289 lines
26 KiB
XML
289 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.419.5739" ID-GBIF-Dataset="3a3e01cb-b600-482e-8963-e4159f54c6f1" ID-PMC="PMC4109453" ID-Pensoft-Pub="1313-2970-419-87" ID-PubMed="25061362" ID-ZBK="2BC0965952D04B5081B9CFBD78EB2DAE" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-419-87" ModsDocOrigin="ZooKeys 419" ModsDocTitle="Typhlonesticus gocmeni sp. n., a new cave-dwelling blind spider species from the Aegean region of Turkey (Araneae, Nesticidae)" checkinTime="1451245707135" checkinUser="pensoft" docAuthor="Ribera, Carles, Elverici, Mert, Kunt, Kadir Bogac & Oezkuetuek, Recep Sulhi" docDate="2014" docId="F8F51A073671C60068A905F0C2C9D15F" docLanguage="en" docName="ZooKeys 419: 87-102" docOrigin="ZooKeys 419" docSource="http://dx.doi.org/10.3897/zookeys.419.5739" docTitle="Typhlonesticus gocmeni Ribera, Elverici, Kunt & Oezkuetuek, 2014, sp. n." docType="treatment" docUuid="E3A3721A-0E71-479B-9EA9-A31A47D6E8C6" docUuidSource="ZooBank" docVersion="4" lastPageNumber="95" masterDocId="FFD820227604F668FFC8A14FFFF7FF8F" masterDocTitle="Typhlonesticus gocmeni sp. n., a new cave-dwelling blind spider species from the Aegean region of Turkey (Araneae, Nesticidae)" masterLastPageNumber="102" masterPageNumber="87" pageNumber="90" updateTime="1668158746226" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Typhlonesticus gocmeni sp. n., a new cave-dwelling blind spider species from the Aegean region of Turkey (Araneae, Nesticidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ribera, Carles</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Elverici, Mert</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kunt, Kadir Bogac</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Oezkuetuek, Recep Sulhi</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>419</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>87</mods:start>
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<mods:end>102</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.419.5739</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.419.5739</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-419-87</mods:identifier>
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<mods:identifier type="ZBK">2BC0965952D04B5081B9CFBD78EB2DAE</mods:identifier>
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<mods:identifier type="ZooBank">2BC0965952D04B5081B9CFBD78EB2DAE</mods:identifier>
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||
</mods:mods>
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<treatment ID-GBIF-Taxon="152053818" LSID="urn:lsid:zoobank.org:act:E3A3721A-0E71-479B-9EA9-A31A47D6E8C6" httpUri="http://treatment.plazi.org/id/F8F51A073671C60068A905F0C2C9D15F" lastPageId="8" lastPageNumber="95" pageId="3" pageNumber="90">
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<subSubSection pageId="3" pageNumber="90" type="multiple">
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<paragraph pageId="3" pageNumber="90">Taxon classification Animalia Araneae Nesticidae</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="90" type="nomenclature">
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<paragraph pageId="3" pageNumber="90">
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<taxonomicName LSID="http://zoobank.org/E3A3721A-0E71-479B-9EA9-A31A47D6E8C6" class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus gocmeni" order="Araneae" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="gocmeni">Typhlonesticus gocmeni</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="90">sp. n.</taxonomicNameLabel>
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Figs 2-21
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="90" type="material examined">
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<paragraph pageId="3" pageNumber="90">Material examined.</paragraph>
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<paragraph pageId="3" pageNumber="90">
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Holotype ♂ (AUZM) Denizli Province,
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<normalizedToken originalValue="Acıpayam">Acipayam</normalizedToken>
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District, Dodurgalar Town,
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<normalizedToken originalValue="Keloğlan">Keloglan</normalizedToken>
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Cave (
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="37.38743">37°23'14.74"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="29.571749">29°34'18.29"E</geoCoordinate>
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), 10.07.2011, leg. M. Elverici. Paratypes 1 ♂ 1 ♀ 9 juveniles (AUZM), 1 ♂ 1 ♀ (CRBA) same data as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="90" type="derivatio nominis">
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<paragraph pageId="3" pageNumber="90">Derivatio nominis.</paragraph>
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<paragraph pageId="3" pageNumber="90">
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The specific name is given in honour of the prominent Turkish biologist, Prof. Dr. Bayram
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<normalizedToken originalValue="Göçmen">Goecmen</normalizedToken>
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(University of Ege,
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<normalizedToken originalValue="İzmir">Izmir</normalizedToken>
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, Turkey). Noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="90" type="diagnosis">
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<paragraph pageId="3" pageNumber="90">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="90">
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Males of the new species differ from
