184 lines
20 KiB
XML
184 lines
20 KiB
XML
<document id="D3E9A77AD28AB4F5F81928DBA4EE4C6F" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. & Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A37BB13FEFAE199E60828DE" docLanguage="en" docName="J_NAT_HIST.53.37–38.2231–2351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) blancoi Nunes & Vaz-de-Mello 2019, new species" docType="treatment" docVersion="18" lastPageNumber="2316" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2315" updateTime="1698782751744" updateUser="ExternalLinkService">
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<mods:titleInfo id="08D6FCB78EBDF316C7EDE38250F2FB81">
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<mods:title id="CE1C2B0E3EC317B72C8B22B8D136CF19">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
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<mods:name id="A298697E0EE39CF74196F5A40A8D0C12" type="personal">
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<mods:roleTerm id="78FBC23D9B57366B5EE482FCA53115B7">Author</mods:roleTerm>
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<mods:namePart id="3F14BE8ACE06AE81C955D45CB1EFFB59">Nunes, Rafael V.</mods:namePart>
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<mods:name id="F7E1ECC3633B5D2CCE5D13BF39661D76" type="personal">
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<mods:namePart id="35BE7013C71D0162447951CAE8ED53D0">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:title id="DC42AB61FC4BA7C3BA416BE66AF73E67">Journal of Natural History</mods:title>
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<mods:part id="3E75B8A96AC434B7E4ECD6ACBCB57301">
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<mods:date id="2D6AE328A8AD9BF2541F49215941EEF8">2019</mods:date>
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<mods:number id="FD3D5AC91D6AB4C7A1A492597EC7896B">2020-01-13</mods:number>
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<mods:number id="A7FEECDED0BB93D6FB4C472AE8BCC55B">53</mods:number>
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<mods:detail id="2365F0C51616FB0C5F125D3F83CAB064" type="issue">
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<mods:number id="5D9A050D16562673A9B9BC88580C6ACD">37</mods:number>
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<mods:classification id="0B6C3C2CEA5B8DCBDC39B380FF7C4867">journal article</mods:classification>
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<mods:identifier id="42C5A25DF176F1A8ADCEDE956D868B25" type="DOI">10.1080/00222933.2019.1692088</mods:identifier>
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<treatment id="C740D6092A37BB13FEFAE199E60828DE" ID-DOI="http://doi.org/10.5281/zenodo.3671922" ID-GBIF-Taxon="162116271" ID-Zenodo-Dep="3671922" LSID="urn:lsid:plazi:treatment:C740D6092A37BB13FEFAE199E60828DE" httpUri="http://treatment.plazi.org/id/C740D6092A37BB13FEFAE199E60828DE" lastPageId="86" lastPageNumber="2316" pageId="85" pageNumber="2315">
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<subSubSection id="07F334942A37BB10FEFAE199E6872900" pageId="85" pageNumber="2315" type="nomenclature">
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<paragraph id="4F56671F2A37BB10FEFAE199E7A02923" blockId="85.[344,972,1255,1315]" box="[344,972,1255,1280]" pageId="85" pageNumber="2315">
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<heading id="141ED0732A37BB10FEFAE199E7A02923" box="[344,972,1255,1280]" centered="true" fontSize="11" level="2" pageId="85" pageNumber="2315" reason="2">
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3.10.:
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<taxonomicName id="88E91C9C2A37BB10FE3EE199E7412923" ID-CoL="35N9P" authority="Nunes & Vaz-de-Mello, 2019" authorityName="Nunes & Vaz-de-Mello" authorityYear="2019" box="[412,813,1255,1280]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="2315" phylum="Arthropoda" rank="species" species="blancoi" status="sp. nov." subGenus="Cephagonus">
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<emphasis id="7D9DBB0D2A37BB10FE3EE199E6402923" box="[412,556,1255,1280]" italics="true" pageId="85" pageNumber="2315">Dichotomius</emphasis>
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||
(
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<emphasis id="7D9DBB0D2A37BB10FD99E199E6A52923" box="[571,713,1255,1280]" italics="true" pageId="85" pageNumber="2315">Cephagonus</emphasis>
