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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.357.5861" ID-GBIF-Dataset="e2125835-030a-4f84-b1e6-8e03cb32fe3e" ID-PMC="PMC3867175" ID-Pensoft-Pub="1313-2970-359-53" ID-PubMed="24363589" ID-ZBK="B3B8D706F2FB45F5B20EEE93BC611AD5" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-359-53" ModsDocOrigin="ZooKeys 359" ModsDocTitle="A taxonomic revision of the silphaeformis species-group of the genus Tachinus Gravenhorst (Staphylinidae, Tachyporinae) from China" checkinTime="1451246595444" checkinUser="pensoft" docAuthor="Feng, Ting, Schuelke, Michael &amp; Li, Li-Zhen" docDate="2013" docId="C99C3DC76484FF0B932F5E3A658D0B7B" docLanguage="en" docName="ZooKeys 359: 53-99" docOrigin="ZooKeys 359" docSource="http://dx.doi.org/10.3897/zookeys.357.5861" docTitle="Tachinus hercules Feng, Li &amp; Schuelke, sp. n." docType="treatment" docUuid="C7AF0BFD-AD27-40BE-AC4A-CCF9A9AE7FB2" docUuidSource="ZooBank" docVersion="4" lastPageNumber="60" masterDocId="FFA0664AFFF6FF852724FF83B7335F5B" masterDocTitle="A taxonomic revision of the silphaeformis species-group of the genus Tachinus Gravenhorst (Staphylinidae, Tachyporinae) from China" masterLastPageNumber="99" masterPageNumber="53" pageNumber="59" updateTime="1668157111880" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A taxonomic revision of the silphaeformis species-group of the genus Tachinus Gravenhorst (Staphylinidae, Tachyporinae) from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Feng, Ting</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schuelke, Michael</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Li-Zhen</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>359</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>53</mods:start>
<mods:end>99</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.357.5861</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.357.5861</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-359-53</mods:identifier>
<mods:identifier type="ZBK">B3B8D706F2FB45F5B20EEE93BC611AD5</mods:identifier>
<mods:identifier type="ZooBank">B3B8D706F2FB45F5B20EEE93BC611AD5</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152049982" LSID="urn:lsid:zoobank.org:act:C7AF0BFD-AD27-40BE-AC4A-CCF9A9AE7FB2" httpUri="http://treatment.plazi.org/id/C99C3DC76484FF0B932F5E3A658D0B7B" lastPageId="7" lastPageNumber="60" pageId="6" pageNumber="59">
<subSubSection pageId="6" pageNumber="59" type="nomenclature">
<paragraph pageId="6" pageNumber="59">
<taxonomicName LSID="http://zoobank.org/C7AF0BFD-AD27-40BE-AC4A-CCF9A9AE7FB2" authority="Feng, Li &amp; Schuelke" class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus hercules" order="Coleoptera" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="hercules">
Tachinus (s. str.) hercules Feng, Li &amp;
<normalizedToken originalValue="Schülke">Schuelke</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="59">sp. n.</taxonomicNameLabel>
Figs 2C, 2D, 9D, 10D, 11D, 13D, 15D, 18D, 20D, 21D
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="59" type="type locality">
<paragraph pageId="6" pageNumber="59">Type locality.</paragraph>
<paragraph pageId="6" pageNumber="59">China, Sichuan Province, Songpan, Huanglongsi.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="59" type="type material">
<paragraph pageId="6" pageNumber="59">Type material.</paragraph>
<paragraph pageId="6" pageNumber="59">
Holotype: ♂, CHINA: Sichuan Prov., Aba A. R., Songpan County, Huanglongsi, 24.vii.2001, LI &amp; ZHAO leg. (SNUC). Paratypes: 25 ♂♂, 29 ♀♀, same label data as holotype (SNUC); 7 ♂♂, 10 ♀♀, CHINA: Sichuan Prov., pass between Songpan and Nanping,
<normalizedToken originalValue="Eside">E-side</normalizedToken>
, 3,450-3,500 m, 21.vi.2002, S. Murzin &amp; I. Shokhin leg. (cSch); ♂, ♀, CHINA: Sichuan Prov., Zhangla env., 4,200-4,700 m, 9-11.vii.1991, J.
<normalizedToken originalValue="Kaláb">Kalab</normalizedToken>
leg. (NHMB, cSch) [labelled as paratypes of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus szechuanensis" order="Coleoptera" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="szechuanensis">Tachinus szechuanensis</taxonomicName>
Campbell in litteris]; ♂, CHINA: Sichuan Prov., Songpan, 2,000 m,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="32.5">32°30'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="103.666664">103°40'E</geoCoordinate>
, 13-17.vii.1990, J.
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
leg. (NHMB); 3 ♂♂, 1 ♀, CHINA: Sichuan Prov., Min Shan,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="33.166668">33°10'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="103.833336">103°50'E</geoCoordinate>
, 2,500-4,500 m, 14-16.vii.1990, J.
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
leg. (NHMB); 5 ♂♂, 1 ♀, CHINA: Sichuan Prov., pass between Pingwu and Jiuzhaigou, 3,000 m, 10-15.vii.2005, V. Patrikeev leg. (cSch); 1 ♂, 3 ♀♀, CHINA:
<normalizedToken originalValue="NSichuan">N-Sichuan</normalizedToken>
[CH12-23] Min Shan, 17 km NE Songpan, E pass,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="32.739723">32°44'23&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="103.74194">103°44'31&quot;E</geoCoordinate>
, 3920 m,
<normalizedToken originalValue="Nslope">N-slope</normalizedToken>
with
<taxonomicName class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhododendron" order="Ericales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="genus">Rhododendron</taxonomicName>
below rocks, litter, moss, and grass roots sifted, 10.viii.2012, leg. M.
<normalizedToken originalValue="Schülke">Schuelke</normalizedToken>
(cSch); 3 ♂♂, 3 ♀♀, CHINA:
<normalizedToken originalValue="NSichuan">N-Sichuan</normalizedToken>
[CH12-24] pass 35 km NNW Songpan,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="32.925556">32°55'32&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="103.43222">103°25'56&quot;E</geoCoordinate>
, 3600 m, moist
<normalizedToken originalValue="Nslope">N-slope</normalizedToken>
with Salix and other shrubs, litter, grass roots, and moss sifted, 11.viii.2012, leg. M.
<normalizedToken originalValue="Schülke">Schuelke</normalizedToken>
(cSch); ♀, CHINA:
<normalizedToken originalValue="NSichuan">N-Sichuan</normalizedToken>
[CH12-19] 47 km N Songpan, road S 301 km 118, N Gongangling pass,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.054165">33°03'15&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="103.72667">103°43'36&quot;E</geoCoordinate>
, 3390 m, spruce forest with shrubs, litter, moss, and mushrooms sifted, 9.viii.2012, leg. M.
<normalizedToken originalValue="Schülke">Schuelke</normalizedToken>
(cSch); ♂♀, CHINA: [19]
<normalizedToken originalValue="NSichuan">N-Sichuan</normalizedToken>
N Songpan,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.054165">33°03'15&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="103.72667">103°43'36&quot;E</geoCoordinate>
, 3390 m, spruce forest, sifted, 9.viii.2012, V. Assing (cAss); ♀, CHINA: [22]
<normalizedToken originalValue="NSichuan">N-Sichuan</normalizedToken>
pass ENE Songpan, 4080 m,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="32.748333">32°44'54&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="103.728615">103°43'43&quot;E</geoCoordinate>
, sifted. 10.viii.2012, V. Assing (cAss).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="60" type="description">
<paragraph pageId="7" pageNumber="60">
<pageBreakToken pageId="7" pageNumber="60" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="7" pageNumber="60">
Measurements of males (holotype): BL 3.61-4.00 (4.00); FL 2.84-3.11 (3.11); PL 0.89-1.00 (1.00); EL 1.45-1.61 (1.61); SEL 0.06-0.11 (0.11); HW 0.88-0.90 (0.89); PW 1.45-1.56 (1.56); EW 1.61-1.78 (1.78); relative length of antennomeres
<normalizedToken originalValue="IXI">I-XI</normalizedToken>
: 23: 15: 15: 10: 17: 14: 16: 15: 15: 14: 29. Measurements of females: BL 3.95-4.39; FL 3.39-3.61 mm; PL 0.99-1.01; EL 1.56-1.73; SEL 0.16-0.18; HW 0.94-0.96; PW 1.50-1.56; EW 1.82-1.84; relative length of antennomeres
<normalizedToken originalValue="IXI">I-XI</normalizedToken>
: 23: 15: 15: 10: 14: 14: 14: 13: 14: 15: 28.
</paragraph>
<paragraph pageId="7" pageNumber="60">Body (Figs 2C, 2D) dark brown to black, head black, mouthparts, basal four antennomeres, elytra, posterior margin of each abdominal tergite and legs reddish brown.</paragraph>
<paragraph pageId="7" pageNumber="60">Head shorter than wide and narrower than pronotum, HW: PW = 0.56-0.64 (0.57), disc of head with dense microsculpture consisting of irregular striae, basal part with microsculpture consisting of transverse meshes, punctation fine and sparse. Antennae moderately short, antennomeres X shorter than wide.</paragraph>
<paragraph pageId="7" pageNumber="60">Pronotum: PL: PW = 0.57-0.69 (0.64), widest at basal third; surface with punctation slightly more distinct than that of head, microsculpture consisting of irregular transverse striae.</paragraph>
<paragraph pageId="7" pageNumber="60">Elytra elongate: EL: PL = 1.45-1.81 (1.60), EL: EW = 0.81-1.00 (0.90), EW: PW = 1.03-1.23 (1.14) in males; EL: PL = 1.54-1.75, EL: EW = 0.85-0.95, EW: PW = 1.17-1.23 in females. Microsculpture consisting of fine, short meshes, punctation coarser and denser than that of head and pronotum.</paragraph>
<paragraph pageId="7" pageNumber="60">Surface of abdomen evenly pubescent and punctate, microsculpture consisting of transverse lines.</paragraph>
<paragraph pageId="7" pageNumber="60">Male. Sternite VII (Figs 9D, 10D) with posterior margin broadly and deeply emarginate, apical margin bent ventrad, with moderately broad area of coarse granules. Tergite VIII (Fig. 11D) with four short lobes, median lobes slightly longer than lateral ones. Sternite VIII as in Fig. 13D. Median lobe of aedeagus (Fig. 15D) broad and with two apical projections, parameres long and directed latero-apicad.</paragraph>
<paragraph pageId="7" pageNumber="60">Female. Antennomeres X only slightly shorter than wide (length: width = 0.93). Pronotum with microsculpture more distinct and denser than in male. Elytra slightly longer than in male; apical margin broadly rounded, with microsculpture more distinct, denser and with shorter meshes than in male. Tergite VIII (Fig. 18D) with lobes almost fused, apical margin of median lobe broadly and triangularly projecting beyond lateral lobes; lateral lobes each with one long seta. Sternite VIII (Fig. 20D) with six lobes, fimbriate median lobes nearly fused, with shallow emargination at middle, sublateral lobes as long as median ones.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="60" type="etymology">
<paragraph pageId="7" pageNumber="60">Etymology.</paragraph>
<paragraph pageId="7" pageNumber="60">The species is named after the Greek divine hero Heracles, whose Roman name is Hercules.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="60" type="remarks">
<paragraph pageId="7" pageNumber="60">Remarks.</paragraph>
<paragraph pageId="7" pageNumber="60">
Males of the new species are similar to those of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus parahercules" order="Coleoptera" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="parahercules">Tachinus parahercules</taxonomicName>
externally, and can be separated only by aedeagal characters, they differ from the other species by the shape of the aedeagal median lobe (two apical projections, long parameres directed lateroapically). Females are distinguished by the shape of the apical margin of tergite VIII, from females of the similar
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus hujiayaoi" order="Coleoptera" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="hujiayaoi">Tachinus hujiayaoi</taxonomicName>
and
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus coronatus" order="Coleoptera" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="coronatus">Tachinus coronatus</taxonomicName>
by the presence of a single pair of long setae on the lateral lobes of tergite VIII.
</paragraph>
</subSubSection>
</treatment>
</document>