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus absoloni" order="Araneae" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="absoloni">Typhlonesticus absoloni</taxonomicName>
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by the shape and length of the tegulum, the shape of p1 and p2 processes, the arrangement of the embolus and the shape and arrangement of the paracymbial apophyses, mainly the ventral one, which is erected as a thin spine in
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus gocmeni" order="Araneae" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="gocmeni">Typhlonesticus gocmeni</taxonomicName>
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sp. n., whereas in
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus absoloni" order="Araneae" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="absoloni">Typhlonesticus absoloni</taxonomicName>
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it consists of a curved lamella. Females differ from
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus absoloni" order="Araneae" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="absoloni">Typhlonesticus absoloni</taxonomicName>
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by the shape of the epigyne and the position of the spermathecae. The dimension and orientation of the insemination and fertilization ducts are also diagnostic. In
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus gocmeni" order="Araneae" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="gocmeni">Typhlonesticus gocmeni</taxonomicName>
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sp. n. the epigyne is scarcely sclerotized and the spermathecae and insemination ducts are visible through the tegument, moreover the spermathecae are nearly spherical and separated by a distance approximately equal to their diameter. In
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus absoloni" order="Araneae" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="absoloni">Typhlonesticus absoloni</taxonomicName>
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the epigyne is strongly sclerotized and the spermathecae are separated by almost twice their diameter. In
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus absoloni" order="Araneae" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="absoloni">Typhlonesticus absoloni</taxonomicName>
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the insemination and fertilization ducts are thicker and almost fill the entire genital area.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="92" pageId="3" pageNumber="90" type="male">
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<paragraph pageId="3" pageNumber="90">Male.</paragraph>
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<paragraph pageId="3" pageNumber="90">Coloration. Carapace whitish, slightly yellowish. Appendages and sternum slightly testaceous. Opisthosoma brownish, with many dark patches (Fig. 20). The specimens preserved in alcohol have a whitish opisthosoma, slightly greyer than the prosoma.</paragraph>
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<paragraph pageId="3" pageNumber="90">Prosoma. Carapace approximately circular in dorsal view. Cephalic region not differentiated from the rest of the carapace. Eyeless (Figs 18, 19).</paragraph>
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<paragraph pageId="3" pageNumber="90">Opisthosoma. Sub-elliptical in dorsal view.</paragraph>
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<paragraph pageId="4" pageNumber="91">
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<pageBreakToken pageId="4" pageNumber="91" start="start">Appendages</pageBreakToken>
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. prolateral margin of the chelicerae with 3 teeth, the central slightly longer. Male palp (Figs 2-13). Paracymbium short, dorsal and ventral processes scarcely developed. The ventral process consists of a short and flattened lamella, curved towards the apex and prolonged into a thin spine. The ventral one consisting of a short laminar apophysis, apically curved toward to the ventral side. Distal, paradistal and dorsomedian paracymbial apophyses absent (Figs 2-3 and 5-7). Tegulum very prominent, consisting of a ventrally directed triangular apophysis. Small inconspicuous median apophysis located behind the tegulum (Figs 2, 4, 6, 8, 12). TTA with two well developed processes (p1 and p2): p1 is saddle-shaped, longer than wide, slightly curved in the middle and directed ventrally; p2 is in an apical position and ends with two convergent apical hooks running as a conductor for the embolus (Figs 2, 4-6, 8, 11, 13). Embolus filamentous following a semicircular course towards the apex and bordering the tegulum.
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</paragraph>
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<caption pageId="4" pageNumber="91">
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<paragraph pageId="4" pageNumber="91">
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Figure 1. ML tree inferred using the concatenated dataset of cox1, rrnL mtDNA and H3 nuDNA gene fragments. Numbers next to nodes correspond to bootstrap support values. The tree was rooted using
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Canarionesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canarionesticus quadridentatus" order="Araneae" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="quadridentatus">Canarionesticus quadridentatus</taxonomicName>
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from the Canary Islands.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="91">
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<paragraph pageId="4" pageNumber="91">
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Figures 2-5.
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus gocmeni" order="Araneae" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="gocmeni">Typhlonesticus gocmeni</taxonomicName>
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sp. n. male palp. 2 ventral view 3 dorsal view 4 prolateral view 5 retrolateral view. Abbreviations: T = tegulum, MA = median apophysis, p1 = process 1 of TTA, p2 = process 2 of TTA, P = paracymbium, vp = ventral process of paracymbium, dp = dorsal process of paracymbium. Scale bars 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="91">
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<paragraph pageId="4" pageNumber="91">
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Figures 6-9.
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus gocmeni" order="Araneae" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="gocmeni">Typhlonesticus gocmeni</taxonomicName>
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sp. n. male palp. 6 ventral view 7 dorsal view 8 retrolateral view 9 prolateral view. Abbreviations: T = tegulum, MA = median apophysis, p1 = process 1 of TTA, p2 = process 2 of TTA, vp = ventral process of paracymbium, dp = dorsal process of paracymbium. Scale bars 0.5 mm.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="91">
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<paragraph pageId="4" pageNumber="91">
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Figures 10-13.
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus gocmeni" order="Araneae" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="gocmeni">Typhlonesticus gocmeni</taxonomicName>
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sp. n. male palp. 10 nearly retrolateral view 11 ditto 12 nearly ventral view 13 ventral view. Abbreviations: E = embolus, T = tegulum, St = subtegulum, MA = median apophysis, p1 = process 1 of TTA, p2 = process 2 of TTA, P = paracymbium, vp = ventral process of paracymbium, dp = dorsal process of paracymbium.
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</paragraph>
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</caption>
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<paragraph pageId="5" pageNumber="92">
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<pageBreakToken pageId="5" pageNumber="92" start="start">Measurements</pageBreakToken>
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(holotype ♂): carapace length 1.15, width 0.88, opisthosoma length 1.60, width 0.84. Total length = 2.75.
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</paragraph>
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<paragraph pageId="5" pageNumber="92">
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<table pageId="5" pageNumber="92">
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<tr pageId="5" pageNumber="92">
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<th colspan="1" pageId="5" pageNumber="92" rowspan="1" style="text-align:left">Leg</th>
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<th colspan="1" pageId="5" pageNumber="92" rowspan="1" style="text-align:left">coxa</th>
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<th colspan="1" pageId="5" pageNumber="92" rowspan="1" style="text-align:left">troc.</th>
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<th colspan="1" pageId="5" pageNumber="92" rowspan="1" style="text-align:left">femur</th>
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<th colspan="1" pageId="5" pageNumber="92" rowspan="1" style="text-align:left">patella</th>
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<th colspan="1" pageId="5" pageNumber="92" rowspan="1" style="text-align:left">tibia</th>
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<th colspan="1" pageId="5" pageNumber="92" rowspan="1" style="text-align:left">meta.</th>
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<th colspan="1" pageId="5" pageNumber="92" rowspan="1" style="text-align:left">tarsus</th>
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<th colspan="1" pageId="5" pageNumber="92" rowspan="1" style="text-align:left">total</th>
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</tr>
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<tr pageId="5" pageNumber="92">
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<td colspan="1" pageId="5" pageNumber="92" rowspan="1">I</td>
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</tr>
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<tr pageId="5" pageNumber="92">
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<td colspan="1" pageId="5" pageNumber="92" rowspan="1">II</td>
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</tr>
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<tr pageId="5" pageNumber="92">
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<td colspan="1" pageId="5" pageNumber="92" rowspan="1">III</td>
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</tr>
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<tr pageId="5" pageNumber="92">
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<td colspan="1" pageId="5" pageNumber="92" rowspan="1">IV</td>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="93" type="female">
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<paragraph pageId="6" pageNumber="93">
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<pageBreakToken pageId="6" pageNumber="93" start="start">Female</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="6" pageNumber="93">All somatic characters as in male but slightly larger in size. Epigyne convex and prominent, without sclerotized plates (Fig. 14). The posterior edge is slightly sclerotized. Spermathecae and insemination ducts can be observed through the tegument. Vulva quite simple (Figs 15-17), consisting of two almost spherical spermathecae, insemination and fertilization ducts. Insemination duct coiled, forming two laps around the fertilization duct before reaching the spermatheca. Vulval pockets absent.</paragraph>
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<caption pageId="6" pageNumber="93">
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<paragraph pageId="6" pageNumber="93">
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Figures 14-17.
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus gocmeni" order="Araneae" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="gocmeni">Typhlonesticus gocmeni</taxonomicName>
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sp. n. epigyne and vulva. 14 epigyne ventral view 15 vulva ventral view, 16 ditto 17 vulva dorsal view. Abbreviations: co = copulatory orifice, id = insemination duct, fd = fertilization duct, S = spermatheca. Scale bars 14-15 0.25 mm 16-17 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="6" pageNumber="93">
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<paragraph pageId="6" pageNumber="93">
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Figures 18-21.
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus gocmeni" order="Araneae" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="gocmeni">Typhlonesticus gocmeni</taxonomicName>
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sp. n. 18-19 male, prosoma 18 dorsal view 19 frontal view 20 male in the web 21 female.
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</paragraph>
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</caption>
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<paragraph pageId="6" pageNumber="93">Measurements (paratype ♀): carapace length 1.20, width 1.00, opisthosoma length 2.04, width 1.28. Total length = 3.24.</paragraph>
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<paragraph pageId="6" pageNumber="93">
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<table pageId="6" pageNumber="93">
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<tr pageId="6" pageNumber="93">
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<th colspan="1" pageId="6" pageNumber="93" rowspan="1" style="text-align:left">Leg</th>
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<th colspan="1" pageId="6" pageNumber="93" rowspan="1" style="text-align:left">coxa</th>
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<th colspan="1" pageId="6" pageNumber="93" rowspan="1" style="text-align:left">troc.</th>
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<th colspan="1" pageId="6" pageNumber="93" rowspan="1" style="text-align:left">femur</th>
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<th colspan="1" pageId="6" pageNumber="93" rowspan="1" style="text-align:left">patella</th>
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<th colspan="1" pageId="6" pageNumber="93" rowspan="1" style="text-align:left">tibia</th>
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<th colspan="1" pageId="6" pageNumber="93" rowspan="1" style="text-align:left">meta.</th>
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<th colspan="1" pageId="6" pageNumber="93" rowspan="1" style="text-align:left">tarsus</th>
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<th colspan="1" pageId="6" pageNumber="93" rowspan="1" style="text-align:left">total</th>
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</tr>
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<tr pageId="6" pageNumber="93">
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<td colspan="1" pageId="6" pageNumber="93" rowspan="1">I</td>
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</tr>
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<tr pageId="6" pageNumber="93">
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<td colspan="1" pageId="6" pageNumber="93" rowspan="1">II</td>
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</tr>
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<tr pageId="6" pageNumber="93">
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<td colspan="1" pageId="6" pageNumber="93" rowspan="1">III</td>
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</tr>
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<tr pageId="6" pageNumber="93">
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<td colspan="1" pageId="6" pageNumber="93" rowspan="1">IV</td>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="93" type="distribution">
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<paragraph pageId="6" pageNumber="93">Distribution.</paragraph>
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<paragraph pageId="6" pageNumber="93">
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus gocmeni" order="Araneae" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="gocmeni">Typhlonesticus gocmeni</taxonomicName>
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sp. n. is only known from the type locality. This new species was previously identified as
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<taxonomicName class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus morisii" order="Araneae" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="morisii">Nesticus morisii</taxonomicName>
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by
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||
<normalizedToken originalValue="Aydın">Aydin</normalizedToken>
|
||
<normalizedToken originalValue="Topçu">Topcu</normalizedToken>
|
||
and collaborators (
|
||
<bibRefCitation author="Topcu, A" journalOrPublisher="Zoology in the Middle East" pageId="10" pageNumber="97" pagination="91 - 92" title="A contribution to the cave spider fauna of Turkey (Araneae)." url="10.1080/09397140.2013.795075" volume="59" year="2013">2013</bibRefCitation>
|
||
). The cave is located at the northern part of the West Taurus karst region; on the east side of the
|
||
<normalizedToken originalValue="Acıpayam">Acipayam</normalizedToken>
|
||
polje, at the south eastern slope of the
|
||
<normalizedToken originalValue="Karadağ">Karadag</normalizedToken>
|
||
hill, about 200 m above the polje level (
|
||
<bibRefCitation author="MTA," journalOrPublisher="Mineral Research & Exploration General Directorate, Ankara" pageId="10" pageNumber="97" title="Dodurgalar Cave (Dodurgalar-Acipayam-Denizli) research report, architectural and electrification as-built project" year="1998">MTA 1998</bibRefCitation>
|
||
). It is a fossil cave, almost horizontal and 145 m long with a roof height varying between 1-9 m with many calcite speleothem formations. It is one of the tourist caves in Turkey, open to public access since 2003, with formed tracks and fixed lighting that extend almost the full length of the cave. Specimens were collected or observed during 3 visits on 20.03.2011, 10.07.2011 and 16.10.2011. Adult specimens from both sexes were only collected in July, but also observed in October; while only subadults could be found in March. Specimens were abundant in the dark zone all along the cave, located on their webs build on the speleothem formations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="8" lastPageNumber="95" pageId="6" pageNumber="93" type="molecular data">
|
||
<paragraph pageId="6" pageNumber="93">Molecular data.</paragraph>
|
||
<paragraph pageId="6" pageNumber="93">Specimens, locality and sequences with corresponding GenBank accession numbers analyzed in the present study are listed in Appendix 1. The final concatenated dataset of the three partial genes sequences includes 13 terminals and 1807 aligned characters (cox1 = 1049, rrnL = 420 and H3 = 338). Primer fidelity across taxa was not always consistent in cox1, consequently some specimens have slightly truncated sequence lengths. Uncorrected cox1 genetic divergences among terminal taxa, and uncorrected genetic cox1 divergences within and between the analyzed genera are summarized in Appendices 3 and 4.</paragraph>
|
||
<paragraph pageId="7" pageNumber="94">
|
||
<pageBreakToken pageId="7" pageNumber="94" start="start">Figure</pageBreakToken>
|
||
1 shows the ML tree inferred using the combined data matrix. The new species groups with
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus absoloni" order="Araneae" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="absoloni">Typhlonesticus absoloni</taxonomicName>
|
||
,
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus morisii" order="Araneae" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="morisii">Nesticus morisii</taxonomicName>
|
||
,
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus obcaecatus" order="Araneae" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="obcaecatus">Nesticus obcaecatus</taxonomicName>
|
||
and
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus idriacus" order="Araneae" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="idriacus">Nesticus idriacus</taxonomicName>
|
||
. These five species constitute a highly supported evolutionary lineage (bootstrap support = 100). The remaining species included in the analysis belong to the genera
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus" order="Araneae" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus">Nesticus</taxonomicName>
|
||
and
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Carpathonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carpathonesticus" order="Araneae" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus">Carpathonesticus</taxonomicName>
|
||
, which constitute independent and highly supported evolutionary lineages as well.
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus gocmeni" order="Araneae" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="gocmeni">Typhlonesticus gocmeni</taxonomicName>
|
||
sp. n. occupies a basal position in the
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus" order="Araneae" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus">Typhlonesticus</taxonomicName>
|
||
clade, and is the sister species of the European representatives. Within this lineage the evolutionary relationships of the species are poorly supported (low bootstrap supports).
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="95">
|
||
<pageBreakToken pageId="8" pageNumber="95" start="start">The</pageBreakToken>
|
||
mean uncorrected p-distances of cox1 between and within taxa analyzed (Appendices 3-4) show high values. The mean p-distance between genera ranges from 11.29% (
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus" order="Araneae" pageId="8" pageNumber="95" phylum="Arthropoda" rank="genus">Nesticus</taxonomicName>
|
||
versus
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Carpathonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carpathonesticus" order="Araneae" pageId="8" pageNumber="95" phylum="Arthropoda" rank="genus">Carpathonesticus</taxonomicName>
|
||
) to 17.19 (
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus" order="Araneae" pageId="8" pageNumber="95" phylum="Arthropoda" rank="genus">Typhlonesticus</taxonomicName>
|
||
versus
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Carpathonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carpathonesticus" order="Araneae" pageId="8" pageNumber="95" phylum="Arthropoda" rank="genus">Carpathonesticus</taxonomicName>
|
||
). Also, the average evolutionary divergence within the representatives of the three Mediterranean genera analyzed ranges from 6.43% (
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Carpathonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carpathonesticus" order="Araneae" pageId="8" pageNumber="95" phylum="Arthropoda" rank="genus">Carpathonesticus</taxonomicName>
|
||
) to 11.11% (
|
||
<taxonomicName class="Arachnida" family="Nesticidae" genus="Typhlonesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhlonesticus" order="Araneae" pageId="8" pageNumber="95" phylum="Arthropoda" rank="genus">Typhlonesticus</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |