org/record/7795240/files/figure.png" pageId="42" pageNumber="43">35A</figureCitation>
): carapace 1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; anterior pair of eyes well developed, posterior pair of eyes reduced into eyespots; anterior margin slightly serrate; epistome small and pointed, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 9–11 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="EDE62A22FFBDFFF1FC155E5B3E17FED6" box="[930,1031,330,357]" captionStart="FIGURE 35" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[179,1396,181,1905]" captionTargetId="figure-18@45.[179,1408,181,1919]" captionTargetPageId="45" captionText="FIGURE 35. Tyrannochthonius duo sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795240" httpUri="https://zenodo.org/record/7795240/files/figure.png" pageId="43" pageNumber="44">Fig. 35C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<caption id="21A2662FFFBDFFF1FF205A1F3F8DFA94" ID-DOI="http://doi.org/10.5281/zenodo.7795236" ID-Zenodo-Dep="7795236" box="[151,1437,1295,1319]" httpUri="https://zenodo.org/record/7795236/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,250,1295,1319]" targetBox="[167,1422,424,1259]" targetPageId="43">
<paragraph id="756236A7FFBDFFF1FF205A1F3F8DFA94" blockId="43.[151,1437,1295,1319]" box="[151,1437,1295,1319]" pageId="43" pageNumber="44">
<emphasis id="47A9EAB5FFBDFFF1FF205A1F3B0EFA94" bold="true" box="[151,286,1295,1319]" pageId="43" pageNumber="44">FIGURE 33.</emphasis>
<taxonomicName id="B2DD4D24FFBDFFF1FE955A1F3814FA94" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[290,516,1295,1319]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="duo" status="sp. nov.">
<emphasis id="47A9EAB5FFBDFFF1FE955A1F3814FA94" box="[290,516,1295,1319]" italics="true" pageId="43" pageNumber="44">Tyrannochthonius duo</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFBDFFF1FDBF5A1F3847FA94" bold="true" box="[520,599,1295,1319]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="5C9A57CEFFBDFFF1FDBF5A1F3847FA94" box="[520,599,1295,1319]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view).
</paragraph>
</caption>
<paragraph id="756236A7FFBDFFF1FF705A403B83F9D4" blockId="43.[151,1437,1360,2034]" pageId="43" pageNumber="44">
Chelicera (
<figureCitation id="EDE62A22FFBDFFF1FEF65A403BBCFAD9" box="[321,428,1360,1386]" captionStart="FIGURE 34" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[179,1408,186,1920]" captionTargetId="figure-20@44.[179,1408,181,1920]" captionTargetPageId="44" captionText="FIGURE 34. Tyrannochthonius duo sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795238" httpUri="https://zenodo.org/record/7795238/files/figure.png" pageId="43" pageNumber="44">Figs 34D</figureCitation>
, 
<figureCitation id="EDE62A22FFBDFFF1FE005A403BFAFAD8" box="[439,490,1360,1387]" captionStart="FIGURE 35" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[179,1396,181,1905]" captionTargetId="figure-18@45.[179,1408,181,1919]" captionTargetPageId="45" captionText="FIGURE 35. Tyrannochthonius duo sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795240" httpUri="https://zenodo.org/record/7795240/files/figure.png" pageId="43" pageNumber="44">35B</figureCitation>
): large, about as long as carapace, 2.08 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation both dorsal and ventral side. Both fingers well provided with teeth, fixed finger with 15 teeth, distal one largest; movable finger with 15 retrorse contiguous small teeth; galea completely vestigial (
<figureCitation id="EDE62A22FFBDFFF1FBA35AF03E6CFA48" box="[1044,1148,1504,1531]" captionStart="FIGURE 35" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[179,1396,181,1905]" captionTargetId="figure-18@45.[179,1408,181,1919]" captionTargetPageId="45" captionText="FIGURE 35. Tyrannochthonius duo sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795240" httpUri="https://zenodo.org/record/7795240/files/figure.png" pageId="43" pageNumber="44">Fig. 35B</figureCitation>
). Serrula exterior with 17 and serrula interior with 13 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="EDE62A22FFBDFFF1FE96595C3B97F9D4" box="[289,391,1612,1639]" captionStart="FIGURE 35" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[179,1396,181,1905]" captionTargetId="figure-18@45.[179,1408,181,1919]" captionTargetPageId="45" captionText="FIGURE 35. Tyrannochthonius duo sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795240" httpUri="https://zenodo.org/record/7795240/files/figure.png" pageId="43" pageNumber="44">Fig. 35E</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFBDFFF5FF7059603E58FEF2" blockId="43.[151,1437,1360,2034]" lastBlockId="47.[151,1437,151,1509]" lastPageId="47" lastPageNumber="48" pageId="43" pageNumber="44">
Pedipalp (
<figureCitation id="EDE62A22FFBDFFF1FE8059603BF0F939" box="[311,480,1648,1674]" captionStart="FIGURE 34" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[179,1408,186,1920]" captionTargetId="figure-20@44.[179,1408,181,1920]" captionTargetPageId="44" captionText="FIGURE 34. Tyrannochthonius duo sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795238" httpUri="https://zenodo.org/record/7795238/files/figure.png" pageId="43" pageNumber="44">Figs 34A, B, E</figureCitation>
, 
<figureCitation id="EDE62A22FFBDFFF1FE5B5960380FF938" box="[492,543,1648,1675]" captionStart="FIGURE 35" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[179,1396,181,1905]" captionTargetId="figure-18@45.[179,1408,181,1919]" captionTargetPageId="45" captionText="FIGURE 35. Tyrannochthonius duo sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795240" httpUri="https://zenodo.org/record/7795240/files/figure.png" pageId="43" pageNumber="44">35D</figureCitation>
, 
<figureCitation id="EDE62A22FFBDFFF1FD9E5960386DF939" box="[553,637,1648,1674]" captionStart="FIGURE 36" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@46.[179,1408,181,1919]" captionTargetPageId="46" captionText="FIGURE 36. Tyrannochthonius duo sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795242" httpUri="https://zenodo.org/record/7795242/files/figure.png" pageId="43" pageNumber="44">36A, B</figureCitation>
): long and slender, trochanter 1.36, femur 5.85, patella 2.27, chela 6.94, hand 2.25 times longer than broad; femur 2.24 times longer than patella; movable chelal finger 2.14 times longer than hand and 0.69 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (
<figureCitation id="EDE62A22FFBDFFF1FF6259CC3B2DF945" box="[213,317,1756,1782]" captionStart="FIGURE 34" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[179,1408,186,1920]" captionTargetId="figure-20@44.[179,1408,181,1920]" captionTargetPageId="44" captionText="FIGURE 34. Tyrannochthonius duo sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795238" httpUri="https://zenodo.org/record/7795238/files/figure.png" pageId="43" pageNumber="44">Figs 34E</figureCitation>
, 
<figureCitation id="EDE62A22FFBDFFF1FEFF59CC3B6CF944" box="[328,380,1756,1783]" captionStart="FIGURE 35" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[179,1396,181,1905]" captionTargetId="figure-18@45.[179,1408,181,1919]" captionTargetPageId="45" captionText="FIGURE 35. Tyrannochthonius duo sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795240" httpUri="https://zenodo.org/record/7795240/files/figure.png" pageId="43" pageNumber="44">35D</figureCitation>
). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, 
<emphasis id="47A9EAB5FFBDFFF1FC29583439A4F88D" box="[926,948,1828,1854]" italics="true" pageId="43" pageNumber="44">ib</emphasis>
and 
<emphasis id="47A9EAB5FFBDFFF1FC5E58343E1AF88D" box="[1001,1034,1828,1854]" italics="true" pageId="43" pageNumber="44">isb</emphasis>
situated close together, submedially on dorsum of chelal hand; 
<emphasis id="47A9EAB5FFBDFFF1FE0958583BC8F8D1" box="[446,472,1864,1890]" italics="true" pageId="43" pageNumber="44">eb</emphasis>
, 
<emphasis id="47A9EAB5FFBDFFF1FE515858381BF8D1" box="[486,523,1864,1890]" italics="true" pageId="43" pageNumber="44">esb</emphasis>
and 
<emphasis id="47A9EAB5FFBDFFF1FDF75859384AF8D1" box="[576,602,1865,1890]" italics="true" pageId="43" pageNumber="44">ist</emphasis>
forming a straight oblique row at base of fixed chelal finger; 
<emphasis id="47A9EAB5FFBDFFF1FAB558593F02F8D1" box="[1282,1298,1865,1890]" italics="true" pageId="43" pageNumber="44">it</emphasis>
slightly distal to 
<emphasis id="47A9EAB5FFBDFFF1FF62587D3AE4F835" box="[213,244,1901,1926]" italics="true" pageId="43" pageNumber="44">est</emphasis>
, situated subdistally; 
<emphasis id="47A9EAB5FFBDFFF1FE52587D3BE9F835" box="[485,505,1901,1926]" italics="true" pageId="43" pageNumber="44">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth; 
<emphasis id="47A9EAB5FFBDFFF1FB55587C3EECF835" box="[1250,1276,1900,1926]" italics="true" pageId="43" pageNumber="44">dx</emphasis>
situated distal to 
<emphasis id="47A9EAB5FFBDFFF1FF0358813AD8F819" box="[180,200,1937,1962]" italics="true" pageId="43" pageNumber="44">et</emphasis>
; 
<emphasis id="47A9EAB5FFBDFFF1FF6158803AFFF819" box="[214,239,1936,1962]" italics="true" pageId="43" pageNumber="44">sb</emphasis>
closer to 
<emphasis id="47A9EAB5FFBDFFF1FEEB58803B7AF819" box="[348,362,1936,1962]" italics="true" pageId="43" pageNumber="44">b</emphasis>
than to 
<emphasis id="47A9EAB5FFBDFFF1FE7358813BC7F819" box="[452,471,1937,1962]" italics="true" pageId="43" pageNumber="44">st</emphasis>
; 
<emphasis id="47A9EAB5FFBDFFF1FE5158803BE4F819" box="[486,500,1936,1962]" italics="true" pageId="43" pageNumber="44">b</emphasis>
and 
<emphasis id="47A9EAB5FFBDFFF1FD9E58813821F819" box="[553,561,1937,1962]" italics="true" pageId="43" pageNumber="44">t</emphasis>
situated subdistally, 
<emphasis id="47A9EAB5FFBDFFF1FCAC58813933F819" box="[795,803,1937,1962]" italics="true" pageId="43" pageNumber="44">t</emphasis>
situated at same level as 
<emphasis id="47A9EAB5FFBDFFF1FB8958813E4DF819" box="[1086,1117,1937,1962]" italics="true" pageId="43" pageNumber="44">est</emphasis>
and distal to 
<emphasis id="47A9EAB5FFBDFFF1FB4458803F11F819" box="[1267,1281,1936,1962]" italics="true" pageId="43" pageNumber="44">b</emphasis>
(
<figureCitation id="EDE62A22FFBDFFF1FAA758803F68F819" box="[1296,1400,1936,1962]" captionStart="FIGURE 36" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@46.[179,1408,181,1919]" captionTargetPageId="46" captionText="FIGURE 36. Tyrannochthonius duo sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795242" httpUri="https://zenodo.org/record/7795242/files/figure.png" pageId="43" pageNumber="44">Fig. 36A</figureCitation>
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to 
<emphasis id="47A9EAB5FFBDFFF1FBB658A53E0BF87D" box="[1025,1051,1973,1998]" italics="true" pageId="43" pageNumber="44">ist</emphasis>
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 32 macrodenticles, slightly retrorse and pointed, plus 
<specimenCount id="63DBFD2EFFB9FFF5FEE85FAB3BEAFF66" box="[351,506,187,213]" pageId="47" pageNumber="48" type="intercalary">21 intercalary</specimenCount>
microdenticles, 
<quantity id="B2259B42FFB9FFF5FD025FAA38FEFF66" box="[693,750,186,213]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" pageId="47" pageNumber="48" unit="in" value="53.0">53 in</quantity>
total; movable chelal finger with 16 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 
<specimenCount id="63DBFD2EFFB9FFF5FC4A5FCF3E8AFF4A" box="[1021,1178,223,249]" pageId="47" pageNumber="48" type="intercalary">10 intercalary</specimenCount>
microdenticles and 14 vestigial, rounded and contiguous basal teeth, 
<quantity id="B2259B42FFB9FFF5FD175E1338CAFEAE" box="[672,730,259,285]" metricMagnitude="0" metricUnit="m" metricValue="1.016" pageId="47" pageNumber="48" unit="in" value="40.0">40 in</quantity>
total (
<figureCitation id="EDE62A22FFB9FFF5FC945E13399CFEAE" box="[803,908,259,285]" captionStart="FIGURE 36" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@46.[179,1408,181,1919]" captionTargetPageId="46" captionText="FIGURE 36. Tyrannochthonius duo sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795242" httpUri="https://zenodo.org/record/7795242/files/figure.png" pageId="47" pageNumber="48">Fig. 36A</figureCitation>
). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="EDE62A22FFB9FFF5FC235E3739EEFEF2" box="[916,1022,295,321]" captionStart="FIGURE 34" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[179,1408,186,1920]" captionTargetId="figure-20@44.[179,1408,181,1920]" captionTargetPageId="44" captionText="FIGURE 34. Tyrannochthonius duo sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795238" httpUri="https://zenodo.org/record/7795238/files/figure.png" pageId="47" pageNumber="48">Figs 34B</figureCitation>
, 
<figureCitation id="EDE62A22FFB9FFF5FBBE5E373E2BFEF2" box="[1033,1083,295,321]" captionStart="FIGURE 36" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@46.[179,1408,181,1919]" captionTargetPageId="46" captionText="FIGURE 36. Tyrannochthonius duo sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795242" httpUri="https://zenodo.org/record/7795242/files/figure.png" pageId="47" pageNumber="48">36B</figureCitation>
).
</paragraph>
<caption id="21A2662FFFBAFFF6FF205887397BF84B" ID-DOI="http://doi.org/10.5281/zenodo.7795238" ID-Zenodo-Dep="7795238" httpUri="https://zenodo.org/record/7795238/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,250,1943,1967]" targetBox="[179,1408,186,1920]" targetPageId="44">
<paragraph id="756236A7FFBAFFF6FF205887397BF84B" blockId="44.[151,1437,1943,2040]" pageId="44" pageNumber="45">
<emphasis id="47A9EAB5FFBAFFF6FF2058873B31F81C" bold="true" box="[151,289,1943,1967]" pageId="44" pageNumber="45">FIGURE 34.</emphasis>
<taxonomicName id="B2DD4D24FFBAFFF6FE9F5888381BF803" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[296,523,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="duo" status="sp. nov.">
<emphasis id="47A9EAB5FFBAFFF6FE9F5888381BF803" box="[296,523,1944,1968]" italics="true" pageId="44" pageNumber="45">Tyrannochthonius duo</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFBAFFF6FDA558883874F803" bold="true" box="[530,612,1944,1968]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="5C9A57CEFFBAFFF6FDA558883874F803" box="[530,612,1944,1968]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view).
</paragraph>
</caption>
<caption id="21A2662FFFBBFFF7FF205887381AF84B" ID-DOI="http://doi.org/10.5281/zenodo.7795240" ID-Zenodo-Dep="7795240" httpUri="https://zenodo.org/record/7795240/files/figure.png" pageId="45" pageNumber="46" startId="45.[151,250,1943,1967]" targetBox="[179,1396,181,1905]" targetPageId="45">
<paragraph id="756236A7FFBBFFF7FF205887381AF84B" blockId="45.[151,1437,1943,2040]" pageId="45" pageNumber="46">
<emphasis id="47A9EAB5FFBBFFF7FF2058873B31F81C" bold="true" box="[151,289,1943,1967]" pageId="45" pageNumber="46">FIGURE 35.</emphasis>
<taxonomicName id="B2DD4D24FFBBFFF7FE905888381BF803" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[295,523,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="duo" status="sp. nov.">
<emphasis id="47A9EAB5FFBBFFF7FE905888381BF803" box="[295,523,1944,1968]" italics="true" pageId="45" pageNumber="46">Tyrannochthonius duo</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFBBFFF7FDA558883873F803" bold="true" box="[530,611,1944,1968]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="5C9A57CEFFBBFFF7FDA558883873F803" box="[530,611,1944,1968]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E).
</paragraph>
</caption>
<caption id="21A2662FFFB8FFF4FF2058873F59F860" ID-DOI="http://doi.org/10.5281/zenodo.7795242" ID-Zenodo-Dep="7795242" httpUri="https://zenodo.org/record/7795242/files/figure.png" pageId="46" pageNumber="47" startId="46.[151,250,1943,1967]" targetBox="[179,1408,181,1919]" targetPageId="46">
<paragraph id="756236A7FFB8FFF4FF2058873F59F860" blockId="46.[151,1436,1943,2004]" pageId="46" pageNumber="47">
<emphasis id="47A9EAB5FFB8FFF4FF2058873B32F81C" bold="true" box="[151,290,1943,1967]" pageId="46" pageNumber="47">FIGURE 36.</emphasis>
<taxonomicName id="B2DD4D24FFB8FFF4FE9D5888381FF803" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[298,527,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="duo" status="sp. nov.">
<emphasis id="47A9EAB5FFB8FFF4FE9D5888381FF803" box="[298,527,1944,1968]" italics="true" pageId="46" pageNumber="47">Tyrannochthonius duo</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFB8FFF4FDAF5888387AF803" bold="true" box="[536,618,1944,1968]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="5C9A57CEFFB8FFF4FDAF5888387AF803" box="[536,618,1944,1968]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm.
</paragraph>
</caption>
<paragraph id="756236A7FFB9FFF5FF705E5B3EBEFE62" blockId="47.[151,1437,151,1509]" pageId="47" pageNumber="48">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 3: 4: 6: 7: 8: 8: 6: 4: T2T: 0, tergites VI–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 12: 12: 9: 9: 9: 10: 10: 8: 0: 2. Anterior genital operculum with 12 setae, genital opening slit-like, with 17–18 marginal setae on each side, 
<quantity id="B2259B42FFB9FFF5FC0B5EA739E5FE62" box="[956,1013,439,465]" metricMagnitude="0" metricUnit="m" metricValue="1.1938" pageId="47" pageNumber="48" unit="in" value="47.0">47 in</quantity>
total (
<figureCitation id="EDE62A22FFB9FFF5FB8A5EA73EB1FE62" box="[1085,1185,439,465]" captionStart="FIGURE 34" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[179,1408,186,1920]" captionTargetId="figure-20@44.[179,1408,181,1920]" captionTargetPageId="44" captionText="FIGURE 34. Tyrannochthonius duo sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795238" httpUri="https://zenodo.org/record/7795238/files/figure.png" pageId="47" pageNumber="48">Fig. 34F</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFB9FFF5FF705ECB3BAEFD1A" blockId="47.[151,1437,151,1509]" pageId="47" pageNumber="48">
Legs (
<figureCitation id="EDE62A22FFB9FFF5FEBB5ECB3B84FE46" box="[268,404,475,501]" captionStart="FIGURE 36" captionStartId="46.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@46.[179,1408,181,1919]" captionTargetPageId="46" captionText="FIGURE 36. Tyrannochthonius duo sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795242" httpUri="https://zenodo.org/record/7795242/files/figure.png" pageId="47" pageNumber="48">Fig. 36C, D</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 2.09 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.13 times longer than tibia. Femoropatella of leg IV 2.91 times longer than deep; tibia 4.80 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.29 times longer than deep (TS= 0.35), tarsus 11.60 times longer than deep and 2.52 times longer than metatarsus (TS= 0.36). Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<paragraph id="756236A7FFB9FFF5FF705DA33B08FC32" blockId="47.[151,1437,151,1509]" pageId="47" pageNumber="48">
Adult females (
<typeStatus id="AA668805FFB9FFF5FECD5DA33BF9FD7E" box="[378,489,691,717]" pageId="47" pageNumber="48" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="EDE62A22FFB9FFF5FDB55DA3387FFD7E" box="[514,623,691,717]" captionStart="FIGURE 33" captionStartId="43.[151,250,1295,1319]" captionTargetBox="[167,1422,424,1259]" captionTargetId="figure-141@43.[151,1436,417,1271]" captionTargetPageId="43" captionText="FIGURE 33. Tyrannochthonius duo sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795236" httpUri="https://zenodo.org/record/7795236/files/figure.png" pageId="47" pageNumber="48">Figs 33B</figureCitation>
, 
<figureCitation id="EDE62A22FFB9FFF5FDC95DA338A2FD7E" box="[638,690,691,717]" captionStart="FIGURE 34" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[179,1408,186,1920]" captionTargetId="figure-20@44.[179,1408,181,1920]" captionTargetPageId="44" captionText="FIGURE 34. Tyrannochthonius duo sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795238" httpUri="https://zenodo.org/record/7795238/files/figure.png" pageId="47" pageNumber="48">34G</figureCitation>
). Mostly same as male, but a little larger; with same chaetotaxy of coxae; tergal chaetotaxy I–XII: 4: 4: 4: 4–6: 7–8: 8: 8: 8–9: 8–9: 4: T2T: 0; sternal chaetotaxy IV–XII: 11–13: 12: 11–12: 12: 12: 11–12: 9: 0: 2; anterior genital operculum with 11 setae, posterior margin with 12–14 marginal setae, 
<quantity id="B2259B42FFB9FFF5FF575C0E3B55FC8A" box="[224,325,798,825]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" metricValueMax="6.35" metricValueMin="5.842" pageId="47" pageNumber="48" unit="in" value="24.0" valueMax="25.0" valueMin="23.0">23–25 in</quantity>
total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.00–3.43 times longer than deep (TS= 0.33), tarsus 11.60–11.80 times longer than deep and 2.42–2.46 times longer than metatarsus (TS= 0.31–0.34).
</paragraph>
<paragraph id="756236A7FFB9FFF5FF705C9B39F5FBEA" blockId="47.[151,1437,151,1509]" pageId="47" pageNumber="48">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Male: body length 1.73. Pedipalps: trochanter 0.19/0.14 (1.36), femur 0.76/0.13 (5.85), patella 0.34/0.15 (2.27), chela 1.11/0.16 (6.94), hand 0.36/0.16 (2.25), movable chelal finger length 0.77. Chelicera 0.52/0.25 (2.08), movable finger length 0.28. Carapace 0.52/0.52 (1.00). Leg I: trochanter 0.15/0.12 (1.25), femur 0.46/0.07 (6.57), patella 0.22/0.07 (3.14), tibia 0.24/0.05 (4.80), tarsus 0.51/0.04 (12.75). Leg IV: trochanter 0.19/0.14 (1.36), femoropatella 0.67/0.23 (2.91), tibia 0.48/0.10 (4.80), metatarsus 0.23/0.07 (3.29), tarsus 0.58/0.05 (11.60).</paragraph>
<paragraph id="756236A7FFB9FFF5FF705B733EE4FAE6" blockId="47.[151,1437,151,1509]" pageId="47" pageNumber="48">Females: body length 1.93–2.12. Pedipalps: trochanter 0.19–0.21/0.14–0.16 (1.19–1.50), femur 0.79–0.81/0.13– 0.14 (5.79–6.08), patella 0.36/0.15–0.16 (2.25–2.40), chela 1.17–1.18/0.18–0.20 (5.85–6.56), hand 0.38–0.39/0.18– 0.20 (1.95–2.11), movable chelal finger length 0.80–0.82. Chelicera 0.57–0.58/0.28–0.29 (2.00–2.04), movable finger length 0.31–0.32. Carapace 0.56/0.61–0.63 (0.89–0.92). Leg I: trochanter 0.13–0.14/0.12–0.13 (1.08), femur 0.47–0.49/0.07–0.08 (5.88–7.00), patella 0.22–0.24/0.07 (3.14–3.43), tibia 0.24/0.05 (4.80), tarsus 0.48–0.54/0.05– 0.06 (9.00–9.60). Leg IV: trochanter 0.20–0.23/0.14–0.15 (1.33–1.64), femoropatella 0.70/0.23 (3.04), tibia 0.52– 0.53/0.10 (5.20–5.30), metatarsus 0.24/0.07–0.08 (3.00–3.43), tarsus 0.58–0.59/0.05 (11.60–11.80).</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFB9FFF5FF705A4F3ACBFA72" pageId="47" pageNumber="48" type="discussion">
<paragraph id="756236A7FFB9FFF5FF705A4F3ACBFA72" blockId="47.[151,1437,151,1509]" pageId="47" pageNumber="48">
<emphasis id="47A9EAB5FFB9FFF5FF705A4F3B2BFACA" bold="true" box="[199,315,1375,1401]" pageId="47" pageNumber="48">Remarks.</emphasis>
<taxonomicName id="B2DD4D24FFB9FFF5FEF35A4F3851FACA" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[324,577,1375,1401]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="duo" status="sp. nov.">
<emphasis id="47A9EAB5FFB9FFF5FEF35A4F3851FACA" box="[324,577,1375,1401]" italics="true" pageId="47" pageNumber="48">Tyrannochthonius duo</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFB9FFF5FDFD5A4F38B6FACA" bold="true" box="[586,678,1375,1401]" pageId="47" pageNumber="48">
<taxonomicNameLabel id="5C9A57CEFFB9FFF5FDFD5A4F38B6FACA" box="[586,678,1375,1401]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from other described Chinese troglomorphic 
<taxonomicName id="B2DD4D24FFB9FFF5FF205A933BA8FA2E" box="[151,440,1411,1437]" pageId="47" pageNumber="48">
<emphasis id="47A9EAB5FFB9FFF5FF205A933B71FA2E" box="[151,353,1411,1437]" italics="true" pageId="47" pageNumber="48">Tyrannochthonius</emphasis>
species
</taxonomicName>
by the presence of a pair of well-developed anterior eyes and more setae on tergites V‒IX (6–9).
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFB9FFF5FF705ADA38E1FA56" box="[199,753,1482,1509]" pageId="47" pageNumber="48" type="distribution">
<paragraph id="756236A7FFB9FFF5FF705ADA38E1FA56" blockId="47.[151,1437,151,1509]" box="[199,753,1482,1509]" pageId="47" pageNumber="48">
<emphasis id="47A9EAB5FFB9FFF5FF705ADA3B71FA57" bold="true" box="[199,353,1482,1508]" pageId="47" pageNumber="48">Distribution.</emphasis>
Known only from the 
<typeStatus id="AA668805FFB9FFF5FDD75ADB3880FA56" box="[608,656,1483,1509]" pageId="47" pageNumber="48">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>