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<mods:title id="C6AB4055328CE4AAA5A410273CA2B12F">A revision of the Palaearctic species of the ant genus Cardiocondyla Emery 1869 (Hymenoptera: Formicidae)</mods:title>
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<treatment id="F8463F1440609C41FF19669AFB56AF24" ID-DOI="http://doi.org/10.5281/zenodo.7888214" ID-GBIF-Taxon="209580621" ID-Zenodo-Dep="7888214" LSID="urn:lsid:plazi:treatment:F8463F1440609C41FF19669AFB56AF24" httpUri="http://treatment.plazi.org/id/F8463F1440609C41FF19669AFB56AF24" lastPageNumber="31" pageId="30" pageNumber="31">
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<paragraph id="70508E0240609C41FF19669AFCBEA9A0" blockId="30.[151,1436,402,575]" box="[151,823,402,429]" pageId="30" pageNumber="31">
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<taxonomicName id="B7EFF58140609C41FF19669AFDDCA9A0" authority="Seifert 2003" authorityName="Seifert" authorityYear="2003" box="[151,597,403,429]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="brachyceps">
<emphasis id="429B521040609C41FF19669AFE4CA9A0" box="[151,453,403,429]" italics="true" pageId="30" pageNumber="31">Cardiocondyla brachyceps</emphasis>
<bibRefCitation id="147EF3F340609C41FE42669AFDDCA9A0" author="Seifert, B." box="[460,597,403,429]" pageId="30" pageNumber="31" pagination="203 - 338" refId="ref43774" refString="Seifert, B. (2003) The ant genus Cardiocondyla (Insecta: Hymenoptera: Formicidae) - a taxonomic revision of the C. elegans, C. bulgarica, C. batesii, C. nuda, C. shuckardi, C. stambuloffii, C. wroughtonii, C. emeryi, and C. minutior species groups. Annalen des Naturhistorischen Museums, Wien, Serie B, 104, 203 - 338." type="journal article" year="2003">Seifert 2003</bibRefCitation>
</taxonomicName>
[
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investigation]
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This taxon has been described from west of Shiraz / Iran. Investigated were the
<typeStatus id="AF5430A040609C41FBBD66BEFB1CA9DC" box="[1075,1173,439,465]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
worker and
<materialsCitation id="C087845F40609C41FAA666BFFF5FAA14" collectorName="Schodl" elevation="1940" pageId="30" pageNumber="31" specimenCount="15" specimenCount-worker="15" typeStatus="paratype">
<specimenCount id="66E9458B40609C41FAA666BFFA15A9DC" box="[1320,1436,438,465]" pageId="30" pageNumber="31" type="worker">15 worker</specimenCount>
<typeStatus id="AF5430A040609C41FF1966D2FE8BA9F8" box="[151,258,475,501]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
labelled “Iran, Fars 1997 (15), 51 road-km W Shiraz, Chehel Chesmeh,
<quantity id="B71723E740609C41FB9266D2FBF8A9F8" box="[1052,1137,475,501]" metricMagnitude="3" metricUnit="m" metricValue="1.94" pageId="30" pageNumber="31" unit="m" value="1940.0">
<elevation id="FBC2693140609C41FB9266D2FBF8A9F8" box="[1052,1137,475,501]" metricMagnitude="3" metricUnit="m" metricValue="1.94" pageId="30" pageNumber="31" unit="m" value="1940.0">1940 m</elevation>
</quantity>
, 17.9.; leg.
<collectorName id="DD1AEBD440609C41FB6166D2FAB4A9F8" box="[1263,1341,475,501]" pageId="30" pageNumber="31">Schödl</collectorName>
“, NHM Wien
</materialsCitation>
;
<specimenCount id="66E9458B40609C41FF6F66F6FECDAA14" box="[225,324,511,537]" pageId="30" pageNumber="31" type="worker">9 worker</specimenCount>
<typeStatus id="AF5430A040609C41FEC466F6FE3CAA14" box="[330,437,511,537]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
from the same sample in SMN Goerlitz.
<materialsCitation id="C087845F40609C41FCF266F6FAA6AA30" collectorName="Schulz" country="Turkey" location="Artvin" pageId="30" pageNumber="31" specimenCount="3" specimenCount-worker="3" stateProvince="Artvin" typeStatus="paratype">
<specimenCount id="66E9458B40609C41FCF266F6FC56AA14" box="[892,991,511,537]" pageId="30" pageNumber="31" type="worker">3 worker</specimenCount>
<typeStatus id="AF5430A040609C41FC6866F6FBD8AA14" box="[998,1105,511,537]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
labelled “
<collectingRegion id="B22B40E040609C41FB3166F6FAA3AA14" box="[1215,1322,511,537]" country="Turkey" name="Artvin" pageId="30" pageNumber="31">ARTVIN</collectingRegion>
—Taslica,
<quantity id="B71723E740609C41FF19652AFF57AA30" box="[151,222,547,573]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="km" value="10.0">10 km</quantity>
W
<location id="7530D8D940609C41FE8A652AFED9AA30" LSID="urn:lsid:plazi:treatment:F8463F1440609C41FF19669AFB56AF24:7530D8D940609C41FE8A652AFED9AA30" box="[260,336,547,573]" country="Turkey" name="Artvin" pageId="30" pageNumber="31" stateProvince="Artvin">Artvin</location>
, 1000 mH, Laubmischwald, Steinmauer, leg.
<collectorName id="DD1AEBD440609C41FCC1652AFC12AA30" box="[847,923,547,573]" pageId="30" pageNumber="31">Schulz</collectorName>
02.07.89
<collectingCountry id="08F8CE9240609C41FB85652DFBFEAA32" box="[1035,1143,548,575]" name="Turkey" pageId="30" pageNumber="31">TÜRKEI</collectingCountry>
”, SMN Goerlitz
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="38F5DD8940609C41FF196563FC12AB18" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="70508E0240609C41FF196563FF36AAC0" blockId="30.[151,1437,618,1833]" pageId="30" pageNumber="31">
<emphasis id="429B521040609C41FF196563FE2DAA89" bold="true" box="[151,420,618,644]" pageId="30" pageNumber="31">All material examined</emphasis>
.
<materialsCitation id="C087845F40609C41FE3D6562FB2DAAA7" country="Afghanistan" location="For" municipality="This" pageId="30" pageNumber="31" specimenCount="13" specimenCount-worker="13">
Numeric phenotypical data were available in 4 samples with
<specimenCount id="66E9458B40609C41FBE46562FB65AA88" box="[1130,1260,619,645]" pageId="30" pageNumber="31" type="worker">13 workers</specimenCount>
.
<location id="7530D8D940609C41FB766562FA96AA88" LSID="urn:lsid:plazi:treatment:F8463F1440609C41FF19669AFB56AF24:7530D8D940609C41FB766562FA96AA88" box="[1272,1311,619,645]" country="Afghanistan" municipality="This" name="For" pageId="30" pageNumber="31">For</location>
details see supplementary information SI1, SI2.
<collectingMunicipality id="9034147840609C41FDB16586FDF8AAA4" box="[575,625,655,681]" pageId="30" pageNumber="31">This</collectingMunicipality>
material originated from
<collectingCountry id="08F8CE9240609C41FC196586FBA9AAA4" box="[919,1056,655,681]" name="Afghanistan" pageId="30" pageNumber="31">Afghanistan</collectingCountry>
(1 sample)
</materialsCitation>
,
<materialsCitation id="C087845F40609C41FB3C6586FABCAAA4" box="[1202,1333,655,681]" country="Georgia" isTailEnumeration="true" location="For" municipality="This" pageId="30" pageNumber="31" specimenCount="1">
<collectingCountry id="08F8CE9240609C41FB3C6586FA85AAA4" box="[1202,1292,655,681]" name="Georgia" pageId="30" pageNumber="31">Georgia</collectingCountry>
(1)
</materialsCitation>
and
<collectingCountry id="08F8CE9240609C41FAE16586FA15AAA4" box="[1391,1436,655,681]" name="Iran" pageId="30" pageNumber="31">Iran</collectingCountry>
(2).
</paragraph>
<paragraph id="70508E0240609C41FF1965DFFC12AB18" blockId="30.[151,1437,618,1833]" pageId="30" pageNumber="31">
<emphasis id="429B521040609C41FF1965DFFEF9AAFC" bold="true" box="[151,368,726,753]" pageId="30" pageNumber="31">Geographic range</emphasis>
. From S
<collectingCountry id="08F8CE9240609C41FE5265DEFDBFAAFC" box="[476,566,727,753]" name="Georgia" pageId="30" pageNumber="31">Georgia</collectingCountry>
and NE
<collectingCountry id="08F8CE9240609C41FD1265DEFD65AAFC" box="[668,748,727,753]" name="Turkey" pageId="30" pageNumber="31">Turkey</collectingCountry>
(41.7°E) eastwards over the
<collectingCountry id="08F8CE9240609C41FBB765DEFBEFAAFC" box="[1081,1126,727,753]" name="Iran" pageId="30" pageNumber="31">Iran</collectingCountry>
to E
<collectingCountry id="08F8CE9240609C41FB2A65DEFAA4AAFC" box="[1188,1325,727,753]" name="Afghanistan" pageId="30" pageNumber="31">Afghanistan</collectingCountry>
(69.5°E). The altitudinal range extends from
<quantity id="B71723E740609C41FDAF65F3FDC5AB18" box="[545,588,762,789]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="m" value="5.0">
<elevation id="FBC2693140609C41FDAF65F3FDC5AB18" box="[545,588,762,789]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="m" value="5.0">5 m</elevation>
</quantity>
in
<collectingCountry id="08F8CE9240609C41FDFE65F2FD43AB18" box="[624,714,763,789]" name="Georgia" pageId="30" pageNumber="31">Georgia</collectingCountry>
to
<quantity id="B71723E740609C41FD6365F2FCCBAB18" box="[749,834,763,789]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="30" pageNumber="31" unit="m" value="2000.0">
<elevation id="FBC2693140609C41FD6365F2FCCBAB18" box="[749,834,763,789]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="30" pageNumber="31" unit="m" value="2000.0">2000 m</elevation>
</quantity>
in
<collectingCountry id="08F8CE9240609C41FCE865F2FC1EAB18" box="[870,919,763,789]" name="Iran" pageId="30" pageNumber="31">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="38F5DD8940609C41FF196417FC6BAF08" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="70508E0240609C41FF196417FC4AAE5C" blockId="30.[151,1437,618,1833]" pageId="30" pageNumber="31">
<emphasis id="429B521040609C41FF196417FE83AB35" bold="true" box="[151,266,798,824]" pageId="30" pageNumber="31">Diagnosis</emphasis>
: --Worker (
<tableCitation id="3D6DBBB940609C41FE1E6416FE50AB34" box="[400,473,799,825]" captionStart="TABLE 1" captionStartId="31.[152,244,151,175]" captionTargetPageId="31" captionText="TABLE 1. Measurements of workers of the Cardiocondyla elegans group. Data are given in the sequence arithmetic mean ± standard deviation [minimum, maximum]. For characters with fewer data than indicated in the head of the columns, the number of measurements is given after the square brackets." httpUri="http://table.plazi.org/id/2490DE8A40619C40FF16679EFC98A8FA" pageId="30" pageNumber="31" tableUuid="2490DE8A40619C40FF16679EFC98A8FA">Tab. 1</tableCitation>
,
<figureCitation id="E8D4928740609C41FE666416FDE6AB34" box="[488,623,799,825]" captionStart="FIGURES 1821" captionStartId="12.[151,264,1970,1994]" captionTargetBox="[261,1327,1147,1946]" captionTargetId="figure-92@12.[261,1327,1147,1946]" captionTargetPageId="12" captionText="FIGURES 1821. Cardiocondyla brachyceps; Fig. 18: head in dorsal view; Fig. 19: lateral view; Fig. 20: dorsal view; Fig. 21: head surface between inner eye margin and paramedian vertex. Iran: Fars, 1997.09.19" figureDoi="http://doi.org/10.5281/zenodo.7888210" httpUri="https://zenodo.org/record/7888210/files/figure.png" pageId="30" pageNumber="31">Figs. 1821</figureCitation>
, key; pictures CASENT0919731 in www.antweb.org). Relatively large, CS 559 µm. Head rather short, CL/CW 1.131. Postocular distance very low, PoOc/CL 0.380. Scape rather long, SL/CS 0.831. Eye rather large, EYE/CS 0.251. Occipital margin suggestively concave. Frons rather broad (FRS/ CS 0.257), frontal carinae slightly converging immediately caudal of FRS level (FL/FR 1.044). Dorsal profile of promesonotum and of propodeum convex with a deep metanotal depression (MGr/CS 4.39 %). Propodeal spines slightly longer and slightly thinner than in related species (SP/CS 0.128), their axis in profile deviating by about 35° from longitudinal axis of mesosoma, their bases more approached (SPBA/CS 0.237). Petiole narrower than in
<taxonomicName id="B7EFF58140609C41FA0D64FEFF64AC38" baseAuthorityName="Seifert" baseAuthorityYear="2018" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="63" phylum="Arthropoda" rank="species" species="elegans">
C.
<emphasis id="429B521040609C41FF196312FF64AC38" box="[151,237,1051,1077]" italics="true" pageId="30" pageNumber="31">elegans</emphasis>
</taxonomicName>
and as high as wide (PeW/CS 0.299, PeH/CS 0.301); in profile with a long peduncle that is about 1.8fold as long as wide and an extremely steep anterior slope of the node (about 85° relative to ventral profile). Postpetiole much narrower and lower than in related species (PpW/CS 0.522, PpH/CS 0.260), in dorsal view heard-shaped, with a concave anterior margin and convex sides; postpetiolar sternite with the anteromedian portion significantly more bulging than its anteroparamedian portion; in lateral view this anteromedian bulge forms a small, obtusely-angled, rounded corner and changes into the helcium with a distinct angle. Frontal laminae, paramedian vertex, and lateral clypeus with weak longitudinal sculpture. Vertex with strongly demarcated, bicoronate foveolae of 1821 µm diameter; interspaces narrower than foveolar diameter, near to eyes larger; the interspaces between foveolae shiny and in places with fragments of a very delicate microreticulum. Mesosoma on most of its surface shining and with very fine microreticulum. Dorsal promesonotum finely longitudinally rugulose and with scattered bicoronate foveolae. Lateral meso- and metapleurae delicately longitudinally striate-reticulate; at lower magnifications appearing duller, contrasting the shining lateral parts of mesonotum and pronotum. Dorsum of waist glabrous except for a very delicate microreticulum. First gaster tergite glabrous. Pubescence on whole body long but less dense than in related species, PLG/CS 7.96 %, sqPDG 4.65. Color variable: head medium brown to blackish brown; mesosoma and petiole usually light orange-brown, in specimens with to dark brown mesosoma an orange-yellowish color component is still visible; postpetiole and gaster dark to blackish brown.
</paragraph>
<paragraph id="70508E0240609C41FF196153FC6BAF08" blockId="30.[151,1437,618,1833]" pageId="30" pageNumber="31">
<emphasis id="429B521040609C41FF196153FD16AE78" bold="true" box="[151,671,1626,1653]" pageId="30" pageNumber="31">Taxonomic comments and clustering results</emphasis>
.
<taxonomicName id="B7EFF58140609C41FD226152FC50AE78" authorityName="Seifert" authorityYear="2003" box="[684,985,1627,1653]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="brachyceps">
<emphasis id="429B521040609C41FD226152FC50AE78" box="[684,985,1627,1653]" italics="true" pageId="30" pageNumber="31">Cardiocondyla brachyceps</emphasis>
</taxonomicName>
is obviously a member of the
<taxonomicName id="B7EFF58140609C41FAA96152FA14AE78" baseAuthorityName="Seifert" baseAuthorityYear="2018" box="[1319,1437,1627,1653]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="429B521040609C41FAA96152FA14AE78" box="[1319,1437,1627,1653]" italics="true" pageId="30" pageNumber="31">C. elegans</emphasis>
</taxonomicName>
species group but differs from
<emphasis id="429B521040609C41FE7B6176FC84AE94" box="[501,781,1663,1689]" italics="true" pageId="30" pageNumber="31">
<taxonomicName id="B7EFF58140609C41FE7B6176FDE7AE94" baseAuthorityName="Seifert" baseAuthorityYear="2018" box="[501,622,1663,1689]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="elegans">C. elegans</taxonomicName>
,
<taxonomicName id="B7EFF58140609C41FDF56176FC84AE94" authorityName="Soudek" authorityYear="1925" box="[635,781,1663,1689]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="dalmatica">C. dalmatica</taxonomicName>
</emphasis>
and
<taxonomicName id="B7EFF58140609C41FCCB6176FB8DAE94" authority="Seifert, 2023" authorityName="Seifert" authorityYear="2023" box="[837,1028,1663,1689]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="dalmaticoides" status="sp. nov.">
<emphasis id="429B521040609C41FCCB6176FB8DAE94" box="[837,1028,1663,1689]" italics="true" pageId="30" pageNumber="31">C. dalmaticoides</emphasis>
</taxonomicName>
<emphasis id="429B521040609C41FB826176FBC5AE94" bold="true" box="[1036,1100,1663,1689]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="59A8EF6B40609C41FB826176FBC5AE94" box="[1036,1100,1663,1689]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by much narrower and lower waist segments, more dilute pubescence and the differing petiole profile (compare
<figureCitation id="E8D4928740609C41FBED61AAFB3CAEB0" box="[1123,1205,1699,1725]" captionStart="FIGURES 69" captionStartId="11.[151,264,1045,1069]" captionTargetBox="[234,1354,181,1021]" captionTargetId="figure-15@11.[234,1354,181,1021]" captionTargetPageId="11" captionText="FIGURES 69. Cardiocondyla elegans; Fig. 6: head in dorsal view; Fig. 7 lateral view; Fig. 8: dorsal view; Fig. 9: head surface between inner eye margin and paramedian vertex. France: Brehemont-1.1 km E, 2010.07.15." figureDoi="http://doi.org/10.5281/zenodo.7888202" httpUri="https://zenodo.org/record/7888202/files/figure.png" pageId="30" pageNumber="31">Figs. 7</figureCitation>
,
<figureCitation id="E8D4928740609C41FB4861AAFB6AAEB0" box="[1222,1251,1699,1725]" captionStart="FIGURES 1013" captionStartId="11.[151,264,1973,1997]" captionTargetBox="[247,1341,1129,1950]" captionTargetId="figure-52@11.[247,1341,1129,1950]" captionTargetPageId="11" captionText="FIGURES 1013. Cardiocondyla dalmatica, Fig. 10: head in dorsal view; Fig. 11: lateral view; Fig. 12: dorsal view; Fig. 13: head surface between inner eye margin and paramedian vertex. Hungary: Balassagyarmat, 2007.06.31" figureDoi="http://doi.org/10.5281/zenodo.7888204" httpUri="https://zenodo.org/record/7888204/files/figure.png" pageId="30" pageNumber="31">11</figureCitation>
,
<figureCitation id="E8D4928740609C41FB7D61ABFA86AEB0" box="[1267,1295,1698,1725]" captionStart="FIGURES 1417" captionStartId="12.[151,264,1027,1051]" captionTargetBox="[246,1342,181,1003]" captionTargetId="figure-31@12.[246,1342,181,1003]" captionTargetPageId="12" captionText="FIGURES 1417. Cardiocondyla dalmaticoides n. sp., holotype; Fig. 14: head in dorsal view; Fig. 15: lateral view, postpetiole and gaster in full lateral view shown upper left; Fig. 16: dorsal view, postpetiole and gaster in full dorsal view shown lower left; Fig. 17: head surface between inner eye margin and paramedian vertex. Turkey: Reyhauli-2 km N, 1993.06.09" figureDoi="http://doi.org/10.5281/zenodo.7888206" httpUri="https://zenodo.org/record/7888206/files/figure.png" pageId="30" pageNumber="31">15</figureCitation>
and
<figureCitation id="E8D4928740609C41FAC361AAFAE5AEB0" box="[1357,1388,1699,1725]" captionStart="FIGURES 1821" captionStartId="12.[151,264,1970,1994]" captionTargetBox="[261,1327,1147,1946]" captionTargetId="figure-92@12.[261,1327,1147,1946]" captionTargetPageId="12" captionText="FIGURES 1821. Cardiocondyla brachyceps; Fig. 18: head in dorsal view; Fig. 19: lateral view; Fig. 20: dorsal view; Fig. 21: head surface between inner eye margin and paramedian vertex. Iran: Fars, 1997.09.19" figureDoi="http://doi.org/10.5281/zenodo.7888210" httpUri="https://zenodo.org/record/7888210/files/figure.png" pageId="30" pageNumber="31">19</figureCitation>
). In numeric terms,
<taxonomicName id="B7EFF58140609C41FECA61CEFE6DAEEC" authorityName="Seifert" authorityYear="2003" box="[324,484,1735,1761]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="brachyceps">
<emphasis id="429B521040609C41FECA61CEFE6DAEEC" box="[324,484,1735,1761]" italics="true" pageId="30" pageNumber="31">C. brachyceps</emphasis>
</taxonomicName>
is strongly separable from the other three species by a PCA of the characters PpH/CS, dFOV, SL/CS, PoOc/CL, sqPDG, SP/CS, PLG/CS and PeW/CS (
<figureCitation id="E8D4928740609C41FCFD61E2FC5FAF08" box="[883,982,1770,1797]" captionStart="FIGURE 135" captionStartId="55.[151,250,1048,1072]" captionTargetBox="[246,1345,193,1014]" captionTargetId="figure-15@55.[234,1354,181,1024]" captionTargetPageId="55" captionText="FIGURE 135. Separation of nest samples of Cardiocondyla brachyceps (black rhombs, n=4), C. dalmatica (black dots, n=32), C. dalmaticoides n. sp. (crosses, n=3) and C. elegans (white dots, n=50) by a principal component analysis considering the morphometric characters PpH/CS, dFOV, SL/CS, PoOc/CL, sqPDG, SP/CS, PLG/CS and PeW/CS." figureDoi="http://doi.org/10.5281/zenodo.7888309" httpUri="https://zenodo.org/record/7888309/files/figure.png" pageId="30" pageNumber="31">Fig. 135</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="38F5DD8940609C41FF196007FB56AF24" box="[151,1247,1806,1833]" pageId="30" pageNumber="31" type="biology_ecology">
<paragraph id="70508E0240609C41FF196007FB56AF24" blockId="30.[151,1437,618,1833]" box="[151,1247,1806,1833]" pageId="30" pageNumber="31">
<emphasis id="429B521040609C41FF196007FF7EAF25" bold="true" box="[151,247,1806,1832]" pageId="30" pageNumber="31">Biology.</emphasis>
The nest sample from
<collectingCountry id="08F8CE9240609C41FE766006FDDBAF24" box="[504,594,1807,1833]" name="Georgia" pageId="30" pageNumber="31">Georgia</collectingCountry>
was collected on a sandy riverbank with scattered bushes.
</paragraph>
</subSubSection>
</treatment>
</document>