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)
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<emphasis id="7D9DBB0D2A37BB10FD7BE199E7412923" box="[729,813,1255,1280]" italics="true" pageId="85" pageNumber="2315">blancoi</emphasis>
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</taxonomicName>
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<emphasis id="7D9DBB0D2A37BB10FC96E199E7A02923" bold="true" box="[820,972,1255,1280]" pageId="85" pageNumber="2315">
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<taxonomicNameLabel id="66AE06762A37BB10FC96E199E7A02923" box="[820,972,1255,1280]" pageId="85" pageNumber="2315" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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<paragraph id="4F56671F2A37BB10FD9AE074E6872900" blockId="85.[344,972,1255,1315]" box="[568,747,1290,1315]" pageId="85" pageNumber="2315">
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<heading id="141ED0732A37BB10FD9AE074E6872900" box="[568,747,1290,1315]" centered="true" fontSize="11" level="2" pageId="85" pageNumber="2315" reason="5">
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(
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<figureCitation id="D7D27B9A2A37BB10FDE2E074E6C32900" box="[576,687,1290,1315]" captionStart="Figure 35" captionStartId="50.[160,225,1682,1705]" captionTargetBox="[163,1153,142,1634]" captionTargetId="figure@50.[163,1153,142,1650]" captionTargetPageId="50" captionText="Figure 35. D. blancoi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view; (e) detail of the carena found on the middle of paramera longitudinal excavation. D. candangoi: (f) male dorsal habitus; (g) paramera dorsal view; (h) paramera ventral view; (i) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665585" httpUri="https://zenodo.org/record/3665585/files/figure.png" pageId="85" pageNumber="2315">Figure 35</figureCitation>
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(a-e))
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="07F334942A37BB10FF02E010E7BE2A31" pageId="85" pageNumber="2315" type="diagnosis">
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<paragraph id="4F56671F2A37BB10FF02E010E7BE2A31" blockId="85.[160,1155,1390,1554]" pageId="85" pageNumber="2315">
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<emphasis id="7D9DBB0D2A37BB10FF02E010E57629A4" bold="true" box="[160,282,1390,1415]" italics="true" pageId="85" pageNumber="2315">Diagnosis.</emphasis>
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this species is separated from other in the group by the following combined characters: males pronotum with pair of weak knobs on the anterior portion of disc; elytral striae and punctures deeply impressed; males paramera with a longitudinal carina inserted on the longitudinal excavation (
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<figureCitation id="D7D27B9A2A37BB10FDDAE0A9E68B29CC" box="[632,743,1495,1519]" captionStart="Figure 35" captionStartId="50.[160,225,1682,1705]" captionTargetBox="[163,1153,142,1634]" captionTargetId="figure@50.[163,1153,142,1650]" captionTargetPageId="50" captionText="Figure 35. D. blancoi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view; (e) detail of the carena found on the middle of paramera longitudinal excavation. D. candangoi: (f) male dorsal habitus; (g) paramera dorsal view; (h) paramera ventral view; (i) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665585" httpUri="https://zenodo.org/record/3665585/files/figure.png" pageId="85" pageNumber="2315">Figure 35</figureCitation>
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(e)). Very similar to many species of the
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<taxonomicName id="88E91C9C2A37BB10FF6EE084E5342A31" authorityName="Felsche" authorityYear="1901" box="[204,344,1530,1554]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="2315" phylum="Arthropoda" rank="species" species="quadraticeps">
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<emphasis id="7D9DBB0D2A37BB10FF6EE084E5342A31" box="[204,344,1530,1554]" italics="true" pageId="85" pageNumber="2315">quadraticeps</emphasis>
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</taxonomicName>
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groupı being differentiated basically by males genitalia.
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</paragraph>
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</subSubSection>
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<subSubSection id="07F334942A37BB10FF1CE33EE52F2A9C" pageId="85" pageNumber="2315" type="materials_examined">
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<paragraph id="4F56671F2A37BB10FF1CE33EE6412A59" blockId="85.[190,1155,1599,1727]" pageId="85" pageNumber="2315">
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<emphasis id="7D9DBB0D2A37BB10FF1CE33EE4A02A77" bold="true" box="[190,204,1600,1620]" pageId="85" pageNumber="2315">●</emphasis>
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<materialsCitation id="FF816D422A37BB10FF7AE341E6492A59" ID-GBIF-Occurrence="2570409554" collectingDate="1993-05" collectionCode="CEMT" country="Venezuela" location="Leg. J. Blanco" municipality="Parque Paramillo. Edo. Tachira" pageId="85" pageNumber="2315" specimenCount="1" specimenCount-male="1" stateProvince="Tachira" typeStatus="holotype">
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<typeStatus id="9052D9BD2A37BB10FF7AE341E53E2A74" box="[216,338,1599,1623]" pageId="85" pageNumber="2315" type="holotype">HOLOTYPE</typeStatus>
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(
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<specimenCount id="59EFAC962A37BB10FEC3E341E5F02A74" box="[353,412,1599,1623]" pageId="85" pageNumber="2315" type="male">male</specimenCount>
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):
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<collectingCountry id="37FE278F2A37BB10FE12E341E6542A74" box="[432,568,1599,1623]" name="Venezuela" pageId="85" pageNumber="2315">VENEZUELA</collectingCountry>
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.
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<emphasis id="7D9DBB0D2A37BB10FDE1E341E6F72A74" bold="true" box="[579,667,1599,1623]" pageId="85" pageNumber="2315">
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<collectingRegion id="8D2DA9FD2A37BB10FDE1E341E6F72A74" box="[579,667,1599,1623]" country="Venezuela" name="Tachira" pageId="85" pageNumber="2315">Tachira</collectingRegion>
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</emphasis>
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.
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<collectingMunicipality id="AF32FD652A37BB10FD08E341E0682A74" box="[682,1028,1599,1623]" pageId="85" pageNumber="2315">Parque Paramillo. Edo. Tachira</collectingMunicipality>
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V32.
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<date id="3B5741DF2A37BB10FBE4E341E57D2A59" pageId="85" pageNumber="2315" value="1993-05">
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<collectingDate id="2B13B8372A37BB10FBE4E341E57D2A59" pageId="85" pageNumber="2315" value="1993-05">Mayo 1993</collectingDate>
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</date>
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||
.
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<location id="4A3631C42A37BB10FEBEE31CE5DF2A59" LSID="urn:lsid:plazi:treatment:C740D6092A37BB13FEFAE199E60828DE:4A3631C42A37BB10FEBEE31CE5DF2A59" box="[284,435,1634,1658]" country="Venezuela" municipality="Parque Paramillo. Edo. Tachira" name="Leg. J. Blanco" pageId="85" pageNumber="2315" stateProvince="Tachira">Leg. J. Blanco</location>
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[at
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<collectionCode id="29F8FFDA2A37BB10FE42E31CE6492A59" box="[480,549,1634,1658]" pageId="85" pageNumber="2315">CEMT</collectionCode>
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</materialsCitation>
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].
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</paragraph>
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<paragraph id="4F56671F2A37BB10FF1CE3FBE52F2A9C" blockId="85.[190,1155,1599,1727]" pageId="85" pageNumber="2315">
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<emphasis id="7D9DBB0D2A37BB10FF1CE3FBE4A02ABA" bold="true" box="[190,204,1669,1689]" pageId="85" pageNumber="2315">●</emphasis>
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<typeStatus id="9052D9BD2A37BB10FF7AE3FBE5302ABE" box="[216,348,1669,1693]" pageId="85" pageNumber="2315" type="paratype">PARATYPES</typeStatus>
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: same data of
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<typeStatus id="9052D9BD2A37BB10FE5FE3FBE60D2ABE" box="[509,609,1669,1693]" pageId="85" pageNumber="2315" type="holotype">holotype</typeStatus>
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[
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<specimenCount id="59EFAC962A37BB10FDD0E3FBE6F52ABF" box="[626,665,1668,1693]" pageId="85" pageNumber="2315" type="female">1 ♀</specimenCount>
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at
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<collectionCode id="29F8FFDA2A37BB10FD1EE3FBE76D2ABE" box="[700,769,1669,1693]" pageId="85" pageNumber="2315">CEMT</collectionCode>
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];
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<collectingCountry id="37FE278F2A37BB10FCB2E3FBE70E2ABE" box="[784,866,1669,1693]" name="Brazil" pageId="85" pageNumber="2315">BRAZIL</collectingCountry>
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.
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<emphasis id="7D9DBB0D2A37BB10FCC9E3FBE7A32ABE" bold="true" box="[875,975,1669,1693]" pageId="85" pageNumber="2315">
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<collectingRegion id="8D2DA9FD2A37BB10FCC9E3FBE7A32ABE" box="[875,975,1669,1693]" country="Brazil" name="Roraima" pageId="85" pageNumber="2315">Roraima</collectingRegion>
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</emphasis>
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. 1997. [1 Ƌı
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<specimenCount id="59EFAC962A37BB10FBF9E3FBE0EF2ABF" box="[1115,1155,1668,1693]" pageId="85" pageNumber="2315" type="female">1 ♀</specimenCount>
|
||
at
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||
<collectionCode id="29F8FFDA2A37BB10FF57E3D9E5562A9C" box="[245,314,1703,1727]" pageId="85" pageNumber="2315">CEMT</collectionCode>
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||
].
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="07F334942A34BB13FF02E5F0E5BC286C" pageId="86" pageNumber="2316" type="description">
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||
<paragraph id="4F56671F2A34BB13FF02E5F0E50E2F7F" blockId="86.[160,1156,142,860]" pageId="86" pageNumber="2316">
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||
<emphasis id="7D9DBB0D2A34BB13FF02E5F0E5952C84" bold="true" box="[160,505,142,167]" italics="true" pageId="86" pageNumber="2316">Description (Holotypeı male).</emphasis>
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BL: 15 mmı PW
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<quantity id="8811CAFA2A34BB13FD6AE5F1E77C2C84" box="[712,784,143,167]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="86" pageNumber="2316" unit="mm" value="9.0">9 mm</quantity>
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. Colour: dark brown and blackı shiny.
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<emphasis id="7D9DBB0D2A34BB13FF51E5CCE55E2CE9" bold="true" box="[243,306,178,202]" pageId="86" pageNumber="2316">Head</emphasis>
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: dorsal surface smooth except for few striations near clypeo-genal groove. Gena borders slightly curved inward. Clypeo-genal junction slightly curved upward. Cephalic carina longer than wideı conical with rounded at apex.
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<emphasis id="7D9DBB0D2A34BB13FBA1E589E0112D2C" bold="true" box="[1027,1149,247,271]" pageId="86" pageNumber="2316">Pronotum</emphasis>
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: borders smoothı not serrated. Pronotal disc shinyı lacking punctures and having two weak central lobes on the anterior portion of disc. Ocellate punctures present along pronotal margins except at anterior portion adjacent to eyes.
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<emphasis id="7D9DBB0D2A34BB13FC37E421E0542D54" bold="true" box="[917,1080,351,375]" pageId="86" pageNumber="2316">Metasternum</emphasis>
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: sides and margins of anterior lobe covered by hair. Anterior portion of anterior lobe glabrous with chagrinated microsculpture. Setigerous punctures evenly spaced at sides.
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<emphasis id="7D9DBB0D2A34BB13FF4CE4B9E5592DFC" bold="true" box="[238,309,455,479]" pageId="86" pageNumber="2316">Elytra</emphasis>
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: striae deeply impressed having fine punctures spaced 2-four times their diameter. Interstriae convex. Interstriae disc with dark brown colourı lacking microsculpture with very fine punctures (20 x).
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<emphasis id="7D9DBB0D2A34BB13FD4CE772E7092E07" bold="true" box="[750,869,524,548]" pageId="86" pageNumber="2316">Abdomen</emphasis>
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: sides of each ventrite excavated with groups of ocellate punctures and 1
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<emphasis id="7D9DBB0D2A34BB13FD4CE751E6972E64" box="[750,763,559,583]" italics="true" pageId="86" pageNumber="2316">–</emphasis>
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4 setae. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite.
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<emphasis id="7D9DBB0D2A34BB13FF02E70AE5752EAF" bold="true" box="[160,281,628,652]" pageId="86" pageNumber="2316">Aedeagus</emphasis>
|
||
(
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<figureCitation id="D7D27B9A2A34BB13FE9BE70AE5D52EAF" box="[313,441,628,652]" captionStart="Figure 35" captionStartId="50.[160,225,1682,1705]" captionTargetBox="[163,1153,142,1634]" captionTargetId="figure@50.[163,1153,142,1650]" captionTargetPageId="50" captionText="Figure 35. D. blancoi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view; (e) detail of the carena found on the middle of paramera longitudinal excavation. D. candangoi: (f) male dorsal habitus; (g) paramera dorsal view; (h) paramera ventral view; (i) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665585" httpUri="https://zenodo.org/record/3665585/files/figure.png" pageId="86" pageNumber="2316">Figure 35</figureCitation>
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(b-e)): dorsallyı paramere apexes roundedı convergent. Longitudinal excavation taking the whole paramera length and with midlongitudinal carina interrupted medially; longitudinal carina with acute apex; basal trapezoidal excavation present. Laterallyı subtle declivity from basis towards apex; widely excavated (mid-longitudinal carina also seen in this view). Ventrallyı basis with two wide elliptical excavations. Subgenital plate widely emarginated at apex and rounded basally.
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</paragraph>
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<paragraph id="4F56671F2A34BB13FF02E6F7E5BC286C" blockId="86.[160,1155,905,1103]" pageId="86" pageNumber="2316">
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<emphasis id="7D9DBB0D2A34BB13FF02E6F7E5A82F81" bold="true" box="[160,452,905,930]" italics="true" pageId="86" pageNumber="2316">Morphological variation.</emphasis>
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females BL: 12
|
||
<emphasis id="7D9DBB0D2A34BB13FDD0E6F4E6132F81" box="[626,639,906,930]" italics="true" pageId="86" pageNumber="2316">–</emphasis>
|
||
15 mmı PW:
|
||
<quantity id="8811CAFA2A34BB13FCABE6F4E7162F81" box="[777,890,906,930]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="86" pageNumber="2316" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7 to 9 mm</quantity>
|
||
and distinguished from males as follows: Head: fronto-clypeal surface striated. Cephalic carina produced in a weak pair of central tubercles. Pronotum: simply convex. Elytra: striae weakly impressed deeply impressedı punctures not distinguishable. Interstriae disc flatı with chagrinated microsculpture. Blue sheen present near basis. Abdomen: sixth ventrite bearing a weak emargination at central portion.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07F334942A34BB13FF02E102E79328B6" box="[160,1023,1148,1173]" pageId="86" pageNumber="2316" type="distribution">
|
||
<paragraph id="4F56671F2A34BB13FF02E102E79328B6" blockId="86.[160,1023,1148,1173]" box="[160,1023,1148,1173]" pageId="86" pageNumber="2316">
|
||
<emphasis id="7D9DBB0D2A34BB13FF02E102E55F28B6" bold="true" box="[160,307,1148,1173]" italics="true" pageId="86" pageNumber="2316">Distribution.</emphasis>
|
||
<collectingCountry id="37FE278F2A34BB13FEE1E103E5DA28B6" box="[323,438,1149,1173]" name="Venezuela" pageId="86" pageNumber="2316">Venezuela</collectingCountry>
|
||
and
|
||
<collectingCountry id="37FE278F2A34BB13FE4DE103E64728B6" box="[495,555,1149,1173]" name="Brazil" pageId="86" pageNumber="2316">Brazil</collectingCountry>
|
||
(RO)ı this region is also known as Tepuís.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07F334942A34BB13FF02E1BFE60828DE" pageId="86" pageNumber="2316" type="etymology">
|
||
<paragraph id="4F56671F2A34BB13FF02E1BFE60828DE" blockId="86.[160,1155,1217,1277]" pageId="86" pageNumber="2316">
|
||
<emphasis id="7D9DBB0D2A34BB13FF02E1BFE54F28F9" bold="true" box="[160,291,1217,1242]" italics="true" pageId="86" pageNumber="2316">Etymology.</emphasis>
|
||
named after Jorge Blanco (Universidad Nacional Experimental del Táchiraı
|
||
<collectingCountry id="37FE278F2A34BB13FF02E19BE57428DE" box="[160,280,1253,1277]" name="Venezuela" pageId="86" pageNumber="2316">Venezuela</collectingCountry>
|
||
)ı who collected the
|
||
<typeStatus id="9052D9BD2A34BB13FE58E19BE60D28DE" box="[506,609,1253,1277]" pageId="86" pageNumber="2316" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |