419 lines
61 KiB
XML
419 lines
61 KiB
XML
<document id="2E4F8AE520A426F4B0D567867F8E539D" ID-CLB-Dataset="56235" ID-DOI="10.11646/zootaxa.5262.1.1" ID-GBIF-Dataset="e24a8032-296e-442d-80ae-e0bf8c7237f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7795161" ID-ZooBank="F05805FD-BC5B-4236-BF04-C6AA5C37F2D7" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680528957709" checkinUser="plazi" docAuthor="Hou, Yanmeng, Feng, Zegang & Zhang, Feng" docDate="2023" docId="FD7487B1FF8EFFC4FF205BBA38E1FB92" docLanguage="en" docName="zootaxa.5262.1.1.pdf" docOrigin="Zootaxa 5262 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5262.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tyrannochthonius babaowanensis Hou & Feng & Zhang 2023, sp. nov." docType="treatment" docVersion="7" lastPageNumber="31" masterDocId="014DFFC9FF96FFDAFFB75F103A10FFB3" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="158" masterPageNumber="1" pageNumber="25" updateTime="1699513313842" updateUser="ExternalLinkService">
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<mods:title id="D4749A25A016CAA008DEDA09398E9CA2">Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="70B2ECBBC052591DFE59FF9BDD5B2F25">Hou, Yanmeng</mods:namePart>
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<mods:roleTerm id="33BFD6D609536E5BE2BA65CAB8A07B40">Author</mods:roleTerm>
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<mods:namePart id="D132F29D5E6CF0F57F0728D12A31353A">Feng, Zegang</mods:namePart>
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<mods:namePart id="D0B048DA5BC7C3391B8B32932FBA9AA0">Zhang, Feng</mods:namePart>
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<mods:title id="FCF8AB478A34DA4BD601E5B43D856091">Zootaxa</mods:title>
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<mods:part id="B9A810C73F1D5898D5152B89FB447998">
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<mods:date id="B8E1CCD409BE1A2A4D2954D447E3CA3D">2023</mods:date>
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<mods:number id="518FFADEB630691C62BF89E4357AADF5">2023-04-03</mods:number>
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<mods:number id="EB7CE31AD71E8FAF19B6D5EE4F88AEEB">5262</mods:number>
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<treatment id="FD7487B1FF8EFFC4FF205BBA38E1FB92" ID-DOI="http://doi.org/10.5281/zenodo.7795401" ID-GBIF-Taxon="209487186" ID-Zenodo-Dep="7795401" LSID="urn:lsid:plazi:treatment:FD7487B1FF8EFFC4FF205BBA38E1FB92" httpUri="http://treatment.plazi.org/id/FD7487B1FF8EFFC4FF205BBA38E1FB92" lastPageId="30" lastPageNumber="31" pageId="24" pageNumber="25">
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<subSubSection id="3DC7652CFF8EFFC2FF205BBA3977FB76" box="[151,871,1194,1221]" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="756236A7FF8EFFC2FF205BBA3977FB76" blockId="24.[151,871,1194,1257]" box="[151,871,1194,1221]" pageId="24" pageNumber="25">
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<heading id="2E2A81CBFF8EFFC2FF205BBA3977FB76" box="[151,871,1194,1221]" fontSize="11" level="2" pageId="24" pageNumber="25" reason="5">
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<emphasis id="47A9EAB5FF8EFFC2FF205BBA388BFB76" bold="true" box="[151,667,1194,1221]" pageId="24" pageNumber="25">
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<taxonomicName id="B2DD4D24FF8EFFC2FF205BBA3826FB76" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[151,566,1194,1221]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="babaowanensis" status="sp. nov.">
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<emphasis id="47A9EAB5FF8EFFC2FF205BBA3826FB76" bold="true" box="[151,566,1194,1221]" italics="true" pageId="24" pageNumber="25">Tyrannochthonius babaowanensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5C9A57CEFF8EFFC2FD895BBB388BFB76" box="[574,667,1195,1221]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
(
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<emphasis id="47A9EAB5FF8EFFC2FD1A5BBB394DFB70" bold="true" box="[685,861,1195,1219]" pageId="24" pageNumber="25">ÃÍẠƟṁae</emphasis>
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)
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="3DC7652CFF8EFFC2FF205BDF3B04FB5A" box="[151,276,1231,1257]" pageId="24" pageNumber="25" type="description">
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<paragraph id="756236A7FF8EFFC2FF205BDF3B04FB5A" blockId="24.[151,871,1194,1257]" box="[151,276,1231,1257]" pageId="24" pageNumber="25">
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<figureCitation id="EDE62A22FF8EFFC2FF205BDF3B04FB5A" box="[151,276,1231,1257]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStart-3="FIGURE 20" captionStart-4="FIGURE 21" captionStartId-0="25.[151,250,1606,1630]" captionStartId-1="26.[151,250,1664,1688]" captionStartId-2="27.[151,250,1939,1963]" captionStartId-3="28.[151,250,1943,1967]" captionStartId-4="29.[151,250,1943,1967]" captionTargetBox-0="[159,1429,181,1582]" captionTargetBox-1="[271,1330,181,1640]" captionTargetBox-2="[172,1435,489,1916]" captionTargetBox-3="[173,1411,181,1903]" captionTargetBox-4="[179,1408,181,1919]" captionTargetId-0="figure-18@25.[159,1429,181,1582]" captionTargetId-1="figure-20@26.[250,1337,181,1640]" captionTargetId-2="figure-193@27.[151,1435,489,1916]" captionTargetId-3="figure-20@28.[173,1414,181,1919]" captionTargetId-4="figure-18@29.[179,1408,181,1919]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="28" captionTargetPageId-4="29" captionText-0="FIGURE 17. Babaowan Cave, type locality of Tyrannochthonius babaowanensis sp. nov., A. Entrance; B. Inside the cave entrance; C. A water tunnel in the cave; D–E. Area where T. babaowanensis sp. nov. specimen was collected (red arrow); F. Live male of T. babaowanensis sp. nov. in its natural environment." captionText-1="FIGURE 18. Tyrannochthonius babaowanensis sp. nov., holotype male, habitus (dorsal view)." captionText-2="FIGURE 19. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Male genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." captionText-3="FIGURE 20. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." captionText-4="FIGURE 21. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7795202" figureDoi-1="http://doi.org/10.5281/zenodo.7795204" figureDoi-2="http://doi.org/10.5281/zenodo.7795206" figureDoi-3="http://doi.org/10.5281/zenodo.7795208" figureDoi-4="http://doi.org/10.5281/zenodo.7795210" httpUri-0="https://zenodo.org/record/7795202/files/figure.png" httpUri-1="https://zenodo.org/record/7795204/files/figure.png" httpUri-2="https://zenodo.org/record/7795206/files/figure.png" httpUri-3="https://zenodo.org/record/7795208/files/figure.png" httpUri-4="https://zenodo.org/record/7795210/files/figure.png" pageId="24" pageNumber="25">Figs 17–21</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3DC7652CFF8EFFC2FF205A073F8DFACA" pageId="24" pageNumber="25" type="materials_examined">
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<paragraph id="756236A7FF8EFFC2FF205A073F8DFACA" blockId="24.[151,1437,1302,2013]" pageId="24" pageNumber="25">
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<emphasis id="47A9EAB5FF8EFFC2FF205A073B55FA83" bold="true" box="[151,325,1302,1329]" pageId="24" pageNumber="25">Type material.</emphasis>
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<materialsCitation id="C5B53CFAFF8EFFC2FEFC5A073F8DFACA" ID-GBIF-Occurrence="4071438333" collectingDate="2020-08-23" collectorName="Zegang Feng & Hongru Xu & Yanmeng Hou" country="China" county="Yinjiang County" elevation="655" latitude="28.064741" location="Tianchi Village" longLatPrecision="1" longitude="108.36469" municipality="Zhongxing Sub-district" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
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<typeStatus id="AA668805FF8EFFC2FEFC5A073BA3FA82" box="[331,435,1303,1329]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
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♂ (Ps.-MHBU-GZYJ-20-14-01):
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<collectingCountry id="0DCA7637FF8EFFC2FC835A07396AFA82" box="[820,890,1303,1329]" name="China" pageId="24" pageNumber="25">China</collectingCountry>
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,
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<collectingRegion id="B719F845FF8EFFC2FC335A073E42FA82" box="[900,1106,1303,1329]" country="China" name="Guizhou" pageId="24" pageNumber="25">Guizhou Province</collectingRegion>
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,
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<collectingCounty id="9C034E2BFF8EFFC2FBEB5A073F07FA82" box="[1116,1303,1303,1329]" pageId="24" pageNumber="25">Yinjiang County</collectingCounty>
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,
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<collectingMunicipality id="9506ACDDFF8EFFC2FA965A073B0DFAE6" pageId="24" pageNumber="25">Zhongxing Sub-district</collectingMunicipality>
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||
,
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||
<location id="7002607CFF8EFFC2FE915A2B3BC3FAE6" LSID="urn:lsid:plazi:treatment:FD7487B1FF8EFFC4FF205BBA38E1FB92:7002607CFF8EFFC2FE915A2B3BC3FAE6" box="[294,467,1339,1365]" country="China" county="Yinjiang County" latitude="28.064741" longLatPrecision="1" longitude="108.36469" municipality="Zhongxing Sub-district" name="Tianchi Village" pageId="24" pageNumber="25" stateProvince="Guizhou">Tianchi Village</location>
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,
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<location id="7002607CFF8EFFC2FE6B5A2B3884FAE6" LSID="urn:lsid:plazi:treatment:FD7487B1FF8EFFC4FF205BBA38E1FB92:7002607CFF8EFFC2FE6B5A2B3884FAE6" box="[476,660,1339,1365]" country="China" county="Yinjiang County" latitude="28.064741" longLatPrecision="1" longitude="108.36469" municipality="Zhongxing Sub-district" name="Babaowan Cave" pageId="24" pageNumber="25" stateProvince="Guizhou">Babaowan Cave</location>
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, on a stone wall in the deep zone (Temperature: 14°C, Humidity: 85%) [
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<geoCoordinate id="10E95060FF8EFFC2FF295A703B2DFAC8" box="[158,317,1376,1403]" degrees="28" direction="north" minutes="3" orientation="latitude" pageId="24" pageNumber="25" precision="1" seconds="53.07" value="28.064741">28°3′53.07″N</geoCoordinate>
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,
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<geoCoordinate id="10E95060FF8EFFC2FEFC5A703813FAC8" box="[331,515,1376,1403]" degrees="108" direction="east" minutes="21" orientation="longitude" pageId="24" pageNumber="25" precision="1" seconds="52.89" value="108.36469">108°21′52.89″E</geoCoordinate>
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],
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<elevation id="FEF0D194FF8EFFC2FDAF5A4E38B1FACA" box="[536,673,1374,1401]" metricMagnitude="2" metricUnit="m" metricValue="6.55" pageId="24" pageNumber="25" unit="m" value="655.0">
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<quantity id="B2259B42FF8EFFC2FDAF5A4E3872FACA" box="[536,610,1374,1401]" metricMagnitude="2" metricUnit="m" metricValue="6.55" pageId="24" pageNumber="25" unit="m" value="655.0">655 m</quantity>
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a.s.l.
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||
</elevation>
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||
,
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||
<date id="01631067FF8EFFC2FD075A4F3979FACA" box="[688,873,1375,1401]" pageId="24" pageNumber="25" value="2020-08-23">
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||
<collectingDate id="1127E98FFF8EFFC2FD075A4F3979FACA" box="[688,873,1375,1401]" pageId="24" pageNumber="25" value="2020-08-23">23 August 2020</collectingDate>
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</date>
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,
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<collectorName id="D8285371FF8EFFC2FCC05A4F3E1FFACA" box="[887,1039,1375,1401]" pageId="24" pageNumber="25">Zegang Feng</collectorName>
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,
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<collectorName id="D8285371FF8EFFC2FBAA5A4F3E8EFACA" box="[1053,1182,1375,1401]" pageId="24" pageNumber="25">Hongru Xu</collectorName>
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&
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<collectorName id="D8285371FF8EFFC2FB715A4F3F79FACA" box="[1222,1385,1375,1401]" pageId="24" pageNumber="25">Yanmeng Hou</collectorName>
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leg.
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||
</materialsCitation>
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DC7652CFF8EFFC2FF705A933976FA2E" box="[199,870,1411,1437]" pageId="24" pageNumber="25" type="etymology">
|
||
<paragraph id="756236A7FF8EFFC2FF705A933976FA2E" blockId="24.[151,1437,1302,2013]" box="[199,870,1411,1437]" pageId="24" pageNumber="25">
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<emphasis id="47A9EAB5FF8EFFC2FF705A933B5EFA2E" bold="true" box="[199,334,1411,1437]" pageId="24" pageNumber="25">Etymology.</emphasis>
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Named after the
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<typeStatus id="AA668805FF8EFFC2FDA75A933850FA2E" box="[528,576,1411,1437]" pageId="24" pageNumber="25">type</typeStatus>
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||
locality, Babaowan Cave.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="3DC7652CFF8EFFC2FF705AB638E9F9E2" pageId="24" pageNumber="25" type="diagnosis">
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||
<paragraph id="756236A7FF8EFFC2FF705AB638E9F9E2" blockId="24.[151,1437,1302,2013]" pageId="24" pageNumber="25">
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||
<emphasis id="47A9EAB5FF8EFFC2FF705AB63B5BFA73" bold="true" box="[199,331,1446,1472]" pageId="24" pageNumber="25">Diagnosis (</emphasis>
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♂
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<emphasis id="47A9EAB5FF8EFFC2FED75AB63B60FA73" bold="true" box="[352,368,1446,1472]" pageId="24" pageNumber="25">).</emphasis>
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Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, pointed, triangular; posterior margin of carapace with 2 setae; tergites I–VI each with 4 setae. Pedipalps slender, femur 7.67 times longer than broad; chela 7.27 times longer than broad; both chelal fingers with intercalary teeth (present in the distal 1/3 of the fingers); chemosensory setae present on dorsum of chelal hand.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="3DC7652CFF8EFFC4FF70594A39F6FD72" lastPageId="30" lastPageNumber="31" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="756236A7FF8EFFC2FF70594A39A4F9C6" blockId="24.[151,1437,1302,2013]" box="[199,948,1626,1653]" pageId="24" pageNumber="25">
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||
<emphasis id="47A9EAB5FF8EFFC2FF70594A3B49F9C7" bold="true" box="[199,345,1626,1652]" pageId="24" pageNumber="25">Description.</emphasis>
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Male (
|
||
<typeStatus id="AA668805FF8EFFC2FE1F594B3800F9C6" box="[424,528,1627,1653]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
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||
), female unknown (
|
||
<figureCitation id="EDE62A22FF8EFFC2FD5A594B3942F9C6" box="[749,850,1627,1653]" captionStart="FIGURE 17" captionStartId="25.[151,250,1606,1630]" captionTargetBox="[159,1429,181,1582]" captionTargetId="figure-18@25.[159,1429,181,1582]" captionTargetPageId="25" captionText="FIGURE 17. Babaowan Cave, type locality of Tyrannochthonius babaowanensis sp. nov., A. Entrance; B. Inside the cave entrance; C. A water tunnel in the cave; D–E. Area where T. babaowanensis sp. nov. specimen was collected (red arrow); F. Live male of T. babaowanensis sp. nov. in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.7795202" httpUri="https://zenodo.org/record/7795202/files/figure.png" pageId="24" pageNumber="25">Figs 17F</figureCitation>
|
||
,
|
||
<figureCitation id="EDE62A22FF8EFFC2FCEA594B39B7F9C6" box="[861,935,1627,1653]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStart-2="FIGURE 20" captionStart-3="FIGURE 21" captionStartId-0="26.[151,250,1664,1688]" captionStartId-1="27.[151,250,1939,1963]" captionStartId-2="28.[151,250,1943,1967]" captionStartId-3="29.[151,250,1943,1967]" captionTargetBox-0="[271,1330,181,1640]" captionTargetBox-1="[172,1435,489,1916]" captionTargetBox-2="[173,1411,181,1903]" captionTargetBox-3="[179,1408,181,1919]" captionTargetId-0="figure-20@26.[250,1337,181,1640]" captionTargetId-1="figure-193@27.[151,1435,489,1916]" captionTargetId-2="figure-20@28.[173,1414,181,1919]" captionTargetId-3="figure-18@29.[179,1408,181,1919]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionText-0="FIGURE 18. Tyrannochthonius babaowanensis sp. nov., holotype male, habitus (dorsal view)." captionText-1="FIGURE 19. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Male genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." captionText-2="FIGURE 20. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." captionText-3="FIGURE 21. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7795204" figureDoi-1="http://doi.org/10.5281/zenodo.7795206" figureDoi-2="http://doi.org/10.5281/zenodo.7795208" figureDoi-3="http://doi.org/10.5281/zenodo.7795210" httpUri-0="https://zenodo.org/record/7795204/files/figure.png" httpUri-1="https://zenodo.org/record/7795206/files/figure.png" httpUri-2="https://zenodo.org/record/7795208/files/figure.png" httpUri-3="https://zenodo.org/record/7795210/files/figure.png" pageId="24" pageNumber="25">18–21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="756236A7FF8EFFC2FF70596F3EFFF92A" blockId="24.[151,1437,1302,2013]" box="[199,1263,1663,1689]" pageId="24" pageNumber="25">Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</paragraph>
|
||
<paragraph id="756236A7FF8EFFC2FF7059B33EA9F86E" blockId="24.[151,1437,1302,2013]" pageId="24" pageNumber="25">
|
||
Cephalothorax (
|
||
<figureCitation id="EDE62A22FF8EFFC2FECB59B33BF7F90E" box="[380,487,1699,1725]" captionStart="FIGURE 19" captionStartId="27.[151,250,1939,1963]" captionTargetBox="[172,1435,489,1916]" captionTargetId="figure-193@27.[151,1435,489,1916]" captionTargetPageId="27" captionText="FIGURE 19. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Male genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795206" httpUri="https://zenodo.org/record/7795206/files/figure.png" pageId="24" pageNumber="25">Figs 19C</figureCitation>
|
||
,
|
||
<figureCitation id="EDE62A22FF8EFFC2FE4259B3383BF90E" box="[501,555,1699,1725]" captionStart="FIGURE 20" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[173,1411,181,1903]" captionTargetId="figure-20@28.[173,1414,181,1919]" captionTargetPageId="28" captionText="FIGURE 20. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795208" httpUri="https://zenodo.org/record/7795208/files/figure.png" pageId="24" pageNumber="25">20A</figureCitation>
|
||
): carapace 1.04 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8–9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
|
||
<figureCitation id="EDE62A22FF8EFFC2FDAA58D33894F86E" box="[541,644,1987,2013]" captionStart="FIGURE 20" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[173,1411,181,1903]" captionTargetId="figure-20@28.[173,1414,181,1919]" captionTargetPageId="28" captionText="FIGURE 20. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795208" httpUri="https://zenodo.org/record/7795208/files/figure.png" pageId="24" pageNumber="25">Fig. 20C</figureCitation>
|
||
); intercoxal tubercle absent; without sub-oral seta.
|
||
</paragraph>
|
||
<caption id="21A2662FFF8FFFC3FF2059563919F915" ID-DOI="http://doi.org/10.5281/zenodo.7795202" ID-Zenodo-Dep="7795202" httpUri="https://zenodo.org/record/7795202/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1606,1630]" targetBox="[159,1429,181,1582]" targetPageId="25">
|
||
<paragraph id="756236A7FF8FFFC3FF2059563919F915" blockId="25.[151,1436,1606,1703]" pageId="25" pageNumber="26">
|
||
<emphasis id="47A9EAB5FF8FFFC3FF2059563B30F9ED" bold="true" box="[151,288,1606,1630]" pageId="25" pageNumber="26">FIGURE 17.</emphasis>
|
||
Babaowan Cave, type locality of
|
||
<taxonomicName id="B2DD4D24FF8FFFC3FDCD595639C4F9ED" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[634,980,1606,1630]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="babaowanensis" status="sp. nov.">
|
||
<emphasis id="47A9EAB5FF8FFFC3FDCD595639C4F9ED" box="[634,980,1606,1630]" italics="true" pageId="25" pageNumber="26">Tyrannochthonius babaowanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="47A9EAB5FF8FFFC3FC6D59573E3BF9EC" bold="true" box="[986,1067,1607,1631]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="5C9A57CEFF8FFFC3FC6D59573E3BF9EC" box="[986,1067,1607,1631]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A. Entrance; B. Inside the cave entrance; C. A water tunnel in the cave; D–E. Area where
|
||
<taxonomicName id="B2DD4D24FF8FFFC3FD7B597B3991F931" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[716,897,1642,1666]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="babaowanensis" status="sp. nov.">
|
||
<emphasis id="47A9EAB5FF8FFFC3FD7B597B3991F931" box="[716,897,1642,1666]" italics="true" pageId="25" pageNumber="26">T. babaowanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="47A9EAB5FF8FFFC3FC30597B39C8F930" bold="true" box="[903,984,1643,1667]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="5C9A57CEFF8FFFC3FC30597B39C8F930" box="[903,984,1643,1667]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
specimen was collected (red arrow); F. Live male of
|
||
<taxonomicName id="B2DD4D24FF8FFFC3FF5D599F3B8FF915" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[234,415,1678,1702]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="babaowanensis" status="sp. nov.">
|
||
<emphasis id="47A9EAB5FF8FFFC3FF5D599F3B8FF915" box="[234,415,1678,1702]" italics="true" pageId="25" pageNumber="26">T. babaowanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="47A9EAB5FF8FFFC3FE11599F3BE6F914" bold="true" box="[422,502,1679,1703]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="5C9A57CEFF8FFFC3FE11599F3BE6F914" box="[422,502,1679,1703]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in its natural environment.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="756236A7FF8FFFC3FF7059C53B84F858" blockId="25.[151,1437,1748,2027]" pageId="25" pageNumber="26">
|
||
Chelicera (
|
||
<figureCitation id="EDE62A22FF8FFFC3FEF659C53BBCF95C" box="[321,428,1749,1775]" captionStart="FIGURE 19" captionStartId="27.[151,250,1939,1963]" captionTargetBox="[172,1435,489,1916]" captionTargetId="figure-193@27.[151,1435,489,1916]" captionTargetPageId="27" captionText="FIGURE 19. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Male genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795206" httpUri="https://zenodo.org/record/7795206/files/figure.png" pageId="25" pageNumber="26">Figs 19D</figureCitation>
|
||
,
|
||
<figureCitation id="EDE62A22FF8FFFC3FE0059C53BFAF95C" box="[439,490,1749,1775]" captionStart="FIGURE 20" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[173,1411,181,1903]" captionTargetId="figure-20@28.[173,1414,181,1919]" captionTargetPageId="28" captionText="FIGURE 20. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795208" httpUri="https://zenodo.org/record/7795208/files/figure.png" pageId="25" pageNumber="26">20B</figureCitation>
|
||
): large, about as long as carapace, 2.37 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 14 teeth, distal one largest; movable finger with 13 retrorse contiguous small teeth; galea completely vestigial (
|
||
<figureCitation id="EDE62A22FF8FFFC3FBA358753E6CF8CC" box="[1044,1148,1893,1919]" captionStart="FIGURE 20" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[173,1411,181,1903]" captionTargetId="figure-20@28.[173,1414,181,1919]" captionTargetPageId="28" captionText="FIGURE 20. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795208" httpUri="https://zenodo.org/record/7795208/files/figure.png" pageId="25" pageNumber="26">Fig. 20B</figureCitation>
|
||
). Serrula exterior with 21 and serrula interior with 14 blades. Rallum with 7 blades, the distal one longer and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
|
||
<figureCitation id="EDE62A22FF8FFFC3FE9658C13B97F858" box="[289,391,2001,2027]" captionStart="FIGURE 20" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[173,1411,181,1903]" captionTargetId="figure-20@28.[173,1414,181,1919]" captionTargetPageId="28" captionText="FIGURE 20. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795208" httpUri="https://zenodo.org/record/7795208/files/figure.png" pageId="25" pageNumber="26">Fig. 20E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="21A2662FFF8CFFC0FF2059903E4DF92B" ID-DOI="http://doi.org/10.5281/zenodo.7795204" ID-Zenodo-Dep="7795204" box="[151,1117,1664,1688]" httpUri="https://zenodo.org/record/7795204/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,250,1664,1688]" targetBox="[271,1330,181,1640]" targetPageId="26">
|
||
<paragraph id="756236A7FF8CFFC0FF2059903E4DF92B" blockId="26.[151,1117,1664,1688]" box="[151,1117,1664,1688]" pageId="26" pageNumber="27">
|
||
<emphasis id="47A9EAB5FF8CFFC0FF2059903B30F92B" bold="true" box="[151,288,1664,1688]" pageId="26" pageNumber="27">FIGURE 18.</emphasis>
|
||
<taxonomicName id="B2DD4D24FF8CFFC0FE915990386FF92B" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[294,639,1664,1688]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="babaowanensis" status="sp. nov.">
|
||
<emphasis id="47A9EAB5FF8CFFC0FE915990386FF92B" box="[294,639,1664,1688]" italics="true" pageId="26" pageNumber="27">Tyrannochthonius babaowanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="47A9EAB5FF8CFFC0FD32599038C6F92B" bold="true" box="[645,726,1664,1688]" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="5C9A57CEFF8CFFC0FD32599038C6F92B" box="[645,726,1664,1688]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male, habitus (dorsal view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="756236A7FF8CFFC1FF7059D639D8FE62" blockId="26.[151,1437,1734,2013]" lastBlockId="27.[151,1437,151,465]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
|
||
Pedipalp (
|
||
<figureCitation id="EDE62A22FF8CFFC0FE8F59D63BCEF952" box="[312,478,1734,1761]" captionStart="FIGURE 19" captionStartId="27.[151,250,1939,1963]" captionTargetBox="[172,1435,489,1916]" captionTargetId="figure-193@27.[151,1435,489,1916]" captionTargetPageId="27" captionText="FIGURE 19. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Male genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795206" httpUri="https://zenodo.org/record/7795206/files/figure.png" pageId="26" pageNumber="27">Figs 19A, B, F</figureCitation>
|
||
,
|
||
<figureCitation id="EDE62A22FF8CFFC0FE5E59D6382DF953" box="[489,573,1734,1760]" captionStart="FIGURE 21" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@29.[179,1408,181,1919]" captionTargetPageId="29" captionText="FIGURE 21. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795210" httpUri="https://zenodo.org/record/7795210/files/figure.png" pageId="26" pageNumber="27">21A, B</figureCitation>
|
||
): long and slender, trochanter 1.22, femur 7.67, patella 2.72, chela 7.27, hand 2.41 times longer than broad; femur 2.35 times longer than patella; movable chelal finger 2.06 times longer than hand and 0.68 times longer than chela. Setae generally long and acuminate; one distal lyrifissures present on patella (
|
||
<figureCitation id="EDE62A22FF8CFFC0FF2858223B17F8FF" box="[159,263,1842,1868]" captionStart="FIGURE 19" captionStartId="27.[151,250,1939,1963]" captionTargetBox="[172,1435,489,1916]" captionTargetId="figure-193@27.[151,1435,489,1916]" captionTargetPageId="27" captionText="FIGURE 19. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Male genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795206" httpUri="https://zenodo.org/record/7795206/files/figure.png" pageId="26" pageNumber="27">Figs 19F</figureCitation>
|
||
,
|
||
<figureCitation id="EDE62A22FF8CFFC0FEA258223B5AF8FF" box="[277,330,1842,1868]" captionStart="FIGURE 20" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[173,1411,181,1903]" captionTargetId="figure-20@28.[173,1414,181,1919]" captionTargetPageId="28" captionText="FIGURE 20. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795208" httpUri="https://zenodo.org/record/7795208/files/figure.png" pageId="26" pageNumber="27">20D</figureCitation>
|
||
). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
|
||
<emphasis id="47A9EAB5FF8CFFC0FC29586A39A4F827" box="[926,948,1914,1940]" italics="true" pageId="26" pageNumber="27">ib</emphasis>
|
||
and
|
||
<emphasis id="47A9EAB5FF8CFFC0FC5E586A3E1AF827" box="[1001,1034,1914,1940]" italics="true" pageId="26" pageNumber="27">isb</emphasis>
|
||
situated close together, submedially on dorsum of chelal hand;
|
||
<emphasis id="47A9EAB5FF8CFFC0FE7D588E3BF4F80B" box="[458,484,1950,1976]" italics="true" pageId="26" pageNumber="27">eb</emphasis>
|
||
,
|
||
<emphasis id="47A9EAB5FF8CFFC0FE43588E3809F80B" box="[500,537,1950,1976]" italics="true" pageId="26" pageNumber="27">esb</emphasis>
|
||
and
|
||
<emphasis id="47A9EAB5FF8CFFC0FDE5588F387CF80B" box="[594,620,1951,1976]" italics="true" pageId="26" pageNumber="27">ist</emphasis>
|
||
forming a straight oblique row at base of fixed chelal finger;
|
||
<emphasis id="47A9EAB5FF8CFFC0FA87588F3F50F80B" box="[1328,1344,1951,1976]" italics="true" pageId="26" pageNumber="27">it</emphasis>
|
||
slightly distal to
|
||
<emphasis id="47A9EAB5FF8CFFC0FF4E58D33B08F86F" box="[249,280,1987,2012]" italics="true" pageId="26" pageNumber="27">est</emphasis>
|
||
, situated subdistally;
|
||
<emphasis id="47A9EAB5FF8CFFC0FDB958D33832F86F" box="[526,546,1987,2012]" italics="true" pageId="26" pageNumber="27">et</emphasis>
|
||
slightly near to tip of fixed chelal finger, very close to chelal teeth;
|
||
<emphasis id="47A9EAB5FF8CFFC0FA9458D23F2DF86F" box="[1315,1341,1986,2012]" italics="true" pageId="26" pageNumber="27">dx</emphasis>
|
||
situated distal to
|
||
<emphasis id="47A9EAB5FF8DFFC1FF4F5F873B1CFF03" box="[248,268,151,176]" italics="true" pageId="27" pageNumber="28">et</emphasis>
|
||
;
|
||
<emphasis id="47A9EAB5FF8DFFC1FEAB5F873B25FF02" box="[284,309,151,177]" italics="true" pageId="27" pageNumber="28">sb</emphasis>
|
||
closer to
|
||
<emphasis id="47A9EAB5FF8DFFC1FE145F873BA1FF02" box="[419,433,151,177]" italics="true" pageId="27" pageNumber="28">b</emphasis>
|
||
than to
|
||
<emphasis id="47A9EAB5FF8DFFC1FDBA5F873830FF03" box="[525,544,151,176]" italics="true" pageId="27" pageNumber="28">st</emphasis>
|
||
;
|
||
<emphasis id="47A9EAB5FF8DFFC1FD985F87382DFF02" box="[559,573,151,177]" italics="true" pageId="27" pageNumber="28">b</emphasis>
|
||
and
|
||
<emphasis id="47A9EAB5FF8DFFC1FDC35F87386CFF03" box="[628,636,151,176]" italics="true" pageId="27" pageNumber="28">t</emphasis>
|
||
situated subdistally,
|
||
<emphasis id="47A9EAB5FF8DFFC1FCD05F873965FF02" box="[871,885,151,177]" italics="true" pageId="27" pageNumber="28">b</emphasis>
|
||
situated at same level as
|
||
<emphasis id="47A9EAB5FF8DFFC1FB235F873EA3FF03" box="[1172,1203,151,176]" italics="true" pageId="27" pageNumber="28">est</emphasis>
|
||
,
|
||
<emphasis id="47A9EAB5FF8DFFC1FB765F873ED9FF03" box="[1217,1225,151,176]" italics="true" pageId="27" pageNumber="28">t</emphasis>
|
||
slightly distal to
|
||
<emphasis id="47A9EAB5FF8DFFC1FA3A5F873F8DFF03" box="[1421,1437,151,176]" italics="true" pageId="27" pageNumber="28">it</emphasis>
|
||
(
|
||
<figureCitation id="EDE62A22FF8DFFC1FF285FAB3B17FF66" box="[159,263,187,213]" captionStart="FIGURE 21" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@29.[179,1408,181,1919]" captionTargetPageId="29" captionText="FIGURE 21. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795210" httpUri="https://zenodo.org/record/7795210/files/figure.png" pageId="27" pageNumber="28">Fig. 21A</figureCitation>
|
||
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
|
||
<emphasis id="47A9EAB5FF8DFFC1FB2E5FAB3EA3FF67" box="[1177,1203,187,212]" italics="true" pageId="27" pageNumber="28">ist</emphasis>
|
||
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 42 macrodenticles, slightly retrorse and pointed, plus
|
||
<specimenCount id="63DBFD2EFF8DFFC1FDAF5E3638A3FEF2" box="[536,691,294,321]" pageId="27" pageNumber="28" type="intercalary">15 intercalary</specimenCount>
|
||
microdenticles (present in the distal 1/3 of the fingers, till
|
||
<emphasis id="47A9EAB5FF8DFFC1FAFE5E373F78FEF3" box="[1353,1384,295,320]" italics="true" pageId="27" pageNumber="28">est</emphasis>
|
||
),
|
||
<quantity id="B2259B42FF8DFFC1FA375E363ABDFED6" metricMagnitude="0" metricUnit="m" metricValue="1.4478" pageId="27" pageNumber="28" unit="in" value="57.0">57 in</quantity>
|
||
total; movable chelal finger with 31 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus
|
||
<specimenCount id="63DBFD2EFF8DFFC1FE145E7F382CFE3A" box="[419,572,367,393]" pageId="27" pageNumber="28" type="intercalary">11 intercalary</specimenCount>
|
||
microdenticles (present in the distal 1/3 of the fingers, till
|
||
<emphasis id="47A9EAB5FF8DFFC1FB705E7F3EC5FE3A" box="[1223,1237,367,393]" italics="true" pageId="27" pageNumber="28">b</emphasis>
|
||
) and 14 vestigial, rounded and contiguous basal teeth,
|
||
<quantity id="B2259B42FF8DFFC1FD9A5E823876FE1E" box="[557,614,402,429]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="27" pageNumber="28" unit="in" value="56.0">56 in</quantity>
|
||
total (
|
||
<figureCitation id="EDE62A22FF8DFFC1FD1B5E833904FE1E" box="[684,788,403,429]" captionStart="FIGURE 21" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@29.[179,1408,181,1919]" captionTargetPageId="29" captionText="FIGURE 21. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795210" httpUri="https://zenodo.org/record/7795210/files/figure.png" pageId="27" pageNumber="28">Fig. 21A</figureCitation>
|
||
). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
|
||
<figureCitation id="EDE62A22FF8DFFC1FCA45EA7396DFE62" box="[787,893,439,465]" captionStart="FIGURE 19" captionStartId="27.[151,250,1939,1963]" captionTargetBox="[172,1435,489,1916]" captionTargetId="figure-193@27.[151,1435,489,1916]" captionTargetPageId="27" captionText="FIGURE 19. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Male genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795206" httpUri="https://zenodo.org/record/7795206/files/figure.png" pageId="27" pageNumber="28">Figs 19B</figureCitation>
|
||
,
|
||
<figureCitation id="EDE62A22FF8DFFC1FC3E5EA739ABFE62" box="[905,955,439,465]" captionStart="FIGURE 21" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@29.[179,1408,181,1919]" captionTargetPageId="29" captionText="FIGURE 21. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795210" httpUri="https://zenodo.org/record/7795210/files/figure.png" pageId="27" pageNumber="28">21B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="21A2662FFF8DFFC1FF2058833B4EF847" ID-DOI="http://doi.org/10.5281/zenodo.7795206" ID-Zenodo-Dep="7795206" httpUri="https://zenodo.org/record/7795206/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,250,1939,1963]" targetBox="[172,1435,489,1916]" targetPageId="27">
|
||
<paragraph id="756236A7FF8DFFC1FF2058833B4EF847" blockId="27.[151,1436,1939,2036]" pageId="27" pageNumber="28">
|
||
<emphasis id="47A9EAB5FF8DFFC1FF2058833B33F818" bold="true" box="[151,291,1939,1963]" pageId="27" pageNumber="28">FIGURE 19.</emphasis>
|
||
<taxonomicName id="B2DD4D24FF8DFFC1FE9B58843897F81F" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[300,647,1940,1964]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="babaowanensis" status="sp. nov.">
|
||
<emphasis id="47A9EAB5FF8DFFC1FE9B58843897F81F" box="[300,647,1940,1964]" italics="true" pageId="27" pageNumber="28">Tyrannochthonius babaowanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="47A9EAB5FF8DFFC1FD26588438F4F81F" bold="true" box="[657,740,1940,1964]" pageId="27" pageNumber="28">
|
||
<taxonomicNameLabel id="5C9A57CEFF8DFFC1FD26588438F4F81F" box="[657,740,1940,1964]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Male genital area (ventral view); F. Left pedipalp (minus chela, dorsal view).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="21A2662FFF8AFFC6FF205887386FF84B" ID-DOI="http://doi.org/10.5281/zenodo.7795208" ID-Zenodo-Dep="7795208" httpUri="https://zenodo.org/record/7795208/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,250,1943,1967]" targetBox="[173,1411,181,1903]" targetPageId="28">
|
||
<paragraph id="756236A7FF8AFFC6FF205887386FF84B" blockId="28.[151,1437,1943,2040]" pageId="28" pageNumber="29">
|
||
<emphasis id="47A9EAB5FF8AFFC6FF2058873B31F81C" bold="true" box="[151,289,1943,1967]" pageId="28" pageNumber="29">FIGURE 20.</emphasis>
|
||
<taxonomicName id="B2DD4D24FF8AFFC6FE9F58883892F803" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[296,642,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="babaowanensis" status="sp. nov.">
|
||
<emphasis id="47A9EAB5FF8AFFC6FE9F58883892F803" box="[296,642,1944,1968]" italics="true" pageId="28" pageNumber="29">Tyrannochthonius babaowanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="47A9EAB5FF8AFFC6FD3D588838CBF803" bold="true" box="[650,731,1944,1968]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="5C9A57CEFF8AFFC6FD3D588838CBF803" box="[650,731,1944,1968]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="21A2662FFF8BFFC7FF2058873F83F860" ID-DOI="http://doi.org/10.5281/zenodo.7795210" ID-Zenodo-Dep="7795210" httpUri="https://zenodo.org/record/7795210/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1943,1967]" targetBox="[179,1408,181,1919]" targetPageId="29">
|
||
<paragraph id="756236A7FF8BFFC7FF2058873F83F860" blockId="29.[151,1437,1943,2004]" pageId="29" pageNumber="30">
|
||
<emphasis id="47A9EAB5FF8BFFC7FF2058873B30F81C" bold="true" box="[151,288,1943,1967]" pageId="29" pageNumber="30">FIGURE 21.</emphasis>
|
||
<taxonomicName id="B2DD4D24FF8BFFC7FE9058883891F803" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[295,641,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="babaowanensis" status="sp. nov.">
|
||
<emphasis id="47A9EAB5FF8BFFC7FE9058883891F803" box="[295,641,1944,1968]" italics="true" pageId="29" pageNumber="30">Tyrannochthonius babaowanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="47A9EAB5FF8BFFC7FD3F588838C9F803" bold="true" box="[648,729,1944,1968]" pageId="29" pageNumber="30">
|
||
<taxonomicNameLabel id="5C9A57CEFF8BFFC7FD3F588838C9F803" box="[648,729,1944,1968]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="756236A7FF88FFC4FF705F873871FEA9" blockId="30.[151,1437,151,1057]" pageId="30" pageNumber="31">
|
||
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 5: 5: 5: 2: T2T: 0. Sternal chaetotaxy III–XII: 14: 12: 7: 7: 8: 9: 9: 7: 0: 2. Anterior genital operculum with 13 setae, genital opening slit-like, with 14–15 marginal setae on each side,
|
||
<quantity id="B2259B42FF88FFC4FED85E103BB8FEA9" box="[367,424,256,282]" metricMagnitude="0" metricUnit="m" metricValue="1.0668" pageId="30" pageNumber="31" unit="in" value="42.0">42 in</quantity>
|
||
total (
|
||
<figureCitation id="EDE62A22FF88FFC4FE585E103845FEA9" box="[495,597,256,282]" captionStart="FIGURE 19" captionStartId="27.[151,250,1939,1963]" captionTargetBox="[172,1435,489,1916]" captionTargetId="figure-193@27.[151,1435,489,1916]" captionTargetPageId="27" captionText="FIGURE 19. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Male genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795206" httpUri="https://zenodo.org/record/7795206/files/figure.png" pageId="30" pageNumber="31">Fig. 19E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="756236A7FF88FFC4FF705E343BAEFE5E" blockId="30.[151,1437,151,1057]" pageId="30" pageNumber="31">
|
||
Legs (
|
||
<figureCitation id="EDE62A22FF88FFC4FEBB5E333B84FE8D" box="[268,404,291,318]" captionStart="FIGURE 21" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@29.[179,1408,181,1919]" captionTargetPageId="29" captionText="FIGURE 21. Tyrannochthonius babaowanensis sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795210" httpUri="https://zenodo.org/record/7795210/files/figure.png" pageId="30" pageNumber="31">Fig. 21C, D</figureCitation>
|
||
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.97 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.43 times longer than tibia. Femoropatella of leg IV 4.21 times longer than deep; tibia 7.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.75 times longer than deep (TS= 0.33), tarsus 14.00 times longer than deep and 2.80 times longer than metatarsus (TS= 0.32). Arolium slightly shorter than the claws, not divided; claws simple.
|
||
</paragraph>
|
||
<paragraph id="756236A7FF88FFC4FF705EE739F6FD72" blockId="30.[151,1437,151,1057]" pageId="30" pageNumber="31">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Male: body length 2.35. Pedipalps: trochanter 0.22/0.18 (1.22), femur 1.15/0.15 (7.67), patella 0.49/0.18 (2.72), chela 1.60/0.22 (7.27), hand 0.53/0.22 (2.41), movable chelal finger length 1.09. Chelicera 0.64/0.27 (2.37), movable finger length 0.35. Carapace 0.59/0.57 (1.04). Leg I: trochanter 0.19/0.15 (1.27), femur 0.69/0.08 (8.63), patella 0.35/0.07 (5.00), tibia 0.30/0.06 (5.00), tarsus 0.73/0.06 (12.17). Leg IV: trochanter 0.27/0.16 (1.69), femoropatella 1.01/0.24 (4.21), tibia 0.70/0.10 (7.00), metatarsus 0.30/0.08 (3.75), tarsus 0.84/0.06 (14.00).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DC7652CFF88FFC4FF705DDA3F7AFC4D" pageId="30" pageNumber="31" type="discussion">
|
||
<paragraph id="756236A7FF88FFC4FF705DDA3915FCC2" blockId="30.[151,1437,151,1057]" pageId="30" pageNumber="31">
|
||
<emphasis id="47A9EAB5FF88FFC4FF705DDA3B2BFD57" bold="true" box="[199,315,714,740]" pageId="30" pageNumber="31">Remarks.</emphasis>
|
||
Female unknown.
|
||
<taxonomicName id="B2DD4D24FF88FFC4FDB85DDA399CFD57" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[527,908,714,740]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="babaowanensis" status="sp. nov.">
|
||
<emphasis id="47A9EAB5FF88FFC4FDB85DDA399CFD57" box="[527,908,714,740]" italics="true" pageId="30" pageNumber="31">Tyrannochthonius babaowanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="47A9EAB5FF88FFC4FC255DDA39FBFD57" bold="true" box="[914,1003,714,740]" pageId="30" pageNumber="31">
|
||
<taxonomicNameLabel id="5C9A57CEFF88FFC4FC255DDA39FBFD57" box="[914,1003,714,740]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="B2DD4D24FF88FFC4FBCD5DDB3F3DFD57" authorityName="Mahnert" authorityYear="2009" box="[1146,1325,714,740]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="antridraconis">
|
||
<emphasis id="47A9EAB5FF88FFC4FBCD5DDB3F3DFD57" box="[1146,1325,714,740]" italics="true" pageId="30" pageNumber="31">T. antridraconis</emphasis>
|
||
</taxonomicName>
|
||
in having intercalary teeth on both chelal fingers and a small, rounded and obtuse epistome, but differs by a larger body size (body length 1.62–1.69 (♂)
|
||
<emphasis id="47A9EAB5FF88FFC4FE645C013BE1FC99" box="[467,497,785,810]" italics="true" pageId="30" pageNumber="31">vs.</emphasis>
|
||
1.29–1.33 (♂) mm), the number of setae on tergite VII (5
|
||
<emphasis id="47A9EAB5FF88FFC4FB355C013EB1FC99" box="[1154,1185,785,810]" italics="true" pageId="30" pageNumber="31">vs.</emphasis>
|
||
4) and tergite Ⅹ (2
|
||
<emphasis id="47A9EAB5FF88FFC4FAC95C013F8CFC99" box="[1406,1436,785,810]" italics="true" pageId="30" pageNumber="31">vs.</emphasis>
|
||
4–5), the proportion of intercalary teeth on the fixed finger (1/3
|
||
<emphasis id="47A9EAB5FF88FFC4FCD15C243994FCFE" box="[870,900,820,845]" italics="true" pageId="30" pageNumber="31">vs.</emphasis>
|
||
2/3), the number of blades of coxal spines (8–9
|
||
<emphasis id="47A9EAB5FF88FFC4FF205C473AA6FCC3" box="[151,182,855,880]" italics="true" pageId="30" pageNumber="31">vs.</emphasis>
|
||
12–15) and the number of blades of rallum (7
|
||
<emphasis id="47A9EAB5FF88FFC4FD755C4738F0FCC3" box="[706,736,855,880]" italics="true" pageId="30" pageNumber="31">vs.</emphasis>
|
||
8).
|
||
</paragraph>
|
||
<paragraph id="756236A7FF88FFC4FF705C6A3F7AFC4D" blockId="30.[151,1437,151,1057]" pageId="30" pageNumber="31">
|
||
<taxonomicName id="B2DD4D24FF88FFC4FF705C6A3856FC27" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" box="[199,582,890,916]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="babaowanensis" status="sp. nov.">
|
||
<emphasis id="47A9EAB5FF88FFC4FF705C6A3856FC27" box="[199,582,890,916]" italics="true" pageId="30" pageNumber="31">Tyrannochthonius babaowanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="47A9EAB5FF88FFC4FDF95C6A38B9FC27" bold="true" box="[590,681,890,916]" pageId="30" pageNumber="31">
|
||
<taxonomicNameLabel id="5C9A57CEFF88FFC4FDF95C6A38B9FC27" box="[590,681,890,916]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="B2DD4D24FF88FFC4FC695C6B3E5DFC27" authorityName="Gao, Wynne & Fe.Zhang" authorityYear="2018" box="[990,1101,890,916]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="chixing">
|
||
<emphasis id="47A9EAB5FF88FFC4FC695C6B3E5DFC27" box="[990,1101,890,916]" italics="true" pageId="30" pageNumber="31">T. chixing</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B2DD4D24FF88FFC4FB315C6B3EC1FC27" authorityName="Gao, Zhang & Chen" authorityYear="2020" box="[1158,1233,890,916]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="zhai">
|
||
<emphasis id="47A9EAB5FF88FFC4FB315C6B3EC1FC27" box="[1158,1233,890,916]" italics="true" pageId="30" pageNumber="31">T. zhai</emphasis>
|
||
</taxonomicName>
|
||
by the larger size (body length 2.35
|
||
<emphasis id="47A9EAB5FF88FFC4FED65C8E3B90FC04" box="[353,384,926,951]" italics="true" pageId="30" pageNumber="31">vs.</emphasis>
|
||
max.
|
||
<quantity id="B2259B42FF88FFC4FE755C8D3838FC0B" box="[450,552,925,952]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" pageId="30" pageNumber="31" unit="mm" value="1.64">1.64 mm</quantity>
|
||
; chela length 1.60
|
||
<emphasis id="47A9EAB5FF88FFC4FD415C8E3904FC04" box="[758,788,926,951]" italics="true" pageId="30" pageNumber="31">vs.</emphasis>
|
||
max.
|
||
<quantity id="B2259B42FF88FFC4FCE05C8D39AEFC04" box="[855,958,925,951]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="30" pageNumber="31" unit="mm" value="1.24">1.24 mm</quantity>
|
||
), the number of setae at the anterior margin of the carapace (6
|
||
<emphasis id="47A9EAB5FF88FFC4FED55CD13B91FC69" box="[354,385,961,986]" italics="true" pageId="30" pageNumber="31">vs.</emphasis>
|
||
4) and the presence of intercalary teeth on both chelal fingers, from
|
||
<taxonomicName id="B2DD4D24FF88FFC4FBC35CD13EF4FC69" authorityName="Mahnert" authorityYear="2009" box="[1140,1252,960,986]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="akaelus">
|
||
<emphasis id="47A9EAB5FF88FFC4FBC35CD13EF4FC69" box="[1140,1252,960,986]" italics="true" pageId="30" pageNumber="31">T. akaelus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B2DD4D24FF88FFC4FB465CD13AA5FC4E" authorityName="Mahnert" authorityYear="2009" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="ganshuanensis">
|
||
<emphasis id="47A9EAB5FF88FFC4FB465CD13AA5FC4E" italics="true" pageId="30" pageNumber="31">T. ganshuanensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B2DD4D24FF88FFC4FF5C5CF43B4BFC4E" authorityName="Gao, Zhang & Chen" authorityYear="2020" box="[235,347,995,1021]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="47A9EAB5FF88FFC4FF5C5CF43B4BFC4E" box="[235,347,995,1021]" italics="true" pageId="30" pageNumber="31">T. harveyi</emphasis>
|
||
</taxonomicName>
|
||
by the number of setae on the tergites I–II (4
|
||
<emphasis id="47A9EAB5FF88FFC4FCEB5CF4396AFC4E" box="[860,890,996,1021]" italics="true" pageId="30" pageNumber="31">vs.</emphasis>
|
||
2) (
|
||
<bibRefCitation id="114C4B56FF88FFC4FC105CF33E41FC4D" author="Mahnert, V." box="[935,1105,995,1022]" pageId="30" pageNumber="31" pagination="185 - 201" refId="ref77141" refString="Mahnert, V. (2009) New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue Suisse de Zoologie, 116 (2), 185 - 201. https: // doi. org / 10.5962 / bhl. part. 79492" type="journal article" year="2009">Mahnert 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="114C4B56FF88FFC4FBEA5CF43F03FC4D" author="Gao, Z. Z. & Wynne, J. J. & Zhang, F." box="[1117,1299,995,1022]" pageId="30" pageNumber="31" pagination="345 - 354" refId="ref76312" refString="Gao, Z. Z., Wynne, J. J. & Zhang, F. (2018) Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones: Neobisiidae, Chthoniidae) from Guangxi, China. Journal of Arachnology, 46, 345 - 354. https: // doi. org / 10.1636 / JoA-S- 17 - 063.1" type="journal article" year="2018">
|
||
Gao
|
||
<emphasis id="47A9EAB5FF88FFC4FB235CF43EDFFC4E" box="[1172,1231,995,1021]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="114C4B56FF88FFC4FAA85CF43F4CFC4D" author="Gao, Z. Z. & Zhang, F. & Chen, H. M." box="[1311,1372,996,1022]" pageId="30" pageNumber="31" pagination="572 - 580" refId="ref76374" refString="Gao, Z. Z., Zhang, F. & Chen, H. M. (2020) Two new cave-dwelling species of Tyrannochthonius Chamberlin 1929 (Pseudoscorpiones: Chthoniidae) from the Guizhou karst, China. Zootaxa, 4853 (4), 572 - 580. https: // doi. org / 10.11646 / zootaxa. 4853.4.6" type="journal article" year="2020">2020</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DC7652CFF88FFC4FF705B1638E1FB92" box="[199,753,1030,1057]" pageId="30" pageNumber="31" type="distribution">
|
||
<paragraph id="756236A7FF88FFC4FF705B1638E1FB92" blockId="30.[151,1437,151,1057]" box="[199,753,1030,1057]" pageId="30" pageNumber="31">
|
||
<emphasis id="47A9EAB5FF88FFC4FF705B163B71FB93" bold="true" box="[199,353,1030,1056]" pageId="30" pageNumber="31">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="AA668805FF88FFC4FDD75B173880FB92" box="[608,656,1031,1057]" pageId="30" pageNumber="31">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |