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<mods:title id="97303EB666EA4E9369A1873AF65C03E4">Redescriptions of three species of freshwater crabs from Angola, southern Africa (Brachyura: Potamoidea: Potamonautidae)</mods:title>
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<mods:namePart id="A049160F4D24A3EB698054413E0556B7">Ahles, Katelyn M.</mods:namePart>
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<emphasis id="7DBDEA926044FFCE6B83FBEAFE63FB97" bold="true" box="[151,422,1050,1077]" italics="true" pageId="1" pageNumber="517">Potamonautes adeleae</emphasis>
<bibRefCitation id="2B584B716044FFCE6ABAFBEAFDE8FB96" author="Bott, R." box="[430,557,1050,1077]" pageId="1" pageNumber="517" pagination="165 - 172" refId="ref9155" refString="Bott, R. (1968) Decapoden aus dem Museu do Dundo (Crust. Decap.). Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 77, 165 - 172." type="journal article" year="1968">Bott, 1968</bibRefCitation>
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<paragraph id="4F7636806044FFCE6B83FBCEFE68FBFB" blockId="1.[151,602,1050,1149]" box="[151,429,1087,1113]" pageId="1" pageNumber="517">Adele’s Freshwater Crab</paragraph>
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(
<figureCitation id="D7F22A056044FFCE6B8BFB92FF28FBDF" box="[159,237,1123,1149]" captionStart="FIGURE 1" captionStartId="7.[151,250,1965,1989]" captionTargetBox="[191,1397,179,1938]" captionTargetId="figure-21@7.[191,1397,179,1938]" captionTargetPageId="7" captionText="FIGURE 1. Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 10.17 mm, B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="1" pageNumber="517">Figs. 1</figureCitation>
, 
<figureCitation id="D7F22A056044FFCE6BEFFB92FEDAFBDF" box="[251,287,1123,1149]" captionStart="FIGURE 4" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[152,1436,309,1723]" captionTargetId="figure-93@10.[152,1436,309,1723]" captionTargetPageId="10" captionText="FIGURE 4. Frontal view of carapace. (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, paratype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph B by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 10.4 mm; B = 5.7 mm; C = 4.7 mm." figureDoi="http://doi.org/10.5281/zenodo.5497107" httpUri="https://zenodo.org/record/5497107/files/figure.png" pageId="1" pageNumber="517">4A</figureCitation>
, 
<figureCitation id="D7F22A056044FFCE6A3FFB93FE70FBDF" box="[299,437,1122,1149]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="1" pageNumber="517">5A, B, G, H</figureCitation>
, 
<figureCitation id="D7F22A056044FFCE6AD4FB92FDCDFBDF" box="[448,520,1123,1149]" captionStart="FIGURE 6" captionStartId="12.[151,250,1828,1852]" captionTargetBox="[197,1391,181,1805]" captionTargetId="figure-17@12.[197,1391,181,1805]" captionTargetPageId="12" captionText="FIGURE 6. Right G1, ventral view (A–C) and dorsal view (D, F). (A, D) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B, E) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photo- graphs B and E by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A= 6.7 mm; B = 2.3 mm; C = 1.13 mm; D = 2.7 mm; E = 2.3 mm; F = 1.2." figureDoi="http://doi.org/10.5281/zenodo.5497111" httpUri="https://zenodo.org/record/5497111/files/figure.png" pageId="1" pageNumber="517">6A, D</figureCitation>
, 
<figureCitation id="D7F22A056044FFCE6907FB92FDFDFBDF" box="[531,568,1123,1149]" captionStart="FIGURE 7" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[264,1323,181,1831]" captionTargetId="figure-21@13.[264,1323,181,1831]" captionTargetPageId="13" captionText="FIGURE 7. Right G2 ventral view (A–C); (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph C by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 5.6 mm; B = 2.3 mm; C = 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5497113" httpUri="https://zenodo.org/record/5497113/files/figure.png" pageId="1" pageNumber="517">7A</figureCitation>
, 
<figureCitation id="D7F22A056044FFCE6957FB92FD94FBDF" box="[579,593,1123,1149]" captionStart="FIGURE 8" captionStartId="14.[151,250,1336,1360]" captionTargetBox="[159,1428,189,1302]" captionTargetId="figure-17@14.[151,1436,181,1312]" captionTargetPageId="14" captionText="FIGURE 8. Map of the Angola showing the known localities of Potamonautes adeleae (black square, not shown to scale), P. kensleyi (black circles, not shown to scale), and P. macrobrachii (black diamond, not shown to scale). Scale bar = 520 km. See text for exact locality coordinates." figureDoi="http://doi.org/10.5281/zenodo.5497115" httpUri="https://zenodo.org/record/5497115/files/figure.png" pageId="1" pageNumber="517">8</figureCitation>
)
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<treatmentCitation id="CE6810916044FFCE6B83FB59FC1CFB1D" author="Bott, R." box="[151,985,1191,1215]" page="169" pageId="1" pageNumber="517" year="1968">
<taxonomicName id="88C94D036044FFCE6B83FB59FC1CFB1D" ID-CoL="6VVMS" authority="Bott, 1968: 168 - 169" authorityName="Bott" authorityPageNumber="168 - 169" authorityYear="1968" box="[151,985,1191,1215]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="517" phylum="Arthropoda" rank="subSpecies" species="lirrangensis" subGenus="Lirrangopotamonautes" subSpecies="adeleae">
<emphasis id="7DBDEA926044FFCE6B83FB59FD3AFB1D" box="[151,767,1191,1215]" italics="true" pageId="1" pageNumber="517">Potamonautes (Lirrangopotamonautes) lirrangensis adeleae</emphasis>
<bibRefCitation id="2B584B716044FFCE6811FB59FC1CFB1D" author="Bott, R." box="[773,985,1191,1215]" pageId="1" pageNumber="517" pagination="165 - 172" refId="ref9155" refString="Bott, R. (1968) Decapoden aus dem Museu do Dundo (Crust. Decap.). Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 77, 165 - 172." type="journal article" year="1968">Bott, 1968: 168–169</bibRefCitation>
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, pl. I, figs. 1–4.
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<taxonomicName id="88C94D036044FFCE6B83FB36FCE2FB7C" ID-CoL="6VVMS" authority=", Cumberlidge &amp; Tavares 2006: 248 - 249" authorityName=", Cumberlidge &amp; Tavares" authorityPageNumber="248 - 249" authorityYear="2006" box="[151,807,1222,1246]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="517" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA926044FFCE6B83FB36FE40FB7C" box="[151,389,1222,1246]" italics="true" pageId="1" pageNumber="517">Potamonautes adeleae,</emphasis>
<bibRefCitation id="2B584B716044FFCE6A99FB37FCE2FB7C" author="Cumberlidge, N. &amp; Tavares, M." box="[397,807,1222,1246]" pageId="1" pageNumber="517" pagination="248 - 257" refId="ref9684" refString="Cumberlidge, N. &amp; Tavares, M. (2006) Remarks on the freshwater crabs of Angola, southwestern Africa, with the description of Potamonautes kensleyi, new species (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 26 (2), 248 - 257. https: // doi. org / 10.1651 / s- 2653.1" type="journal article" year="2006">Cumberlidge &amp; Tavares 2006: 248–249</bibRefCitation>
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, 254–256, table 1; 
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<bibRefCitation id="2B584B716044FFCE68FAFB37FB2AFB7C" author="Cumberlidge, N." box="[1006,1263,1222,1246]" pageId="1" pageNumber="517" pagination="134068" refId="ref9459" refString="Cumberlidge, N. (2008 a) Potamonautes adeleae. The IUCN Red List of Threatened Species, 2008, e. T 134068 A 3906631. https: // doi. org / 10.2305 / IUCN. UK. 2008. RLTS. T 134068 A 3906631. en" type="journal article" year="2008">Cumberlidge 2008a: 1–7</bibRefCitation>
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, fig. 1; 
<treatmentCitation id="CE6810916044FFCE6E55FB37FEF5FB5F" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." page="170" pageId="1" pageNumber="517" year="2008">
<bibRefCitation id="2B584B716044FFCE6E55FB37FEF5FB5F" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." pageId="1" pageNumber="517" pagination="1 - 286" refId="ref10320" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyuorum: part I. An annotated checklist of extant Brachyuran crabs of the world. Raffles Bulletin of Zoology, 17, 1 - 286." type="journal article" year="2008">
Ng 
<emphasis id="7DBDEA926044FFCE6E7CFB36FA58FB7C" box="[1384,1437,1222,1246]" italics="true" pageId="1" pageNumber="517">et al.</emphasis>
2008: 170
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.
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of 
<collectingRegion id="8D0DF8626044FFCE6FBDFADAFAFEFAE7" box="[1193,1339,1323,1349]" country="Angola" name="Lunda Norte" pageId="1" pageNumber="517">Lunda Norte</collectingRegion>
, 
<location id="4A16605B6044FFCE6E51FADAFF12FACB" LSID="urn:lsid:plazi:treatment:C76087966044FFCC6B83FBEAFDC7FEC7:4A16605B6044FFCE6E51FADAFF12FACB" country="Angola" county="Adult" latitude="-8.808363" longLatPrecision="1" longitude="17.97665" municipality="Province" name="Cuango River" pageId="1" pageNumber="517" stateProvince="Lunda Norte">Cuango River</location>
, 
<location id="4A16605B6044FFCE6BF6FABEFEFBFACB" LSID="urn:lsid:plazi:treatment:C76087966044FFCC6B83FBEAFDC7FEC7:4A16605B6044FFCE6BF6FABEFEFBFACB" box="[226,318,1359,1385]" country="Angola" county="Adult" latitude="-8.808363" longLatPrecision="1" longitude="17.97665" municipality="Province" name="Cafunfo" pageId="1" pageNumber="517" stateProvince="Lunda Norte">Cafunfo</location>
(
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, 
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), collector and date unrecorded.
</materialsCitation>
</paragraph>
<paragraph id="4F7636806044FFCE6BD3FA82FE60FA5B" blockId="1.[151,1437,1322,2033]" pageId="1" pageNumber="517">
<materialsCitation id="FFA13CDD6044FFCE6BD3FA82FE64FA5B" collectionCode="SMF" country="Angola" county="Adult" latitude="-8.808363" location="Cuango River" longLatPrecision="1" longitude="17.97665" municipality="Province" pageId="1" pageNumber="517" specimenCode="SMF 4444" specimenCount="75" specimenCount-female="9" specimenCount-male="28" stateProvince="Lunda Norte" typeStatus="paratype">
<emphasis id="7DBDEA926044FFCE6BD3FA82FE83FA2F" bold="true" box="[199,326,1395,1421]" pageId="1" pageNumber="517">
<typeStatus id="907288226044FFCE6BD3FA82FE84FA2F" box="[199,321,1395,1421]" pageId="1" pageNumber="517" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="1F6F9EFB6044FFCE6A5AFA82FE0EFA2F" box="[334,459,1395,1421]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="517">SMF 4444</specimenCode>
. 
<collectingCounty id="A6174E0C6044FFCE6AC1FA82FDD0FA2F" box="[469,533,1395,1421]" pageId="1" pageNumber="517">Adult</collectingCounty>
male (CW 55.3, CL 41.1, CH 17.8, FW 
<quantity id="88319B656044FFCE68FDFA83FB97FA2F" box="[1001,1106,1394,1421]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="1" pageNumber="517" unit="mm" value="15.7">15.7 mm</quantity>
), 
<collectingCountry id="37DE76106044FFCE6F73FA82FB79FA2F" box="[1127,1212,1395,1421]" name="Angola" pageId="1" pageNumber="517">Angola</collectingCountry>
, 
<collectingMunicipality id="AF12ACFA6044FFCE6FDDFA82FAE8FA2F" box="[1225,1325,1395,1421]" pageId="1" pageNumber="517">Province</collectingMunicipality>
of 
<collectingRegion id="8D0DF8626044FFCE6E41FA82FF1FFA13" country="Angola" name="Lunda Norte" pageId="1" pageNumber="517">Lunda Norte</collectingRegion>
, 
<location id="4A16605B6044FFCE6BFCFA66FE4CFA13" LSID="urn:lsid:plazi:treatment:C76087966044FFCC6B83FBEAFDC7FEC7:4A16605B6044FFCE6BFCFA66FE4CFA13" box="[232,393,1431,1457]" country="Angola" county="Adult" latitude="-8.808363" longLatPrecision="1" longitude="17.97665" municipality="Province" name="Cuango River" pageId="1" pageNumber="517" stateProvince="Lunda Norte">Cuango River</location>
, 
<location id="4A16605B6044FFCE6A82FA66FE37FA13" LSID="urn:lsid:plazi:treatment:C76087966044FFCC6B83FBEAFDC7FEC7:4A16605B6044FFCE6A82FA66FE37FA13" box="[406,498,1431,1457]" country="Angola" county="Adult" latitude="-8.808363" longLatPrecision="1" longitude="17.97665" municipality="Province" name="Cafunfo" pageId="1" pageNumber="517" stateProvince="Lunda Norte">Cafunfo</location>
(
<geoCoordinate id="2AFD50476044FFCE6910FA66FDBFFA13" box="[516,634,1431,1457]" degrees="8.808363" direction="south" orientation="latitude" pageId="1" pageNumber="517" precision="1" value="-8.808363">-8.808363</geoCoordinate>
, 
<geoCoordinate id="2AFD50476044FFCE699CFA67FD30FA13" box="[648,757,1430,1457]" degrees="17.97665" direction="east" orientation="longitude" pageId="1" pageNumber="517" precision="1" value="17.97665">17.97665</geoCoordinate>
), collector and date unrecorded. 
<bibRefCitation id="2B584B716044FFCE6F64FA66FB30FA10" author="Bott, R." box="[1136,1269,1431,1458]" pageId="1" pageNumber="517" pagination="165 - 172" refId="ref9155" refString="Bott, R. (1968) Decapoden aus dem Museu do Dundo (Crust. Decap.). Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 77, 165 - 172." type="journal article" year="1968">Bott (1968)</bibRefCitation>
described this taxon from 
<specimenCount id="59CFFD096044FFCE6A0CFA4AFE6AFA77" box="[280,431,1467,1493]" pageId="1" pageNumber="517" type="generic">38 specimens</specimenCount>
all from the same locality, and designated the remainder of these specimens (
<specimenCount id="59CFFD096044FFCE6E1DFA4AFAA8FA77" box="[1289,1389,1467,1493]" pageId="1" pageNumber="517" type="male">28 males</specimenCount>
and 
<specimenCount id="59CFFD096044FFCE6B83FA2EFEC0FA58" box="[151,261,1503,1530]" pageId="1" pageNumber="517" type="female">9 females</specimenCount>
) as 
<typeStatus id="907288226044FFCE6A26FA2EFE64FA5B" box="[306,417,1503,1529]" pageId="1" pageNumber="517" type="paratype">paratypes</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="4F7636806044FFCE6BD3F9F3FE35F853" blockId="1.[151,1437,1322,2033]" pageId="1" pageNumber="517">
<emphasis id="7DBDEA926044FFCE6BD3F9F3FE9BF9BE" bold="true" box="[199,350,1538,1564]" pageId="1" pageNumber="517">Rediagnosis.</emphasis>
Carapace surface mostly smooth except for distinct fields of carinae at anterolateral corners; postfrontal crest distinct, complete, traversing entire carapace to meet anterolateral margins at epibranchial teeth; cervical grooves deep, long, ending just before reaching postfrontal crest; semicircular, urogastric, cardiac, branchial grooves all deep, distinct (
<figureCitation id="D7F22A056044FFCE6913F99EFDA7F92B" box="[519,610,1647,1673]" captionStart="FIGURE 1" captionStartId="7.[151,250,1965,1989]" captionTargetBox="[191,1397,179,1938]" captionTargetId="figure-21@7.[191,1397,179,1938]" captionTargetPageId="7" captionText="FIGURE 1. Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 10.17 mm, B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="1" pageNumber="517">Fig. 1A</figureCitation>
). Exorbital tooth large, pointed, directed forward, outer margin lined by small granules; epibranchial tooth small, low, blunt; anterolateral margin lined by rounded denticles; postfrontal crest, suborbital margin, vertical sulcus on branchiostegite, all granulated (
<figureCitation id="D7F22A056044FFCE68DBF946FB97F973" box="[975,1106,1719,1745]" captionStart="FIGURE 1" captionStartId="7.[151,250,1965,1989]" captionTargetBox="[191,1397,179,1938]" captionTargetId="figure-21@7.[191,1397,179,1938]" captionTargetPageId="7" captionText="FIGURE 1. Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 10.17 mm, B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="1" pageNumber="517">Figs. 1A, B</figureCitation>
). Thoracic sternal suture S3/4 incomplete, separated centrally, comprising 2 shortened sutures that begin at sides of sternum, fade towards middle (
<figureCitation id="D7F22A056044FFCE6BB4F90EFF3DF8BB" box="[160,248,1791,1817]" captionStart="FIGURE 1" captionStartId="7.[151,250,1965,1989]" captionTargetBox="[191,1397,179,1938]" captionTargetId="figure-21@7.[191,1397,179,1938]" captionTargetPageId="7" captionText="FIGURE 1. Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 10.17 mm, B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="1" pageNumber="517">Fig. 1B</figureCitation>
). Third maxilliped exopod with long flagellum, ischium with distinct vertical sulcus (
<figureCitation id="D7F22A056044FFCE6FDCF90EFAE5F8BB" box="[1224,1312,1791,1817]" captionStart="FIGURE 1" captionStartId="7.[151,250,1965,1989]" captionTargetBox="[191,1397,179,1938]" captionTargetId="figure-21@7.[191,1397,179,1938]" captionTargetPageId="7" captionText="FIGURE 1. Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 10.17 mm, B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="1" pageNumber="517">Fig. 1B</figureCitation>
). Cheliped carpus inner margin with 2 large pointed teeth, distal tooth larger than proximal tooth (
<figureCitation id="D7F22A056044FFCE6F6DF8D2FB10F89F" box="[1145,1237,1826,1853]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="1" pageNumber="517">Fig. 5H</figureCitation>
). Cheliped merus lower margin with conspicuous large pointed distal meral tooth (
<figureCitation id="D7F22A056044FFCE6893F8B6FC21F8C3" box="[903,996,1862,1889]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="1" pageNumber="517">Fig. 5G</figureCitation>
). G1 
<collectionCode id="29D8AE456044FFCE6F33F8B6FB8CF8C3" box="[1063,1097,1863,1889]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="1" pageNumber="517">TA</collectionCode>
long (
<collectionCode id="29D8AE456044FFCE6F82F8B6FB79F8C3" box="[1174,1212,1863,1889]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="1" pageNumber="517">TA</collectionCode>
/SS 0.54), midpoint angled outward at 64° to longitudinal axis of G1 SS, distal third tapering to pointed, slightly upcurved tip; G1 
<collectionCode id="29D8AE456044FFCE6E6FF89AFA58F827" box="[1403,1437,1899,1925]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="1" pageNumber="517">TA</collectionCode>
midsection widened by high mesial fold, lateral fold lower; deep longitudinal groove running along whole length of G1 
<collectionCode id="29D8AE456044FFCE6BC9F842FF3AF86F" box="[221,255,1971,1997]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="1" pageNumber="517">TA</collectionCode>
between mesial, lateral folds (
<figureCitation id="D7F22A056044FFCE6947F842FD1CF86F" box="[595,729,1971,1997]" captionStart="FIGURE 6" captionStartId="12.[151,250,1828,1852]" captionTargetBox="[197,1391,181,1805]" captionTargetId="figure-17@12.[197,1391,181,1805]" captionTargetPageId="12" captionText="FIGURE 6. Right G1, ventral view (A–C) and dorsal view (D, F). (A, D) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B, E) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photo- graphs B and E by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A= 6.7 mm; B = 2.3 mm; C = 1.13 mm; D = 2.7 mm; E = 2.3 mm; F = 1.2." figureDoi="http://doi.org/10.5281/zenodo.5497111" httpUri="https://zenodo.org/record/5497111/files/figure.png" pageId="1" pageNumber="517">Figs. 6A, D</figureCitation>
). G2 
<collectionCode id="29D8AE456044FFCE6800F842FCF3F86F" box="[788,822,1971,1997]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="1" pageNumber="517">TA</collectionCode>
long, flagellum-like (G2 
<collectionCode id="29D8AE456044FFCE6F46F842FBB3F86F" box="[1106,1142,1971,1997]" country="0" name="Timescale Adventures Research and Interpretive Center" pageId="1" pageNumber="517">TA</collectionCode>
/ 
<collectionCode id="29D8AE456044FFCE6F6AF842FB5AF86F" box="[1150,1183,1971,1997]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="1" pageNumber="517">SA</collectionCode>
0.65) (
<figureCitation id="D7F22A056044FFCE6FFBF842FA8DF86F" box="[1263,1352,1971,1997]" captionStart="FIGURE 7" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[264,1323,181,1831]" captionTargetId="figure-21@13.[264,1323,181,1831]" captionTargetPageId="13" captionText="FIGURE 7. Right G2 ventral view (A–C); (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph C by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 5.6 mm; B = 2.3 mm; C = 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5497113" httpUri="https://zenodo.org/record/5497113/files/figure.png" pageId="1" pageNumber="517">Fig. 7A</figureCitation>
). Large species, mature at CW 
<quantity id="88319B656044FFCE6A8EF827FE29F853" box="[410,492,2006,2033]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="517" unit="mm" value="55.0">55 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="07D3650B6047FFCD6BD3FF67FD60FB4B" pageId="2" pageNumber="518" type="description">
<paragraph id="4F7636806047FFCD6BD3FF67FE14FD27" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="518">
<emphasis id="7DBDEA926047FFCD6BD3FF67FEB2FF12" bold="true" box="[199,375,150,176]" pageId="2" pageNumber="518">Redescription:</emphasis>
Carapace moderate height (
<collectingCountry id="37DE76106047FFCD69BBFF66FD13FF13" box="[687,726,151,177]" name="Switzerland" pageId="2" pageNumber="518">CH</collectingCountry>
/FW 1.13), width (CW/FW 3.5), length (CL/ FW 2.62). Surface of anterior carapace mostly smooth except for distinct fields of carinae at anterolateral corners; postfrontal crest distinct, complete, traversing entire carapace to meet anterolateral margins at epibranchial teeth; cervical grooves deep, long, ending just before postfrontal crest; semicircular, urogastric, cardiac, branchial grooves all deep, distinct; postfrontal crest, suborbital margin, vertical sulcus on branchiostegite, all granulated; epigastric crests separated by clear, short, forked mid-groove (
<figureCitation id="D7F22A056047FFCD6914FEBAFD9CFEC7" box="[512,601,331,357]" captionStart="FIGURE 1" captionStartId="7.[151,250,1965,1989]" captionTargetBox="[191,1397,179,1938]" captionTargetId="figure-21@7.[191,1397,179,1938]" captionTargetPageId="7" captionText="FIGURE 1. Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 10.17 mm, B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="2" pageNumber="518">Fig. 1A</figureCitation>
). Exorbital tooth large, pointed, directed forward, outer margin granulated; epibranchial tooth small, low, blunt; anterolateral margin between exorbital, epibranchial teeth granulated, curving slightly outward, lacking intermediate tooth; anterolateral margin posterior to epibranchial tooth raised, lined by rounded denticles, posterior end curving inward, not continuous with posterolateral margin. Branchiostegite with distinct granulated vertical sulcus, subhepatic region heavily granulated, suborbital, pterygostomial regions smooth (
<figureCitation id="D7F22A056047FFCD6B8BFE0EFEE1FDBB" box="[159,292,511,537]" captionStart="FIGURE 1" captionStartId="7.[151,250,1965,1989]" captionTargetBox="[191,1397,179,1938]" captionTargetId="figure-21@7.[191,1397,179,1938]" captionTargetPageId="7" captionText="FIGURE 1. Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 10.17 mm, B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="2" pageNumber="518">Figs. 1A, B</figureCitation>
, 
<figureCitation id="D7F22A056047FFCD6A24FE0EFE93FDBB" box="[304,342,511,537]" captionStart="FIGURE 4" captionStartId="10.[151,250,1747,1771]" captionTargetBox="[152,1436,309,1723]" captionTargetId="figure-93@10.[152,1436,309,1723]" captionTargetPageId="10" captionText="FIGURE 4. Frontal view of carapace. (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, paratype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph B by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 10.4 mm; B = 5.7 mm; C = 4.7 mm." figureDoi="http://doi.org/10.5281/zenodo.5497107" httpUri="https://zenodo.org/record/5497107/files/figure.png" pageId="2" pageNumber="518">4A</figureCitation>
). Third maxilliped exopod with long flagellum, ischium with deep vertical sulcus (
<figureCitation id="D7F22A056047FFCD6FECFE0EFA94FDBB" box="[1272,1361,511,537]" captionStart="FIGURE 1" captionStartId="7.[151,250,1965,1989]" captionTargetBox="[191,1397,179,1938]" captionTargetId="figure-21@7.[191,1397,179,1938]" captionTargetPageId="7" captionText="FIGURE 1. Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 10.17 mm, B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="2" pageNumber="518">Fig. 1A</figureCitation>
). Thoracic sternal sulcus S2/3 deep, completely traversing sternum; S3/4 incomplete, deep at sides, absent in middle, sides slanted inward toward anterior margin of sternopleonal cavity; thoracic episternal sulci S4/E4, S5/E5, S6/ E6, S7/E7 all obscure (
<figureCitation id="D7F22A056047FFCD6A78FD9AFE06FD27" box="[364,451,619,645]" captionStart="FIGURE 1" captionStartId="7.[151,250,1965,1989]" captionTargetBox="[191,1397,179,1938]" captionTargetId="figure-21@7.[191,1397,179,1938]" captionTargetPageId="7" captionText="FIGURE 1. Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 10.17 mm, B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="2" pageNumber="518">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="4F7636806047FFCD6BD3FD7EFD2CFC07" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="518">
Male pleon, telson together forming slim triangle, pleon edges slightly indented; telson triangular, apex round- ed, base broadest, sides outwardly sloping; pleomeres PL1–6 rectangular, wider than long, PL 6 longest, more than 1/2 as long as wide (
<figureCitation id="D7F22A056047FFCD6A6AFD26FE10FD53" box="[382,469,727,753]" captionStart="FIGURE 1" captionStartId="7.[151,250,1965,1989]" captionTargetBox="[191,1397,179,1938]" captionTargetId="figure-21@7.[191,1397,179,1938]" captionTargetPageId="7" captionText="FIGURE 1. Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 10.17 mm, B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="2" pageNumber="518">Fig. 1B</figureCitation>
); remaining pleomeres short, less than 1/3 as long as wide. G1 TA long (TA/SA 0.54), midpoint angled outward at 64° to longitudinal axis of G1 SA, distal 1/3 tapering to pointed, slightly upcurved tip; G1 TA midsection widened by high mesial fold, lateral fold lower; deep longitudinal groove running along whole length of G1 TA between mesial, lateral folds (
<figureCitation id="D7F22A056047FFCD6989FCB2FCE7FCFF" box="[669,802,835,861]" captionStart="FIGURE 6" captionStartId="12.[151,250,1828,1852]" captionTargetBox="[197,1391,181,1805]" captionTargetId="figure-17@12.[197,1391,181,1805]" captionTargetPageId="12" captionText="FIGURE 6. Right G1, ventral view (A–C) and dorsal view (D, F). (A, D) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B, E) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photo- graphs B and E by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A= 6.7 mm; B = 2.3 mm; C = 1.13 mm; D = 2.7 mm; E = 2.3 mm; F = 1.2." figureDoi="http://doi.org/10.5281/zenodo.5497111" httpUri="https://zenodo.org/record/5497111/files/figure.png" pageId="2" pageNumber="518">Figs. 6A, D</figureCitation>
); G1 SA distal dorsal side with conspicuous dorsal membrane widest at mesial margin, narrowest at lateral margin (
<figureCitation id="D7F22A056047FFCD6820FC96FC4BFC23" box="[820,910,870,897]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="2" pageNumber="518">Fig. 5D</figureCitation>
). G2 TA long (G2 TA/SA 0.65), flagellum-like (
<figureCitation id="D7F22A056047FFCD6BB4FC7AFF3CFC07" box="[160,249,907,933]" captionStart="FIGURE 7" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[264,1323,181,1831]" captionTargetId="figure-21@13.[264,1323,181,1831]" captionTargetPageId="13" captionText="FIGURE 7. Right G2 ventral view (A–C); (A) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (B) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (C) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892). Photograph C by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 5.6 mm; B = 2.3 mm; C = 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5497113" httpUri="https://zenodo.org/record/5497113/files/figure.png" pageId="2" pageNumber="518">Fig. 7A</figureCitation>
). Adult size ranging from CW 
<quantity id="88319B656047FFCD6947FC7BFD20FC07" box="[595,741,906,933]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="2" pageNumber="518" unit="mm" value="57.5" valueMax="60.0" valueMin="55.0">55 to 60 mm</quantity>
.
</paragraph>
<paragraph id="4F7636806047FFCD6BD3FC5EFC1FFB67" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="518">
Chela unequal, right (major) longer than left (minor) (
<figureCitation id="D7F22A056047FFCD682CFC5EFC50FC6B" box="[824,917,942,969]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="2" pageNumber="518">Fig. 5A</figureCitation>
). Fixed finger (pollex) of propodus, movable finger (dactylus) each with 2 to 3 large teeth interspersed with smaller teeth; dactylus straight, slim, not arched, tips of fingers touching enclosing long thin interspace when closed (
<figureCitation id="D7F22A056047FFCD684BFC06FC26FBB3" box="[863,995,1014,1041]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="2" pageNumber="518">Figs. 5A, B</figureCitation>
); propodi of right, left chelae elongated (inferior margin 
<quantity id="88319B656047FFCD6A46FBEBFE7DFB97" box="[338,440,1050,1077]" metricMagnitude="-2" metricUnit="m" metricValue="5.51" pageId="2" pageNumber="518" unit="mm" value="55.1">55.1 mm</quantity>
, 
<quantity id="88319B656047FFCD6AD0FBEBFDECFB97" box="[452,553,1050,1077]" metricMagnitude="-2" metricUnit="m" metricValue="5.029999999999999" pageId="2" pageNumber="518" unit="mm" value="50.3">50.3 mm</quantity>
respectively), almost as long as CW (
<quantity id="88319B656047FFCD68C7FBEBFBF8FB97" box="[979,1085,1050,1077]" metricMagnitude="-2" metricUnit="m" metricValue="5.529999999999999" pageId="2" pageNumber="518" unit="mm" value="55.3">55.3 mm</quantity>
); palms of right, left chelae not enlarged (height 
<quantity id="88319B656047FFCD6A4DFBCEFE07FBFB" box="[345,450,1087,1113]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="2" pageNumber="518" unit="mm" value="18.2">18.2 mm</quantity>
, 
<quantity id="88319B656047FFCD6AC4FBCFFDF3FBFB" box="[464,566,1086,1113]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="2" pageNumber="518" unit="mm" value="15.5">15.5 mm</quantity>
respectively). Cheliped carpus inner margin with large pointed distal tooth, smaller pointed proximal tooth (
<figureCitation id="D7F22A056047FFCD6917FB92FD9BFBDF" box="[515,606,1122,1149]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="2" pageNumber="518">Fig. 5H</figureCitation>
); cheliped merus lower margins both lined by large granules; distal meral tooth large, pointed; superior surface of cheliped merus granulated (
<figureCitation id="D7F22A056047FFCD68B0FB76FBC4FB03" box="[932,1025,1158,1185]" captionStart="FIGURE 5" captionStartId="11.[151,250,1588,1612]" captionTargetBox="[159,1428,181,1565]" captionTargetId="figure-21@11.[159,1428,181,1565]" captionTargetPageId="11" captionText="FIGURE 5. (A–F) Frontal view of right and left chela; (G–L) cheliped merus and carpus. (A, B) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (C, D) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province, Angola (AMG GEN 1887A); (E, F) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892); (G, H) Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444); (G) ventral view of right chela merus; (H) dorsal view of right chela carpus and merus; (I, J) Potamonautes kensleyi Cumberlidge &amp; Tavares, 2006, adult male CW 35.4 mm, from the Cuanza River, Soxujwa, Kekere, Malanje Province,Angola (AMG GEN 1887A) (I) ventral view of right chela merus; (J) dorsal view of right chela merus and carpus; (K, L) Potamonautes macrobrachii Bott, 1953, holotype, adult ♂ CW 32.9 mm, from Huamba Province, Angola (SMF 1892): (K) ventral view of right chela merus; (L) dorsal view of right chela merus and carpus;. Photographs C, D, I, J by Theo Busschau, University of Stellenbosch, South Africa. Scale bar: A = 18.4 mm; B = 17.9 mm; C, D = 8.9 mm; E = 8.33 mm; F = 5.81 mm; G = 17.9 mm; H = 7.3 mm; I = 8.9 mm; J = 5.9 mm; K = 5.81 mm; L = 5.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5497109" httpUri="https://zenodo.org/record/5497109/files/figure.png" pageId="2" pageNumber="518">Fig. 5G</figureCitation>
). P2–5 slender, long. For additional characters and photographs of this species see 
<bibRefCitation id="2B584B716047FFCD69B5FB5AFCDBFB67" author="Bott, R." box="[673,798,1195,1221]" pageId="2" pageNumber="518" pagination="165 - 172" refId="ref9155" refString="Bott, R. (1968) Decapoden aus dem Museu do Dundo (Crust. Decap.). Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 77, 165 - 172." type="journal article" year="1968">Bott (1968</bibRefCitation>
: pl. 1, figs. 1–4).
</paragraph>
<paragraph id="4F7636806047FFCD6BD3FB3FFD60FB4B" blockId="2.[151,1437,150,2013]" box="[199,677,1230,1257]" pageId="2" pageNumber="518">
<emphasis id="7DBDEA926047FFCD6BD3FB3FFF32FB4A" bold="true" box="[199,247,1230,1256]" pageId="2" pageNumber="518">Size</emphasis>
. Large species, adult at CW 55.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="07D3650B6047FFCD6BD3FB02FA98FA3F" pageId="2" pageNumber="518" type="materials_examined">
<paragraph id="4F7636806047FFCD6BD3FB02FA9FFAAF" blockId="2.[151,1437,150,2013]" box="[199,1370,1266,1293]" pageId="2" pageNumber="518">
<emphasis id="7DBDEA926047FFCD6BD3FB02FEA3FAAE" bold="true" box="[199,358,1266,1293]" pageId="2" pageNumber="518">Type locality.</emphasis>
<materialsCitation id="FFA13CDD6047FFCD6A7FFB02FA93FAAF" box="[363,1366,1266,1293]" country="Angola" county="Province" latitude="-8.808363" location="Cafunfo" longLatPrecision="1" longitude="17.97665" municipality="Cuango River" pageId="2" pageNumber="518" specimenCount="1" stateProvince="Lunda Norte">
<collectingCountry id="37DE76106047FFCD6A7FFB02FE04FAAF" box="[363,449,1267,1293]" name="Angola" pageId="2" pageNumber="518">Angola</collectingCountry>
. 
<collectingCounty id="A6174E0C6047FFCD6ADFFB02FDEAFAAF" box="[459,559,1267,1293]" pageId="2" pageNumber="518">Province</collectingCounty>
of 
<collectingRegion id="8D0DF8626047FFCD6940FB02FD23FAAF" box="[596,742,1267,1293]" country="Angola" name="Lunda Norte" pageId="2" pageNumber="518">Lunda Norte</collectingRegion>
, 
<collectingMunicipality id="AF12ACFA6047FFCD69E4FB02FC4BFAAF" box="[752,910,1267,1293]" pageId="2" pageNumber="518">Cuango River</collectingMunicipality>
, 
<location id="4A16605B6047FFCD688EFB02FC33FAAF" LSID="urn:lsid:plazi:treatment:C76087966044FFCC6B83FBEAFDC7FEC7:4A16605B6047FFCD688EFB02FC33FAAF" box="[922,1014,1267,1293]" country="Angola" county="Province" latitude="-8.808363" longLatPrecision="1" longitude="17.97665" municipality="Cuango River" name="Cafunfo" pageId="2" pageNumber="518" stateProvince="Lunda Norte">Cafunfo</location>
(
<geoCoordinate id="2AFD50476047FFCD6F10FB02FBBEFAAF" box="[1028,1147,1267,1293]" degrees="8.808363" direction="south" orientation="latitude" pageId="2" pageNumber="518" precision="1" value="-8.808363">-8.808363</geoCoordinate>
, 
<geoCoordinate id="2AFD50476047FFCD6F92FB03FB34FAAF" box="[1158,1265,1266,1293]" degrees="17.97665" direction="east" orientation="longitude" pageId="2" pageNumber="518" precision="1" value="17.97665">17.97665</geoCoordinate>
) (
<figureCitation id="D7F22A056047FFCD6E1CFB02FA89FAAF" box="[1288,1356,1267,1293]" captionStart="FIGURE 8" captionStartId="14.[151,250,1336,1360]" captionTargetBox="[159,1428,189,1302]" captionTargetId="figure-17@14.[151,1436,181,1312]" captionTargetPageId="14" captionText="FIGURE 8. Map of the Angola showing the known localities of Potamonautes adeleae (black square, not shown to scale), P. kensleyi (black circles, not shown to scale), and P. macrobrachii (black diamond, not shown to scale). Scale bar = 520 km. See text for exact locality coordinates." figureDoi="http://doi.org/10.5281/zenodo.5497115" httpUri="https://zenodo.org/record/5497115/files/figure.png" pageId="2" pageNumber="518">Fig. 8</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="4F7636806047FFCD6BD3FAE7FC69FADB" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="518">
<emphasis id="7DBDEA926047FFCD6BD3FAE7FEEFFA92" bold="true" box="[199,298,1302,1328]" pageId="2" pageNumber="518">Habitat.</emphasis>
<taxonomicName id="88C94D036047FFCD6A25FAE6FDEAFA93" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[305,559,1303,1329]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="518" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA926047FFCD6A25FAE6FDEAFA93" box="[305,559,1303,1329]" italics="true" pageId="2" pageNumber="518">Potamonautes adeleae</emphasis>
</taxonomicName>
is a large species that is endemic to 
<collectingCountry id="37DE76106047FFCD68D1FAE6FBDEFA93" box="[965,1051,1303,1329]" name="Angola" pageId="2" pageNumber="518">Angola</collectingCountry>
. It is known only from a single locality in the forested 
<collectingRegion id="8D0DF8626047FFCD6A97FACAFDBAFAF7" box="[387,639,1339,1365]" country="Angola" name="Lunda Norte" pageId="2" pageNumber="518">Lunda Norte Province</collectingRegion>
. It was collected from the Cuango River, a northward-flowing tributary of the 
<collectingCountry id="37DE76106047FFCD6BCBFAAEFEEFFADB" box="[223,298,1375,1401]" name="Democratic Republic of the Congo" pageId="2" pageNumber="518">Congo</collectingCountry>
River which is part of the central African region (
<figureCitation id="D7F22A056047FFCD684EFAAEFC5BFADB" box="[858,926,1375,1401]" captionStart="FIGURE 8" captionStartId="14.[151,250,1336,1360]" captionTargetBox="[159,1428,189,1302]" captionTargetId="figure-17@14.[151,1436,181,1312]" captionTargetPageId="14" captionText="FIGURE 8. Map of the Angola showing the known localities of Potamonautes adeleae (black square, not shown to scale), P. kensleyi (black circles, not shown to scale), and P. macrobrachii (black diamond, not shown to scale). Scale bar = 520 km. See text for exact locality coordinates." figureDoi="http://doi.org/10.5281/zenodo.5497115" httpUri="https://zenodo.org/record/5497115/files/figure.png" pageId="2" pageNumber="518">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="4F7636806047FFCD6BD3FA73FA98FA3F" blockId="2.[151,1437,150,2013]" box="[199,1373,1410,1437]" pageId="2" pageNumber="518">
<emphasis id="7DBDEA926047FFCD6BD3FA73FEE5FA3E" bold="true" box="[199,288,1410,1436]" pageId="2" pageNumber="518">Colour.</emphasis>
The colour of living specimens is unknown, but specimens preserved in ethanol are light brown.
</paragraph>
</subSubSection>
<subSubSection id="07D3650B6047FFCD6BD3FA57FD8FF93B" pageId="2" pageNumber="518" type="distribution">
<paragraph id="4F7636806047FFCD6BD3FA57FD25FA47" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="518">
<emphasis id="7DBDEA926047FFCD6BD3FA57FEA4FA62" bold="true" box="[199,353,1446,1472]" pageId="2" pageNumber="518">Distribution.</emphasis>
This species is only known from a single locality in 
<collectingCountry id="37DE76106047FFCD68BAFA56FBC5FA63" box="[942,1024,1447,1473]" name="Angola" pageId="2" pageNumber="518">Angola</collectingCountry>
in the Province of 
<collectingRegion id="8D0DF8626047FFCD6FC2FA56FAADFA63" box="[1238,1384,1447,1473]" country="Angola" name="Lunda Norte" pageId="2" pageNumber="518">Lunda Norte</collectingRegion>
, Cuango River, Cafunfo (
<geoCoordinate id="2AFD50476047FFCD6A9EFA3AFDC4FA47" box="[394,513,1483,1509]" degrees="8.808363" direction="south" orientation="latitude" pageId="2" pageNumber="518" precision="1" value="-8.808363">-8.808363</geoCoordinate>
, 
<geoCoordinate id="2AFD50476047FFCD6918FA3BFDB2FA47" box="[524,631,1482,1509]" degrees="17.97665" direction="east" orientation="longitude" pageId="2" pageNumber="518" precision="1" value="17.97665">17.97665</geoCoordinate>
) (
<figureCitation id="D7F22A056047FFCD699AFA3AFD17FA47" box="[654,722,1483,1509]" captionStart="FIGURE 8" captionStartId="14.[151,250,1336,1360]" captionTargetBox="[159,1428,189,1302]" captionTargetId="figure-17@14.[151,1436,181,1312]" captionTargetPageId="14" captionText="FIGURE 8. Map of the Angola showing the known localities of Potamonautes adeleae (black square, not shown to scale), P. kensleyi (black circles, not shown to scale), and P. macrobrachii (black diamond, not shown to scale). Scale bar = 520 km. See text for exact locality coordinates." figureDoi="http://doi.org/10.5281/zenodo.5497115" httpUri="https://zenodo.org/record/5497115/files/figure.png" pageId="2" pageNumber="518">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="4F7636806047FFCD6BD3FA1FFD8FF93B" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="518">
<emphasis id="7DBDEA926047FFCD6BD3FA1FFE78F9AB" bold="true" box="[199,445,1518,1545]" pageId="2" pageNumber="518">Conservation status.</emphasis>
<taxonomicName id="88C94D036047FFCD6AD1FA1EFD00F9AB" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[453,709,1519,1545]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="518" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA926047FFCD6AD1FA1EFD00F9AB" box="[453,709,1519,1545]" italics="true" pageId="2" pageNumber="518">Potamonautes adeleae</emphasis>
</taxonomicName>
was listed in the IUCN Red List of Threatened Species as Data Deficient (
<bibRefCitation id="2B584B716047FFCD6A1AF9E2FE2AF98F" author="Cumberlidge, N." box="[270,495,1555,1581]" pageId="2" pageNumber="518" pagination="134068" refId="ref9459" refString="Cumberlidge, N. (2008 a) Potamonautes adeleae. The IUCN Red List of Threatened Species, 2008, e. T 134068 A 3906631. https: // doi. org / 10.2305 / IUCN. UK. 2008. RLTS. T 134068 A 3906631. en" type="journal article" year="2008">Cumberlidge 2008a</bibRefCitation>
) because it is known to occur in just one locality and there is very little information available on its population trends, habitat, or threats. This makes it a rare species of high research interest. The diamond mines at Cafunfo have resulted in a great deal of habitat disturbance and pollution of the Cuango River, that may prove to be threats to this species.
</paragraph>
</subSubSection>
<subSubSection id="07D3650B6047FFCC6BD3F952FDC7FEC7" lastPageId="3" lastPageNumber="519" pageId="2" pageNumber="518" type="discussion">
<paragraph id="4F7636806047FFCC6BD3F952FDC7FEC7" blockId="2.[151,1437,150,2013]" lastBlockId="3.[151,1436,151,357]" lastPageId="3" lastPageNumber="519" pageId="2" pageNumber="518">
<emphasis id="7DBDEA926047FFCD6BD3F952FEF1F91F" bold="true" box="[199,308,1699,1725]" pageId="2" pageNumber="518">Remarks</emphasis>
. 
<bibRefCitation id="2B584B716047FFCD6A57F952FE03F91C" author="Bott, R." box="[323,454,1699,1726]" pageId="2" pageNumber="518" pagination="165 - 172" refId="ref9155" refString="Bott, R. (1968) Decapoden aus dem Museu do Dundo (Crust. Decap.). Publicacoes culturais da Companhia de Diamantes de Angola, Lisboa, 77, 165 - 172." type="journal article" year="1968">Bott (1968)</bibRefCitation>
described 
<taxonomicName id="88C94D036047FFCD6956F952FD73F91F" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[578,694,1699,1725]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="518" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA926047FFCD6956F952FD73F91F" box="[578,694,1699,1725]" italics="true" pageId="2" pageNumber="518">P. adeleae</emphasis>
</taxonomicName>
as a subspecies of 
<taxonomicName id="88C94D036047FFCD6886F952FB32F91F" authority="(Rathbun, 1904)" baseAuthorityName="Rathbun" baseAuthorityYear="1904" box="[914,1271,1699,1725]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="518" phylum="Arthropoda" rank="species" species="lirrangensis">
<emphasis id="7DBDEA926047FFCD6886F952FBF3F91F" box="[914,1078,1699,1725]" italics="true" pageId="2" pageNumber="518">P. lirrangensis</emphasis>
(
<bibRefCitation id="2B584B716047FFCD6F52F952FB2AF91F" author="Rathbun, M. J." box="[1094,1263,1699,1725]" pageId="2" pageNumber="518" pagination="255 - 312" refId="ref10467" refString="Rathbun, M. J. (1904) Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle, Paris, 6, 255 - 312." type="journal article" year="1904">Rathbun, 1904</bibRefCitation>
)
</taxonomicName>
from Liranga, 
<collectingCountry id="37DE76106047FFCD6B83F936FE57F943" box="[151,402,1735,1761]" name="Republic of the Congo" pageId="2" pageNumber="518">Republique du Congo</collectingCountry>
, in the 
<collectingCountry id="37DE76106047FFCD6AFCF936FD4CF943" box="[488,649,1735,1761]" name="Democratic Republic of the Congo" pageId="2" pageNumber="518">middle Congo</collectingCountry>
basin but 
<bibRefCitation id="2B584B716047FFCD6815F936FBA3F940" author="Cumberlidge, N. &amp; Tavares, M." box="[769,1126,1735,1762]" pageId="2" pageNumber="518" pagination="248 - 257" refId="ref9684" refString="Cumberlidge, N. &amp; Tavares, M. (2006) Remarks on the freshwater crabs of Angola, southwestern Africa, with the description of Potamonautes kensleyi, new species (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 26 (2), 248 - 257. https: // doi. org / 10.1651 / s- 2653.1" type="journal article" year="2006">Cumberlidge &amp; Tavares (2006)</bibRefCitation>
, 
<bibRefCitation id="2B584B716047FFCD6F67F936FAA2F943" author="Cumberlidge, N." box="[1139,1383,1735,1761]" pageId="2" pageNumber="518" pagination="64386" refId="ref9561" refString="Cumberlidge, N. (2008 c) Potamonautes macrobrachii. The IUCN Red List of Threatened Species, 2008, e. T 64386 A 12766440. https: // doi. org / 10.2305 / IUCN. UK. 2008. RLTS. T 64386 A 12766440. en" type="journal article" year="2008">Cumberlidge (2008c)</bibRefCitation>
, and 
<bibRefCitation id="2B584B716047FFCD6B83F91AFE89F8A7" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[151,332,1771,1797]" pageId="2" pageNumber="518" pagination="1 - 286" refId="ref10320" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyuorum: part I. An annotated checklist of extant Brachyuran crabs of the world. Raffles Bulletin of Zoology, 17, 1 - 286." type="journal article" year="2008">
Ng 
<emphasis id="7DBDEA926047FFCD6BD5F91AFF36F8A7" box="[193,243,1771,1797]" italics="true" pageId="2" pageNumber="518">et al</emphasis>
. (2008)
</bibRefCitation>
recognised 
<taxonomicName id="88C94D036047FFCD6AC1F91AFD8DF8A7" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[469,584,1771,1797]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="518" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA926047FFCD6AC1F91AFD8DF8A7" box="[469,584,1771,1797]" italics="true" pageId="2" pageNumber="518">P. adeleae</emphasis>
</taxonomicName>
as a valid species. 
<bibRefCitation id="2B584B716047FFCD6837F91AFB46F8A4" author="Cumberlidge, N. &amp; Tavares, M." box="[803,1155,1771,1798]" pageId="2" pageNumber="518" pagination="248 - 257" refId="ref9684" refString="Cumberlidge, N. &amp; Tavares, M. (2006) Remarks on the freshwater crabs of Angola, southwestern Africa, with the description of Potamonautes kensleyi, new species (Brachyura: Potamoidea: Potamonautidae). Journal of Crustacean Biology, 26 (2), 248 - 257. https: // doi. org / 10.1651 / s- 2653.1" type="journal article" year="2006">Cumberlidge &amp; Tavares (2006)</bibRefCitation>
cited morphological differences between the adult male 
<typeStatus id="907288226047FFCD6919F8FEFDA8F88B" box="[525,621,1807,1833]" pageId="2" pageNumber="518" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="88C94D036047FFCD6982F8FEFCCFF88B" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[662,778,1807,1833]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="518" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA926047FFCD6982F8FEFCCFF88B" box="[662,778,1807,1833]" italics="true" pageId="2" pageNumber="518">P. adeleae</emphasis>
</taxonomicName>
from 
<collectingCountry id="37DE76106047FFCD6844F8FEFC67F88B" box="[848,930,1807,1833]" name="Angola" pageId="2" pageNumber="518">Angola</collectingCountry>
and the dried adult female type of 
<taxonomicName id="88C94D036047FFCD6E23F8FEFD4DF8EF" authority="Rathbun, 1904" authorityName="Rathbun" authorityYear="1904" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="518" phylum="Arthropoda" rank="species" species="lirrangensis" subGenus="Potamonautes">
<emphasis id="7DBDEA926047FFCD6E23F8FEFE1CF8EF" italics="true" pageId="2" pageNumber="518">Potamon (Potamonautes) lirrangensis</emphasis>
<bibRefCitation id="2B584B716047FFCD6AF6F8C2FD4DF8EF" author="Rathbun, M. J." box="[482,648,1843,1869]" pageId="2" pageNumber="518" pagination="255 - 312" refId="ref10467" refString="Rathbun, M. J. (1904) Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle, Paris, 6, 255 - 312." type="journal article" year="1904">Rathbun, 1904</bibRefCitation>
</taxonomicName>
(CW 
<quantity id="88319B656047FFCD69DAF8C3FCF1F8EF" box="[718,820,1842,1869]" metricMagnitude="-2" metricUnit="m" metricValue="5.39" pageId="2" pageNumber="518" unit="mm" value="53.9">53.9 mm</quantity>
) which was photographed by 
<bibRefCitation id="2B584B716047FFCD6F9EF8C2FAF3F8EF" author="Rathbun, M. J." box="[1162,1334,1843,1869]" pageId="2" pageNumber="518" pagination="255 - 312" refId="ref10467" refString="Rathbun, M. J. (1904) Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle, Paris, 6, 255 - 312." type="journal article" year="1904">Rathbun (1904</bibRefCitation>
: pl. XIV, fig. 8), and illustrated by 
<bibRefCitation id="2B584B716047FFCD6AA5F8A6FD8DF8D3" author="Capart, A." box="[433,584,1878,1905]" pageId="2" pageNumber="518" pagination="819 - 847" refId="ref9369" refString="Capart, A. (1954) Revision des types des especes de Potamonidae de l'Afrique Tropicale conserves au Museum d'Histoire Naturelle de Paris. Volume Jubilaire de Victor Van Strallen, Director de l'Institut royal des Sciences naturelles de Belgique, 1925 - 1934, 2, 819 - 847." type="journal article" year="1954">Capart (1954</bibRefCitation>
: fig. 28). 
<taxonomicName id="88C94D036047FFCD69A6F8A6FB9BF8D3" authorityName="Rathbun" authorityYear="1904" box="[690,1118,1879,1905]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="518" phylum="Arthropoda" rank="species" species="lirrangensis" subGenus="Potamonautes">
<emphasis id="7DBDEA926047FFCD69A6F8A6FB9BF8D3" box="[690,1118,1879,1905]" italics="true" pageId="2" pageNumber="518">Potamon (Potamonautes) lirrangensis</emphasis>
</taxonomicName>
was recently redescribed by 
<bibRefCitation id="2B584B716047FFCD6B83F88AFE70F837" author="Cumberlidge, N. &amp; Johnson, E. &amp; Clark, P. F. &amp; Genner, M. J." box="[151,437,1915,1941]" pageId="2" pageNumber="518" pagination="145 - 178" refId="ref9749" refString="Cumberlidge, N., Johnson, E., Clark, P. F. &amp; Genner, M. J. (2021) Unravelling the Potamonautes lirrangensis (Rathbun, 1904) species complex (Potamoidea: Potamonautidae), with the description of two new species. European Journal of Taxonomy, 744 (1), 145 - 178. https: // doi. org / 10.5852 / ejt. 2021.744.1309" type="journal article" year="2021">
Cumberlidge 
<emphasis id="7DBDEA926047FFCD6A24F88AFEA7F837" box="[304,354,1915,1941]" italics="true" pageId="2" pageNumber="518">et al</emphasis>
. (2021
</bibRefCitation>
: figs. 1, 4A, 7A) as 
<taxonomicName id="88C94D036047FFCD6982F88AFC01F837" baseAuthorityName="Rathbun" baseAuthorityYear="1904" box="[662,964,1915,1941]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="518" phylum="Arthropoda" rank="species" sensu="stricto" species="lirrangensis">
<emphasis id="7DBDEA926047FFCD6982F88AFC01F837" box="[662,964,1915,1941]" italics="true" pageId="2" pageNumber="518">Potamonautes lirrangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="668E57E96047FFCD68DFF88AFC2AF837" box="[971,1007,1915,1941]" pageId="2" pageNumber="518" sensu="stricto">s.s.</taxonomicNameLabel>
who restricted that species to the type specimen from Liranga, 
<collectingCountry id="37DE76106047FFCD6AB9F86EFD62F81B" box="[429,679,1951,1977]" name="Republic of the Congo" pageId="2" pageNumber="518">Republique du Congo</collectingCountry>
, in the 
<collectingCountry id="37DE76106047FFCD69E8F86EFC59F81B" box="[764,924,1951,1977]" name="Democratic Republic of the Congo" pageId="2" pageNumber="518">middle Congo</collectingCountry>
basin. Unfortunately, the fact that the female 
<typeStatus id="907288226047FFCD6B83F832FF3CF87F" box="[151,249,1987,2013]" pageId="2" pageNumber="518" type="holotype">holotype</typeStatus>
of this species is poorly preserved excludes most of the taxonomic characters that are useful to distinguish between species. Despite this, differences in the anterolateral margin of 
<taxonomicName id="88C94D036046FFCC68D2FF66FBFCFF13" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[966,1081,151,177]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="519" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA926046FFCC68D2FF66FBFCFF13" box="[966,1081,151,177]" italics="true" pageId="3" pageNumber="519">P. adeleae</emphasis>
</taxonomicName>
(
<figureCitation id="D7F22A056046FFCC6F5DFF66FB07FF13" box="[1097,1218,151,177]" captionStart="FIGURE 1" captionStartId="7.[151,250,1965,1989]" captionTargetBox="[191,1397,179,1938]" captionTargetId="figure-21@7.[191,1397,179,1938]" captionTargetPageId="7" captionText="FIGURE 1. Potamonautes adeleae Bott, 1968, paratype, adult ♂ CW 55.3 mm, from Lunda Norte Province, Angola (SMF 4444). Whole animal, dorsal view (A), whole animal, ventral view (B). Scale bars: A = 10.17 mm, B = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5497101" httpUri="https://zenodo.org/record/5497101/files/figure.png" pageId="3" pageNumber="519">Fig. 1A, B</figureCitation>
) and 
<taxonomicName id="88C94D036046FFCC6E17FF66FA59FF13" box="[1283,1436,151,177]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="519" phylum="Arthropoda" rank="species" sensu="stricto" species="lirrangenis">
<emphasis id="7DBDEA926046FFCC6E17FF66FA59FF13" box="[1283,1436,151,177]" italics="true" pageId="3" pageNumber="519">P. lirrangenis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="668E57E96046FFCC6B83FF4AFF7EFF77" box="[151,187,187,213]" pageId="3" pageNumber="519" sensu="stricto">s.s.</taxonomicNameLabel>
(cf. 
<bibRefCitation id="2B584B716046FFCC6BE7FF4AFDD0FF77" author="Cumberlidge, N. &amp; Johnson, E. &amp; Clark, P. F. &amp; Genner, M. J." box="[243,533,187,213]" pageId="3" pageNumber="519" pagination="145 - 178" refId="ref9749" refString="Cumberlidge, N., Johnson, E., Clark, P. F. &amp; Genner, M. J. (2021) Unravelling the Potamonautes lirrangensis (Rathbun, 1904) species complex (Potamoidea: Potamonautidae), with the description of two new species. European Journal of Taxonomy, 744 (1), 145 - 178. https: // doi. org / 10.5852 / ejt. 2021.744.1309" type="journal article" year="2021">
Cumberlidge 
<emphasis id="7DBDEA926046FFCC6A9AFF4AFE07FF77" box="[398,450,187,213]" italics="true" pageId="3" pageNumber="519">et al</emphasis>
., 2021
</bibRefCitation>
: fig. 4A) can be used to separate these two taxa. 
<taxonomicName id="88C94D036046FFCC6F59FF4AFA88FF77" authorityName=", Cumberlidge &amp; Tavares" authorityYear="2006" box="[1101,1357,187,213]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="519" phylum="Arthropoda" rank="species" species="adeleae">
<emphasis id="7DBDEA926046FFCC6F59FF4AFA88FF77" box="[1101,1357,187,213]" italics="true" pageId="3" pageNumber="519">Potamonautes adeleae</emphasis>
</taxonomicName>
shares a number of characters (e.g., a complete postfrontal crest, a reduced epibranchial tooth, a granulated anterolateral margin, and the general form of the G1 TA) with species from the 
<collectingCountry id="37DE76106046FFCC6894FEF2FBE5FEBF" box="[896,1056,259,285]" name="Democratic Republic of the Congo" pageId="3" pageNumber="519">middle Congo</collectingCountry>
River basin (such as 
<taxonomicName id="88C94D036046FFCC6E05FEF2FE9BFEE3" authority="(Rathbun, 1904))" baseAuthorityName="Rathbun" baseAuthorityYear="1904" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="519" phylum="Arthropoda" rank="species" species="dybowskii">
<emphasis id="7DBDEA926046FFCC6E05FEF2FA59FEBF" box="[1297,1436,259,285]" italics="true" pageId="3" pageNumber="519">P. dybowskii</emphasis>
(
<bibRefCitation id="2B584B716046FFCC6B8BFED6FE8EFEE3" author="Rathbun, M. J." box="[159,331,295,321]" pageId="3" pageNumber="519" pagination="255 - 312" refId="ref10467" refString="Rathbun, M. J. (1904) Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle, Paris, 6, 255 - 312." type="journal article" year="1904">Rathbun, 1904</bibRefCitation>
))
</taxonomicName>
, and the affinities of this Angolan species lie with the 
<collectingCountry id="37DE76106046FFCC68CDFED6FBE1FEE3" box="[985,1060,295,321]" name="Democratic Republic of the Congo" pageId="3" pageNumber="519">Congo</collectingCountry>
basin fauna rather than with the southern African regional fauna.
</paragraph>
</subSubSection>
</treatment>
</document>